Export of a Checklist of the Terrestrial Arthropods of Alaska
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Export of a checklist of the terrestrial arthropods of Alaska Derek Sikes Carolene Coon Matthew Bowser December 17, 2009 i Contents Contents ii Acknowledgments iv 1 Introduction 1 2 List of Species 1 Arachnida . 1 Acari .............................................. 1 Araneae . 10 Opiliones . 26 Pseudoscorpiones . 26 Chilopoda . 27 Geophilomorpha . 27 Lithobiomorpha . 27 Scolopendrida . 28 Crustacea . 28 Anostraca . 28 Cladocera . 28 Copepoda . 29 Isopoda . 29 Notostraca . 30 Diplopoda . 30 Chordeumatida . 30 Polydesmida . 30 Spirobolida . 31 Hexapoda . 31 Archaeognatha . 31 Blattodea . 31 Coleoptera . 31 Collembola . 79 Dermaptera . 86 Diptera . 86 Ephemeroptera . 139 Hemiptera . 141 Hymenoptera . 160 Isoptera . 200 Lepidoptera . 200 Mecoptera . 211 Neuroptera . 211 Odonata . 212 Orthoptera . 213 Phthiraptera . 214 Plecoptera . 215 Protura . 218 Psocoptera . 219 Siphonaptera . 219 Thysanoptera . 221 ii iii Thysanura . 222 Trichoptera . 222 Pauropoda . 226 Pauropoda . 226 3 List of Sources 227 References 234 Index of Taxa 235 iv ACKNOWLEDGMENTS Acknowledgments Many have contributed to this project. John S. Ascher provided an unpublished checklist of Alaskan bees. Valerie M. Behan-Pelletier sent us a number of papers on Oribatida and made helpul recommendations as to other literature to review. Ernest C. Bernard sent us recent literature on Onychiuridae containing Alaskan records. Ken Christiansen reviewed the list of Collembola. Frans Janssens provided corrections on the Collembola. Jan Klimaszewski mailed us copies of several of his works on the Hemerobiidae of Alaska. Eric Maw sent us an electronic version of his checklist of Hemiptera. David Nicolson extracted and sent to us all of the Alaskan records from the the Krombein et al.’s catalog of Nearctic Hymenoptera. Cheryle O’Donnell shared a list of thrips species that she had collected from Alaska. James O’Hara provided us an electronic version of his catalog of Tachinidae. Ken Philip shared his preliminary checklist of Alaskan butterflies. ARACHNIDA: ACARI: ASCIDAE 1 1 Introduction This is simply a re-formatted version of data exported from Derek Sikes’ database at the University of Alaska Museum. It is an export of all of the Alaska records as of September 1, 2009. There are many omissions and errors. This is a draft document intended only for use in improving the checklist. It should not be distributed. The data for this particular draft of the exported data had many errors due to faulty handling of special characters, particularly in author names. These errors were entirely due to problems in the export and do not exist in the original data. When offering comments, please disregard incorrect rendering of special characters in author names. The taxonomic hierarchy of the Collembola follows Bellinger et al. (1996–2009). 