IV Eurasian Symposium on Hymenoptera | Programme
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(Hymenoptera) from the Middle Jurassic of Inner Mongolia, China
European Journal of Taxonomy 733: 146–159 ISSN 2118-9773 https://doi.org/10.5852/ejt.2021.733.1229 www.europeanjournaloftaxonomy.eu 2021 · Zheng Y. et al. This work is licensed under a Creative Commons Attribution License (CC BY 4.0). Research article urn:lsid:zoobank.org:pub:043C9407-7E8A-4E8F-9441-6FC43E5A596E New fossil records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China Yan ZHENG 1,*, Haiyan HU 2, Dong CHEN 3, Jun CHEN 4, Haichun ZHANG 5 & Alexandr P. RASNITSYN 6,* 1,4 Institute of Geology and Paleontology, Linyi University, Shuangling Rd., Linyi 276000, China. 1,4,5 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, East Beijing Road, Nanjing 210008, China. 2 School of Agronomy and Environment, Weifang University of Science and Techonoly, Jinguang Road, Shouguang, 262700, China. 3 School of Environmental and Municipal Engineering, Qingdao University of Technology, Qingdao 266033, China. 6 Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia. 6 Natural History Museum, London SW7 5BD, UK. * Corresponding authors: [email protected], [email protected] 2 Email: [email protected] 3 Email: [email protected] 4 Email: [email protected] 5 Email: [email protected] 1 urn:lsid:zoobank.org:author:28EB8D72-5909-4435-B0F2-0A48A5174CF9 2 urn:lsid:zoobank.org:author:91B2FB61-31A9-449B-A949-7AE9EFD69F56 3 urn:lsid:zoobank.org:author:51D01636-EB69-4100-B5F6-329235EB5C52 4 urn:lsid:zoobank.org:author:8BAB244F-8248-45C6-B31E-6B9F48962055 5 urn:lsid:zoobank.org:author:18A0B9F9-537A-46EF-B745-3942F6A5AB58 6 urn:lsid:zoobank.org:author:E7277CAB-3892-49D4-8A5D-647B4A342C13 Abstract. -
Managing Alternative Pollinators a Handbook for Beekeepers, Growers, and Conservationists
Managing Alternative Pollinators A Handbook for Beekeepers, Growers, and Conservationists ERIC MADER • MARLA SPIVAK • ELAINE EVANS Fair Use of this PDF file of Managing Alternative Pollinators: A Handbook for Beekeepers, Growers, and Conservationists, SARE Handbook 11, NRAES-186 By Eric Mader, Marla Spivak, and Elaine Evans Co-published by SARE and NRAES, February 2010 You can print copies of the PDF pages for personal use. If a complete copy is needed, we encourage you to purchase a copy as described below. Pages can be printed and copied for educational use. The book, authors, SARE, and NRAES should be acknowledged. Here is a sample acknowledgement: ----From Managing Alternative Pollinators: A Handbook for Beekeepers, Growers, and Conservationists, SARE Handbook 11, by Eric Mader, Marla Spivak, and Elaine Evans, and co- published by SARE and NRAES.---- No use of the PDF should diminish the marketability of the printed version. If you have questions about fair use of this PDF, contact NRAES. Purchasing the Book You can purchase printed copies on NRAES secure web site, www.nraes.org, or by calling (607) 255-7654. The book can also be purchased from SARE, visit www.sare.org. The list price is $23.50 plus shipping and handling. Quantity discounts are available. SARE and NRAES discount schedules differ. NRAES PO Box 4557 Ithaca, NY 14852-4557 Phone: (607) 255-7654 Fax: (607) 254-8770 Email: [email protected] Web: www.nraes.org SARE 1122 Patapsco Building University of Maryland College Park, MD 20742-6715 (301) 405-8020 (301) 405-7711 – Fax www.sare.org More information on SARE and NRAES is included at the end of this PDF. -
Genomes of the Hymenoptera Michael G
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Digital Repository @ Iowa State University Ecology, Evolution and Organismal Biology Ecology, Evolution and Organismal Biology Publications 2-2018 Genomes of the Hymenoptera Michael G. Branstetter U.S. Department of Agriculture Anna K. Childers U.S. Department of Agriculture Diana Cox-Foster U.S. Department of Agriculture Keith R. Hopper U.S. Department of Agriculture Karen M. Kapheim Utah State University See next page for additional authors Follow this and additional works at: https://lib.dr.iastate.edu/eeob_ag_pubs Part of the Behavior and Ethology Commons, Entomology Commons, and the Genetics and Genomics Commons The ompc lete bibliographic information