B5. Mediterranean and Black Sea, FAO Statistical Area 37

Total Page:16

File Type:pdf, Size:1020Kb

B5. Mediterranean and Black Sea, FAO Statistical Area 37 49 B5. MEDITERRANEAN AND BLACK SEA FAO Statistical Area 37 Figure B5.1 - The Mediterranean and Black Sea (Area 37) by Jordi Lleonart * INTRODUCTION The Mediterranean (Figure B5.1) is a semi-enclosed sea with a surface of about 3.3 million km2, contributing 0.8 percent to the total world marine surface. Due to its geographical position, being placed at a relatively narrow range of latitudes (from 30ºN to 46ºN) in the temperate zone of the Northern Hemisphere, the Mediterranean Sea shows a marked seasonal cycle. Water masses are Atlantic through the Strait of Gibraltar proceeds stratified in summer, but the deep-water (below to the eastern Mediterranean progressively losing 400 m) temperature is 13±0.3ºC throughout the nutrients and increasing in salinity through year. The low precipitation in the summer period evaporation. It eventually goes back out to the is the main characteristic of the Mediterranean Atlantic to form the high saline component of the climate. The Mediterranean has a negative water Atlantic circulation. This makes for a large budget: the loss of water through evaporation is number of habitats with many small local stocks greater than the inputs due to rain and river and species whose stock trends cannot be runoff, hence the contribution of about individually described here but show up as 1 700 km3/year of Atlantic water trough the Strait dissimilarities between sub-areas (Garibaldi and of Gibraltar balances these losses (Oliver, 2003). Caddy, 1998). The system of basins includes water ranging from The Mediterranean has been globally considered Atlantic origin in the Western Mediterranean to as an oligotrophic sea (Margalef, 1985; Estrada, warm-temperate hypersaline water in the Levant 1996; Stergiou et al., 1997b). Normally, the basin, hyposaline waters in the Black Sea, and gradual decrease in nutrient would result in a cold-temperate estuarine conditions in the boreal west to east decrease in productivity, but there Azov Sea. Surface water entering from the are local exceptions resulting from a north-south Figure B5.2 - Annual nominal catches ('000t ) by ISSCAAP species groups in the Mediterranean and Black Sea (Area 37) 2 000 35- Herrings, sardines, anchovies 24- Shads Other ISSCAAP Groups 1 500 34- M iscellaneous demersal fishes 33- M iscellaneous coastal fishes 32- Cods, hakes, haddocks 1 000 500 0 1950 1955 1960 1965 1970 1975 1980 1985 1990 1995 2000 Source FAO * FAO, Marine Resources Service, Fishery Resources Division 50 productivity gradient evident also from satellite subsequently increased to 1.5 million tonnes in imagery of chlorophyll A, due to incoming 2001 and 2002, after some small fluctuations. nutrients from northern rivers (notably the Small pelagics account for approximately Rhone, Po and Black Sea inflows into the 50 percent of total Mediterranean catches. Aegean). These anthropogenic effects on Anchovy (Engraulis encrasicolus) with fisheries production have been described by 59 percent of small pelagics catch, and sardine Caddy, Refk and Do Chi (1995). (Sardina pilchardus) with 16 percent are the The continental shelf is mostly a narrow coastal most abundant. Other small pelagics are sprat fringe with the exceptions of the Adriatic Sea, (Sprattus sprattus), sardinella (Sardinella aurita) Gulf of Gabès, northern Black Sea south of Sicily and Azov sea sprat (Clupeonella cultriventris). and gulf of Lions, and represents only 23 percent The substantial increase in European anchovy of the total area. catches from 1960 to the mid 1980s (Figure The fisheries of the Mediterranean have shown a B5.3) partly reflects increased effort from Turkey surprising resilience to fishing compared with in the Black Sea that was added to that of the some areas of the Atlantic. This is especially former Soviet Union fleet previously dominating noteworthy