2 List of Species Collections, published works, and other sources are cited as numbers following species names, which corre- spond to entries in the List of Sources (page 227). Class Arachnida Order Acari Family Achipteriidae Genus Achipteria 1. curta Aoki 137 Genus Parachipteria 2. nivalis (Hammer) 137, 216 Family Alloptidae Genus Alloptes 3. conurus (Trouessart) 38, 38 4. obtusolobus Dubinin 38 Genus Brephosceles 5. parvatus Peterson 165 Family Ameronothridae Genus Alaskozetes 6. coriaceus Hammer 137 Genus Ameronothrus 7. lineatus (Thorell) 137 8. nigrofemoratus (Koch) 137 Family Analgidae Genus Strelkoviacarus 9. critesi Spory 169 10. quadratus (Haller) 169 Family Ascidae Genus Arctoseius 2 2. LIST OF SPECIES 11. multidentatus Evans 63 12. ornatus Evans 63 13. robustus Lindquist 63 Genus Cheiroseius 14. groenlandicus Haarlov 63 Family Astegistidae Genus Cultroribula 15. dentata Willmann 137 Family Autognetidae Genus Autogneta 16. longilamellata (Michael) 137 Family Avenzoariidae Genus Laronyssus 17. martini (Trouessart) 37, 38, 39 Family Bdellidae Genus Bdella 18. muscorum Ewing 9 Genus Cyta 19. coerulipes (DugOs)˝ 9 20. latirostris (Hermann) 9, 63 Genus Neomolgus 21. littoralis (Linnaeus) 155, 156 22. pallipes (Koch) 63 Genus Octobdellodes 23. hurdi Atyeo 9 Family Brachychthoniidae Genus Eobrachychthonius 24. borealis Forsslund 137 25. latior (Berlese) 137 26. oudemansi van der Hammen 137 Genus Liochthonius 27. brevis (Michael) 137 28. forsslundi (Hammer) 137 29. scalaris (Forsslund) 63 30. sellnicki (Thor) 137 Genus Synchthonius 31. elegans Forsslund 137 Family Caleremaeidae Genus Veloppia 32. pulchra Hammer 137 Family Camisiidae Genus Camisia 33. biurus (Koch) 137 34. biverrucata (Koch) 137 35. foveolata Hammer 137 ARACHNIDA: ACARI: CERATOZETIDAE 3 36. horrida (Hermann) 137 37. lapponica (Tragardh) 137 38. segnis (Hermann) 137 39. spinifer (Koch) 137 Genus Heminothrus 40. longisetosus Willmann 137 Genus Neonothrus 41. humicolus Forsslund 137 Genus Platynothrus 42. peltifer (Koch) 137 43. punctatus (Koch) 137 44. sibiricus Sitnikova 137 Family Carabodidae Genus Carabodes 45. labyrinthicus (Michael) 137 Family Cepheidae Genus Cepheus 46. corae Jacot 137 Family Ceratozetidae Genus Ceratozetes 47. borealis Behan-Pelletier 137 48. gracilis (Michael) 137 49. parvulus Sellnick 137 50. spitsbergensis Thor 137 51. thienemanni Willmann 137 Genus Cyrtozetes 52. denaliensis Behan-Pelletier 137 Genus Diapterobates 53. humeralis (Hermann) 137 54. notatus (Thorell) 137 55. rotundocuspidatus Shaldybina 137 56. sitnikovae Shaldybina 137 57. variabilis Hammer 137 Genus Fuscozetes 58. fuscipes (Koch) 137 59. sellnicki Hammer 63, 137 Genus Iugoribates 60. gracilis Sellnick 137 Genus Melanozetes 61. longisetosus Hammer 137 62. meridianus Sellnick 137 63. mollicomus (Koch) 137 Genus Neogymnobates 64. lutens (Hammer) 137 Genus Oromurcia 65. lucens (Koch) 137 Genus Sphaerozetes 66. arcticus Hammer 137 4 2. LIST OF SPECIES 67. castaneus Hammer 137 Genus Svalbardia 68. paludicola Thor 137 Genus Trichoribates 69. copperminensis Hammer 137 70. lucens (Koch) 63 71. obesus (Banks) 137 72. polaris Hammer 137 Family Chamobatidae Genus Chamobates 73. pusillus (Berlese) 137 74. schuetzi (Oudemans) 137 Family Damaeidae Genus Caenobelba 75. alleganiensis Norton 137 Genus Epidamaeus 76. arcticolus (Hammer) 137 77. bakeri (Hammer) 137 78. coxalis (Hammer) 137 79. floccosus Behan-Pelletier Norton 137 80. fortispinosus Hammer 137 81. gibbofemoratus (Hammer) 137 82. hastatus Hammer 137 83. kodiakensis Hammer 