for this item can be found at https://lib.dr.iastate.edu/ eeob_ag_pubs/269. For information on how to cite this item, please visit http://lib.dr.iastate.edu/ howtocite.html. This Article is brought to you for free and open access by the Ecology, Evolution and Organismal Biology at Iowa State University Digital Repository. It has been accepted for inclusion in Ecology, Evolution and Organismal Biology Publications by an authorized administrator of Iowa State University Digital Repository. For more information, please contact [email protected]. Genomes of the Hymenoptera Abstract Hymenoptera is the second-most sequenced arthropod order, with 52 publically archived genomes (71 with ants, reviewed elsewhere), however these genomes do not capture the breadth of this very diverse order (Figure 1, Table 1). These sequenced genomes represent only 15 of the 97 extant families. Although at least 55 other genomes are in progress in an additional 11 families (see Table 2), stinging wasps represent 35 (67%) of the available and 42 (76%) of the in progress genomes. -
Evolution of the Insects
CY501-C11[407-467].qxd 3/2/05 12:56 PM Page 407 quark11 Quark11:Desktop Folder:CY501-Grimaldi:Quark_files: But, for the point of wisdom, I would choose to Know the mind that stirs Between the wings of Bees and building wasps. –George Eliot, The Spanish Gypsy 11HHymenoptera:ymenoptera: Ants, Bees, and Ants,Other Wasps Bees, and The order Hymenoptera comprises one of the four “hyperdi- various times between the Late Permian and Early Triassic. verse” insectO lineages;ther the others – Diptera, Lepidoptera, Wasps and, Thus, unlike some of the basal holometabolan orders, the of course, Coleoptera – are also holometabolous. Among Hymenoptera have a relatively recent origin, first appearing holometabolans, Hymenoptera is perhaps the most difficult in the Late Triassic. Since the Triassic, the Hymenoptera have to place in a phylogenetic framework, excepting the enig- truly come into their own, having radiated extensively in the matic twisted-wings, order Strepsiptera. Hymenoptera are Jurassic, again in the Cretaceous, and again (within certain morphologically isolated among orders of Holometabola, family-level lineages) during the Tertiary. The hymenopteran consisting of a complex mixture of primitive traits and bauplan, in both structure and function, has been tremen- numerous autapomorphies, leaving little evidence to which dously successful. group they are most closely related. Present evidence indi- While the beetles today boast the largest number of cates that the Holometabola can be organized into two major species among all orders, Hymenoptera may eventually rival lineages: the Coleoptera ϩ Neuropterida and the Panorpida. or even surpass the diversity of coleopterans (Kristensen, It is to the Panorpida that the Hymenoptera appear to be 1999a; Grissell, 1999). -
1 Universidade Federal Do Ceará Centro De Ciências
1 UNIVERSIDADE FEDERAL DO CEARÁ CENTRO DE CIÊNCIAS DEPARTAMENTO DE GEOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM GEOLOGIA LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL FORTALEZA 2019 2 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Orientador: Prof. Dr. Geraldo Jorge Barbosa de Moura. Coorientador: Prof. Dr. César Ulisses Vieira Veríssimo. FORTALEZA 2019 3 4 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Aprovada em: 18/01/2019. BANCA EXAMINADORA ________________________________________ Prof. Dr. Geraldo Jorge Barbosa de Moura (Orientador) Universidade Federal Rural de Pernambuco (UFRPE) _________________________________________ Prof. Dr. Marcio Mendes Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dr. Marcos Antônio Leite do Nascimento Universidade Federal do Rio Grande do Norte (UFRN) _________________________________________ Prof. Dr Kleberson de Oliveira Porpino Universidade do Estado do Rio Grande do Norte (UERN) ________________________________________ Dra Pâmela Moura Universidade Federal do Ceará (UFC) 5 A Deus. -
Hymenoptera: Symphyta): Evidence for Multiple Increases in Sperm Bundle Size