since formal and coordinated the fishery. As with sprat catches, it could reflect measures for fisheries management are largely eutrophication of the Black Sea environment by absent in most Mediterranean countries, though incoming rivers (Mee, 1992; Zaitsev, 1993). The more frequent activities and a revision of the collapse of anchovy, sprat and Azov sea sprat in terms of reference of the GFCM (General 1990 is the result of the introduced ctenophore Fisheries Commission for the Mediterranean) and species Mnemiopsis leidy in the Black Sea. A its Scientific Advisory Committee (SAC) is subsequent increase of catches did not reach the intended to remedy this from 2000. Farrugio et former levels. The sardine series shows a smooth al. (1993), Farrugio (1996), Anon. (2001), Bas increasing trend to late 1980s (282 000t in 1988) (2002), Oliver (2003) and Lleonart and Maynou and a subsequent moderate decrease to about (2003), among others, have reviewed 187 000t in 2002. Mediterranean fisheries. Mixed bottom fisheries using small mesh trawls, gillnets, trammel nets, traps, pots and dredges PROFILE OF CATCHES capture a large suite of demersal fish and invertebrates of high value for the fresh fish Nominal catches in Area 37 increased from market, with no single species making up more slightly over 0.7 million tonnes in 1950, to near than 10 percent of the total demersals. However, 2 million tonnes between 1982 and 1988 (Figure in those areas where trawl fleets operate B5.2 and table D5). Catches subsequently regularly, despite the inherent complexity of declined steeply to 1.3 million tonnes as a result multispecies catches, there is an identifiable of the collapse of the sprat and anchovy fishery in series of target species which in biomass or in the Black Sea (Figure B5.3). Total catches have economic terms, constitute an important basis of production. The species considered demersals Figure B5.3 - Annual nominal catches ('000t) of (more than 100, although some of them show a selected species in ISSCAAP Groups 24 & 35, pelagic behaviour) represent around 40 percent of Mediterranean and Black Sea (Area 37) total reported catches in the Mediterranean and 800 European pilchard (=Sardine) Black Sea. Hake (Merluccius merluccius), red 700 European anchovy mullets (Mullus spp.), blue whiting European sprat (Micromesistius poutassou), whiting (Merlangius 600 Sardinellas nei Azov Sea sprat merlangus), anglerfishes (Lophius spp.), Pagellus 500 spp., bogue (Boops boops), picarels (Spicara 400 spp.) striped venus (Chamelea gallina), Octopus 300 spp., cuttlefish (Sepia officinalis), red shrimp 200 (Aristeus antennatus), Norway lobster (Nephrops norvegicus) and deepwater rose shrimp 10 0 (Parapenaeus longirostris), are the main 0 demersals. 1950 1960 1970 1980 1990 2000 Source FAO 51 By-catch species include species not registered catches, but their economic value is far greater. individually in official catches or not always Catches of bluefin tuna increased from the mid present but that together can represent a 1960s to the mid 1990s, but then significantly significant proportion of catches such as small dropped (Figure B5.7). The bluefin tuna fishery sharks (Carbonell et al., 2003). in the Mediterranean is a matter of concern because of the marked expansion of the But there are also other local and/or well priced species caught in relatively small quantities like some species of flatfishes, Scorpaenidae, Figure B5.4 - Annual nominal catches ('000t) of Sparidae, Triglidae, Mugilidae, Serranidae, and selected species in ISSCAAP Group 56, some invertebrates. The last by-catch group is Mediterranean and Black Sea (Area 37) from time to time also relatively abundant but not Clams, cockles, arkshells, etc. always high priced species such as some 60 Scombridae and Carangidae, or from species with Striped venus low market acceptability like bogue or picarels (Spicara spp.). Discards are also important, and 40 the reasons for discarding diverse: illegal sizes, non-commercial species or sizes, or market reasons (Carbonell et al., 1998; Lleonart and 20 Maynou, 2003). It is worth noting the spectacular increase in 0 catches of striped venus, Chamelea gallina, 1950 1960 1970 1980 1990 2000 mainly in the Adriatic, reaching a maximum in Source FAO 1993 (Figure B5.4). This could reflect the Figure B5.5 - Annual nominal catches ('000t) of interaction of fishing with the transition from selected species in ISSCAAP Groups 45 & 57, oligotrophic conditions in the early period to Mediterranean and Black Sea (Area 37) close to mesotrophic conditions. 