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Estudio De Los "Buccinacea" "(Neogastropoda: Gastropoda)"

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Estudio De Los Butll. lust. Cat. Hist. Nat., 48 (Sec. Geol., 3): 61-90. 1982 ESTUDIO DE LOS BUCCINACEA (NEOGASTROPODA, GASTROPODA) DEL PLIOCENO DEL EMPORDA ( CATALUNYA). DESCRIPTIVA Y SISTEMATICA Jordi Martinell ;; Rebut: setembre 1981 RESUM Estudi dels Buccinacea (Neogastropoda, Gastropoda) del Plioce de I'Emporda. Descriptiva i sistematica En el present treball es descriuen i figuren 13 especies de Buccinacea ( Neogas- tropoda , Gastropoda ) procedents del Plioce de I'Emporda. Per a realitzar aquest estudi s'han tingut en compte tant les troballes de camp noves com les ja existents i dipositades be a colleccions particulars be a oficials. Ouan el nombre d'individus ho ha permes, la descriptiva ha estat acompanyada de les seguents dades numeriques: mida minima (m), mida maxima (M), mida mitjana (x), desviacio tipica (s), variancia (s2), intervals de la mitjana i de la variancia, aixi com el coeficient de correlacio (r), per a cada estadi de creixement. A causa de la confusie que s'ha creat amb la proliferacio de generes i subgeneres dels Nassariidae , molts dels quals no estan ben definits, s'ha optat per considerar algunes especies corn a pertanyents al genere Nassarius en el seu sentit mes ampli. INTRODUCCION logfa de Barcelona (Museo Martorell), y entre ]as segundas, la coleccion del doctor En el prescntc trabajo se describer ]as J. F. dc Villalta, de Barcelona. especies de Buccinacea (Neogastropoda, La localizacion geografica de los dife- Gastropoda) que sc han encontrado en los rentes yacimientos esta indicada en la figu- diferentes yacimientos pliocenicos del Em- ra 1; las caracteristicas litologicas de los porda (Catalunya). Para ello se ha tenido mismos han sido descritas por MARTINELL en cuenta tanto el material encontrado en (1973, 1976, 1977). los diferentes yacimientos como el proce- Cuando el numero do individuos lo ha dente de colecciones, ya sean oficiales o permitido, la descriptiva se acompana por particulares. Entre las primeras cabe citar una scric de datos numericos cuyas varia- las de Almera, Bofill, Colominas y Moragas bles estan representadas en la fig. 2. To- depositadas en cl Museo Municipal de Geo- dos los individuos mcdidos proceden del * Departament de Paleontologia. Facultat de Geologia. Universitat de Barcelona. Gran Via de les Corts Catalanes, 585. Barcelona-7. 61 0 10 20 Km. FIG. 1. Localizacion geografica de los yacimientos de los cuales procede la fauna cstudiada. 1) Pue- blo de Siurana, 2) Mas Siurana, 3) Cementerio de Siurana, 4) Baseia, 5) Sant Miquel de Fluvia. mismo nivel, formando parte a su vez de un muestreo aleatorio. En las especies en que hemos contado con un ndmero suficiente de individuos, los datos numericos se han calculado se- paradamente para cada estadio de creci- miento, dando solamente las caracteristi- cas muestrales de aquellos estadios sufi- cientemente representados. Se ha tornado convencionalmente como estadio de creci- miento el ndmero de vueltas de espira, si- guiendo el metodo utilizado por DE RENZI (1967) y MARTINELL & CUADRAS (1977). Los valores calculados han lido: valor maximo (M) y minimo (m) de cada para- metro, media (x), desviacion tfpica (s), va- rianza (s2), intervalos de la media y de la varianza, estimadas todas e1las cn las mag- nitudes