DOCSLIB.ORG

Habitat Selection of Dusky Grouse on a Biosolids-Remediated

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Habitat Selection of Dusky Grouse on a Biosolids-Remediated HABITAT SELECTION OF DUSKY GROUSE ON A BIOSOLIDS-REMEDIATED CATTLE RANCH IN BRITISH COLUMBIA by Kirstie Jaylene Lawson B.S., University of Montana, 2012 A THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE in THE COLLEGE OF GRADUATE STUDIES (Biology) THE UNIVERSITY OF BRITISH COLUMBIA (Okanagan) July 2018 © Kirstie Jaylene Lawson, 2018 The following individuals certify that they have read, and recommend to the College of Graduate Studies for acceptance, a thesis/dissertation entitled: HABITAT SELECTION OF DUSKY GROUSE ON A BIOSOLIDS-REMEDIATED CATTLE RANCH IN BRITISH COLUMBIA submitted by Kirstie Jaylene Lawson in partial fulfillment of the requirements of the degree of Master of Science . Dr. Karen Hodges, Irving K. Barber School of Arts and Sciences Supervisor Frank Doyle, Wildlife Dynamics Consulting Supervisory Committee Member Dr. Bob Lalonde, Irving K. Barber School of Arts and Sciences Supervisory Committee Member John Lavery, SYLVIS Environmental Services Supervisory Committee Member Dr. Trudy Kavanaugh, Irving K. Barber School of Arts and Sciences University Examiner ii Abstract In North America, habitat loss and fragmentation have caused declines in many gallinaceous species. Before we can understand how these changes to a landscape affect an individual species, we must first understand how that species uses their habitat. Our understanding of dusky grouse (Dendragapus obscurus) habitat selection is lacking and I focus my research on increasing our knowledge of this forest grouse species. In this thesis, I assess microhabitat selection at nesting and brood-rearing sites, as well as patch-level habitat selection and the effects of biosolids on amendments on habitat selection of dusky grouse. I trapped and radio-collared 26 dusky grouse hens during 2016-2017 and tracked individuals throughout the summer months (May-July). I used model selection in an AIC framework to determine which vegetation variables affect dusky grouse choice of nesting and brood-rearing sites at the microhabitat level. Dusky grouse are habitat generalists. However, hens selected nest sites with high visual cover, and nest success was positively influenced by visual obstruction of the nest bowl. This suggests that cover from predators is important when hens select nest sites, and that predation may be influenced by the predator’s ability to see the nest. Brood sites were more variable, and brood hens appeared to choose sites based on non-vegetative variables. Furthermore, the use of biosolids, treated human waste, on the landscape allowed me to look at dusky grouse use these amendments. I assess broad-scale habitat select through patch-level use of available habitat types and the distance dusky grouse traveled into biosolids-amended and untreated grasslands. I found drastic year differences in selection and use of grasslands, and weather or food abundance may play a role in site selection. In 2017, a much drier, warmer year, dusky grouse hens traveled significantly further into biosolids-amended grasslands than untreated grasslands. In 2016, hens traveled much further into grasslands overall, showing iii similar distributions in both biosolids-amended and untreated grasslands. My research suggests that biosolids may be beneficial for dusky grouse in drought years, by enhancing plant growth and increasing insect densities. iv Lay Summary Little is known about how dusky grouse, a forest grouse species found in Western North America, use the habitat available to them. I evaluated use of this habitat at a small scale, choice of nest sites and sites to raise their chicks, and at a large scale, choice of forest and grassland patches. I found that the amount of visual cover is important to the choice of nest sites and to nest success, providing cover from animals that might attack the nest. Dusky grouse prefer conifer forests, but females will travel further into grasslands during years with more precipitation and cooler summers. The use of biosolids, treated human waste, on grasslands at the ranch may be beneficial to dusky grouse during drought years, decreasing the rate at which the vegetation dries out and increasing food available to grouse. v Preface For this research, I was responsible for designing the study, conducting fieldwork, analyzing the data, and writing and editing this thesis. Dr. Karen Hodges and Frank Doyle provided feedback throughout the process and guided me on which techniques to use. The original research idea was requested by the Upland Bird Society, and Dr. Hodges and Frank Doyle came up with the initial concepts of the research. Frank Doyle assisted with fieldwork, training me on trapping techniques for dusky grouse, and provided field support throughout the study. Dr. Bob Lalonde and John Lavery provided comments on the original research proposal and the written thesis. Fieldwork