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Xerox University Microfilms 300 North Zeeb Road Ann Arbor, Michigan 48106 76-3520 INFORMATION TO USERS This material was produced from a microfilm copy of the original document. While the most advanced technological means to photograph and reproduce this document have been used, the quality is heavily dependent upon the quality of the original submitted. The following explanation of techniques is provided to help you understand markings or patterns which may appear on this reproduction. 1. The sign or "target" for pages apparently lacking from the document photographed is "Missing Page(s)". If it was possible to obtain the missing page(s) or section, they are spliced into the film along with adjacent pages. This may have necessitated cutting thru an image and duplicating adjacent pages to insure you complete continuity. 2. When an image on the film is obliterated with a large round black mark, it is an indication that the photographer suspected that the copy may have moved during exposure and thus cause a blurred image. You will find a good image of the page in the adjacent frame. 3. When a map, drawing or chart, etc., was part of the material being photographed the photographer followed a definite method in "sectioning" the material. It is customary to begin photoing at the upper left hand corner of a large sheet and to continue photoing from left to right in equal sections with a small overlap. If necessary, sectioning is continued again — beginning below the first row and continuing on until complete. 4. The majority of users indicate that the textual content is of greatest value, however, a somewhat higher quality reproduction could be made from "photographs" if essential to the understanding of the dissertation. Silver prints of "photographs" may be ordered at additional charge by writing the Order Department, giving the catalog number, title, author and specific pages you wish reproduced. 5. PLEASE NOTE: Some pages may have indistinct print. Filmed as received. Xerox University Microfilms 300 North Zeeb Road Ann Arbor, Michigan 48106 76-3520 PASCHQAL, Adi Ison Dias, 1941- A REVISION OF TOE GYMNODAMAEIDAE (ACARI - ORIBATEI): A NUMERICAL TAXONCMIC STUDY. The Ohio State University, Ph.D., 1975 Zoology Xerox University Microfilms, Ann Arbor, Michigan 4sioe © 1975 ADILSON DIAS PASCHQAL ALL RIGHTS RESERVED THIS DISSERTATION HAS BEEN MICROFILMED EXACTLY AS RECEIVED. A REVISION OF THE GYMNODAMAEIDAE (ACARI - ORIBATEI): A NUMERICAL TAXONOMIC STUDY DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By Adilson Dias Paschoal Doctor in Agronomy, University of Sao Paulo, Brazil * * * * * * * The Ohio State University 1975 Reading Committee: Approved by Prof. Donald E. Johnston Prof. Tony J. Peterle Prof. George ¥. Wharton Ldviser Department of Zoology DEDICATORY # A l A t Aik A A A AIk AIk AIk AIk AIk AIk AIk AIk This work is dedicated, with honor, to the memory of FRANCOIS GRANDJEAN, deceased on January 22, 1975. it it it it it it it it it it it it it it , 0 It is also dedicated, with hope, to my son ANDRE, born in this country on April 20, 197 3. ACKNOWLEDGMENTS I wish to express my appreciation to those persons 9 who have intellectually or materially contributed to this work and to those institutions which provided scientific materials and grants-in-aid. I am particularly grateful to my adviser, Prof. Donald E. Johnston, for his constant enthusiasm for my work and for his valuable assistance, suggestions and criticism during all stages of the preparation of this dissertation. I am equally thankful to Prof. George W. Wharton, Director of the Acarology Laboratory, for having accepted me in his Laboratory and for his significant contribution to my scientific development. Grateful acknowledgment is made to Dr. Roy A. Norton, State University of New York, who sent his entire Gymnodamaeidae collection for my study; Dr. Edward W. Baker, USDA, for his assistance during my visit to the Acari collection of the USNM, and for the loan of type species; Dr. Herbert W. Levi, Harvard University, for the loan of type materials and for his assistance during my visit to the Museum of Comparative Zoology; Dr. Norman Platnick, for his assistance during my visit to the American Museum iii of Natural History; Dr. Evert E. Lindquist, Canada Department of Agriculture; Dr. G. F. Knowlton, Utah State University; Dr. Joseph Trave, Laboratoire Arago; Dr. S. Mahunka, Zoologische Abteilung des Ungarischen National 9 Museums; Dr. John A. Wallwork, University of London; and Dr. Carlos Perez-Inigo, Instituto Espanol de Entomologia, for providing me with important specimens for this revision. Acknowledgment is also made to Prof. Tony J. Peterle, Chairman of the Zoology Department, The Ohio State University, for providing funds for computer time and for photographic materials. Mr. Guilford S. Ide, curator of the Acarology Laboratory helped me in many aspects of the laboratory work and I am very grateful to him. This investigation was supported by the Agency for International Development, through the contract USDA-OSU- ESALQ, and also by the "Conselho Nacional de Pesquisas" (CNPq, Rio de Janeiro) and "Coordenadoria do Aperfeicoamento de Pessoal de Nivel Superior” (CAPES, Brasilia). VITA November S, 1 9 4 1 ........ Born - Casa Branca, S. Paulo, Brazil » 1963-1967.. ............... University of Sao Paulo, Superior School of Agricul­ ture "Luiz de Queiroz", Piracicaba 196 4 ..................... Scholarship from the Ministry of Agriculture of Brazil 1966-1969.. ............... Research Assistant, Department of Zoology, University of Sao Paulo, Piracicaba. Professor of Physical and Biological Sciences of a State College of Education, Piracicaba 1967 ..................... Agricultural Engineer, Univer­ sity of Sao Paulo, Piracicaba 196 5 ..................... Member of the "Sociedade Brasileira para o Progresso da Ciencia" (SBPC), Brazil 196&-1969 ............... Graduate Scholarship from the "Conselho Nacional de Pesquisas" (CNPq), Rio de Janeiro, Brazil. Instructor of the Department of Zoology of the Agronomy and Veterinary School of Jaboticabal, S. Paulo. M.S. course on Ento­ mology, University of Sao Paulo, Piracicaba 1969-1970 Instructor of the Department of Zoology, University of Sao Paulo, Superior School of Agriculture "Luiz de Queiroz", Piracicaba v VITA, Continued 1970 ..................... Member of the "Associacion Latinoamericana de Fitotecnia", Bogota, Colombia * 1970 ..................... Doctor in Agronomy, University of Sao Paulo, Piracicaba 1970-present Assistant Professor of the Department of Zoology, Univer­ sity of Sao Paulo, Piracicaba 1971 ..................... Charter member of the Acarologi- cal Society of America, USA. Member of the "Sociedade Brasileira de Fruticultura" and "Sociedade Brasileira de Entomologia", Brazil 1972-1975 ............... Scholarship from the contract USDA-OSU-Superior School of Agriculture "Luiz de Queiroz", at the Ohio State University, Department of Zoology, Acarology Laboratory, Columbus, Ohio. Scholarship from CAPES, Rio de Janeiro, Brazil, at the Ohio State University 1974-1975 ............... Scholarship from CNPq, Rio de Janeiro, Brazil, at the Ohio State University 1975 .............Member of the Entomological Society of America and the Ohio Academy of Sciences PUBLICATIONS Paschoal, A. D. 1967. "Some phytophagous mites and their hosts in the State of Sao Paulo." (in Portuguese) Rev. Agric., Piracicaba, 42(4):146. vi VITA, Continued PUBLICATIONS Paschoal, A. D. 1967. "Mites on citrus plants." (in Portuguese). Solo, Piracicaba, 59(£):53-56. ______________ . 1963. "On the biology of Tetranychus mexicanus (Acarina, Tetranychidae). Previous notes." (in Portuguese). Solo, Piracicaba, 60(l):67-70. 1963. "Mites on plants in the State of Sao Paulo." (in Portuguese). Rev. Agric., Piracicaba 54(2-3):76-7B. _______ . 1963, "A fruit-plant parasitic mite, TetranycTius mexicanus (Acarina, Tetranychidae)." (in Portuguese). Solo, Piracicaba, 60(2):75-77. Reis, P. R. and A. D. Paschoal, 1963. "Some storage- product mites in the State of Sao Paulo." (in Portuguese). Solo, Piracicaba, 60(2):73-74» Paschoal, A. D. 1969. "Mononychus spp. (Acarina, Tetranychidae) causing damage to cassava in the State of Sao Paulo." (in Portuguese). Cienc. Cult. 20(2); 334. ______________ . 1969. "Description of the male of Allonychus braziliensis (Acarina, Tetranychidae)." (in Portuguese). Cienc. Cult. 20(2):333» 1969. "Tetranychus marianae (Acarina, Tetranychidae), harmful to solanaceous plants of Brazil." (in Portuguese). Soc. Bras. Ent., Anals I Meeting:33-34. ______________ . 1969. "List of mites found on plants in Brazil." (in Portuguese). Rev. Agric., Piracicaba, 43(3-4):137-139* ___________ and R. P. L. Carvalho, 1969. "Chemical control of the mite Tetranychus evansi (Acarina, Tetranychidae) on eggplants." (in Portuguese). Solo, Piracicaba, 61(1):75-77. VITA, Continued PUBLICATIONS Paschoal, A. D. 1970. "Contribution to the knowledge of the family Tetranychidae (Acarina) *in Brazil." (in Portuguese). Doctoral Dissertation, University of S. Paulo, ll6p., 7 fig. 1970. New Brazilian spider mites (Acarina, Tetranychidae). Anais E.S.A. "Luiz de Queiroz" 27:439-455. 1970. "Revision of the family Tetranychidae in Brazil (Arachnida, Acarina)." (in Portuguese). Anais E.S.A. "Luiz de Queiroz" 27: 457-483. 1971. "The Tetranychus telarius complex in Brazil." (in Portuguese). Rev. Agric.,"Piracicaba, 46(1 ):3-8 . _______________. 1971* A revision of the Caribbeanae group (Acarina, Tetranychidae). First Latin-Amer. Congr Entomol.,
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