30 Cephalopod catches (Figure B5.5) show a 25 Deepwater rose shrimp maximum around 1990 that could in part reflect Common octopus Common cuttlefish 20 reduced abundance of large predatory fish as Common squids nei speculated by Caddy and Rodhouse (1998), but in most cases these are components in a mixed 15 groundfish fishery. The onset of deep water 10 trawling off the slope areas in the mid-1980s appears to explain the sharp rise in deepwater 5 rose shrimp catches, which are speculated to have benefited from a reduction in the biomass of 0 large hake in slope areas. 1950 1960 1970 1980 1990 2000 Source FAO Most of the gadoids, and other semi-pelagic Figure B5.6 - Annual nominal catches ('000t) of species such as bogue and picarel (Spicara spp.), selected species in ISSCAAP Groups 32 & 33, as well as red mullets (Mullus spp.), have shown 60 Mediterranean and Black Sea (Area 37) steady increasing trends over the whole time Surmullets (=Red mullets) nei 50 period, except for declines of several species in European hake the last few years. More remarkable is the case of Whiting hake which after reaching historic maxima in the 40 Bogue Picarels nei mid 1990 (more than 52 000 t), catches have dropped by more than half in 2002. The whiting, 30 only present in the Eastern Mediterranean, shows 20 also this general pattern with maximum around 1990 (Figure B5.6). 10 The prominent large pelagics are bluefin tuna 0 (Thunnus thynnus) and swordfish (Xiphias 1950 1960 1970 1980 1990 2000 gladius). They represent 3 percent of total Source FAO 52 farming/fattening activities of wild specimens, japonicus), and Atlantic mackerel (Scomber which contributes to increase the fishing pressure scombrus).
Recommended publications
  • Striped Red Mullet (Mullus Surmuletus)
    MarLIN Marine Information Network Information on the species and habitats around the coasts and sea of the British Isles Striped red mullet (Mullus surmuletus) MarLIN – Marine Life Information Network Marine Evidence–based Sensitivity Assessment (MarESA) Review Morvan Barnes 2008-09-02 A report from: The Marine Life Information Network, Marine Biological Association of the United Kingdom. Please note. This MarESA report is a dated version of the online review. Please refer to the website for the most up-to-date version [https://www.marlin.ac.uk/species/detail/81]. All terms and the MarESA methodology are outlined on the website (https://www.marlin.ac.uk) This review can be cited as: Barnes, M.K.S. 2008. Mullus surmuletus Striped red mullet. In Tyler-Walters H. and Hiscock K. (eds) Marine Life Information Network: Biology and Sensitivity Key Information Reviews, [on-line]. Plymouth: Marine Biological Association of the United Kingdom. DOI https://dx.doi.org/10.17031/marlinsp.81.1 The information (TEXT ONLY) provided by the Marine Life Information Network (MarLIN) is licensed under a Creative Commons Attribution-Non-Commercial-Share Alike 2.0 UK: England & Wales License. Note that images and other media featured on this page are each governed by their own terms and conditions and they may or may not be available for reuse. Permissions beyond the scope of this license are available here. Based on a work at www.marlin.ac.uk (page left blank) Date: 2008-09-02 Striped red mullet (Mullus surmuletus) - Marine Life Information Network See online review for distribution map Mullus surmuletus foraging in sand.