was completed with the help of Megan Buers and Emma Gaudreault. Both data chapters will be submitted to peer-reviewed journals. The publications arising from Chapter 2 and Chapter 3 will be co-authored by myself, Dr. Karen Hodges, and Frank Doyle. This research will also be published in concert with a larger study on biosolids impacts on wildlife. All animal handling techniques used in this research were approved by the University of British Columbia Animal Care Committee (Certificate # A15-0216) and the British Columbia Ministry of Forests, Lands, and Natural Resource Operations (Permit # WL17-263857). vi Table of Contents Abstract .......................................................................................................................................... iii Lay Summary .................................................................................................................................. v Preface............................................................................................................................................ vi Table of Contents...........................................................................................................................vii List of Tables ................................................................................................................................. ix List of Figures ................................................................................................................................. x Acknowledgements ........................................................................................................................ xi Chapter 1: Introduction ................................................................................................................... 1 1.1 Focal species: Dusky Grouse ................................................................................................ 1 1.2 Speciation of Dusky Grouse .................................................................................................. 3 1.3 Landscape Restoration and Wildlife Habitat ........................................................................ 3 1.4 What are Biosolids? .............................................................................................................. 4 1.5 Regulation of Biosolids ......................................................................................................... 6 1.6 Wildlife Response to Biosolids ............................................................................................. 7 1.7 Research Objectives .............................................................................................................. 9 Chapter 2: Microhabitat selection of nest and brood sites by Dusky grouse (Dendragapus obscurus) in Southern Interior British Columbia .......................................................................... 10 2.1 Background ......................................................................................................................... 10 2.2 Methods ............................................................................................................................... 13 2.3 Results ................................................................................................................................. 18 2.3.1 Nest Sites .......................................................................................................................... 18 2.3.2 Nest Success ..................................................................................................................... 19 2.3.3 Brood Sites ....................................................................................................................... 21 2.4 Discussion ........................................................................................................................... 22 Chapter 3: Patch-level habitat selection by Dusky Grouse (Dendragapus obscurus) in a biosolids- amended landscape ....................................................................................................................... 34 3.1 Background ......................................................................................................................... 34 3.2 Methods ............................................................................................................................... 38 3.3 Results ................................................................................................................................. 41 3.4 Discussion ..........................................................................................................................
Load more

Recommended publications
  • WATERFOWL ___Cackling Goose
