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(8861)l,rt'60t i(t)9 ersl^nN 310 NuytsiaVol. 6, No. 3 (1988)

Typus:Banhsi.a blueri R. Br. In lhesystematic sequence in.my rev.ision,t.he species mal retainits posilion berween seriesOuercinop s. qlr.and serles Url,'los/y1,s {uenln., A.). L'eorge' Banksia epica A.S. George,sp. nov. (series Cyrtostylis)(Figure 1) Affinis B. praemorsaeAndrews et B mediaeR Br., a quibus perianthio majore et praebitore pollinis longiore praecipuediffert. Folia obovataad angustecuneata, 15 50 mm longa,m-arginibus planis vel leviter recu*is, b.eniterserratis. Perianthium'10 44 mm longumlimbum 4.5-5'5mm longum i""lud""", infra glabrum, supra extus pubescenslimbo g-labro.Pistillum 39-49 mm ioneu-. elabr"m;praebitor pbllinis t.5 1.8mm longus Folliculiubi exserti13 20 mm io"Ei. l-iO -,'' alti, 6-9 mn; Iali, colliculati,parce hirsuti; floresveteres persistentes, stylibus curvatis. ?vpus. Point Culver, Great Australian Bight, Western Aqstrql!1-c, 32" 55'5,724" 42' nlt frl"u 1986.J. Filconer s.n. (holo:PERTH; iso: CANB, MEL).

Fig'ure 1. BonAsia epica A. S. George holotype Bar scale is 5 cm -f

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tlL: P^-rarrsgr8rort S V (1988) ;il2 \uvtsia Vol. 6, No.:l

When the Banksia Atlas (a joint project of the Australian Biolo-gicalResources Study and the Western Australian'Department of Conservation and Land Management) beean in 1SSa,a request was mad; for participants to collecl good specim-en-s.John al-r-d iaiage Falcone. o1'Esperance,Westein Auslralia, made two trips in 1985 and 1986' John- eventually *u. ibl" to collect tlowering material a1t-era solo expedition irom W;;;tt"" (wliere they were then stati,,nedl across lhe Great Victoria Desert and tlutlu.bot ptii". It is in;ecognition of Eyre's epicjourney of 1841and the Falconers'ef forts to collect specimensthat the new speciesis named eplco. Banksia oligantha, A. S. George,sp. nov (subgenuslsosiy.lrs) (Figure 2)' Soeciesinter L i.Liciftti'amR. Br. et B cuneatam.A. S Georgeintermedia. sed ab illa foliis et floribus minoribus, ab hac lbliis nitentibus concavioribus et floribus parum minoribus. et ab ambabusinflorescentia pauciflora, differt. Foliarum lamina 1'5 3 ? cm longa; perianthium 21 23 mrrr longrtm; inflorescentia 20 ll5-flora

& Figure2. Barlrsia oligarljla A. S. George holotype.

?\ ous;Nature Reserve 9098, 28 km NW of Wagin,Western Australia, 33' 10'S, 117'04' Fli't8 Nov. 1981.A.'l'avlor s.n. (holo:PER'I'H; iso: CANB, K, NSW). 'JncJo aurll bulra,uou puE rpalou azrs ur ?lBrpouralu! sruJol l.Bql'Ja^a,uoq aq lsnlll lI suosPasluaraJJlp lP rar\ol' pLrecd8uer tPJlqdeJiodE luala;1rp {dn;..ro,{aq1 arual'arogul puP sJd.\\ollJo dzrsuo .illelrxoloqcliourpa leredas aq ue; ! luelJeAd:dq I suIJollEJld.{ IJ Idq I ul luELl€^xulJJrAu[t jirurnrite-. aqi.{11eur.rr1dtuEuolIn.J ,Ltrdq Pln'r^\ ll JdplsuuJ^\ou I luelJl]\6ulJd^\')Ll -.reruulni' '1ertdi1 aLitrelnir t.redrrli1a11 oql J r -ar.rarlssiq t pdulure\d-rJ 1r'ru Sutre;1

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Map 1. Distribution of Banhsia oli,gantha(^), B. epica (.), B. IeptophyllavaL melteticd(.) and.B. spinulosa vaf. neoanquco\t),

Banksia leptophylla A.S. Georgevar. leptophylla Perianth 42-47 mm long. Pistil 56-62 mm long. Flowers mainly in summer. Distribution. Occursbetween Tathra National Park arld Mogumber,Western Australia (Map 1). Habitat. On sandy rises in shrubland. Flowering period. Summer to autumn.

