Bul l. Br. a rachnol. Soc. (2006) 13 (7), 233- 248 233
Taxonomic notes on nine Aellirililis species of the demonstrate rather strong variation of the embolus and western Mediterranean (Araneae: Salticidae) epigyne. for instance, the embolic tips of A. luctuosus (Figs. 46- 55) and A. monarc/i (Figs. 90- 97) vary signi fi Galina N. Azarkina cantly with regard to the ventral membranous projection Siberian Zoological Muse um , (arrowed in Fig. 55), which can be either elongated Institute for Systematics and Ecology of Animals, (Figs. 53, 55, 90, 92, 96) or rounded (F igs. 51 , 52, 54, 94). Siberian D ivision o f the R uss ia n Academy of Sciences, Of the variants seen, those shown in Figs. 92 and 96 Frunze Street II , Novosibirsk 63 009 1, Russia were the most common and were recorded in most of the studied males of both species. The same range of and embolic variation was demonstrated by Cantarell a (1 982) for A. schembrii Cantarella, 1982 and A. monarch; Dmitri V. Logunov she studied eight males of the latter species from Algeria Manchester M use um, University of Mancheste r, (Biskra and Menuen). The epigynal structure of A. Oxford Road, Manchester, M 13 9PL 11iC/UOSUS , for example, shows strong variation both of the insemination ducts (Figs. 74, 76, 78, 8 1) and of the epigynal flaps, which can be almost subparallel to the Summar), epigastric furrow (Figs. 73 , 77) or form a sort of wide F ive poorl y known species of Aeluril!us from the western median septum varying in length (Figs. 75, 79, 80). It is Medi terranea n are diagnosed, tigured and redescribed: A . well understood that strong variation of genital charac basseleli (Lucas, 1846), A. fl irlipes Denis, 1960, A. IUI'luosuS ters is a common phenomenon in Salticidae, and it has (Lucas, 1846), A. mOllardi (Lucas, 1846) and A. plumipes (T horell , 1875); neotypes are desi gnated for three of them: been clearly demonstrated fo r Pelegrina insignis (Banks, A. basseleli, A. lucluosuS and A. monarcli. Six new synony 1892) (see Cutler, 1980: sub M etaphidippus i.), and in the mies are established : Sallicus offinis Lucas, 1846, A III ls genera Ruophrys and Phlegret (Logunov et al., 1993 ; ogieri Simon, 18 68 a nd Aelurillus pollidemocu!alU., Denis, Logunov, 1996), etc. Taxonomic conclusions in these 1937 are synonymised with Aelurillus lucluosus (Lucas, and many other salticid groups based upon sin gle sexes 1846); Yllenlls cervinus Thorell , 1875 and Aelurillus moyeli Simon, 1885 a re synonymised with Aelurillus plumipes (especially single females) are either poorly grounded or (Thorell , 1875), and Aelurillus sinaicus Proszyil ski , 2000 even impossible; e. g., fe males of Euophrys (s. str.) are wit h A. cOl/veniens (0. Pickard-Cambridge , (872). The virtually indistinguishable (see Logunov et al., 1993). female of A. iIirlipes is described for the tirst time. Com Therefore, in the present paper we pay no attention to ments on the taxonomic status of Aclurillus gesliculator (Lucas, 1846), A. nico leli (L ucas, (846) and A. l1 umidicus tiny differences in the embolic tips and the epigynal fl aps (Lucas, 1846) a re provided, and A. nicoleli is removed from seen in the studied A elurillus species. synonymy with A. monardi. Aelurillus lopadusae Ca nta rella, The aims of the present paper are (I) to diagnose 1983 is recorded from Algeria for the ti rst time. Dist ributio n and redescribe fi ve Aelurillus species from the western maps a re gi ve n for a ll va lid species. Mediterranean, (2) to clarjfy the taxonomic status of the Aelurillus species described by Lucas (1846) from Al geria, (3) to provide evidence for six new synonymies Introduction revealed during the course of this study, and (4) to refi ne The genus A elurillus Simon, 1884, which is currently and map the distribution of all the studied Aelurillus under revi sion by one of us (G.A.), remains insufficiently speCIes. studied, fo r until now many of its species have been known only from old original descriptions based on Material and methods single or a few speci mens of one sex. The most significant case of this kind is the work by Lucas (1846), who T his work is based on both museum collections and described seven species currently assigned to Aelurillus, new material from the western Mediterranean, mainly four from male(s) and three from female(s). The types of from Algeria, Tunisia, Spain and Portugal. A total of all of them are inaccessible and considered lost, and their 195 specimens have been (re)examined . Specimens for taxonomic status remains uncertain. r ortunatcly, Lucas this study were borrowed from or are distributed among published quite detailed descriptiuns and also provided the following museums and personal co ll ections: colour fi gures for a ll of hi s species. Judging from his C BA=Centro de Biologia Ambiental, Faculdade de accounts, along with the study of numerous newly col Ciencias de Lisboa, Baixa da Banheira, Portugal (Dr P. lected Aelurilfus specimens from Algeria, taken by Dr Cardoso); DBAU =Dipartimento di Biologia Animale Robert Bosmans (Gent, Belgium) ;~om t!1.e type localities dell'U niversita, Catania, Italy (Prof. T. Cantarell a); of Lucas' species, we have been able to draw reliable HECO=Hope Entomological Collection, Oxford, U K conclusions on many of these speCIes. I'urthermore, hav (Mr J. Hogan); H UJ=Hebrew U ni versity, Jerusalem ing studied extensive recent salticid collections by vari (Dr G. Levy); ISEA=Siberian Zoological Museum of ous people from Tunisia , Algeria, Spain and Portclgal, the Institute for Systematics and Ecology of Ani mals, we are able to clarify the taxonomic status of some other N ovosibirsk, Russia (Miss G . N. Azarkina); Aeluri//us species described from s;ngle sexes by Sirr.un IZWP=Insti tute of Zoology, Warsaw, Poland (Dr T. (18 68 , 1885), Denis (1937,1960) and Thorell (IR75). Huflejt); MCZ=Museum of Comparative Zoology, Most of the Aelurillus specics known to us, and for Harvard U niversity, Cambridge, USA (M rs L. which we have been able to study sufficient matl.:ricil , Leibensperger); MNHN= M useum N ational d'Histoire 234 /Jell/rilll/s species of the W. Mediterranean
6 8
Figs. 1- 16: Male " faces" and palpal femora of Aelurillus spp, 1-8 A, rnol1ardi; 9-12 A, lucruosl/s; 13-16 A, blm'e/eli, Specimens: (1 - 2) Algeria, Tolga; (3-4) T unisia. Hammamet; (5- 6) Algeria, EI Kala, Cap Rosa; (7- 8) Al ge ria, Ojebel Babor; (9- 10) Algeria, Blida, Ojebel Mouzai'a; (\1- 12) Al ge ri a, Chleff" foret de Taeheta; (13 - 14) Algeria, Tlemcen . Zarifete; (15- 16) Algeria, Alger, EI Harrach.
Naturelle, Paris, France (Dr C. Rollard); MRAC= pr= prolaleral, rt= retrolateral, v= ventral. For the leg Musee Royal de I' Afrique Centrale, Tervuren, Belgium spination the system adopted is that used by Ono (1988). (Dr R. Jocque); PCJM=personal collection of Mr J. The sequence of leg segments in measurement data is as Murphy, Hampton, UK; PCRB=personal collectio n of follows: femur + patella + tibia + metatarsus + tarsus. All D r R , Bosmans, Gent, Be lgium; PCRS=personal collec measurements are in mm. tion of Mr R. Snazell, Swanage, Dorset, UK; PCYM = personal collection of Mr Y. Montardi, Paris, France; Survey of species SMNH =Swedish Museum of Natural History, Stockholm, Sweden (Dr T. Kronestedt); SMFM = A ell/filius basseleti (Lucas, 1846) (Figs. 13- 16, 17- 32) Senckenberg Natural History Museum, Frankfurt am Sa/lin ls basse/elii Lucas. 1846: 158- 159, pI. 7, fig, I (Do; 0 holotype Main, Germany (Dr P. Jager); ZMCD=Zoological lost), A 0 neotype is designated here, Museum, Copenhagen, Denmark (Dr N. Scharff); /Je/urops basse/eli: Simon, 1876: 139, ZMMU=Zoological Museum of Moscow University, IClidops basse/eli: Pavesi, 1880: 383, Moscow, Russia (Dr K. G, Mikhailov), Ae/uri//lls basse/eli: Simon. 1885: 4; Roewer, J955: 1113 ; Bon net, 1955: 165; Proszyn ski, 2003b, The term " face" is used to describe the entire frontal part of the carapace (as seen from in front), the term Type : Neotype 0 of Aelurillus basseleti (MNHN; " cheek" describes the area below and slightly laterad of designated here), Algeria, Wilaya Alger, EI Harrach the PLE, and the term "cilia" is adopted for scales [c. 36°43' N, 3°07'E], 26 November 1985 (RB). surrounding the eyes of the first row. A generalised Diagnosis: This species is most similar to A. luctuosus " face " was used to show male face coloration of and A. monardi, but its males can easily be separated by di ffe rent species in Figs. 1, 3, 5, 7, 9, II, 13 and 15, the presence of a wide, white or yellowish (sometimes The name of the main collector, whose collections tawny), transverse band on the clypeus (Figs, 13, 15) were used for thi s study, is abbreviated as follows: [vs, uniformly black or brown clypeus in A, luctuosus RB =Dr R. Bosmans. Other abbreviations used in the (Figs. 9, II) and the " face" with three narrow vertical tex t: AME=anterior median eye, ALE=anterior lateral lines in A. monardi (Figs. I, 3, 5, 7)]. Females of these eye, PLE=posterior lateral eye; Fm=femur, Pt=patella, species are practically indistinguishable (cf. Figs. 29- 31 , 'Tb=tibia, Mt=metatarsus; ap=apical, d =dorsal, 73- 81 and CantareIla, 1982: fig. 19), Galina N. Azarkina & D. V. Logunol' 235
