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San Ramón, Alajuela, of the San Ramón Cloud Forest 1 Brayan H. Morera-Chacón Gestión de los Recursos Naturales, Universidad de Costa Rica

All photos by Brayan H. Morera-Chacón Produced by: Brayan H. Morera-Chacón © Brayan H. Morera-Chacón [[email protected]]. Thanks to: César L. Barrio-Amorós (Doc Expeditions). (M) Male, (F) Female and (A) [fieldguides.fieldmuseum.org] [855] version 1 01/2017

1 coniferus (M) 2 3 Incilius melanochlorus 4 horribilis

BUFONlDAE BUFONlDAE BUFONlDAE BUFONlDAE

5 Cochranella granulosa 6 prosoblepon (M) 7 Espadarana prosoblepon (F) 8 Sachatamia ilex

CENTROLENIDAE CENTROLENIDAE CENTROLENIDAE CENTROLENIDAE

9 Teratohyla pulverata 10 bransfordii 11 12

CENTROLENIDAE CRAUGASTORIDAE CRAUGASTORIDAE

13 Craugastor fitzingeri 14 C.crassidigitus (above); C. fitzingeri 15 16 Craugastor podiciferus (below) CRAUGASTORIDAE CRAUGASTORIDAE CRAUGASTORIDAE CRAUGASTORIDAE

17 Craugastor stejnegerianus 18 altae 19 Pristimantis caryophyllaceus 20 (M)

CRAUGASTORIDAE CRAUGASTORIDAE CRAUGASTORIDAE CRAUGASTORIDAE San Ramón, Alajuela, Costa Rica Amphibians of the San Ramón Cloud Forest 2 Brayan H. Morera-Chacón Gestión de los Recursos Naturales, Universidad de Costa Rica

All photos by Brayan H. Morera-Chacón Produced by: Brayan H. Morera-Chacón © Brayan H. Morera-Chacón [[email protected]]. Thanks to: César L. Barrio-Amorós (Doc Frog Expeditions). (M) Male, (F) Female and (A) Amplexus [fieldguides.fieldmuseum.org] [855] version 1 01/2017

21 Pristimantis cruentus (F) 22 Pristimantis cruentus (M) 23 Pristimantis cruentus (F) 24 (A)

CRAUGASTORIDAE CRAUGASTORIDAE CRAUGASTORIDAE CRAUGASTORIDAE

25 Pristimantis ridens 26 diastema 27 Diasporus hylaeformis 28 Agalychnis callidryas

CRAUGASTORIDAE ELEUTHERODACTYLIDAE

29 ebraccatus 30 Dendropsophus ebraccatus 31 Duellmanohyla rufioculis 32 Duellmanohyla rufioculis

HYLIDAE HYLIDAE HYLIDAE HYLIDAE

33 Duellmanohyla rufioculis (A) 34 Isthmohyla pseudopuma 35 Scinax elaeochrous 36 Smilisca phaeota

HYLIDAE HYLIDAE HYLIDAE HYLIDAE

37 Smilisca sordida 38 Smilisca sordida (A) 39 savagei 40 Lithobates warszewitschii (M)

HYLIDAE HYLIDAE RANIDAE

San Ramón, Alajuela, Costa Rica Amphibians of the San Ramón Cloud Forest 3 Brayan H. Morera-Chacón Gestión de los Recursos Naturales, Universidad de Costa Rica

All photos by Brayan H. Morera-Chacón Produced by: Brayan H. Morera-Chacón © Brayan H. Morera-Chacón [[email protected]]. Thanks to: César L. Barrio-Amorós (Doc Frog Expeditions). (M) Male, (F) Female and (A) Amplexus [fieldguides.fieldmuseum.org] [855] version 1 01/2017

41 Lithobates warszewitschii (F) 42 Lithobates warszewitschii (A) 43 gamezi 44 Oedipina uniformis

RANIDAE RANIDAE PLETHODONTIDAE

The cloud forest of San Ramón is located in the mountains near the continental divide at an elevation of between 1100 and 1500 meters above sea level. The herpetofauna of this forest has been little studied, with whose is not clear as well as species under threat of Map of Costa Rica showing the San Ramón area extinction.

Map of San Ramón showing the cloud forest

Brayan H. Morera-Chacón is a manager of natural resources with a special interest in amphibians and reptiles. In his few years of research experience he has dedicated his work to the premontane forests of San Ramón.