J. HYM. RES. Vol. 10(Z), 2001, pp. 119-125 Spermatodesmata of the Sawflies (Hymenoptera: Symphyta): Evidence for Multiple Increases in Sperm Bundle Size NATHAN SCHIFF, ANTHONY J. FLEMMINC, AND DONALD L. J. QUICKE (NS) U.S. Forest Service, Southern Research Station, Center for Bottomland Hardwoods Research, P.O. Box 227, Stoneville, MS 38776, USA; (AJF) Department of Biology, Imperial College at Silwood Park, Ascot, Berkshire SL5 7PY, U.K.; (DLJQ) Unit of Parasitoid Systematics, Department of Biology, Imperial College at Silwood Park, Ascot, Berkshire SL5 7PY, UK, and Department of Entomology, The Natural History Museum, London SW7 5BD, UK Abstract.-We present the first survey of spermatodesmata (bundles of spermatozoa connected at the head by an extracellular ‘gelatinous’ matrix) across the sawfly superfamilies. Spermatodes- mata occur in all examined taxa within the sawfly grade (Xyelidae-Orussidae inclusive), but are not found in the Apocrita. Using DAPI staining, the numbers of individual sperm per sperma- todesm were calculated and the values obtained are mapped on to the current phylogenetic hy- pothesis. The plesiomorphic spermatodesm in the Hymenoptera, based on that observed in the putatively basal family Xyelidae, contains relatively few sperm, approximately 16. However, in the Tenthredinoidea and in the Siricidae, far larger numbers are found, reaching up to 256 in the Cimbicidae. In many insects, mature sperm released Until now, spermatodesmata have only from testicular follicles are neither free in- been characterised in a few sawflies, al- dividuals nor packaged into variously most entirely as part of ultrastructural in- complex spermatophores, but are ar- vestigations using transmission electron ranged in organised bundles with their microscopy (Quicke et al. -
The Origins of Species Richness in the Hymenoptera: Insights from a Family-Level Supertree BMC Evolutionary Biology 2010, 10:109
Davis et al. BMC Evolutionary Biology 2010, 10:109 http://www.biomedcentral.com/1471-2148/10/109 RESEARCH ARTICLE Open Access TheResearch origins article of species richness in the Hymenoptera: insights from a family-level supertree Robert B Davis*1,2, Sandra L Baldauf1,3 and Peter J Mayhew1 Abstract Background: The order Hymenoptera (bees, ants, wasps, sawflies) contains about eight percent of all described species, but no analytical studies have addressed the origins of this richness at family-level or above. To investigate which major subtaxa experienced significant shifts in diversification, we assembled a family-level phylogeny of the Hymenoptera using supertree methods. We used sister-group species-richness comparisons to infer the phylogenetic position of shifts in diversification. Results: The supertrees most supported by the underlying input trees are produced using matrix representation with compatibility (MRC) (from an all-in and a compartmentalised analysis). Whilst relationships at the tips of the tree tend to be well supported, those along the backbone of the tree (e.g. between Parasitica superfamilies) are generally not. Ten significant shifts in diversification (six positive and four negative) are found common to both MRC supertrees. The Apocrita (wasps, ants, bees) experienced a positive shift at their origin accounting for approximately 4,000 species. Within Apocrita other positive shifts include the Vespoidea (vespoid wasps/ants containing 24,000 spp.), Anthophila + Sphecidae (bees/thread-waisted wasps; 22,000 spp.), Bethylidae + Chrysididae (bethylid/cuckoo wasps; 5,200 spp.), Dryinidae (dryinid wasps; 1,100 spp.), and Proctotrupidae (proctotrupid wasps; 310 spp.). Four relatively species-poor families (Stenotritidae, Anaxyelidae, Blasticotomidae, Xyelidae) have undergone negative shifts. -
Ebook Download the Life Cycle of an Ant Ebook, Epub