    WATERFOWL ____ Least Sandpiper FALCONS BUSHTITS ____ Pine Grosbeak ____ Cackling Goose ____ Wilson's Snipe ____ American Kestrel ____ Bushtit ____ House Finch ____ Canada Goose ____ Spotted Sandpiper ____ Merlin ____ Purple Finch NUTHATCHES ____ Wood Duck ____ Solitary Sandpiper ____ Peregrine Falcon ____ Red-breasted Nuthatch ____ Cassin’s Finch ____ Blue-winged Teal ____ Greater Yellowlegs ____ Prairie Falcon ____ White-breasted Nuthatch ____ Common Redpoll ____ Cinnamon Teal ____ Red Crossbill CORMORANTS FLYCATCHERS ____ Pygmy Nuthatch ____ Northern Shoveler ____ Double-crested Cormorant ____ Western Kingbird ____ Pine Siskin CREEPERS ____ Gadwall ____ Eastern Kingbird ____ American Goldfinch ____ American Wigeon PELICANS, HERONS, IBIS ____ Brown Creeper ____ Great Blue Heron ____ Olive-sided Flycatcher SPARROWS ____ Mallard ____ Western Wood-Pewee WRENS ____ Chipping Sparrow ____ Green-winged Teal VULTURES, RAPTORS ____ Willow Flycatcher ____ Rock Wren ____ Fox Sparrow ____ Ring-necked Duck ____ Turkey Vulture ____ Least Flycatcher ____ Canyon Wren ____ American Tree Sparrow ____ Harlequin Duck ____ Osprey ____ Hammond’s Flycatcher ____ House Wren ____ Dark-eyed Junco ____ Bufflehead ____ Golden Eagle ____ Gray Flycatcher ____ Pacific Wren ____ White-crowned Sparrow ____ Common Goldeneye ____ Northern Harrier ____ Dusky Flycatcher ____ Marsh Wren ____ Golden-crowned Sparrow ____ Barrow's Goldeneye ____ Sharp-shinned Hawk ____ Pacific-slope Flycatcher ____ Bewick’s Wren ____ White-throated Sparrow ____ Hooded Merganser ____ Cooper’s
    [Show full text]
  • 2020 Upland Game Bird Hunting Season Frameworks
    Oregon Upland Game Bird Hunting Season Framework Effective dates: September 1, 2015 through August 31, 2020 Prepared by Wildlife Division Oregon Department of Fish and Wildlife 4034 Fairview Industrial Dr SE Salem, Oregon 97302 Proposed 2015 -2020 Upland Game Bird Framework Executive Summary Oregon’s diverse habitats support 12 species of upland game birds, 8 of which are native, and 10 of which have hunting seasons. This document contains the proposed framework for upland game bird hunting seasons for the next five years. The seasons are designed to provide recreational hunting opportunities compatible with the overall status of upland game bird populations. The multi-year framework approach for setting upland game bird regulations was first adopted by the Oregon Fish and Wildlife Commission in 1996 for a 3-year period and again in 1999, 2004, and 2009 for 5-year periods. The role of regulations in game bird management has many functions including protection of a species, providing recreational opportunities, and in consultation with hunters to provide bag limits and seasons. Regulations should be as simple as possible to make them easy to understand. These frameworks are also based on the concept that annual fluctuations in upland bird numbers, which can vary greatly and are normal, should not be the basis for setting hunting seasons year by year. Standardized frameworks are biologically sound management tools that help the Oregon Department of Fish and Wildlife (Department) provide consistent, stable regulations that reduce confusion, assist hunters with planning trips, and lower administrative costs. In an effort to stabilize hunting regulations, the following concepts are the foundation for the frameworks offered in this document: Many upland game bird populations exhibit a high annual death rate and cannot be stockpiled from year to year Similar annual death rates occur in most upland game bird populations whether they are hunted or not.
    [Show full text]
  • BLUE GROUSE and RUFFED GROUSE (Dendragapus Obscurus and Bonasa Umbellus)
    Chapter 13 BLUE GROUSE AND RUFFED GROUSE (Dendragapus obscurus and Bonasa umbellus) Harry Harju I. CENSUS – A. Production Surveys – 1. Rationale – Random brood counts are a survey method used to assess the reproductive success of blue or ruffed grouse. Unfortunately, these counts are done too late in the year to be considered in setting or adjusting hunting seasons. If the sample size is large enough, the counts can help identify important habitats used by broods and can provide insight to the potential quality of hunting in the fall. 2. Application – Random brood counts can be conducted on foot, from horseback or from a vehicle and should cover all portions of the brood-rearing area. A good pointing dog is invaluable to locate broods. If a flushing dog is used, it should be trained to walk very close to the observer. Each time a grouse is seen, record the species, location, age, sex and habitat on a wildlife observation form. If a count is incomplete, circle the number of birds recorded. The time frame for these surveys is July 15 to August 31. Warm, clear days are best for brood counts. The best results are obtained by searching for broods in the first two and last three hours of daylight. When a well-trained dog is used, counts can be conducted throughout the day. 3. Analysis of Data – Refer to chapter 12 (Sage-grouse), Section II.B (Brood Production). 4. Disposition of Data – All records of brood observations are forwarded to Regional Wildlife Management Coordinators for proofing, and then entered into the Wildlife Observation System Database.