Comments: This species was described by Lucas (1846) designate a 0 neotype collected from A lger (El Harrach), from a single male from Constantine (Algeria). The 0 a locality close to the original one (Constantine). holotype of A. basseleli is considered lost. However. In his extensive work on the Salticidae of Levant, matching the studied males of A. hasselcli t() Lucas' Proszyl'lski (2003a: figs . 92- 93) also illustrated the male description and colour illustration (1846: 158 159, ()f A. hasseleti, which was loaned to him from the MCZ pI. 7, fig. I) seems to cause no problems. While describ (Proszyl'lski, pers. comm.). We have also re-examined ing the carapace colour (op. cit.: p. 158), Lucas clearly this male and found that it belongs to A. luctuosus (see stated that " ... les bords lateraux sont entierement below). Therefore, Proszyl'lski's record is excluded from entoures de poils d'un bl anc tn':s-legerement jaunatn:, the synonymy list of A. basseleli. ..." and later " ... la partie anterieure ou Ie front Distribution: Algeria and Tunisia (Lucas, 1846: fer rugineux." Of the three related species (see " Diagno sub Salticus b. ; Simon, 1885; present data) (Fig. 32). sis" above), oniy one possesses the wide transverse The record from Tunisia (Ain-D raham) (Simon, 1885) white/tawny band on the clypeus and lateral bands on remains unrevised, as we have been unable to re-examine the carapace (Figs. 13, IS); both characters correspond Simon's specimen(s); it is even unclear which sex Simon to Lucas' description. studied. As the 0 holotype ot A. basseleli is considered lost. Description: Male (from Algeria, Alger, EI Harrach): to stabilise the taxonomic status of this species we Carapace 2.R long, 2.0 wide, 1.5 high at PLE. Ocular
628
23
,1<2) I ( .' _':. : . ( ~~.,,~~ 30 Figs. 17- 3 I: Aelurillus basse/eli (Lucas. 1846) 17, 23 Male. body colour pattern, 18, 24 Male palp, ve ntral view: 19, 25 Ditto, retrolateral view: 20,26 Embolic dIvIsIon, dorsal view; 21, 27 Ditto, relrolatnul vIew ; 22, 28 D itto, ventral vIew; 29, 31 Epigyne, ventral view; 30 Spermathecae, dol' al vI ew. Specimens (17- 22) Algeria. Tlemccn. Zarlftte. (23- 31) Algeria. Blida, Ohell.lI. Scale lines = I mm (1 7, 23), 01 mm (18- 22, 24-31). 236 Ilelufillus species of the W. Mediterranean area 1.05 long, 1.5 wide anteriorly and 1.4 wide posteri spination: 1: Fm d 1-1-3: Tb pr 1-0, v 2-2-2ap; Mt pr I-I , orly. Diameter of AME 0.4. A.bdomen 2.3 long, 1.7 rt 0-1, v 2-2ap. II: Fm d 1-2-4; Tb pr 1-1 , v 1-1-2ap; M t wide. Chelicerallength 0.9. Clypeal height 0.3. Length or pr and rt I-I, v 2-2ap. III: 1m d 1-2-4; Pt pr and rt I; T b leg segments: 1 1.2+0.8+0.8+0.6+0.5; II 1.2+0.8+0.7+ d 1-0-0, pr and rt 1-1-1, v 1-0-2ap; Mt d 1-1-0, pr and rt 0.6+0.5; III 1.7+0.9+0.9+ 1.0+0.8; IV 1.6+0.9+ 1.0+ 1-0-2, v 1-1-2ap. IV: Fm d 1-1-2; Pt pr and rt 1; Tb d 1.2+0.75. Leg spination: I: Fm d 1-1-4; Pt pr and rt 1; 1-0-0, pr and rt I-I-I, v l-0-2ap; M t d 1-1-0, pr 1-1-2, rt Tb d 1-0-0, pr I-I-I, v 2-2-2ap; Mt pr and rt I-I, v 2-2ap. 1-0-2, v 1-1-2ap. Coloration: carapace dark brown, II: Fm d 1-2-5; Pt pr and rt I; Tb d 1-0-0, pr I-I-I, v covered with white and yellowish scales, sometimes 1-1-2ap; Mt pr and rt I-I, \ 2-2ap. III: Fm d 1-3-5; Pt pr arranged in a poorly marked reticulate pattern. Clypeus and rt I; T b d 1-0-0, pr and rt 1-1-1-1, v 1-0-2ap; Mt d and "cheeks" brown, covered with white hairs. ".Cilia" 1-1-0, pr and rt 1-0-2, v 2-2ap. IV: Fm d 1-1-5; Pt pr and white or brown. Chelicerae dark brown. Abdomen rt I; Tb d 1-0-0, pr and rt 1-1-1-1, v 2-2ap; Mt d 1-1-0, yellow-grey, with brown dorsum covered with brown pr 1-1-2, rt 1-0-2, v 1-1-2ap. Coloration: carapace dark and white scales forming reticulated pattern. Book-lung brown, with black eye field and dorsally with two white covers yellow-grey. Spinnerets brown-grey. Legs brown longitudinal stripes reaching approximately halfway ish, with dark brown patches and semi-annulations. across eye field (Figs. 17, 23), and with white or more Epigyne and spermathecae as in Figs. 29- 31. often tawny la teral borders. Clypeus and "cheeks" Other maleria/ examined: ALG ERIA: A/ger / =£/ Djazai"r j: 20 densely white/tawny haired (F igs. 13, 15). Chelicerae (PC RB), Les Eucalyptus, c. 36 ° 39' N , 3°09' E, 35 m a.s.l. , on ground, brown, covered wi th short brown hairs. Abdomen grey 17 .\pril 1989-6 April 1990 (RB); 10 12 (PCRB) , EI Harrach, yellow, but dorsum brown, with white longitudinal c. 36"43'l\!, 3'O:'E, 9 \1ay· 22 June 1983 (RB); 50 (PC RB), sa me locality, 25 m a.o.1., pitfall in open grassland with Ox alis pes-cap rae, median band. Book-lung covers grey-yellow. Spinnerets 30 January 198)-8 \1arch 1986 (RB); 10 12 (PCRB), sa me loca li ty, grey-brown. Legs yellow. Palps yellow, covered with 27 April 30 Jun~ 1988 (RB). Bechar: 10 (PC RB), Taghit, c. 30 0 54' N , wh ite hairs. Palpal femora without ventral knob. cov 2'01 'W, 630 m a.s.l.. palm grove, 3 April 1989 (RB). Blida [ =£/ ered dorsally with brown/black hairs and prolaterally BOll/aida} ncO 39 (PCRB). Atlas Bl ideen, Djebel Ferroukha , with white hairs (Figs. 14, 16). Palpal structure as in (' 36°29'i\, Y5S' E. Uhellai. 1:150 m a.s. 1., cedar plantations, 19 Ap ril 1987 (RB). BOlldouaou: Ici (PCRB), Bo'u- Merdes [= Boumerdas], Figs. 18- 28 . 7~mmouri, 10 m a.s.l., dunes (between lo w herbs a long a pool), Female (from Alge ria, Blida, Ghella'i): Carapace. 3.4 36'47'N, J"35'1, 27 April 1984 (R B); 10 (PCR B), same locality, lo ng, 2. 6 wide, 1.6 high at PLE. Ocular area 1.4 long, maquis of CiS/liS so/vi/olills (pitfall traps), 12 April 1985 (R B); 12 1. 85 wide anteriorly and 1.75 wide posteriorly. Diameter (PC RB). BOll-:vterde'. [= Boumerdas], Regha'ia , c. 36 °29 ' N, 2 0 55 ' E, 4 of AME 0.5. Abdomen 4.0 long, 2.9 wide. Cheliceral April 1988 (RB), Skikda: 1',' (PC RB), W of Collo, T amanart, c. 3rOO' N , 6°33'1::, 25 III a. s.l., three pitfall tra ps in Q. suber le ngth 1.0. Clypeal height 0.3. Length of leg segmems: I forest, 3- 6 J une 1987 (RB). Tlemcen: 20 (PCRB), Col de Zarifete, 1. 5+ 1.1 +0.9+0.6+0.5; II 1.5+ 1.0+ 1.0+0.6+0.5: III 1150 m a.s.l. , young Q dex fo rest (pitfall traps), 6 May 1984 2. 2+ 1.2 + 1.2+ 1. 4 +0.8; IV 2.0+ 1.2+ :.3+ 1.4+0.9. Leg (RB)
40°
'. ' . 32 I, 10° 20° r r.g. 32: Oistriblllion of '!e/llrillus hasse/eli (l.ucas, 1046) (circles) a nd A. hinipes Deni s, 1960 (squa res). One dot may represent more tha n one close locality. Galina N. Azarkina & D. V. Loguno. 237
331!! 4~
Figs. 33--45: A,dllrilllls liirlipes Denis, 1960. 33 Ma le. body colour pa ttern ; 34, 40 Male palp, ventral view; 35, 41 Di tto, retrola tera l view; 3 6 , 4 2 Embolic division, dorsal view; 37 Uitto, retro latera l view; 38 Ditto, prola teral view; 43 Ditto, ve ntral view; 39 Leg I, pro la tera l view; 44 Epigyne, ventral view; 45 Sperma thecae, dorsa l view. Specimens: (33- 38, 44, 45) Morocco, l:r R achida; (39--43) Egypt, Sinai. Va tiya Pass, paratypc of /l eiliriflus sinaieus ProszY li ski , 2000. Scale lines = I mm (33 ), 0 I mm (~ 4 45).