brutas, asi como el coeficiente de correlacion (r). Debido a la confusion que existe aun FIG. 2. Medidas tomadas para el estudio de los actualmente en la posicion especifica de Buccinacea. H=altura total de la concha, hv=al- tura de la ultima vuelta, bh=altura de la boca, Nassarius, se ha creido preferible, tal Co- A = anchura. mo ya propusieron ADAM & GLIBERT (1974), 62 no tend' en cucnta la gran cantidad tic DF. SC RIPCION. Concha fusiforme do es- gencros y subgeneros a quc han sido atri- piral concca salicnte . Ornamentacion muy buidas algunas especies por diferentes au- suave formada por las lineas de crecimien- tores, a menudo de mancra contradicto- to, las cuales son del tipo opistocirtico. Las ria. Por el momento se han considerado lineas de sutura son lineales y poco pro- Como pertenecientes al genero Nassarius fundas. en su sentido mas amplio. La altura total de la concha del ejemplar mas adulto es de 1,237 cln y la anchura, 0,477 cm. CLASE: GASTROPODA Ornamentacion. La protoconcha es pau- SUBCLASE: PROSOBRANCHIA cispiral, sumcrgida y algo desviada estan- ORDEN: NEOGASTROPODA do formada por dos vueltas v media. SUPERFAMILIA: BUCCINACEA La ornamentacion es constante a to lar- FAMILIA: PYRENIDAE go dc toda ]a concha, y esta formada ex- Genero Mitrella Risso, 1826 clusivamcntc por la suave estriacion axial Subgenero: Columbellopsis Bucquoy, Daut- quc forman ]as lineas de crecimiento. A zenbcrg & Dollfus, 1882 simple vista, las conchas son completa- mentc lisas. La ultima vuelta es algo mayor que la Mitrella ( Columbellopsis) minor mitad de la altura total de la concha. La (Scacchi, 1833) ornamentacion es Lisa en la mitad adapical (Lam. I, figs. 1 y 2) de la vuelta, apareciendo una scrie de cor- doncs cspirales quc Ilegan hasta el final 1887 Columbella minima Scacchi. BELLARDI, de ]a base de ]a minma. p. 46, lam. II, fig. 54. La abertura es mas o menos romboidal. 1890 Macizirella minima Sacco. SACCO, p. 46, El labro es algo sinuoso y en su interior lam. II, fig. 54. se obscrvan algunas veces pequenos re- 1898 Columbella minima Scacchi, var. an- pliegucs parictales. La columcla gulata ALMERA & BOFILL, p. 32. esta fuer- 1907 Columbella minima Scacchi et al. AL- temente excavada; el canal sifonal es cor- MERA, p. 145, lam. V, fig. 26, lam. VI, to. El horde columelar es suave, poco ex- fig. 1. pansionado, sin rcpliegues, formando una 1911 Columbella (Macrurella) minima Sac- pcquena callosidad en su parte abapical. co. CERULLI-IRELLI, p. 257, lam. XXIII, figs. 73-74. OBSERVACIONES. Esta especie es bas- 1944 Mitrella (Columbellopsis) minor (Scac- tante parccida a Mitrella (Columbellopsis) chi). WENZ, p. 1138, fig. 3232. subulata (Brocchi), diferenciandose 1952 Columbella (Atilia) minima Sacco. LE- basi- COINTRE., p. 126. camente por ]a mayor altura de la ultima 1963 Mitrella (Columbellopsis) minor (Scac- vuclta con respecto a ]a altura total de la chi ). GLIBERT, p. 44. concha, la forma de ]a abertura y por tenor 1970 Mitrella minor (Scacchi). PARENZAN, pa- la columcla mas fuertemente excavada. gina 171, fig. 668. Tambicn se asemeja bastante a Mitrella 1974 Mitrella (Clinurella) minima (Sacco). (Columbellopsis) nassoides (Grateloup), di- MALATESTA, p. 315, lam. 25, fig. 7. ferenciandose por tencr esta ultima ]a 1976 Mitrella (Columbellopsis) minor (Scac- abertura mas alargada y Ifneas chi). MARTINELL, p. 194, lam. 25 figs. 1-2. ]as de su- 1977 Mitrella (Columbellopsis) minor (Scac- tura mucho mejor marcadas. chi). MARTINELL, pp. 97-102. 