    [Show full text]
  • 11 Blue Grouse
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Grouse and Quails of North America, by Paul A. Johnsgard Papers in the Biological Sciences May 2008 11 Blue Grouse Paul A. Johnsgard University of Nebraska-Lincoln, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/bioscigrouse Part of the Ornithology Commons Johnsgard, Paul A., "11 Blue Grouse" (2008). Grouse and Quails of North America, by Paul A. Johnsgard. 13. https://digitalcommons.unl.edu/bioscigrouse/13 This Article is brought to you for free and open access by the Papers in the Biological Sciences at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Grouse and Quails of North America, by Paul A. Johnsgard by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Blue Grouse Dendragapw obsctlrus (Say) 182 3 OTHER VERNACULAR NAMES @USKY grouse, fool hen, gray grouse, hooter, mountain grouse, pine grouse, pine hen, Richardson grouse, sooty grouse. RANGE From southeastern Alaska, southern Yukon, southwestern Mackenzie, and western Alberta southward along the offshore islands to Vancouver and along the coast to northern California, and in the mountains to southern California, northern and eastern Arizona, and west central New Mexico (A.O.U. Check-list). SUBSPECIES (ex A.O.U. Check-list) D. o. obscurus (Say): Dusky blue grouse. Resident in the mountains from central Wyoming and western South Dakota south through eastern Utah and Colorado to northern and eastern Arizona and New Mexico. D. o. sitkensis Swarth: Sitkan blue grouse. Resident in southeastern Alaska south through the coastal islands to Calvert Island and the Queen Charlotte Islands, British Columbia.
    [Show full text]
  • Observation on a Sooty Grouse Population at Sage Hen Creek
    THE CONDOR--- VOLUME 58 SEPTEMBER-OCTOBER, 1956 NUMBER 5 OBSERVATIONS ON A SOOTY GROUSE POPULATION AT SAGE HEN CREEK, CALIFORNIA By ROBERT S. HOFFMANN In recent years there has been considerable interest in two separate aspects of the biology of Blue Grouse (Dendragapus) . The first of these is the controversy concerning their taxonomy. The genus is widely distributed in the coniferous forests of the western states and consists of two groups of races recognized by some as separate species: the fuliginosus group, or Sooty Grouse, along the Pacific coast, and the obscures group, or Dusky Grouse, in the Great Basin and Rocky Mountain areas. Originally these two groups were placed under the name Dendragapus obscurus on the basis of supposed intergradation (Bendire, 1892:41,44, SO). The work of Brooks (1912,1926,1929) and Swarth (1922, 1926), however, led to their separation in the fourth edition of the A.O.U. Check-list (1931) into coastal and interior species. This split stood until the publication of the 19th Supplement to the A.O.U. Check-list (1944) when, following Peters (1934), D. fuliginosus and its subspecieswere replaced under D. obscurus. Al- though at that time some doubt was still expressedabout the correctnessof this merger (Grinnell and Miller, 1944: 113), intergradation in northern Washington and southern British Columbia between the races fuliginosus and pallidus has now been reported by several authors (Munro and Cowan, 1947:89; Carl, Guiguet, and Hardy, 1952:86; Jewett, Taylor, Shaw, and Aldrich, 1953 : 200). The taxonomy of these grouse must rest upon the fact of intergradation between the two groups of races.
    [Show full text]
  • Dusky Grouse Tend to Be Adapted Dusky and Sooty Grouse Were Considered Distinct
    Volt.un£ 14 .Numkt 3 9all2006 Blue Grouse divided by two equals Dusky and Sooty Grouse Michael A. Schroeder, Upland Bird Research Biologist Washington Department of Fish and Wildlife P.O. Box 1077 Bridgeport, W A 98813 email: [email protected] The state of Washington has a new species of grouse, at least forested habitats throughout the year, it appears to be according to the American Ornithologists Union (Auk, 2006, vulnerable to variations in forest practices. For example, Pages 926-936). Dendragapus obscurus (formerly known as research on Sooty Grouse in British Columbia indicated that the Blue Grouse) has now been split into 2 species based on their populations fluctuated dramatically depending on the genetic, morphological, and behavioral evidence; the Dusky age of the forest following clear-cutting. Unfortunately, there Grouse (Dendragapus obscurus) and the Sooty Grouse has been little effort to evaluate the relationship between (Dendragapus fuliginosus). This split is actually a reversion forest management practices and Sooty Grouse populations. to the previous situation during the early 1900s when the In contrast to Sooty Grouse, Dusky Grouse tend to be adapted Dusky and Sooty Grouse were considered distinct. to relatively open habitats in forest openings or close to forest edges. Because these open habitats are preferred areas for As a group, Dusky and Sooty Grouse are widely distributed livestock production and development, it is necessary to in the mountainous portions of western North America. understand the relationships between land use and grouse Although they generally winter in coniferous forest, their populations. The human population increase in the breeding breeding habitats are quite varied.