Aellll'illus hil'tipes Denis, 1960 (Figs. 32 45) CommcnlS: We have he<.:n unable to obtain and Aelurilllls liirripes D enis, 1960: 163 , fi gs. 2 3 (Do: 0 holotype in re-examine the 0' holotype of A. hirtipes from Chad M N H N; not examined). (Qued ue Djanet; Denis, 1960). Instead, we studied a Aellirilllls hirlipes: Proszyri ski, 1971 : 375; Brignoli. 19X3: fl30: male from Tibesti (NW Chad), which was collected Proszynski, 2003b; Platnick, 2004. during the same expedition to the Sahara (known now Aelurillus sinaieLis Proszyri ski, 2000 (p ro parte; the para type ft'om Egypt): 236, figs . 16- 19. as "the missions Berliet-Tenere" ) where the holotype of Aellirillus sinuiells (nee ProszYli ski , 2000, mi sidentified): Proszynski, A. hirlipe.\' was collected; this male was also examined by 2003a: 35- 36, figs. 19, 25 , 31. r. Denis and is kept in thi: collection of MRAC (N o. Diagnosis: This species is similar to A. plumipes 119358). Thus, there are no doubts that we correctly (Figs. 99- 108), but males difl~r in the presence of a dense recognised this species. yellow hair cover of the carapace (Fig. 33), legs I and II Aeluri/lus sinaicus was described by ProszYllski (2000) densely covered with yellow hairs, and in the absence from two males, the 0' holotype from Israel (Ein Duyuk) of a pronounced tegular knob (cf. Figs. 34, 40 and 100) and a 0' paratype from Egypt (Sinai); the description and and of a colour pattern on the dorsum (cf. Figs. 33 and illustrations were made from the paratype. Having 99). F emales of A. hir/ipes have a clearly different, fe-examined both specimens, we come to the conclusion S-shaped form of the epigynal flaps (cf. Figs. 44 and L06) that these males a re not conspecific. T he 0' holotype and shorter ducts of the spermathecae (cf. figs. 45 belongs to A. conveniens (0. Pickard-Cambridge, 1872) and 107). and has to be synonymised with it (syn. n.); the matter 238 A elllrilllls species of the W. M editerranean
wi ll be disc ussed further by one of us (G.A.) elsewhere. c 36 ° 12' N, 4 °23' E, 13 May 1988 (RB); I'? (PC RB), Ill izi. c. 26 °29' N. The 0' paratype of A. sinaicus (Figs. 40 43) should 8 °28 ' E, Guelta Ba llo uaki, I June 1978 (1. Mertens); 10 (PCRB), Tamanrasset, Hirafok, c. 23 ° 39 ' N, 5°45' E, 2 Februa ry 2003 (K. De beyond doubt be assigned to A. hirtipes. Smet); 10 (PCJM, 18734), Qued Djera t, Elieh, date') (1. Murphy). Distribution: Algeria, Moroccu. W Chad and Egypt EGYPT: 10 (H U1, 15202; paratype of Aelurillus sinaicus, Figs. 39-43), (Fig. 32); in Egypt, the species was hitherto reported Si nai [= Shibh Jazi rat Sina'] Governorate, Vatiya [=Watia] Pass, under the name A. sinaicus (Proszynski, 2000). 28 °41 ' N, 33 ° 59' E, c. 11 26 m a. s. l., 18 July 1968 (A. Shulov). Description: Male (from Algeria, road Illizi- El Bhaktou): Carapace 3.b long, 2.5 wide, 2.0 high at PLE. AelllrillllS lopadllsae CantareIla, 1983 (F ig. 82) Ocular area 1. 5 long, 1.9 wide anteriorly and 1.8 wide posteriorly. D iameter of AME 0.6. Abdomen 3.0 long, For a complete reference li st see Pl atnick (2004). 2.3 wide. Cheliceral length 1.1. Clypeal height 0.4. Types: Holotype 0' and paratype ajJinis: Simon, 1864: 3 16; 1868: 7 1; 187 1: 154 (0 ). 2-2ap. III: F m d 1-2-4; Pt pr and rt I; Tb d 1-0-0, pr and Iclidups a[finis: Karsch, 188 1: 5. rt I-I-I , v 1-0-2ap; Mt d 1-1-0, pr and rt 1-0-2, v 1- 1-2ap Aelurillus ajJinis: Pl anet, 1905: 269; Stra nd, 1906: 98; 1909: 5; G alia no, or 2-2ap. IV: Fm d I-I-I; Pt pr and rt I; Tb d 1-0-0, pI' 19 10: 349, 409; Reim oser, 19 19: III ; Simon. 1937: 1228, and rt I-I-I , v 1-0-2ap; Mt d 1-1 -0, pr 1- 1-2, rt 1-0-2, v 1268 (o'?); Bodenhei mer, 1937: 239; Denis, 1937: 1056 ('?); 1-1-2ap. Coloration: carapace brown, with dark brown Caporiacco, 1948: 16 1; Bo nnet, 1955: 165; Wunderlich, 1995: fig. 42 (0 ); Metzner, 1999: 73, 192- 193, figs. 38a- j, 43j, map 40 eye field , covered with white scales, with no colour (o,?); Ali cata & Cantarella, 2000: 486; Cardoso, 2000: 26; pattern. Clypeus yellow, " cheeks" and chelicerae brown. Logunov & Chatzaki, 2003: 95; ProszYJi sk i, 2003b; Pl a tnick. Clypeus covered with white hairs. "Cilia" around 2004. anterior eyes white. Abdomen, book-lung covers and Aelurillus basse/eli (/le e Lucas, 1846: mi sidenti fied): ProszYliski, 200 3'1: spinnerets yellow, but dorsum brown and covered with fi gs. 92- 93 (0). AI/us monardi (n ee Lucas, 1846, misidentified ): Simon, 1868: 66, pI. 5, white hai rs . All legs and palpi yellow, covered wi th white fig. 17 (o'?). hairs. Epigyne and spermathecae as in Figs. 44-45. Aelurillus monardi (nee Lucas, 1846; misid entified): Cardoso, 2000: 26 . Malerial examilled: CHA D: 10 (MRAC, 11 9358; del. J. Denis), A ltus ogieri Simon, 1868: 534 (Do ; 0 holotype in MNHN; ex amined). Trou au Natron, Mission Beige a u Tibesti [NW part of Tibesti region New sy nonymy. in NW ti p of C had], 10 June 196 1 (coll.?). MOROCCO: 10 I'? Aelurops agieri: Si mon, 1876: 139. (MRAC, 159548), Ksar es Souk [= Er Rachida, = Er Rachidia], YllenllS ugieri : Pa v..;s i, 1878: 396. c. 310 55'N, 4 ° 26 ' W, Stuuwmeer, 28 July 197 1 (R. Jocque). Aelilrillus M.ieri: Si mon, 1884: 315; Gali a no, 19 10: 375 , 409; Reim ose r, A LGERIA: 10 (PCRB), road Illi zi- EI Bh aktou, 30 May 1978 (1 . 19 19: 11 2; Bristowe, 1935: 775; D rensky. 1936: 189; ProszYJlski, Mertens); 10 (PCRB), Wilaya Bordj Bo u Arreridj, Portes de Feu, 2003 b; Platnick. 2004. Galina N. Azarkina & D. V. Logunov 239
Aelurillw' pallidell1{1c ulalus Denis, 193 7: 1056, pI. 5, fig . 12 (02; 2 Comments: Salticus luctuosus was described from a holotype in MNHN; not examined). New synonymy. single male from Las Tonga (envi rons of El Kala; Types: Neotype 0 of Ae!urillus lUCluosus (MNHN; Alge ria) (Lucas, 1846). The 0 holotype of A . lu ctuosus designated here), Algeria, Wilaya Blida, Atlas de Blida is considered lost. Matching of the newly collected [=Atlas Blideen], Djebel Mouzala, [c . 36°28' N, material to the original description of A. luctuosus is not 2°40' E], 1450 m a. s. l. , li tter of Q. faginea forest, 22 May easy. However, of the three closely related Aeluril1us 1985 (RB). Holotype 0 of A ttus ogieri (MNHN, 790), species collected by R. Bosmans in Algeria (see above [Spain] , 'Sierra Nevada 651'. " Diagnosis"), this is the only one having a unicolorous Diagnosis: This species is most similar to A. basseleli clypeus cove red wi th black or brownish hairs/bristles and A. monarch, but males can easily be separated by the and having the lateral margins of the carapace with a uniformly black or dark brown c1ypeus (and the entire thin edging of white hairs (cf. Lucas, 1846: 139; " ... il "face"), which is densely covered with black/brownis h present, a sa partie anterieure, quelques poi Is noirs hai rs (Figs. 9, 11) [a wide transverse clypeal band of assez allonges, et, de plus, ses parties laterales sont white/yellowish hai rs is present in A . basseleti (Figs. 13 , entourees de poils blancs, lesquels torment une bordure 15) and there are three narrow vertical lines on the tres-fine."). F urthermore, Lucas clearly stated that "Les "face" in A. monardi (Figs. I, 3, 5, 7)] . Also, the male palpes sont courts, noirs, avec la partie superieure des pal ps of A. luctuosus have well-pronounced, contrasting troisieme, quatrieme et cinquieme articles d'un blanc yellow palpal patellae and tibiae as compared wi th A. jaunatre" . In other wo rds, palps have contrastingly monardi, which possesses much less contrasting yelJ ow coloured segments. As we found, the male palps in brown patellae and tibiae as against brown femora, A . luctuosus have cymbium, bulbus and femur brownish cymbium and bulbus. Females of these three species are (Figs. 10, 12), whereas the patella and tibia are contrast practically indistinguishable (cf. Figs. 73- 81 , 29- 31 and ingly yell ow and covered wi th white hairs (see under Cantarella, 1982: fig. 19). "Description" below). Thus , it is safe to conclude that