THE LIFE CYCLE OF AN ANT PDF, EPUB, EBOOK Hadley Dyer,Bobbie Kalman | 32 pages | 28 Feb 2006 | Crabtree Publishing Co,Canada | 9780778707004 | English | New York, Canada The Life Cycle of an Ant PDF Book Worker Ants Worker ants have no wings and typically survive several months with some exceptions of workers who can live up to seven years. Some ant species, such as those that use natural cavities, can be opportunistic and make use of the controlled micro-climate provided inside human dwellings and other artificial structures to house their colonies and nest structures. Back to previous page. They also are mentioned in religious texts. Like ants, termites are eusocial , with sterile workers, but they differ greatly in the genetics of reproduction. A wide range of reproductive strategies have been noted in ant species. In Central and South America, Eciton burchellii is the swarming ant most commonly attended by " ant- following " birds such as antbirds and woodcreepers. In South Africa , ants are used to help harvest the seeds of rooibos Aspalathus linearis , a plant used to make a herbal tea. Since the weather is warmer out it is a great time for children to observe ants firsthand. Austrocynipidae Cynipidae gall wasps Figitidae Ibaliidae Liopteridae. Their success in so many environments has been attributed to their social organisation and their ability to modify habitats, tap resources, and defend themselves. Journal of Experimental Biology. The American Naturalist. The marked male is interpreted as an invader by worker ants and is killed. However, they need only a couple hours for their exoskeletons to harden and for them to get stronger and change their appearance by getting darker. -
New Fossil Records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China
European Journal of Taxonomy 733: 146–159 ISSN 2118-9773 https://doi.org/10.5852/ejt.2021.733.1229 www.europeanjournaloftaxonomy.eu 2021 · Zheng Y. et al. This work is licensed under a Creative Commons Attribution License (CC BY 4.0). Research article urn:lsid:zoobank.org:pub:043C9407-7E8A-4E8F-9441-6FC43E5A596E New fossil records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China Yan ZHENG 1,*, Haiyan HU 2, Dong CHEN 3, Jun CHEN 4, Haichun ZHANG 5 & Alexandr P. RASNITSYN 6,* 1,4 Institute of Geology and Paleontology, Linyi University, Shuangling Rd., Linyi 276000, China. 1,4,5 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, East Beijing Road, Nanjing 210008, China. 2 School of Agronomy and Environment, Weifang University of Science and Techonoly, Jinguang Road, Shouguang, 262700, China. 3 School of Environmental and Municipal Engineering, Qingdao University of Technology, Qingdao 266033, China. 6 Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia. 6 Natural History Museum, London SW7 5BD, UK. * Corresponding authors: [email protected], [email protected] 2 Email: [email protected] 3 Email: [email protected] 4 Email: [email protected] 5 Email: [email protected] 1 urn:lsid:zoobank.org:author:28EB8D72-5909-4435-B0F2-0A48A5174CF9 2 urn:lsid:zoobank.org:author:91B2FB61-31A9-449B-A949-7AE9EFD69F56 3 urn:lsid:zoobank.org:author:51D01636-EB69-4100-B5F6-329235EB5C52 4 urn:lsid:zoobank.org:author:8BAB244F-8248-45C6-B31E-6B9F48962055 5 urn:lsid:zoobank.org:author:18A0B9F9-537A-46EF-B745-3942F6A5AB58 6 urn:lsid:zoobank.org:author:E7277CAB-3892-49D4-8A5D-647B4A342C13 Abstract. -
Big and Beautiful: the Megaxyela Species (Hymenoptera, Xyelidae) of East Asia and North America
European Journal of Taxonomy 348: 1–46 ISSN 2118-9773 https://doi.org/10.5852/ejt.2017.348 www.europeanjournaloftaxonomy.eu 2017 · Blank S.M. et al. This work is licensed under a Creative Commons Attribution 3.0 License. Research article urn:lsid:zoobank.org:pub:56D8BC29-3C29-4EE8-8633-B997784CA00A Big and beautiful: the Megaxyela species (Hymenoptera, Xyelidae) of East Asia and North America Stephan M. BLANK 1,*, Katja KRAMP 2, David R. SMITH 3, Yuri N. SUNDUKOV 4, Meicai WEI 5 & Akihiko SHINOHARA 6 1,2 Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Müncheberg, Germany. 3 Systematic Entomology Laboratory, Agricultural Research Service, U.S. Department of Agriculture, c/o National Museum of Natural History, P.O. Box 37012, MRC 168, Washington, DC 20013-7012, USA. 4 State Nature Reserve “Kurilskiy”, Zarechnaya str. 5, Yuzhno-Kurilsk, Sakhalinskaya oblast, 694500 Russia. 5 Key Laboratory of Cultivation and Protection for Non-Wood Forest Trees (Central South University of Forestry and Technology), Ministry of Education, Changsha 410004, China. 6 Department of Zoology, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba, Ibaraki, 305-0005 Japan. * Corresponding author: [email protected] 2 Email: [email protected] 3 Email: [email protected] 4 Email: [email protected] 5 Email: [email protected] 6 Email: [email protected] 1 urn:lsid:zoobank.org:author:0E65D322-6E6B-489E-9243-921D28E0472D 2 urn:lsid:zoobank.org:author:72579D81-D3A8-4803-91AC-4C928C83EBEB 3 urn:lsid:zoobank.org:author:B25C3A30-9EF6-4561-8DCE-C95869DFD7E8 4 urn:lsid:zoobank.org:author:0D46A0A2-6555-4045-A276-B9539EC54088 5 urn:lsid:zoobank.org:author:E687AE23-588C-4866-8C14-19B3112BB4BA 6 urn:lsid:zoobank.org:author:C7382A9B-948F-479B-BEE7-848DAFECD3BA Blank S.M., Kramp K., Smith D.R., Sundukov Y.N., Wei M. -