    [Show full text]
  • Alpha Codes for 2168 Bird Species (And 113 Non-Species Taxa) in Accordance with the 62Nd AOU Supplement (2021), Sorted Taxonomically
    Four-letter (English Name) and Six-letter (Scientific Name) Alpha Codes for 2168 Bird Species (and 113 Non-Species Taxa) in accordance with the 62nd AOU Supplement (2021), sorted taxonomically Prepared by Peter Pyle and David F. DeSante The Institute for Bird Populations www.birdpop.org ENGLISH NAME 4-LETTER CODE SCIENTIFIC NAME 6-LETTER CODE Highland Tinamou HITI Nothocercus bonapartei NOTBON Great Tinamou GRTI Tinamus major TINMAJ Little Tinamou LITI Crypturellus soui CRYSOU Thicket Tinamou THTI Crypturellus cinnamomeus CRYCIN Slaty-breasted Tinamou SBTI Crypturellus boucardi CRYBOU Choco Tinamou CHTI Crypturellus kerriae CRYKER White-faced Whistling-Duck WFWD Dendrocygna viduata DENVID Black-bellied Whistling-Duck BBWD Dendrocygna autumnalis DENAUT West Indian Whistling-Duck WIWD Dendrocygna arborea DENARB Fulvous Whistling-Duck FUWD Dendrocygna bicolor DENBIC Emperor Goose EMGO Anser canagicus ANSCAN Snow Goose SNGO Anser caerulescens ANSCAE + Lesser Snow Goose White-morph LSGW Anser caerulescens caerulescens ANSCCA + Lesser Snow Goose Intermediate-morph LSGI Anser caerulescens caerulescens ANSCCA + Lesser Snow Goose Blue-morph LSGB Anser caerulescens caerulescens ANSCCA + Greater Snow Goose White-morph GSGW Anser caerulescens atlantica ANSCAT + Greater Snow Goose Intermediate-morph GSGI Anser caerulescens atlantica ANSCAT + Greater Snow Goose Blue-morph GSGB Anser caerulescens atlantica ANSCAT + Snow X Ross's Goose Hybrid SRGH Anser caerulescens x rossii ANSCAR + Snow/Ross's Goose SRGO Anser caerulescens/rossii ANSCRO Ross's Goose
    [Show full text]
  • Blue Grouse Ecology and Habitat Requirements in North Central Utah
    Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 5-1972 Blue Grouse Ecology and Habitat Requirements in North Central Utah David A. Weber Utah State University Follow this and additional works at: https://digitalcommons.usu.edu/etd Part of the Life Sciences Commons Recommended Citation Weber, David A., "Blue Grouse Ecology and Habitat Requirements in North Central Utah" (1972). All Graduate Theses and Dissertations. 3512. https://digitalcommons.usu.edu/etd/3512 This Thesis is brought to you for free and open access by the Graduate Studies at DigitalCommons@USU. It has been accepted for inclusion in All Graduate Theses and Dissertations by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. ii ACKNOWLEDGMENTS I would hke to express my sincere appreciation to Dr. J. B. Low, leader of the Utah Cooperative Wildlife Research Unit, for giving me the opportunity to work on this project and for his continuing guidance and patience during the study. I would also like to thank the members of my graduate committee: Dr. Donald Davis, Dr. Arthur Holmgren, and Dr. William Sigler for their advice and assis­ tance in many ways during the two years of the research. Frank Gunnell and Bruce Reese of the U. S. Forest Service deserve special thanks for many hours of help in the field and for continuing constructive criticism. Jack Rensel, Clair Huff, and John Kimball of the Utah Division of Wildlife Resources also provtded field assistance and suggestions during the course of the study. I very much appreciate the help and advice of my co-worker during 19 71, J:!arry l:larnes, and wish him well in his continuing studies of the blue grouse.