Banksia leptophylla var. mellitica A.S. George,var. nov. Ab Banhsia leptophyLlavar. leptophylla floribus minoribus (perianthiis 80-36 mm longo,pistillo 34-44mm longo).hieme florentibus, differt. ?yp-us:.c.^13 km N of Gingin.tur_lgff,Perth-Lancelin road, Western Australia, 10 June 1966,A. S. George7767 (holo: PERTH; iso: CANB, K, MEL, NSW). Distribution. Occursbetween the lower Murchison River and Guilderton.Western Aus- tralia, mostly within 30 km of the coast (Map 1). Habitat..ln -deepsand. in sarrdover limestone and in depressions(not permanently damp), in shrubland. Often locally common. Flowering period. Winter. 'BxE? snoraqnl.ou.SrT-uouqlr,$ aJ€ds ldau JoJSutladuroJ ur a8€luE^p€srp B l€ ar" raqlo aq1 uo lnq'oJrJ l.surP;B pJenSaJESe padola^op a^€q {aq] pu€q auo aqJ.uo snqJ slupld pxel 'arU r$au Jo Juaurltnr)ar alllrl 8uI^Eq ur alqEls JeaddE qrns Jo suorlBlndod lnq ()1 asuodsal€ {lluapr^a sr sJupld usrlprlsnv ur Jaqnlou8lTaql Jo uorlnlo^a aqJ slupld snoraqnlouSrl-uou Io asoql u€rll dl{Jrnb ssal qf,nul ,{\oJF{ll€nsn ^aq} a^l^rns op slueld ',\\ol snoraqnl.ou8rl Jo sturlpaas uaq^\ sr poas Jo uorlcnpord aql ( rg U pdJD)onDqds Dls4uDg'E a) ?x€l auros ur pup'plr,u aqJ ur arr?JaJe slu€ld snoJaqn?ou;rllo sSuqpaas 'paasruoq.{luo sa}BrauaBarpup pa[r{ sr Jaq}oaql alq,\\ sI ulo4 slnords uaq] 'arg {Jolsloor ur.ro;snoraqnlou8rT aqt lnq ,{q pallr{ arp srlrEdpunorB a^oqB aql aspr q)ea uI (e6t d 'A, 8IJ 'I86I ?8roat aas 3 a) rall€t qcnu sd\orgpue ulals alSurs€ seq dllEnsnraqntou8q .lueld B lnoqllr auo s€aJaqd\surals urBur lBralas seq ^ll€nsn Jaqnlou8ll e qll^\ auO € Jo ruroJ ql$oJ8 punod-a^oqe aql ul palcaUarsr raqnlou8rl e fo oJuasqBJo acuasardaqJ 'uorlnqrrlsrpJo rpuEIalqBJ arluaD aql pupl5ug 'ldaNaq? ol sratar laqlrda IelauBA aqJ 'uox€t aql asru8orar fllEurJoJ 01 suaas 111'tLuD113u1uunJ 'JE,\ InJasn uro{ paqsrn-8urlsrp aq louu€c ,{aql ^Soloqdroru pup a^rlPla8a^ ur qFnoqlly 'Jaqnlou8rl l€rou E Sul^pq ur ]ua]srsuoc ar€ puPlsuaano uJalspa-qlnos pue salel qlnos d\aN urarlFou ur uuaq8uluun) J€A ur papnlcur .{lsnouaJd suorlplndod lBq} paurJrJ 'slu€dlJrlipd 'suorJ?olloJ -uo) a^Bq sEIIV r{q j{ll€rJadsa puE suorlE^rasqoluaraE (l6g prsl'{nN) 96t :t s.rolJEJEq)JEal puE runluaulnpur ,{q pue raqnlou8rl e 1o aJuasqe aql ,{q pasualJ€reqJ sr ?rruDrlduTuun) f,E^Dsolnulds D,tslaD€r'salErpaurralul ql+\ aslJe uolleJg -rluapr 'osp 'pasruSoDaJ Jo suralqoJd?nq sarlauE^aql ^ll€nsn puE ,{ ppal srJlasl.r sorJads aqJ Ia^al lBlauEA aql lp s?tc?os auo ulqllr paur€lal lsaq aJe ex€l asoql ?€ql pauJrJ uoc s€q's€lly €rs{uBg oql ur sluBdrJrlJBdfq ,{llBrJadsa'uorlpJrlrss€p srrll Sursn ul af,uauadxg Dso?ru|d.^DlsTuDg urqT ^\ sarlaue^ aaJql pasru8of,aJI uorsuar Ig6I .{trl uI