Zootaxa,Phylogeny and Classification of Hymenoptera
Zootaxa 1668: 521–548 (2007) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2007 · Magnolia Press ISSN 1175-5334 (online edition) Phylogeny and Classification of Hymenoptera* MICHAEL J. SHARKEY S-225 Ag. Sci. Building-N, Department of Entomology, University of Kentucky, Lexington, KY 40546-0091, USA [email protected] *In: Zhang, Z.-Q. & Shear, W.A. (Eds) (2007) Linnaeus Tercentenary: Progress in Invertebrate Taxonomy. Zootaxa, 1668, 1–766. Table of contents Abstract . 521 Introduction . 521 Hymenopteran Diversity . 522 Phylogenetic Position Within Holometabola . 523 Monophyly of Hymenoptera . 523 Review of Classical Hymenopteran Classification and Phylogeny . 524 Phylogenetic (Cladistic) Studies of Hymenoptera . 528 Symphytan Studies . 532 Apocritan Analyses . 534 Morphologcial Character Systems . 536 What We Think We Know . 536 Best Guess . 539 Current and Future Research . 539 Key Innovations and the Evolution of life history traits, a phylogenetic approach . 539 Acknowledgements . 543 References . 543 Abstract An overview of recent advances in our understanding of the higher phylogeny and classification is presented. Molecular and morphological cladistic and pre-cladistic studies are summarized. A superfamily-level classifi- cation of the Hymenoptera is offered to reflect recent advances in our understanding of the phylogenetic rela- tionships of the Hymenoptera. It differs from most recent classifications in the recognition of the Diaprioidea, to include Diapriidae, Monomachidae, and Maamingidae. Key words: Diaprioidea, taxonomy, cladistics, life history, Insecta Introduction Much progress has been made in our understanding of the phylogeny of Hymenoptera since the advent of cla- distic methods. Here I summarize recent influential studies and pre-cladistic studies are also treated, at least cursorily. -
Hymenoptera of Canada 311 Doi: 10.3897/Zookeys.819.28510 RESEARCH ARTICLE Launched to Accelerate Biodiversity Research
A peer-reviewed open-access journal ZooKeys 819: 311–360 (2019) Hymenoptera of Canada 311 doi: 10.3897/zookeys.819.28510 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Hymenoptera of Canada Andrew M.R. Bennett1, Cory S. Sheffield2, Jeremy R. deWaard3 1 Canadian National Collection of Insects, Arachnids and Nematodes, K.W. Neatby Bldg., 960 Carling Ave- nue, Ottawa, Ontario, K1A 0C6, Canada 2 Royal Saskatchewan Museum, 2340 Albert Street, Regina, Sa- skatchewan, S4P 2V7, Canada 3 Centre for Biodiversity Genomics, University of Guelph, 50 Stone Road East, Guelph, Ontario, N1G 2W1, Canada Corresponding author: Andrew M.R. Bennett ([email protected]) Academic editor: D. Langor | Received 20 July 2018 | Accepted 21 November 2018 | Published 24 January 2019 http://zoobank.org/3C7E5BD2-1B3E-4B06-ACC4-9A2C83EF3FF4 Citation: Bennett AMR, Sheffield CS, deWaard JR (2019) Hymenoptera of Canada. In: Langor DW, Sheffield CS (Eds) The Biota of Canada – A Biodiversity Assessment. Part 1: The Terrestrial Arthropods. ZooKeys 819: 311–360. https://doi.org/10.3897/zookeys.819.28510 Abstract A summary of the numbers of species of the 83 families of Hymenoptera recorded in Canada is provided. In total, 8757 described species are recorded compared to approximately 6000 in 1979, which is a 46% increase. Of the families recognized in 1979, three have been newly recorded to Canada since the previous survey: Anaxyelidae (Anaxyleoidea), Liopteridae (Cynipoidea), and Mymarommatidae (Mymarommatoidea). More than 18,400 BINs of Canadian Hymenoptera are available in the Barcode of Life Data Systems (Ratnasing- ham and Hebert 2007) implying that nearly 9650 undescribed or unrecorded species of Hymenoptera may be present in Canada (and more than 10,300 when taking into account additional species that have not been DNA barcoded).