    [Show full text]
  • (Mullus Surmuletus) in Bottom Trawl Fisheries Enis Noyan Kostak1 , Adnan Tokaç2
    EISSN 2602-473X AQUATIC SCIENCES AND ENGINEERING Aquat Sci Eng 2018; 33(3): 90-95. • DOI: 10.26650/ASE201817 Original Article Predicting the Size Selectivity Based on the Striped Red Mullet Morphology (Mullus surmuletus) in Bottom Trawl Fisheries Enis Noyan Kostak1 , Adnan Tokaç2 Cite this article as: Kostak, E.N., Tokaç, A. (2018). Predicting the Size Selectivity Based on the Striped Red Mullet Morphology (Mullus surmule- tus) in Bottom Trawl Fisheries. Aquatic Sciences and Engineering, 33(3): 90-95. ABSTRACT The striped red mullet (Mullus surmuletus) is a commercially important demersal species caught by mixed trawl fishing in the Mediterranean Sea. Although it is not among the target species of bottom trawl operations, the striped red mullet is an important species due to its high commercial value in trawl catch composition. The pri- mary target is to catch adult individuals while allowing the immature and juvenile fish to escape for sustainable fisheries using selective fishing gears. Various selectivity studies regarding trawl codends aimed at improving selectivity have been carried out in the past two decades. However, the selectivity sea trials generally require a great deal of work, time, and labor on the deck and are also expensive. Therefore, instead of experimental- based selectivity studies, simulation-based studies (i.e., the FISHSELECT methodology) were recently started to predict the size selectivity of species in bottom trawl fisheries. In this study, sampled individuals of the striped red mullet were used in morphological measurements, fall-through experiments, and simulation phas- es. Diamond mesh sizes of 40, 44, and 50 mm and a square mesh size of 40 mm of bottom trawl codends were simulated, and the L50 values were calculated as 9.87, 10.75, 12.19, and 12.3 cm for the aforementioned mesh sizes, respectively.
    [Show full text]
  • Parupeneus Forsskali (Fourmanoir & Guézé, 1976) in the Mediterranean, with Preliminary Information on Its Diet Composition in Cyprus
    BioInvasions Records (2020) Volume 9, Issue 2: 209–222 CORRECTED PROOF Research Article Progress of the dispersal of the alien goatfish Parupeneus forsskali (Fourmanoir & Guézé, 1976) in the Mediterranean, with preliminary information on its diet composition in Cyprus Athanasios Evagelopoulos1,*, Andreas Nikolaou1, Nikolas Michailidis2,3, Thodoros E. Kampouris1 and Ioannis E. Batjakas1 1Department of Marine Sciences, University of the Aegean, University Hill, 81100 Mytilene, Greece 2Department of Fisheries and Marine Research, 101 Vithleem Str., 1416 Strovolos, Nicosia, Cyprus 3Department of Biological Sciences, University of Cyprus, 1 Panepistimiou Str., 2109 Aglantzia, Nicosia, Cyprus Author e-mails: [email protected] (AE), [email protected] (AK), [email protected] (NM), [email protected] (TEK), [email protected] (IEB) *Corresponding author Citation: Evagelopoulos A, Nikolaou A, Michailidis N, Kampouris TE, Batjakas IE Abstract (2020) Progress of the dispersal of the alien goatfish Parupeneus forsskali Parupeneus forsskali has been the latest Indo-Pacific goatfish species to expand its (Fourmanoir & Guézé, 1976) in the range into the Mediterranean. It is the least studied alien mullid in the Eastern Mediterranean, with preliminary Mediterranean, and specific information on its diet is generally lacking in the information on its diet composition in literature. The objectives of this paper are (1) to comprehensively document the Cyprus. BioInvasions Records 9(2): 209– 222, https://doi.org/10.3391/bir.2020.9.2.06 progress of its invasion in the Mediterranean through a systematic literature review to retrieve all published records of the species in the region, and (2) to present Received: 15 October 2019 preliminary quantitative information on its diet in its non-native range.