137 84. koyukon Behan-Pelletier Norton 137 85. longitarsalis (Hammer) 137 86. mackenziensis (Hammer) 137 87. nasutus Behan-Pelletier Norton 137 88. tenuissimus Hammer 137 89. tritylos Behan-Pelletier Norton 137 Family Demodicidae Genus Demodex 90. brevis Akbulatova 91. folliculorum Simon Family Dinychidae Genus Dinychus 92. micropunctatus Evans 63 Family Epidermoptidae Genus Promyialges 93. uncus (Vizthum) 169 Family Eremaeidae Genus Eremaeus 94. foveolatus Hammer 137 95. grandis Hammer 19, 137 96. quadrilamellatus (Hammer) 137 97. translamellatus Hammer 19 ARACHNIDA: ACARI: HYDROZETIDAE 5 Genus Eueremaeus 98. foveolatus (Hammer) 19 99. marshalli Behan-Pelletier 19 Family Eriophyidae Genus Cocceupodes 100. breweri Strandtmann 63 Genus Nanorchestes 63 Genus Penthalodes 101. ovalis (Duges) 63 Genus Protereunetes 102. boerneri SigThor 63 Family Erythraeidae Genus Balaustium 103. sp. 210, 210 Family Eulohmanniidae Genus Eulohmannia 104. ribagai (Berlese) 137 Family Eupalopsellidae Genus Eupalopsellus 105. olandicus Sellnick 216 Family Euphthiracaridae Genus Rhysotritia 106. ardua (Koch) 137 Family Galumnidae Genus Pilogalumna 107. tenuiclava (Berlese) 137 Family Gymnodamaeidae Genus Nortonella 108. gildersleeveae (Hammer) 137 Family Haplozetidae Genus Peloribates 109. alaskensis Hammer 137 110. canadensis Hammer 137 111. pilosus Hammer 137 Family Hermanniidae Genus Hermannia 112. gibba (Koch) 137 113. reticulata Thorell 137 114. scabra (Koch) 137 115. subglabra Berlese 137 Family Histiostomatidae Genus Anoetus 6 2. LIST OF SPECIES 116. varia 209 Family Hydrozetidae Genus Hydrozetes 117. thienemanni Strenzke 137 Family Hypochthoniidae Genus Hypochthonius 118. rufulus Koch 137 Family Ixodidae Genus Amblyomma 119. dissimile Koch 208 Genus Dermacentor 120. albipictus 209 121. occidentalis Marx 210 122. variabilis (Say) 210 Genus Haemophysalis 123. leporispalustris Packard 166, 208, 209 Genus Ixodes 124. angustus Neumann 146 125. auritulus Neumann 208 126. howelli Cooley Kohls 208 127. signatus Birula 208 128. uriae (White) 153, 208 Family Kodiakellidae Genus Kodiakella 129. lutea Hammer 137 Family Liacaridae Genus Dorycranosus 130. abdominalis (Banks) 137 131. acutidens (Aoki) 137 132. altaicus Krivoluckij 137 133. parallelus (Hammer) 137 Genus Liacarus 134. bidentatus Ewing 137 Family Limnozetidae Genus Limnozetes 135. canadensis Hammer 137 Family Malaconothridae Genus Malaconothrus 136. mollisetosus Hammer 137 Genus Trimalaconothrus 137. foveolatus Willmann 137 138. novus (Sellnick) 137 Family Mesoplophoridae Genus Mesoplophora ARACHNIDA: ACARI: ORIBATELLIDAE 7 139. japonica Aoki 137 Family Metrioppiidae Genus Ceratoppia 140. bipilis (Hermann) 137 141. quadridentata (Haller) 137 142. sexpilosa Willmann 137 143. sphaerica (Koch) 137 Genus Metrioppia 144. helvetica Grandjean 137 Genus Pyroppia 145. dentata Krivoluckij 137 146. lanceolata Hammer 137 147. serrifrons (Banks) 137 Family Mycobatidae Genus Mycobates 148. conitus Hammer 137 149. consimilis Hammer 137 150. incurvatus Hammer 137 151. punctatus Hammer 137 Genus Punctoribates 152. hexagonus Berlese 137 153. quadrivertex (Halbert) 137 Family Nanhermanniidae Genus Nanhermannia 154. coronata Berlese 137 Family Nothridae Genus Nothrus