Figs. 46--60: Aeluri/lus lucluosus (Lucas, 1846). 46-50 Embolic division, retrolateral vi ew; 51-55 Ditto, dorsal view; 56-59 Male. body colour pattern; 60 Female, body colo ur pattern. Specimens: (46. 51, 56) Al geri a, Meurdja, A tl as Blid een; (47, 52, 57) Algeria, Chleff, foret de Tacheta; (4 8, 53 , 58) F rance, Corsica, Propriano ; (49, 54) Spain , "Sierra Nevada", holotype of Aelurillus ogicri Simon. 1868; (50, 55, 59) Portugal. Mertola; (60) Spain. Malaga. Scale lin es= O.1 mm (46- 55), I mm (56- 60). 24U Aelllrilllls species of the W. Mediterranean
we have correctly identified specimens of A. luctuosus. and Yugoslavia, which he thought belonged to A. affinis. Finally, A. luctuosus was described by Lucas (1846) on However, beyond any doubt all his illustrations should page 139, i.e. its description precedes those of the other be assigned either to A. v-insignilus (Clerck) or to a Aelurillus species described in the same work. Therefore, related species. Fet (1985), Logunov (1993), M ikhailov if any of the subsequent descriptions, especially those & Fet (1994) and Mikhailov (1997) reported on the made from single females, belong to the same species, occurrence of A. affinis in Central Asia. Having re the name A. luctuosus will stand as the valid one for the examined their material, we have concluded that all species anyway. these records belong to a new species closely rel ated to As the 0 holotype of A. luctuosus is inaccessible and A. v-insignitus; its description will be provided elsewhere almost surely lost, we designate a 0 neotype collected by one of us (G.A.). Thus, all the above references are from Blida (Djebel Mouzai'a), a locality close to the omitted from the synonymy list of A. /uctuosus. More original one (Las Tonga). over, as we nov. know, females in this species group (i.e. We re-examined the 0 of Aelurillus ogieri (Figs. 49, A. /uctuOSlIS, A. basse/eli and A. monardi) cannot be 54) studied by Simon (I 868), which seems to be the properly separated and it remains unclear how most of holotype. Although the specimen is old, it is obvious the authors who used the name A. ciffinis could identify that it 'las no colour pattern either on the carapace or on this species. Aelurillus affinis was described by Lucas its "face". The structure of the embolus of A. ogieri is (1846) from females taken from three localities: Bejaia, identical to that of A. luctuosus (cf. Figs. 46-48, 51 - 53). Constantine and El Kala, of which one (EI Kala) The 0 holotype came from Spain, where the latter corresponds to the locality of A. /u ctuosus described species is quite common. Thus, we have come to from a male. Reasoning from the above discussion, we the conclusion that the name A. ogieri should be are of the opinion that A. affinis is a junior synonym of synonymised with A. lUClu osuS. A. /u eluosus. The name Ae/urillus afjinis has beel! used to date by The species A('/urillus pallidemaculatus D enis, 1937 many authors (see above), but none of them seems to was described from females from Algeria (Djebel Daya); have examined properly identified specimens (or at a holotype was not designated. U nfortunately, we have least those taken from the Lype local ity). For instance, been unable to obtain and re-examine the « syntypes, of the contemporary authors, Pr6szynski (1976: but in spite of this it is certain that this specific name is figs . 323- 325) first iIIustrated females from Poland, Italy not valid. It is obvious that Denis (1937) hardly had a
~69 -
/\ I
Figs. 61 - 72: Aeluriflus lucluosus (Lucas, 1846).61,63,65,67 Male palp, ve ntral view; 62, 64, 66, 68 Ditto, rett'O laleral view; 69-72 Embolic division, ventral view. Specimens: (6 1, 62, 69) Algeria, Meu rdja. Atlas Blideen; (63, 64 70) Algeria, ChleA'. foret de Tacheta; (65, 66, 7 1) Portugal. Mertola; (67, 68. 72) France. Corsica, Propria no. Scale lines =O.1 mm.
------~------Galina N. Azarkina & D. V. Logunov 241 clear idea how to distinguish Aelurillus monardi, A. In the original description of Aelurillus ogieri, besides a!finis and A. gesticulator at the time when he described Spain, Simon (1868) reported this species also from A. pallidemaculatus. All these four species are mentioned Tino::; Island (Greece). H owever, he did not provide in hi s paper, and all of them were collected from exactly exact data on particular specimens collected from T inos the same locality, Zouagha Forest (Djebel Daya), during and s~ udied by him. Therefore, we re-examined the 0 the period 9-15 May. Denis ( 1937) reported on 6 males specimL:n of A. ogieri described by Simon (1868), which of A. monardi, a single female of both A. affinis and A. we consider its holotype. Pavesi (1878) and Drensky gesticulator, and 4 fe males of A. pallidcmaculatus. We (l 9~6) recorded A. ogieri from G reece reasoning are of the opinion that Denis simply treated males and from Simon' s information alone. Metzner (1999) in his different « variants of the same species under four thorough taxonomic-faunistic revision of the Greek different names. Judging from the illustrated epigynes of Salticidae did not mention this species. Perhaps, both his females (Denis, 1937: pI. V, Egs. i 0, i I. 12), it is Pa vesi and Drensky dealt with A. luctuosus, of which certain that all of them corresrond to ex isting «variants A. ogieri is a junio r synonym (see above). of A. luctuosus (cf. Figs. 80. 75 and 77 respectively) . Description: Male (from A lgeria, Blida, Djebel Therefore, the name A. pallidell1aculatus is here treated Mouzai' a): Carapace 3.4 long, 2.4 wide, 1.6 high at PLE. as a junior synonym of A. luctuosus. Ocular area 1.2 long, 1.6 wide anteriorly and 1. 5 wide Distribution : This is a widespread Mediterranean posteriorly. Diameter of AME 0.5. Abdomen 2.8 long, species (Fig. 82) known from Portugal a nd \1orocco in 2.1 wid e. Chelicera I length 0.9. Clypeal height 0.3. the west (Simon, 1909; Cardoso, 2000; present data), Length of leg segments: I 1.5+ 1.0+0.9+0.7+0.6; II throughout Algeria, T unisia, Italy (Sici ly) (Simon, 1.5+ 1.0+0.9 +0.7+0.6; III 2.0+ 1.1 + 1.0+ 1.3+0.8; IV 1937), G reece and Crete (Metzner, 1999) to Rhodos 1.9 + 1.0 + 1.1 + l.3 + 0.8. Leg spination: I: Fm d 1-1-5; Pt (S imon, 1937; Caporiacco, 1948) in the east. In most pr and rt I; Tb d 1-0-0, pr \-1-1 , rt I-I, v 2-2-2ap; Mt pr cases, the species was recorded under the name Aelurillus and rt \- 1, v 2-2a p. II: Fm d 1-2-5; Pt pr and rt 1; Tb d affinis, which is synonymised herein with A. luctuosus 1-0-0, pr and rt I-I-I , v 1-1-2ap; Mt pr and rt I-I , v (see above). 2-2ap. III: Fm d 1-3-5; Pt pr and rt I; T b d 1-0-0, pr and This species has also been reported from Syria and rt 1-1-\-1. v 1-0-2ap; Mt d 1-1-0, pr and rt 1-0-2, v Israel (Simon, 1937; Bodenheimer, 1937; Metzner, 1999; 1-1-2ap or 2-2ap. IV: Fm d 1-2-5; Pt pr and rt I; T b d all under A. a.!finis). However, in his recent extensive 1-0-0, pr and rt \-1-1-1 , v 2-2ap or 1-2ap; M t d 1-1-0, pr review of the Salticidae of Levant, Proszyt1ski (2003a) 1-1-2, rt 1-0-2, v 1-1-2ap. Coloration: carapace dark rejected these records as being not supported by the brown, alm ost black. with black around eyes and two pertinent specimens. Thus, we have disregarded them as white longi tudinal stripes running from eye fie ld to well and not mapped them. thoracic part (Figs. 56-59); lateral margins with thin
\ "-..C 79 , \