    [Show full text]
  • (Mullus Surmuletus) and Striped Red Mullet (M. Barbatus) an Exchange for a New Set of M
    Red mullet ( Mullus surmuletus ) and striped red mullet ( M. barbatus ) otolith and scale exchange 2011 Red mullet ( Mullus surmuletus ) and striped red mullet ( M. barbatus ) otolith and scale exchange 2011 Mahé, K., Elleboode, R., Charilaou, C., Ligas, A., Carbonara, P. & Intini, S., 2012. Red mullet ( Mullus surmuletus ) and striped red mullet ( M. barbatus ) otolith and scale exchange 2011, 30pp. Table of contents 1. Introduction..................................................................................4 2. Participants ..................................................................................4 3. Material .........................................................................................4 4. Reading procedure ......................................................................6 5. Results..........................................................................................8 5.1. Precision.............................................................................................................9 5.2. Relative bias (Accuracy).....................................................................................10 5.3. Age reading quality.............................................................................................12 6. Executive Summary.....................................................................13 7. References ...................................................................................15 8. Appendix 1 : Details results of Mullus surmuletus Otolith Exchange (VIIIab)...............................................................................................16
    [Show full text]
  • Feeding Habits of the Striped Red Mullet, Mullus Surmuletus in the Eastern Adriatic Sea
    ISSN: 0001-5113 ACTA ADRIAT., ORIGINAL SCIENTIFIC PAPER AADRAY 59 (1): 123 - 136, 2018 Feeding habits of the striped red mullet, Mullus surmuletus in the eastern Adriatic Sea Mišo PAVIČIĆ, Jasna ŠILJIĆ, Dajana BRAJČIĆ JURICA and Sanja MATIĆ-SKOKO* Institute of Oceanography and Fisheries, Šetalište Ivana Meštrovića 63, P.O. Box. 500, 21000 Split, Croatia *Corresponding author: [email protected] Feeding habits of the striped red mullet, Mullus surmutetus in the eastern Adriatic Sea were investigated. Stomach contents of 203 specimens (11.5 - 32.9 cm TL) collected by bottom trawling were analyzed. Commonly accepted procedures were followed during the diet composition inspec- tion and standard keys were used for food items determination. Examinations showed that 39 iden- tified prey taxa belong to 11 major systematic groups: Crustacea, Gastropoda, Bivalvia, Pisces, Cnidaria, Echinodermata, Polychaeta, Nematoda, Bryozoa, Algae and Rhizaria. The predominant and preferred prey category was decapod crustaceans within all size categories and only in the largest individuals, the dominance of Bivalvia followed by Polychaeta and Crustacea were found. The largest individuals also showed more variety in consumption of different prey categories and a higher mean number of prey items in comparison with smaller fish. No significant differences in prey foraging between males and females (p>0.05) and regarding to season (p>0.05) were found. Feeding on such wide spectra of prey without significant variations regarding to sex and season suggests that the striped red mullets are able to adapt to shifts in spatio-temporal variations in the abundance of potential prey. Presence of detritus in the digestive tracts is highly related to its forag- ing behaviour on muddy detritic bottoms.