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'uox€l aqlJo snlEls oqlJo^o uorlEJaqqapToporradSuol aqJ ol aJualofoJur,(Suroplo 'op {urql dluo ol pua}ur ll1) ollaur qral {aarC aql ruoq pa^rrap sr laqtrda 1e1arre,raqj 'aJrI Jalle paas ur04 {lsnoJo:r^ se1e.raue.8a.r pup saAJasoJuorJBAJasuo, Iera^as ur sJntf,o,{lou€AaqJ'arEJ loN snJpJSua4palasuoi) tIl P/srfuPg airoa) S.V 316 NuvtsiaVol. 6, No. 3 (1988)

^ Three other species of BonAslo appe_arto show rhis divergence of habitat but require f^urther st-udy. They are B. ashbyi'8.L. Burtt, B. marginita Cav. and B. uiolaiia C. Gardner.The situation alsoexists in somespecies of the"genusDryanjro ,..g. i.;;";. R. Br., D. /raserl R. Br.

Presumed natural hybrid While recording Banksias for the Banksia Atlas, Mr G. Schmidt of Kalamunda, West em Audralia. d.iscovered_a-plant which appears to be a hybrid between B. prionotes LinctleyandB. /indlej cna Meissner.In most charactersit is intermediate betwien these twn. both of which were present witlr the presumedhybrid and which belong to closely relatedseries within the genus.The leaves,however, are similarto those ofBlprionotei. _ Spreading shrub 2.5 m tall. Leaves to 22 cm long, 13 15 mm wide; margins ilat, shallowly lobed almost to base; Iobes t triangular, to 2 mm high, obtusely and shortly mucronate. Inflorescence12 13 cm long. Perianth 3? mm tong'includinglimb of 8 md; claws closelytomentose and shortly hirsute; limb minutelv pubescent.Pistil 40-45 mm long, bowed, glabrous;pollen presenter c.3.5 mm long, swollen at base, fusiform and ribbed above. OId flowers persistent on infructescence, the pistils wiry. Follicles elliptic in plan view, 12 13 mm long, 7 8 mm wide, 4 mm high, shortly beakedat stylar point, denselv hirsute.

Specimen examined. 4.5 km N of double gate in State Barrier fence, N edse of Murchison House Stn, Western Australia, 27" 34' S, 114'03'E, 8 Sept. 1985,G. Scdmldt (PERTH). A singleplant ofthe presumedhybrid was sighted,and the description is basedon two fruiting specimens,probably developedfrom the previous season'silowering. The per--has sistent perianths and pislils have retained their morphology. Each infl orescence produced a number of follicles that are fully grown but were probably not fully mature when collected. Only two other presumed natural hybrids in Banlisla in Western Australia have been report€d, one brtween B. hctokerianaMeissner and B. prionotes (Keighery 1985),the other betweenB. hooheri.clneandB. menzjeslj R.Br. (Dixon 1986).In Contiast, naiural hybrids appear to be frequent in eastern Australia.

Further names in Bqnksia .. The following names were fourd by Mr Arthur D. Chapman while searching literature lbr entries for the Australian Plant Name Index. None aflects the accepted nomencla- ture of Bonftslo. Banksia lamberti Hort. ex Courtois, Magasin d'Horticulture, Suppl. 1; 295 (1883). Type citation: none given. No_sheet bearing this name has been seen. From the description it appears that this may be synonymous rith B. spinuLosaSmith var. cunningEamii (Sieber ex Reichb.) A.S. Georse.

- Banhsia intermedla Sweet ex Courtois, Magasin d'Horticulture, Suppl. l: 295 (1g33); Banhsia.intermedjo Sweet,Hort. Brit., 2nd ecin,2;849 (182?), "in nomernudum. ?1pe cj- tation: Nova Hollandia [Australia], introduced to Britain in 1824,,(Sweeti' No sheet bearing tl.ris name has been seen. The description is of leaves only and is in sufficient to allow definite application of the name. It is possibly a synonym of B. oou)n{tloll0 l-av. Banhsia langifolia van.pubescens (Willd.) Breiter, Hortus Breiterianus 2g2 (1817). is based pubescens .,This on Co

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