Figs. 73- 8 1 Aefunffw f{{e{l/ow .I (Lucas. 1846) 73, 75, 77, 79, 80 Epigyne; 74, 76, 78, 81 Spermathecae, dor:al vi ew. Specimens: (73 , 74) Algeria, Blida, DJebel Mouza'ia; (75, 76) Algeri a, Chlcff, roret de Tachela: (77. 78) France, Corsica. Propnano; (79) Tunisia, Zagho uan. (80. 8 1) Portuga l, Mertola . Scale hnes= O I mm 242 A elll,.illlls species of the W. Mediterranean edging of white hairs. Eye field covered with aupressed spicuous reticulated pattern. Legs yellow, with brown dark brown scales. C lypeus, chelicerae and "cheeks" patches and semi-annulations. Palps yellow, covered dark brown, densely covered with black or dark brown with white hairs. Epigyne and spermathecae as in hairs and sparse bristles (Figs. 9, II). A bdomen brown Figs. 73- 81. yellow, dorsum dark brown, with white longitudinal medial stripe. Book-lung covers yellowish. Spinnerets lV/alai,,/ ex amined: MOROCCO: Kheniji"a: 2<;? (PCRB), Aguelman brownish. Legs brown-yellow, each femur dorsally with (Lake), 33°02'N, 5°01'W, Azigza, c. SE side, 1575 m a. s.1., mixed blackish longitudinal stripe. Palp: cymbium, bulbus and Cedrus, Q. ilex. Q. ji:lgill(,{{ and Fraxinus forest (among stones and sieving litter), 13 May 1984 (R B) . ALGERIA: 30 (PCRB) (no locality femur brownis h, whereas palpal patella and tibia con and date data) (RB). Aiil Dejla: 10 (PCRB), Bethia , c. 120 km Wof trastingly yellow a nd covered with white hairs (contrast Alger, 45 km from coast. c. 36 ° 16 'N, 1058'E, 800m as!., among ing yellow colour sometimes not well pronounced). stones in Pinus ha/epensis forest, 6 M ay 1989 (RB); 2<;? (PC RB), Palpal femur lacking a ventral knob, dorsa lly covered Miliana, Djebel Zaccar [='Ain Torki] , c. 36° 19' N, 2° 13' E, 1000 m with black/brown hairs and prolaterally with yellow a.s. l. , steppe and Q. i/ex forest, 23 April 1989 (RB). A frw [= £/ Dja:;oii"j: 10 (MCZ, 28448; hitherto determined as Aelurops hosse/eli), hairs (Figs. 10, 12). Palpal structure as in Figs. 46- 55 , " Algiers (G. W. & E. G. Peckham coll.)". BO/l1a : 10 I <;? (PCRB) . 61 - 72. M assif de [,Aures, Belezma Mts., Col Telmet, 1820 m a .s.l.. pitfall Female (from Algeria, Blida, Djebel Mouzaia): traps in Cedrus a/lanlica forest, 6 November 1987-11 June 1988 (RB); Carapace 3.3 long, 2.4 wide, 1.4 high at PLE. Ocular I <;? (PC RB), Massif de [,Aures, Monts de Belezma, Col T elmet, 1800 m area 1.3 long, 1.7 wide anteriorly and 1.6 wide pos a .s. l. , pitfall traps in Cedrus forest, S November 1987- 9 April 1988 (RB); 10 (PCRB), Oued Taga, c. 35°24' N, 6° 22 'E, 1200 m a .s.l., teriorly. D iameter of AM E 0.5. Abdomen 4.0 long, degraded Q. ilex forest, II February 1988 (RB). Blida [ =£1 Boulaida]: 3.3 wide. C heliceraI length 0.9. C lypeal height 0.3. I <;? (PCRB), Atlas Blidee n, Djebel Mouzai"a (together with the neo Length of leg segments: I 1.4+0.8+0.8+0.n+05; II type), c. 36° 28' N , 2 °40'£, 22 May 1985 (RB); 10 (PCRB). A tl as 1.4+0.8+0.8+0.6+0.5; III 2.0+ 1.0+ 1.0+ 1.2+0.9; IV Blideen, Chrea (50 km SW of Alger) , c. 36 ° 25' E, 2 ° 52' E, Pic 1.9+ 1.0 + 1.2+ 1.3+0.9. Leg spination: 1: Fm d 1-1-4; Tb Abdelkader, 1520 m a .s. l. , pitfa ll traps in Cedrus forest. 27 June 1989 (RB); I <;? (PCR B) , C hrea, c. 36 ° 25 ' £ , 2 ° 52' £ ,1300 m a.s. !. , on rock. pr I-I , v 2-2-2ap; Mt pr and rt I-I , v 2-2ap. II: Fm d 3 March 1987 (RB); 2 <;? (PCRB), Chrea, c. 36 ° 25' E, 2 ° 52'E. 1550m 1-2-4; Tb pr I-I , v 1-1-2ap; Mt pr and rt I-I , v 2-2ap. III: a.s. l. , montane grassla nd, 10 April 1985 (RB); 10 (PCRB), Atlas Fm d 1-2-4; Pt pr and rt 1; Tb d ;-0-0, pr and rL I-I-I, Blideen, Meurdja [= Merdja], c. 36 ° 08 ' , 3° 18' E, 900 m a.s.l. , sweep v 1-0-2ap; Mt d 1-1-0, pr and rt 1-0-2, v 1-1-2ap. IV : Fm ing tall herbs in flooded garden, 30 May 1987 (RB); 10 (PCRB), d 1-1-2; Pt pr and rt I; Tb d 1-0-0, pr and rt 1-1-1 , v Meftah, Djebel Zerouela , 450 m a.s. l. , among stones in mixed Q. suber and Pinus ha/epensis forest, II April 1988 (RB). Cldeff [ = £ ch Chelijjj: 2-2ap; Mt d 1- 1-0, pr and rt 1-1-2, v 1-1-2ap. Coloration: lo I <;? (PCR B), foret de Tacheta , c. 110 km W of Alger, 30 km from carapace brown, with dark brown eye field and mono coast, c. 36 ° 22' N, 1° 36' E, 850 m a.s.I. , in litter of Q. jClginea forest, 29 chromously coloured body (Fig. 60). Clypeus, " cheeks" Sc:ptember 1989 (RB). 5uic/a: 2" (PC RB), c. 40 km SE of Sa'ida, Oum and chelicerae yellow-brown. Clypeus sparsely covered Djcrane, c. 34 °49' E, 0 19' C, 1300 m a s l. , pitfall trap in m aquis of Q. with short white hairs and brownish bristles. Abdomen, ilex, I October 1981 (RB). Tamanrussel: I <;? (PCRB) , Mertoutek [= Ain Mertoutek], c. 24 ° 15' , 5°3 1'E, 1350m a .s. l., 15 February 1990 spinnerets and book-lung covers brownish jl!llow. (RB) T~b ess([ [ = Tbessilf: · 10 I <;? (PCRB). F oret de Bekkaria, Dorsum grey-brown, densely covered with light hairs c. 35°2 1' N, 8° 15' E, 15 June 1989 (R B). Tipas([: 10 (PCRB). Sidi and brownish brist'""s, sometimes (rarely) with Il1con F redj [= Sidi Ferruch], c. 36 °45' N, 2° 50'E, 10 m a .s.!. , sweeping
40°
'. ~ , ,.~. --J-----~~----~~~~./------~--~~~~--~:------~~~~~_;~--1_--__i 30° '. ,.. ,,1 I i i
-" 'Il 82>" _ T - - /'- I .-. - ..- ' 1 - \ 10° 20° Fig. 82: Distribution of Aell/rillus luClu()sUS (Lucas, 1846) (circles) and A. lopaduso ~ Ca ntarclla, 1983 (square). One dot may represent more than one close locality. Galina N. Azarkina & D. V. Loguno\, 243 vegetation a nd among stones in dunes around hotel E I Riadch, c.40036' , 6 °53 ' W, 700 m a.s. l. , stones in burnt fo rest, 9 A pril 1996 20 March 1983 (RB); 19 (PC RB), same loca lit y, 15 April 1987 (R B) (RB). Leiria Dis/I'. 19 (CBA), Va le da Garsia , c.39°44 'N, 8 °45' W , Ti:i OU: OII: 10 (PCRB), Massif du Djurdjura, Ait Ouhalane, pitfall traps, 2 July 2002 (P. Cardoso); 19 (CBA), sa me locality, pitfall c. 36 ° 31 ' N, 4 ° 10 ' E, 1410 m a.s. l. , pitfall traps in open Cedrus forest, tra ps, 23 April 2002 (P. Cardoso). Viseu Dis/I'.: 32 ( BA, 1.641), Sao 6 October 1987- 1 June 1988 (RB); 19 (PC RB), NE 'Ain el Hamll1am, Joao da Pesqueira, 41 °08' N, r24 ' W, 4 May 1943 (F . Magalh iies). Oued Boubehir [= Oued Boubckir], c. 36 °34 'N, 4 ° 18' E, stones F RANCE: Corsica [ = Corse-du-Slld): 10 19 (PCYM, # T 331), along te mpora ry pool, 10 October 1987 (RB); 10 (PCRB), Sebaou-el Propria no, c.41 ° 39' ,8° 53 ' E, 29 April 2000 (Y. Monta rdi). [T ALy . Kedim [= Sebaou el Kdim], c. 36 °47'N, 3° 52 ' E, 50 m a.s.l. , dry Sicilia : 10 (ZMC D), Paterno Fiume Simeto, c.3 r33' N, 14 °54 ' E. steppe, 10 May 1988 (RB); 19 (PC RB), Massif du Djurdjura, Tala 7 May 1968 (S. Lengcmark). G uilef, montane grassland, 2000 m a. s. l. , 2 May 1985 (RB); 19 (PC RB), Ain eI Hammam, c. 36 ° 34 'N, 4 ° 18'E, 1080 m a.s.l. , a mong stones . 2 October 1987 (R B). T U ISlA: 10 19 (ISEA), Zaghouan A e1urilllls m Ollardi (Lucas, 1846) (Figs. 1- 8, 83- 98) [= Zaghwan), c. 36 °23' N, 10 0 08' E, 17 May 1997 (R . DolanskY) . SPAI N : 10 (H ECO , 2160), Andalusia, no date (Dr D. R. Dorylas): 10 Sall/cLlllI1onardi Lucas, 1846: 15 6, pI. 7, fi g. 2 (Do; 0 ho lotype lost). A (SM N H , 1746; hitherto determined as A. 1110r/urc!i), 'Spanien (Simon), -) ncotype is designated here. T . Tho rell coli .'. AllI1eria Prov.: 10 (PC RB), Carboneras , Ilia monol'di: Simon, 1864: 316; 1868: 66. c. 36° 59 ' N, 1° 54 ' W, 10 m, maquis and stones near beach, 7 April .-ielll J'(J/I .I m Ol/lin": Simon, 1876: 139 ; Pavesi, 1895: 10 . 1997 (RB); 19 (PC RB), Cabo de G ata [= EI Cabo de Gata], lclidops /lIO/lordi : Pavesi, 1880: 383. c. 36 °46 ' N, 2 ° 14 ' W, 10 m a.s.l., in salt marsh, 7 April 1997 (R B) . . Ieluril/I/O I11UII({r£i/: Simon, 190 I 664; 1908: 54: Reimose r, 191 9: 11 2: Caceres Prov.: 10 (PCRB) , Zahara de la Sierra, grassland along Denis, 19 37: 1056; Cantarella , 1982: 241 - 243, figs. 5 9, 19 (09): emba lse , 8 April 1993 (R B) . COdi: Pro v.: 19 (PC RB), Ta rifa, Pr6szYl1 Ski , 2003b; Plat nick, 2004. c. 36° 01 ' N, 5° 35' W, April 1990 (P. Poot); 20 19 (PCJM, 3373), Zahara de los A tunes, c. 36 °08' N, 5° 50 ' W, 0 m a.s.l., sa nd dunes, Type: Neotype 0 of A e/uriflus monardi (M NHN; 18 April 1974 (1. Murphy); 10 (PC RS, 1788 ), Algar, Embalse des designated here; Figs. 83, 86, 87, 90, 91), Algeria, Wilaya Hurones, c. 36 ° 39'N, 5° 39' W, 3 April 1985, R. Snazell; 10 (PCRS, Biskra [=Beskra], Tolga [c. 34°43'N, 5°22'E], 125 m (789), Al geciras, Sierra de Luna, c. 36 °08' N, 5°27'W, 31 March 1985, a.s.l., in litter and among Gramineae in palm grove, R. Snaze ll ; 10 (PCRS, 1790). N W Jimena de la Frontera, c. 36 ° 25' N. 10 April 1982 (R B). 