    [Show full text]
  • Trophic Diversity of a Fish Community Associated with a Caulerpa Prolifera (Forsskål) Meadow in a Shallow Semi-Enclosed Embayment
    Journal of Marine Science and Engineering Article Trophic Diversity of a Fish Community Associated with a Caulerpa prolifera (Forsskål) Meadow in a Shallow Semi-Enclosed Embayment Maria Maidanou 1,2, Panayota Koulouri 1,* , Paraskevi K. Karachle 3 , Christos Arvanitidis 1, Drosos Koutsoubas 2 and Costas Dounas 1 1 Institute of Marine Biology, Biotechnology and Aquaculture, Hellenic Centre for Marine Research, Gournes Pediados, P.O. Box 2214, 71003 Heraklion, Greece; [email protected] (M.M.); [email protected] (C.A.); [email protected] (C.D.) 2 Faculty of Environment, Department of Marine Sciences, University of the Aegean, 81100 Mytilene, Greece; [email protected] 3 Institute of Marine Biological Resources and Inland Waters, Hellenic Centre for Marine Research, Athens-Sounio Ave, P.O. Box 712, 19013 Anavyssos, Greece; [email protected] * Correspondence: [email protected]; Tel.: +30-2810-337716 Abstract: This study investigates the trophic diversity of fishes living in a meadow of Caulerpa prolifera on a bimonthly basis between May 2006 and April 2007 in a semi-enclosed coastal marine ecosystem of the Mediterranean Sea (Elounda Bay, Crete Island). The study area is shallow and protected from waves, and it is covered by a C. prolifera bed, characterized by high organic input and a highly diverse macrobenthic community. Feeding patterns of the fish, investigated on the basis of stomach content analyses, were described in terms of numerical abundance and frequency of occurrence of prey taxa. A total of 1642 fish individuals, belonging to 17 species, were examined. In total, 45,674 prey Citation: Maidanou, M.; Koulouri, P.; individuals were identified belonging to 110 prey taxa, most of which were Malacostraca including Karachle, P.K.; Arvanitidis, C.; their larvae and Copepoda (41,175 individuals identified to 71 taxa).
    [Show full text]
  • Investigation of Microplastics Ingestion and Effects in Striped Red Mullet in East Mediterranean
    Investigation of Microplastics Ingestion and Effects in Striped Red Mullet in East Mediterranean Sea Danae Patsiou∗1, Nikoletta Digka1, and Catherine Tsangaris1 1Hellenic Centre for Marine Research { Street Address : 46,7 km Athens Sounio ave., P.O. Box 712, P.C. 19013 Anavyssos Attiki, Greece Abstract Marine plastic debris is increasing and so does the microplastic (MP) ingestion by marine organisms. Detection of MPs in the marine environment is ongoing, however, there is lack of consensus regarding the effects of MP ingestion in marine organisms. The aim of the present study is to assess MP ingestion together with cellular and biochemical biomarkers in native Mediterranean fish species proposed as bioindicators of MPs, at coastal sites under different human pressure intensity in the Ionian Sea, and to investigate whether biomarker variations can be related to the occurrence of MPs in their gastro-intestinal (GI) tract. Striped red mullets Mullus surmuletus, were sampled along the tourist coast of Zakynthos island and inside the Zakynthos National Marine Park. The fish GI tracts were digested using 15% H2O2 at oC and then filtered to detect ingested MPs. The polymer identification analysis using Fourier-Transform Infrared Spectroscopy analysis of the samples is ongoing. Blood samples were collected from alive M. surmuletus to evaluate DNA damage, and the results so far indicate no differences in the micronuclei occurrence in fish blood cells among sampling sites. The investigation of biochemical biomarkers includes: acetylocholinesterase in muscle tissues to evaluate neurotoxicity, and catalase and glutathione S-transferase in the liver tissues to assess antioxidant defence and biotransformation responses. The present study carried out in the framework of the Plastic Busters MPAs project intends to measure the extend of MPs pollution by in situ investigation of cellular and biochemical parameters of native Mediterranean species, to attribute the animal health status directly to the presence of MPs in the animal.