1979 Mitrella (Columbellopsis) minor (Scac- chi). MARTINELL & VILLALTA, pp. 23-25. MATERIAL RECOGIDO Y CITADO. Dos FAMILIA: BUCCINIDAE ejemplares procedentes de la capa U, del Genero: Parvisipho Cossmann, 1889 yacimiento del cementerio dc Siurana y Subgenero: Andonia Harris & Burrows, un ejemplar encontrado en las arcillas azu- 1891 les dc Sant Miquel de Fluvia. ALMERA & BOFILL (1898) citan esta especie Como en- Parvisipho ( Andonia) bonellii contrada en el dcsaparecido yacimiento de (Bellardi & Michelotti, 1840) Baseia. (Lam. I, figs. 3, 4) 63 1840 Fusses bonellii Gene. BELLARDI & Mi- a base de cordones espirales v otra a base CHELOTTI , p. 20, 1am . II, fig. 5. de costulas axiales. 1882 Genea bonellii (Gene ). BELLARDI , p. 205, En la ultima vuelta en que se observa lam. XI, fig. 10 a, b. bien la ornamentacion , vemos que esta la 1890 Fusus ( Genea ) bonellii Gene . HoRNES & forman nueve cordones espirales muy bien lam. XXXII , fig. 8. AVINGER , p. 260 , siendo espacio entre cordo- 1898 Fusus bonellii Gene . ALMERA & BoFILL, marcados, el p. 22. nes mucho mas ancho que ellos mismos. 1898 Fusus chiae Almera & Bofill. ALMERA & Asimismo , existen sicte fuertes costulas BOFILL, p. 22 , 1am. II, fig. 3. axiales cortando los cordones espirales. En 1907 Fusus ( Genea ) bonellii Gene . ALMERA, el espacio intercostular se pueden obser- p. 138. var muy Wen las lineas de crecimiento. 1907 Fusus chiae Almera & Bofill . ALMERA, Toda la ultima vuclta de nuestro ejem- p. 138, 1am. VI, fig. 3. plar esta rota. 1935 Parvisipho (Andonia ) bonellii Gene. MONTANARO , p. 70, lam. VI , fig. 4. 1944 Parvisipho ( Andionia) bonellii ( Gene). OBSERVACIONES. La determinacion de WENZ , p. 1157, fig. 3288. este raro ejemplar ha lido realizada gra- 1975 Parvisipho (Andonia ) bonellii (Bellardi cias a que se ha podido comparar direc- & Michelotti ). PAVIA, p. 43, lam. VI, tamente con los ejemplares de la colec- figs . 11-12, 14, 16. cion Pavia del Plioceno inferior de Monte 1976 Parvisipho ( Andonia ) bonellii ( Bellardi Rocro (Italia). & Michelotti ). MARTINELL , p. 197, lami- La especie figurada por ALMERA & Bo- na 25 , figs. 3-4. FILL (1884, II, fig. 4) y por ALMERA Bellardi lam. 1977 Parvisipho ( Andonia) bonellii ( dcscrita bajo el & Michelotti ). MARTINELL, pp. 98-102. (1907, lam. VI, fig. 3), y 1979 Parvisipho ( Andonia) cf. bonellii (Be- nombre de Fusus chia Almera & Bofill, llardi & Michelotti ). D'ALESSANDRO, LA- parece ser sinonima de P. (A.) bonellii. Al VIANO, RlccIlETTI & SARDELLA, p. 80. realizar la revision de la coleccion Almera 1979 Parvisipho ( Andonia ) bonellii ( Bellardi depositada en el Museo Municipal de Geo- & Michelotti ). PAVIA & ROBBA, p. 554. logia de Barcelona (Musco Martorell), no 1979 Parvisipho (Andonia) bonellii ( Bellardi se ha podido cncontrar cstc ejemplar, por & Michelotti ). MONTEFAMEGLIO , PAVIA & lo cual, atendiendonos a las liguras dadas ROSA , p. 189. por estos autores, se ha considerado quc la especie de Almera & Botill no es mas MATERIAL RECOGIDO Y CITADO. Pro- que un sinonimo de P. (A.) bonellii. cede de la capa U, del cementerio de Siu- Como muy bien hace resaltar PAVIA rana. ALMERA & BOFILL (1898) citan esta (1975), aFusus bonellii» es women ttduni, re- especie como encontrada en el desapare- ya que su autor, Gene, no cumplio las cido yacimiento de Baseia.
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