    [Show full text]
  • Ruffed and Hazel Grouse
    Grouse Bibliography Revision date: 6 December 2013 Check for newer version Compiled by Donald H. Wolfe George M. Sutton Avian Research Center Organized by genus Bonasa and Tetrastes Centrocercus Dendragapus and Falcipennis Lagopus Tetrao and Lyrurus Tympanuchus general grouse related topics Note: I would welcome any other citations not listed here or links to other grouse- related bibliographical sources. Also, please alert me of any errors that are detected. This bibliography is always a “work in progress”, and is updated as new information is available and time allows. Bonasa and Tetrastes (Ruffed, Hazel, and Chinese Grouse) Aberg, J. 1996. Effects of habitat fragmentation on Hazel Grouse (Bonasa bonasia) in boreal landscapes. Swedish University of Agricultural Sciences, Department of Wildlife Ecology, Report 32. 69pp. Aberg, J. 2000. The occurrence of Hazel Grouse in the boreal forest; effects of habitat composition at several spatial scales. Ph. D. dissertation. Swedish University of Agricultural Science, Uppsala. 108pp. Aberg, J., G. Jansson, J. E. Swenson, and P. Angelstam. 1995. The effect of matrix on the occurrence of Hazel Grouse (Bonasa bonasia) in isolated habitat fragments. Oecologia 103:265-269. 1 Aberg, J., G. Jansson, J. E. Swenson, and P. Angelstam. 1996. The effect of matrix on the occurrence of Hazel Grouse in isolated habitat fragments. Grouse News 11:22. Aberg, J., G. Jansson, J. E. Swenson, and G. Mikusinski. 2000. Difficulties in detecting habitat selection by animals in generally suitable areas. Wildlife Biology 6:89- 99. Aberg, J, J. E. Swenson, and H. Andren. 2000. The dynamics of Hazel Grouse (Bonasa bonasia L.) occurrence in habitat fragments.
    [Show full text]
  • Forest Grouse
    Volume 34 | Issue 2 October 2020 FOREST GROUSE Dusky GrouseCCBY Paul Hurtado Flickr Inside: Meet the Grouse Moving Up Leaving Home Hey Baby... Be Outside! idfg.idaho.gov Meet the Grouse If so, Have you ever had a quiet hike interrupted by an exploding bird? you have met a member of the grouse family. While they do not really explode, their sudden get-away right at your feet seems like a feathered explosion. In fact, this sudden rapid flight is common in this group of birds. Their short, rounded wings can beat very fast. This lets the bird suddenly fly into the air. Just like it startles you, it can also startle a predator just long enough for the bird to fly away to safety. Grouse belong to a large group of birds called “galliforms.” This group is often referred to as upland game birds. It includes birds like turkeys, quail, and grouse. They are all ground-dwelling birds that feed mostly on plants and insects. About 183 species of these birds live around the world. They are found everywhere except Antarctica and South America. Twelve species of galliforms are native to North America. Here in Idaho, we have five native grouse species. Several other species of galliform birds live in Idaho, but they were introduced as game birds for hunting. Upland game birds are well camouflaged. Their colors are mainly brown, tan, buff, gray, black, and white. Because they spend a lot of time on the ground, being camouflaged is very important. Three members of this group, called ptarmigan (TAR-mi-gan), even turn white in the winter.