5°27 ' W. 7 April 1985, R. Snazell ; 10 (PCRS, 1791), Sierra del Rctin Ta hi villa, 13 April 1985, R. Snazell ; 20 (PCRS, 1795), near Diagnosis: [n the structure of the copulatory organs, Embalse del Charco Red ., 14 April 1985, R . Snazell. Jalm Prol'.: 10 this species is most similar to A. /u ctuosus and A. (PCRS, 1738), Sierra de Cazorla , c. 3r 54 'N, 3°00'W, Cagasebo, basseleli. The males of A. monardi can easily be separ 22 September 1980, R. Snazell; 10 19 (PC RS, 1727, 1737), same ated by the presence of three narrow vertical lines above locality. 9- 12 September 1982. R . Snazell . Malaga Prov.: 40 19 the eyes of the first row (Figs. I, 3, 5, 7) [monochro (PCJM. 143(6). Maro, c. 40 km N o f Malaga, 30 April 1990 (1 . Murphy); 19 (PCJM, 14362), same locality, stones and litter. 100 m mOllsly black or dark brown " face " in A. /uctuosus a.s. l., March- April 1987 (J . Murphy); 10 (PC RS, 1796), c. 7 km S\V of (Figs. 9, II), and a wide white/tawny transverse band on Ca rra traca, c. 36 ° 51' N, 4 °49'W, 18 April 1985, R . Sn azell: 19 the clypeus in A. hasse/ell (Figs. 13 , 15)]. Also, the males (PC RB), Casares, c. 36 °26'N, 5° 16'W, pitfall traps in maquis, 5 May of A. monardi have less contrasting yellow paJpal patel 1996 (1. de Beir). Salamal/ ca Prov.: 10 (PC RS, (797), near La Aberca, lae and tibiae as compared with A. /UCIUOSUS , which c. 40 °28 ' , 6 °06' W, rocky hillside, 6 May 1986, R. Snaze ll . Mallorc;a : 30 12 (PCJM, 131001), Fonnentor, Casas Veyas, 100 m a.s. l. , wooded possesses brightly contrastipg yellow patellae and tibiae area, 12 A pril 1985 (1 . Murphy); 10 (PCJ M, 13114), Sa Meleta, 700 m as against brown femora, cymbium and bulbus. Females a. s.l. , 8 April 1985 (J Murphy); 20 39 (PCJM, 13029, 13221), Cala of these three species are practically indistinguishable Estreta, stony, pine scrub, 20 m a. s. l. , 7 April 1985 (1. Murphy): 10 (cf. Figs. 29- 31, 73- 81 and Cantarella, 1982: fig. 19). (PCJM , 4614), P. Pollensa, grassy, thin wood, 20 m a.s.l., J7 April Commenls: Sa/ficus monarcli was described by Lucas 1990 (J . Murphy); 10 (PCJM , 18504), Cabo de Gata, stones and low plants, Om a.s.l., 30 April 1990 (1. Murphy); 10 (PCJM, 18501), C. (1846) from a single male from Alger (" dans Ie jardin Trafalga r, pin ewoods on sa nd, 200 m a. s.l .. 16 April 1974 (1. Murphy); de J'h6pital du Dey" ; Algeria). The 0 holotype of A. 10 (PCJM, 4549), Ternell es, wooded streamside, 150 m a. s.l., 14 April monarcli is considered lost. However, matching the 1975 (1 . Murphy). lbi:a: 19 (PCJM, 5608), Santa Eulalia de Rio, studied males of A. monardi to Lucas' description and e. 38 ° 58'N, 1 ° 31 ' E, 100 m a.s. l. , 9 September 1976 (1 . Murphy). colour illustration (1846: 156, pI. 7, fig . 2) caused no PORTUGAL: 19 (CBA). no loca lit y data, 16 August 1998 (P Cardoso); 10 (CBA, PMC0338c), no locality data, 2 October 2000 (P. problems, as among other characters Lucas clearly stated Cardoso). Beja Dis/r.: 10 (C BA, PMC0338b), c. 15 km ES E of that " ... et en dessus il presente trois bandes longitudi Merlola, near Minas de Sao Domingos, c. 37 °40'N, r29'W, pitfall nales de cette couleur, dont la mediane tres-courte, les tra ps. 25 September 2000 (P. Cardoso); 19 (C BA , PMC0330b), same autres larges, partant de la partie anterieure et atteignant locality, 31 July 2000 (P. Cardoso); 10 (CBA, PMC0338), sa me presque la base du cephalotorax" . All the males we locality, pitfall traps, 4 September 2000 (P. Cardoso); 19 (C BA , 5576) , same locality, 16 July 2003; 19 (CBA, 5583), Cerro das Antenas, assigned to this species possess the striped carapace (Figs. c. 20 km NW of Minas de Sao Domingos, pitfall traps, 16 July 2003 (P 1, 3, 5, 7), with one short stroke on the eye field and two Cardoso); 19 (CBA, 5389), Mertola (monado). pitfa iJ traps, 29 May long longitudinal stripes across the carapace, all stripes of 2003 (P. Cardoso); 10 (C BA, 5673 ), same locality, pitfall traps, a white or yellowish colour (or even tawny in da rker 17 August 2003 (P. Cardoso); 19 (CBA, PMC0330), Monte Moreanes, specimens). Of the Aefurilfus material collected by R. c. 37 ° 38'N, r34 ' W , pitfa ll traps, 24 July 2000 (P. Cardoso); 10 (CBA, PMC0338F), Ribeira das Limas [= Limas Rive r) , in ' Parque Bosmans in Algeria, there are three closely related species Natural do Va le do G uardiana', pitfall traps, 16 October 2000 (P. (see " Diagnosis" above), of which only one possesses the Cardoso); 10 (CBA, PMC0338 D), same loca lity, pitfall traps, striped carapace and "face" in males which precisely 9 October 2000 cP. Cardoso). Faro Dis/r.: 10 (C BA, 1. 561), near corresponds to Lucas' description. Moncarapacho. Olhiio, c.3rOI ' N , 7°51 ' W, 3 April 1942, colP; 20 As the 0 holotype of A monardi is lost, to stabilise 19 (PCJM, 24, 40, 243). Monte Gordo, c.3r 10 ' N, r 27' W, marsh, pine wood and garrigue, 0- 10 m a .s .l. , 13-14 August 197 I (1. Murphy); the taxonomic status of this species we designate a 0 10 (PCJ M, 103(0), . St. Vincent , 50 m a .s. l., litter, 14 April 1982 neotype collected from Tolga, not a great distance from (J. Murphy). Guarda Disll': 19 (PC RB), Castelo Bom, Rio Coa, the original locality (Alger). 244 . Idllrilills species or the W. Mediterranean
\1ost of thc subscquent records of A. monarc/i made 67) clearly stated that "des yeux de la quatrieme paire au by va rious aut:lO~s after Lucas' description may be pedicule s'etendent deux larges bandes de poils blancs un doubtfuL Simon (1868) and Cantarella (1982) reported peu dores", but he did not mention a whiteltawny band on A. monardi i'rom Hiskra; both examineo males and on the carapace and/or a striped eye field (both are apparently matched them correctly to the original unmissable characters). Instead, he further menLioned a description. Denis' (1937) record from Djebcl Daya thin white margin on the caiapacc (d'um: fine ligne is disregarded (see above under "Comments" on A. blanche). Thus, we haw concluded that when stUdying pallidemaculatus). The rest of the known records of A. the specimens from Spain and Italy SimoD was actually l110nardi should also be disregarded. dealing with A. luctuosus (cL Figs. 58, 59), which is The name Salticus nicoletii Lucas, '846 was syno common there (see above). nymised with A. tnonardi by Simon (1868; see also Although A. monardi has also been recorded from Platnick, 2004 and previous catalogues by Bonnet and Israel, Syria and Egypt (Cairo) (Pickard-Cambridge. Roewer). However, we are of the opinion that S. nicoletii 1872: sub Saltieus monardi; 1876: sub Altus monardi: may be con specific with Salticus nwnidicus and appar Pavesi, 1895; Bodenheimer, 1937), all these record ~ ently does not belong in Aelurillus (see below). There are disregarded herc. We have re-examined Pickard fore, the name S nicoletii is removed from synonymy Cambridge's specimens of Altus/Saltieus monarrli kept in with A. ll1onardi. the HECO and found that the records from brad were Distribution: Algeria and Tunisia (Fig. 98). based on immaturcs [I subadult 0 (IIECO, botlic 1833, Simon (1868: sub Altus tn.) reported A. monarch from tube 13) , 'Allus monardi Luc.lPalestine'; 5 juveniles Spain (Andalucia) and Italy (Sicily), but we have disre (HECO, bottle 1837, tube 8), 'Saltieus monarch (Luc.)/ garded these records. In his description, Simon (1868: Palestine/)3 Jericho (Luc.)'] and cannot be confirmed.