    [Show full text]
  • Improvement of the Fishery Knowledge of Striped Red Mullet of the Bay of Biscay
    Working Document for the Working Group on Widely Distributed Stocks (WGWIDE) 30 August - 5 September 2017, Copenhagen (Denmark) Improvement of the fishery knowledge of striped red mullet of the Bay of Biscay Nathalie Caill-Milly1, Muriel Lissardy1 Jean-Pierre Léauté2 1 Ifremer, Laboratoire Environnement Ressources d'Arcachon, UFR Côte Basque, 1, allée du Parc Montaury, 64600 Anglet, France 2 Ifremer, Laboratoire Ressources Halieutiques La Rochelle, place Gaby Coll, B.P.7 17137 L’Houmeau, France Introduction Striped red mullet (Mullus surmuletus) belongs to the species for which individualization of stocks is advanced by ICES in western Europe for areas including the Bay of Biscay and the areas bordering the Iberian peninsula. Since 2012, ICES has provided recommendations with regards to this stock. These recommendations are given for two years and are based on an approach adopted by ICES in 2012 in the case of insufficient data for an analytical evaluation (Data Limited Stocks, DLS). For 2013 and 2014, ICES advice consisted in a 20% reduction in catches (average of the last 3 years:- 2009-2011) as a precautionary measure. Since that, no new advice has been formulated, the advice remains the same. Considering the impacts of such measurements on the French fishing fleet, improvement of the data available for stocks in the DLS category is therefore a priority. The project ROMELIGO aims to change this situation by contributing to the improvement of the knowledge on three stocks (striped red mullet, whiting and pollack) on the basis of the available data (declaring landing data or sampling data for French fishermen, data from scientific campaigns, etc.) or data to be collected (biological parameters).
    [Show full text]
  • Report of the Workshop on Age Reading of Red Mullet and Striped Red Mul- Let, 2–6 July 2012, Boulogne-Sur-Mer, France
    ICES WKACM2 REPORT 2012 ICES ADVISORY COMMITTEE ICES CM 2012 / ACOM: 60 REF. WGNEW, PGMED, PGCCDBS Report of the second Workshop on Age Reading of Red Mullet and Striped Red Mullet 2 - 6 July 2012 Boulogne-sur-Mer, France International Council for the Exploration of the Sea Conseil International pour l’Exploration de la Mer H. C. Andersens Boulevard 44–46 DK–1553 Copenhagen V Denmark Telephone (+45) 33 38 67 00 Telefax (+45) 33 93 42 15 www.ices.dk [email protected] Recommended format for purposes of citation: ICES. 2012. Report of the workshop on age reading of red mullet and striped red mul- let, 2–6 July 2012, Boulogne-sur-Mer, France. ICES CM 2012/ACOM:60. 52 pp. For permission to reproduce material from this publication, please apply to the Gen- eral Secretary. The document is a report of an Expert Group under the auspices of the International Council for the Exploration of the Sea and does not necessarily represent the views of the Council. © 2012 International Council for the Exploration of the Sea ICES WKACM2 REPORT 2012 | i Contents Summary .................................................................................................................................. 1 1 Review on the biology .................................................................................................. 2 1.1 Mullus barbatus in the Mediterranean Sea ......................................................... 2 1.2 Mullus surmulutus in the Northeast Atlantic and the Mediterranean Sea ..........................................................................................................................
    [Show full text]
  • Agnatha, Chondrichthyes and Osteichthyes
    See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/311861377 Agnatha, Chondrichthyes and Osteichthyes Chapter · November 2016 CITATIONS READS 0 1,531 2 authors: Antonis Petrou Charitos Zapitis AP Marine Env.Consultancy Ltd & Enalia Physis Environmental Research Centre University of Derby 29 PUBLICATIONS 131 CITATIONS 1 PUBLICATION 0 CITATIONS SEE PROFILE SEE PROFILE Some of the authors of this publication are also working on these related projects: Lionfish in the Mediterranean View project PCY1 - Investigating the effects of recreational diving on the macroalgal communities of the 'Zenobia' shipwreck (Cyprus) View project All content following this page was uploaded by Charitos Zapitis on 24 December 2016. The user has requested enhancement of the downloaded file. Chapter 26 - Agnatha CHAPTER 26: AGNATHA, CHONDRICHTHYES AND OSTEICHTHYES FISHES Antonis Petrou and Charitos Zapitis 1. INTRODUCTION The classification of fishes is not straightforward since they do not form a natural scientific grouping like the other vertebrate classes, i.e. the amphibians, reptiles, birds and mammals. Indeed, fishes can be considered by exclusion to be vertebrates that are not tetrapods (see Chapter 25: Introduction to Vertebrates). They are aquatic, gill- bearing, ectothermic ('cold-blooded') animals with a distinguishable head and, when present, digit-less limbs. Traditionally, fish have been arranged into three groups: ➵ Agnatha 1, the jawless fish (Myxini [hagfishes] and Hyperoartia [lampreys]); ➵ Chondrichthyes, the cartilaginous fish (sharks, skates and rays); ➵ Osteichthyes, the bony fish (Actinopterygii [ray-finned fishes] and Sarcopterygii [lobe-finned fishes]). This classification is adequate for general purposes, although Agnatha is paraphyletic and includes several groups of extinct jawless fishes.