    [Show full text]
  • Birds of the Ahwahnee Hills Regional Park
    Birds of the Ahwahnee Hills Regional Park Snow Goose American Coot Say's Phoebe Prairie Warbler* Ross's Goose* Sandhill Crane* Ash-throated Flycatcher Blackpoll Warbler* Cackling Goose* Semipalmated Plover Cassin's Kingbird* Cerulean Warbler* Canada Goose Killdeer Western Kingbird Black-and-white Warbler* Wood Duck Black-necked Stilt Eastern Kingbird* American Redstart* Green-winged Teal American Avocet Horned Lark Ovenbird* Mallard Greater Yellowlegs Purple Martin* MacGillivray's Warbler Northern Pintail Solitary Sandpiper* Tree Swallow Common Yellowthroat Blue-winged Teal* Willet Violet-green Swallow Wilson's Warbler Cinnamon Teal Spotted Sandpiper Northern Rough-winged Swallow Yellow-breasted Chat Northern Shoveler Long-billed Curlew Bank Swallow Western Tanager Gadwall Western Sandpiper Cliff Swallow Rose-breasted Grosbeak* American Wigeon Least Sandpiper Barn Swallow Black-headed Grosbeak Canvasback Wilson's Snipe Steller's Jay Blue Grosbeak* Redhead Wilson's Phalarope Western Scrub Jay Lazuli Bunting Ring-necked Duck Red-necked Phalarope Pinyon Jay* Green-tailed Towhee* Greater Scaup* Red Phalarope* Clark's Nutcracker* Spotted Towhee Lesser Scaup Ring-billed Gull Black-billed Magpie* California Towhee Harlequin Duck* California Gull Yellow-billed Magpie Rufous-crowned Sparrow Common Goldeneye Herring Gull American Crow Chipping Sparrow Barrow's Goldeneye* Caspian Tern Common Raven Clay-colored Sparrow* Bufflehead Forster's Tern Mountain Chickadee Brewer's Sparrow* Hooded Merganser Rock Pigeon (I) Chestnut-backed Chickadee* Black-chinned
    [Show full text]
  • Colorado Birds the Colorado Field Ornithologists’ Quarterly
    Vol. 45 No. 3 July 2011 Colorado Birds The Colorado Field Ornithologists’ Quarterly Remembering Abby Modesitt Chipping Sparrow Migration Sage-Grouse Spermatozoa Colorado Field Ornithologists PO Box 643, Boulder, Colorado 80306 www.cfo-link.org Colorado Birds (USPS 0446-190) (ISSN 1094-0030) is published quarterly by the Colo- rado Field Ornithologists, P.O. Box 643, Boulder, CO 80306. Subscriptions are obtained through annual membership dues. Nonprofit postage paid at Louisville, CO. POST- MASTER: Send address changes to Colorado Birds, P.O. Box 643, Boulder, CO 80306. Officers and Directors of Colorado Field Ornithologists: Dates indicate end of current term. An asterisk indicates eligibility for re-election. Terms expire 5/31. Officers: President: Jim Beatty, Durango, 2013; [email protected]; Vice President: Bill Kaempfer, Boulder, 2013*; [email protected]; Secretary: Larry Modesitt, Greenwood Village, 2013*; [email protected]; Treasurer: Maggie Boswell, Boulder, 2013; trea- [email protected] Directors: Lisa Edwards, Falcon, 2014*; Ted Floyd, Lafayette, 2014*; Brenda Linfield, Boulder, 2013*; Christian Nunes, Boulder, 2013*; Bob Righter, Denver, 2012*; Joe Roller, Denver, 2012* Colorado Bird Records Committee: Dates indicate end of current term. An asterisk indicates eligibility to serve another term. Terms expire 12/31. Chair: Larry Semo, Westminster; [email protected] Secretary: Doug Faulkner, Arvada (non-voting) Committee Members: John Drummond, Monument, 2013*; Peter Gent, Boulder, 2012; Bill Maynard, Colorado Springs, 2013; Bill Schmoker, Longmont, 2013*; David Silver- man, Rye, 2011*; Glenn Walbek, Castle Rock, 2012* Colorado Birds Quarterly: Editor: Nathan Pieplow, [email protected] Staff: Glenn Walbek (Photo Editor), [email protected]; Hugh Kingery (Field Notes Editor), [email protected]; Tony Leukering (In the Scope Editor), GreatGrayOwl@aol.
    [Show full text]