Figs. 83- 97: Aelurillus monaI'd; (Lucas, 1846). 83-85 Male body colour pattern; 86, 88 Male palp, ventral view; 87, 89 Ditto, retrolateral view; 90,92,94,96 Embolic division, dorsal view: 91, 93, 95, 97 Ditto, retrolateral view. Specimens: (1)3 , 86, 87, 90. 91) Algeria, Biskra, Tolga, neo type of Ae/uri//us Illol/ordl; (84, 96, 9~) Tunisia, Hammamet; (85, 94, 95) Algeria, Setif, Djcbcl Babor; (88, 89, 92, 93) Algeria, EI Kala, Ca p Rosa. Sca le linc :; = 1 mm (83 -85), 0.1 mm (8&-97) . (;alina 1'1;. Azarkina & D. V. Loguno" 245
The record from Egypt was based on a single male [! d' Pal pal femur brown, without ventral knob, dorsally (HECO, hottk 1832, tub~ 113), 'At/us monarch Luc./ covered with brown/black hairs, but prolaterally with Egypt'], which should in fact be assigned eithn to bunch of white hairs (l"igs. 2. 4, 6, 8). Palpal patella and Aeiuril/lfs kochi Roewer, 1951 or to A. bokerinus tibia yellow, coven~d with whik hairs. Cymbium brown. Proszynski. 2003. The taxonomic status of the latter Pal pal structure as in Figs. 86-\)7. sp~cies is unfortunately obscure, as it may be a junior Female: Absent from the material available to us. For synonym of the former. The diagnosis provided by illustrations of the epigyne see Cantarella (1982). Proszynski (2003a) is inadequate to resolve this problem Malerial examined: ALGERIA: IJ Tall 3a (PCRB), El Ka la, Cap now: the matter requires the re-examination of the Rosa, c. 30 0 57'N, 8° 14 ' E, 50m a.s.l., maquis on dunes, 29 Ma rch 1988 (RB). Selij. 10 (PCRB), Ojebel Babor [=Babor Mt. Range, perLin~nt materia!. No specimens of A. monardi from =Massifdes Babol's], c. 36 °32'N, 5°25' E, 1920m a.s.l., liller in Q. Syria were found in Pickard-Cambridge's collection kept faginea forest, 15 June 1987 (RB). TUNISIA: 2a (lSEA), Hammamet, in the HECO. c. 36 °23 ' N, loo38'E, 8- 19 May 1997 (J, Oolansky) Description: Male (Tolga, Algeria): Carapace 2.7 long, 1.9 wide, 1.4 high at PLE. Ocular area 1.1 long, 1.5 wide Aeilirilllls pilimipes (Thorell, 1875) (Figs. 98- 108) anteriorly and 1.45 wide posteriorly. Diameter of AME 0.4. Abdomen 2.4 long, 1.7 wide. Chelicera I length n.8. Yllenus plwl/ipes Thorell, 1875: 195 (0,,'; cc' holo[JP ~ in 7 i\1Cf>: Clypeal height 0.3. Length of leg segments: I ex.amined). Aelurops plul1lipes: Simon, 1870: 139. 1.1 +0.85+0.8+0.6+0.6; II 1.1 +0.8+0.7+0.65+0.6; 1II Aelurillus plumipes: Simon, 190 I: 665; R eim()~.~r, 1919: 112; Roewer, ;.7+0.9 +0.9+ 1.0+0.7; IV 1.6+ 1.0+ 1.1 + 1.2+0.7. Leg 1955: 1116; Bonnet, 1955: 170; Proszyriski. 19~6: fi gs. 319- 322 spination: 1: I,'m d 1-1-5; Pt pr and rt I; Tb d 1-0-0, pr (ci); 2003b; Platnick, 2004. 1-1-1 , rt I, v 2-2-2ap; Mt pr and rt I-I , v 2-2ap. II : Fm Yllel/us cerl'inu< Thorell, 1875: 196 (002; 2 syntype in SMN I-I; d 1-2-5: Pt pr and rt :; Tb d 1-0-0, pr 1-1-1, rt 1-1-0, v examined). "'lew synonymy. Aclurops cal'inus: Simon, 1876: 139. 1-1-2ap; Ml pr and rt I-I, v 2-2ap. Ill: Fm d 1-3-5; Pt pr Aelurillll.l ('nil/liS: Reimo,.cr, 1919: 111, Roewer, 1955: 111 3; Bonnet, and r'~ 1; Tb d 1-0-0. pr and rt 1-1-1-1, v I-0-2ap; Mt d 1955: 166 ; Proszynski, 1976: fi gs. 313-318 (02); 2003b; i-I-O, pr and rt 1-0-2, v 1-1-2ap. IV: Fm d 1-2-5; Pt pr Cantarella, 1983: 55 , fig :.. 10-12, 17 (02); Platnick, 2004. LLnd r( 1: Tb d 1-0-0, pr and rt 1-1-1-1, v 1-0-2ap; Mt d Aelurillus mayeli Simon, 1885 3, 4 (Do; a holotype in MNH N; 1-1-0, pr :-1-2. rt 1-0-2, v 1-I-2ap. Coloration: carapace examined). New synonymy. Ae/urillus mayeli: Simon, 1901 665; Reimose r, 1919: 112 ; Roewer, dark b"own, with black eye field and dorsally with two 1955: 1115; Bonnet, 1955: 169; Proszy nski, 2003a: 36, fi gs. \\,hitt:-ydlowisn longitudinal stripes running posteriorly 91 - 93 (02): 2003b; Platnick, 2004. from ALE (Figs. 83 85). Eye field with short median white-yellowish Jine running from AME to middle of eye Types: Holotype 0 of Yllenus plumipes (ZMCD, 185), field. Clypeus and "cheeks" brown to brown-yellow, !\lgeria, Bona [=Annaba], [c. 36 ° 54' N , 7°45' EJ, date sparsely covered with white hairs (Figs. 1, 3, 5, 7). unknown (Meinert). Syntype 9 of Yllenus cervinus "Cilia" around anterior eyes white, but brown above (SMNH), Algeria, Bi skra [=Beskra], [c. 34°51'N, them. Lateral margins of carapace covered with white 5 ° 43' EJ, date unknown (Meinert). Hulotype 0 of hairs. Abdomen gr~y-ycllow, with dark brown dorsum /lelurillus mayeti (MNHN; Figs. 99- 105), Tunisia, ' Iles and a white longitudinal median stripe. Book-lung de Kerkena' [=Kerkenah Island, =]uzur Qarqanah, covc:rs brownish yellow. Spinnerets brown. Legs yellow, =Jazlrat ash Sharql, c. 34°44'N, Il o 13 ' EJ, date all femora dorsally with a black longitudinal strip~. unknown (V. Mayet).
Fi g> . 98: Oistrihution of Aelurillus mUl1arrii (lucas, 1846) (circles) and A. plw11ljJes (Thorell, 1875) (squares). One dot may represent more than one clo:;e locality, 246 A elUl'illus species of the W. Mediterranean
Diagnosis: This species is similar to A. hirlipes ody. Diameter of AME 0.50. Abdomen 3.2 long, 2.6 (F igs. 33-41), but males differ in having femora I-II and wide. Chelicerallength 1.0. Clypeal height 0.3 . Length of the anterio r part of the carapace densely covered wi th leg segments: I 1.7+ 1.2+ 1.1 +0.9+0.7; II 1.8+0.9+ dark brown hairs, and in the presence of ~ pronounced 0.7+10+0.85; 1lI 2.3+ 1.3 +1.3+1.7+0.9; IV 2.2+1.2+ tegular knob (cf. Figs. 100 and 34, 40) and of a colour 1.4+ 1. 85+0.85. Leg spination: I : Fm d 0-1-1-4; Pt pr pattern on the dorsum (cf. Figs. 99 and 33). Females of and rt I ; Tb d 1-0-0, pr 1-1-1, rt 1-1-0, v 1-1-2ap; M t pr A. plumipes have C-shaped epigynal flaps (cf. Figs. 106 1-1 , rt 0-1, v 2-2ap. II: Fm d 0-1-2-5; Pt pr and rt I ; T b and 44) and longer and more winding ducts of the d 1-0-0, pr and rt 1-1-1 , v 1-2-2ap; Mt pr and rt I-lap, v spermathecae (cf. Figs. 107 and 45). 2-2ap. III: Fm d 0-1-3-5; Pt pr and rt I ; T b d 1-0-0, pr Comments: We have re-examined the type specimens and rt 1- 1-1-1 , v 2-0-2ap; Mt d 1-1-0, pr and rt 1-0-2, v of Yllenus plumipes (0), Aelurillus mayeti (0'; Figs. 99 1-1-2ap. IV: Fm d 1-2-4; Pt pr and rt I; T b d 1-0-0, pr 105) and Yllenus cervinus (2) and compared them to the and rt 1-1-1-1 , v 2-0-2ap; Mt d 1-1-0, pr 1-1-2, rt 1-0-2, samples containing both sexes (see below). As a ll the v l-1-2ap. Coloration: carapace dark brown, with studied specimens have identical copulatory organs almost black eye field, sparsely covered with white hairs. and match each other perfectly, the names A. mayeti and Carapace margins covered with white hairs, but on both Y. cervinus should be synonymised with A. plumipes . sides of eye field densely covered with long brown/black Although Y. plumipes was described in the same work as hairs. Eye field with white patches, sometimes poorly Y. cervinus, its description precedes that of the latter visible, and anteriorly with long black bristles . Eyes of species and hence the name Aelurillus p/ul11ipes stands as first row bordered by white hairs and scales. Clypeus the valid name for the species. brownish yellow, densely covered with white hairs. Distribution: Algeria and T unisia (Fig. 98). Chelicerae yellow-brown. Abdomen grey-yellow, Description: Male (holotype of AelurilLus l71ayeti): dorsum dark brown, with pa ttern of white patches and Carapace 3.8 long, 2.85 wide, 1.7 high a t PLE. Ocular densely covered with long brownish and white hairs area 1A long, 1.8 wide anteriorly and 1. 7 wide posteri- (Fig. 99). Legs I-[J brown, legs III-IV yellow-brownish.