    [Show full text]
  • Feeding Ecology of Mullus Barbatus and Mullus Surmuletus Off The
    International Journal of Fisheries and Aquatic Studies 2017; 5(6): 321-325 E-ISSN: 2347-5129 P-ISSN: 2394-0506 (ICV-Poland) Impact Value: 5.62 Feeding ecology of Mullus barbatus and Mullus (GIF) Impact Factor: 0.549 IJFAS 2017; 5(6): 321-325 surmuletus off the Egyptian mediterranean coast © 2017 IJFAS www.fisheriesjournal.com Received: 12-09-2017 Hatem H Mahmoud, Reda M Fahim, Tark M Srour, Nagy El-Bermawi Accepted: 14-10-2017 and Mohamed A Ibrahim Hatem H Mahmoud College of fisheries technology Abstract and aquaculture, Arab Academy The present work describes the feeding habits of the two-dominant species of family Mullidae; red for science, technology and mullet, Mullus barbatus barbatus Linnaeus, 1758 and striped red mullet, Mullus surmuletus Linnaeus, maritime transport, Egypt 1758 caught off the Egyptian Mediterranean coast throughout the period from January to December 2016. A total of 322 specimens of M. barbatus and 289 of M. surmuletus were investigated. The index of Reda M Fahim Fisheries Biology Laboratory, relative importance (IRI) indicated that Polychaeta was, for M. barbatus, the most important food item in Fisheries Division, National all seasons with an average of 26.86%, followed by Decapoda, Amphipoda and Mysidacea. While, in the Institute of Oceanography and case of M. surmuletus, Mysidacea was the most preferred food item in all seasons with an average of Fisheries, Alexandria, Egypt 34.02%, then Amphipoda and Decapoda. According to the previous results, it can be inferred that the two Mullid species; M. barbatus and M. surmuletus from the Egyptian Mediterranean waters are specialist Tark M Srour zooplanktivorous.
    [Show full text]
  • HELCOM Red List of Threatened and Declining Species of Lampreys and Fishes of the Baltic Sea
    Baltic Sea Environment Proceedings No. 109 HELCOM Red list of threatened and declining species of lampreys and fishes of the Baltic Sea Helsinki Commission Baltic Marine Environment Protection Commission Baltic Sea Environment Proceedings No. 109 HELCOM Red list of threatened and declining species of lampreys and fishes of the Baltic Sea Helsinki Commission Baltic Marine Environment Protection Commission Editor: Dr. Ronald Fricke, Curator of fishes, Ichtyology Contact address: Staatliches Museum für Naturkunde Stuttgart Rosenstein 1, 70191 Stuttgart, Germany E-mail: [email protected] Photographs © BfN, Krause & Hübner. Cover photo: Gobius niger For bibliographic purposes this document should be cited to as: HELCOM 2007: HELCOM Red list of threatened and declining species of lampreys and fish of the Baltic Sea. Baltic Sea Environmental Proceedings, No. 109, 40 pp. Information included in this publication or extracts there of is free for citing on the condition that the complete reference of the publication is given as stated above. Copyright 2007 by the Baltic Marine Environment Protection Commission - Helsinki Commission ISSN 0357-2944 Table of Contents 1 Introduction .......................................................................................................................6 2 Species and area covered.............................................................................................7 2.1 Species covered..............................................................................................................7
    [Show full text]