100 1~ 104 1088B .~ 102
Figs. 99- 108 Aelllltflu.1 plul11/ jJi'\ (Thorell , 1875) 99 Male. body pattern; 100 Ma le palp, ventral view; 101 Ditto, retrolaleral view; 102 Emboltc di VISIo n, pro lateral View, 103 DillO. dorsa l view. 104 Dltlo, ventral view; 105 Ditto, rellolateral view; 106 Epigyne. ventra l Vie w, 107 Spermathecae, dorsal view; 108 Diagramma tic course of the inse mination ducts. Specimens: (99- 105) Tunisia, Kerkenah Island, holo lype of A. 111O \'(' l i Simon. 1885; ( 106-108) A igend, Blskra Sca le 11l1 es= I mill (99), 0.1 mm (100- 107) Galina N. Azarkina & D. V. Loguno\' 247
Femora I-II prolatera ll y densely covered with long Aelul'illus nicoleti (Lucas, 1846) dark brown hairs, retrolatera ll y with dense white hairs. Sa/lieus Ilico/etii Lucas, 1846: 160, pI. 7, fi g. 5 (lY?; 2 holotype lost; not Patellae, tibiae and metatarsi I-II pro- and retrolaterally examined). Synonymi sed wit h A. IIwl llmli by Si mon (1868: sub densel y covered w ith long dark brown hairs. Palps A IIIIS 11.) . brownish yellow, covered wi th long white hairs. Palpal For a complete reference list see Pla tnick (2004) . femur without ve ntral knob. Pal pal structure as in Comments: Salficus nicoleti was described by Lucas Figs. 100- 105. (1 846) from severa l fem ales from Hippone and EI Kala. Female (from Tunisia, no exact locality; M NHN, Lucas even recogni sed two colour morphs of this 6474): Carapace 3.65 long, 2.65 wide, 1.8 high at PLE. species. but provided a colour fig ure of only a single Ocular area 1.45 lo ng, 1. 8 wide anteriorly and 1.7 wide fe male (op. cit. : pI. 7, fig . 5) . The colo ur pattern of this posteriorly. Diameter of AME 0.50. Abdomen 4.1 long. fem ale and its general appearance are very simila r to 2.75 wide. Chelicera I length 1. 1. Clypeal ht: igh t 0.3. those of the male of A. num idicus (op. cit. : pI. 7, fig. 10) Length of leg segments: I 1.5+ 1.2+ 1.1 +0.7+0.55; II and it is likely that it belongs to this species . Thus , 1.5 + 1.0+ 1.0+0.75+0.6; III 2.4+ 1.25 + 1.4+ 1.6+0.8; IV although A. nicoleti was once treated as a j unior 2,2 + 1. 2 + 1.6+ 2. 0+0.9. Leg spination: I: Fm d 1-1-3; T b synonym of A. monardi (Si mon, 1868: 66), we are of the pr I-I, v 2-2-2ap; M t v 2-2ap. II: Fm d 1-2-3; T b pr I-I , opinion that its status remains uncertain and needs v 1-1-2ap; Mt pr I-lap, v 2-2ap. III: Fm d 1-2-4; Pt pr further attention in the future. The name A. nicoleti is and rt I; Tb d 1-0-0, pr and rt I-I- I, v 1-0-2ap; Mt d therefore removed from synonymy with A. munardi (see 1- 1-0, pr and rt 1-0-2, v 1- 1-2ap. IV: Fm d 1- 1-2 ; Pt pr also under that species) . and rt I; T b d 1-0-0, pr and rt 1- 1-1 , v 2-0-2ap; M t d 1-1-0, pr 1-1-2, rt 1-0-2, v 1-1-2ap. Coloration: carapace A eiul'illus lIumiciicus (Lucas, 1846) brown, wit h dark brown eye field , densely covered wi th whi te scaly hairs and wi thout colour pattern. Clypeus Sa/ficus I1wnidicus Luca s, 1846: 152- 153, p!. 6, fi g. LO (D o; 0 ho lotype yellow, " cheeks" brownish; both clypeus and " cheeks" lost; not exa mined ). densely covered with white hairs. Eyes of first row Fo r a complete reference li st see Platnick (2004). bordered by white hairs, Chelicerae yellow-brownish. Comments: Salticus num idicus was described by Luca o. Abdomen ye llow-grey, dorsum with poorly marked (1846) from a single male from Setif. The 0 holotype of spotty pattern, densely covered with yellowish, browni sh this species is considered lost. We have been unable to and white hairs. All legs yellow, w ith brownish pa tches. match our material to Lucas' descriptio n and moreover Palps yellow, covered with white hairs and brown came to the conclusion that this species is un likely to be bristles. Epigyne and spermathecae as in Figs . 106- 107. congeneri c with A elurilfus. The main reason is the part of M atl!l"ia/ examil1 ed: ALG ERI A: 30 12 (ZMCD, 227. 230; hitherto Lucas' description (op. ci t.: p. 152) devoted to leg charac determined as A. cerrillus), Biskra [= Beskra] , c. 34° 51' N, 5 °43' E, ters: " Les pattes sont courtes, greles, al'exception cepend date unk nown (Meinert); 20 (MCZ, 28547; hitherto determi ned by E. Simon as A. lII ayeli), 'Sahara (607)" da te unknown (G. W. & E. G . ant de la premiere paire, qui est un peu plus robuste que Peckham col!.) ; 50 (I ZWP, 3, 21, 23 ), 'Algeria , 1866- 1867' (W. T . les autres; ..." . All the A elurillus species known to us Taczanowski). TUN ISIA: 12 (MCZ. 28549; hitherto determined as A. have all legs equally developed, but Phlegra species always moyen") , T uni s. Ex. Mus. His!. Na t. Pa ri s, 56 '. date un known (G. W. have the fi rst pair of legs much stronger and more robust. & E. G. Peckha m col!.); 20 (S N H M. 1233/ 1905), 'T unisia. Sfax, Thus, we are in favour of the opini nn of Simon (1876), c. 34°44' N, 10 045'E, 1904 (Boerio Consta nce),; 10 (ZM MU, hitherto determined as A . m ayeti), La-Gali t [= La Gali te, = Jazira t Jali tah], who considered Salticus nUl11idicus a member of Phlegra. c. 37° 31'N, 8° 55' E, 9 October 1983 (collector un known); 70 22 The pro blem needs ~ pccial atlcntio [1 in the future. (MNH N. 6474), 'Tunisia' (da te a nd collecto r unknown); 10 (MRAC, 124.249; hitherto determined by O. 1. Clark as A. hirl ipes). T unisie: Reg. Sahel, K hniss' [Tunisia, Monasti r [= AI Munastir] G overnora te. Acknowledgements Khniss [Khunays], c. 35 °43'N, 10 049'E], 9 April 1963 (Ph. Lebru n) We wish to express our warmest thanks to Dr R . Bosmans (Gent, Belgium), Prof. T. Cantarella (DBAU), Comments on the remaining Aelurillus species of Lucas Mr P. Cardoso (CBA), M r J. Hogan (HECO), Dr T. Hufiejt (IZWP), D r P. Jager (SM FM), Dr R . Jocque A eiurillus gesticulator (Lucas, 1846) (M RAC), Dr T. K ronestedt (SMNH), Mrs L. Sa/ticus gesticulator Lucas, 1846: 150- L5 1, pI. 6, fi g. 9 (02; 2 holotypc Leibensperger (MCZ), Dr G. Levy (HUJ), Dr K . G . lost; not exa mined). Mikhailov (ZMMU), Mr J. Murphy (Hampton, UK), For a complete reference list see Pla tniek (2004). Dr C. Rollard (M N H N), Dr N . Scharff (ZMCD) and CO/11lJ1 ents: Salticus gesticulator was described by Mr R , Snazell (Swanage, D orset, UK) for giving access Lucas (1846) from females taken from a number or to their collections. We wish to thank M iss Anna D. localities (Alger, Mila, Consta ntine and Setif) and thus it Logunova (Manchester, U K) for kind help in translating was a rather abundant species. The 2 holotype is con Lucas' and Simon's descriptions from the French. The sidered lost. Neither the description nor the colour fi gure researches by the fi rst author (G.A.) were in part spon of A. gesticulator provi ded by Lucas (1846: pI. 6, fig . 9) sored by grant No. 01-04-48989 from the Russian Fund were sufficient to identify the species. A lthough we think of Basic Research J.nd by the grant fo r postgraduates this may be one of the junior synonyms of A. luctuusus and students N o. 03-04-07008 from the Russian F und of (see above), currently it seems worth consid ering thi s Basic Research. Two anonymous referees are thanked species name a nom en dubium , fo r their critical comments helping to improve the ms. 248 A "/I/rillus species of the W, Mediterranean
References P'\VISI, P 1880: Studi sugh Araenidi africani. J. Aracnidi di Tunisia. Alllwli :vlus. civ. Slor. nal. Giacomo Doria 15: 283-388. ALICATA, P. & CANTARELLA,'I 2000: I ~alticidi di Sicilia: ~ lato PA VESI, P 1895 .\racnidi. In : Viaggio del Dr E. Festa in Palestina, della conoscenza e descrizionc di due nuove sp~cic ( / \ran~ac nel Libano e regione vicine. Boll. Musei Zool. Anal. compo R. Salticidae). MemO/'ie Soc. el1l. i/{J!. 78(2) 484- 49~. Univ. Torino 10(216): I- II . BODENHEIMER, F. S. 1937: Prodromus fauna: Palestinu:. !'vJeil1. PICKARD-CAMBRIDGE, O. 1872: G eneral list of th e spiders of Insl. EgYPle 33: 1- 286. Palestine and Syria, with descriptions of numerous new species, BON NET, P. 1955: Bibliographia Aral1eorum 2(1): I· 91X . Toulouse. a nd characters of two new genera. Proc. ::001. Soc. Land. 1872: BRIGNOLL P. M. 1983: A catalogue oIthe Arwu!({e descrihed betlreell 212- 354. 1940 alld 1981 . 1- 755 . Manchester Univ. Pre,,:;. Manchester. PICKARD-CAMBRIDGE, O. 1876: Catalogue of a collection of BRISTOWE, W. S. 1935: T he spiders of Gr~ece and the adjacent spiders made in Egypt, with descriptions of new species and islands. Pmc. zool Soc. Land. 1934(4): 733~ 788. characters of a new genus. Proc. zool. Soc. Lond. 1876: CANTARELLA, T. 1982: Salticidae (Araneac) delle Isole Maltesi. 54 1- 630. Al1imalia 9(113): 239- 252 . PI.A'\ leT, L 1')05: Araignees (A.-aignees Chernetes, Scorpions, CANTARELLA, T. 1983: Una nuova specie di Aelurillus (Araneae, Opilions). In: Hi