BULLETIN DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE ENTOMOLOGTE, 68: 131-142, 1998 BULLETIN VAN HET KONINKL!JK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN ENTOMOLOGIE, 68: 1 1 ~ 1-142 , 1998

A Red list for the of Flanders by Jean-Pierre MAELFAIT, Leon BAERT, Marc JANSSEN & Mark ALDERWEIRELDT

Abstract logical Society (ARABEL). Distribution data were gathered per and published in catalogues per family (ALDER­ Of the 604 species mentioned as occurring in Flanders only wELRELDT & MAELFA1T, 1990; BAERT, 1996; DE BLAUWE & about half is at moment safe or at low risk of extinction. The extinct and BAERT, 1981; JACOBS , 1993 ; JANSSEN, 1993 ; JANSSEN & threatened species occur( -red) in oligotrophic grasslands, deciduous RAERT, 1987; RANSY & BAERT, 1985, 1987a,b, 1991a,b; forests, heathlands and several kinds of wetland habitats. RANSY et a!., 1987, SEGERS & BAERT, 1991 ; VAN KEER & Key words: Red list, spiders, Flanders, , habitats V ANUYTVEN, 1993). The catalogue treating the subfamily of the Linyphiinae is in preparation (BAERT, in prep). These ca­ talogues and more recent published and unpublished observa­ tions were used to assess the status of each Flemish spider Resume species.

II n'y a, aujourd'hui, que pres de Ia moitie des 604 especes d'araignees connues deFlandre que !'on peut considerer comme etant al 'abri ou ne presentant qu'un faible risque de disparition. Les especes eteintes ou Material and methods menacees colonisaient ou colonisent des prairies oligotrophes, des forets decidues, des Iandes a bruyere et differents types d'habitats gorges d' eau. Each species mentioned as occurring in Flanders was as­ signed one of the following categories. These categories Mots-clefs: Liste rouge, araignees, Flandre, Belgique, biotopes agree with the classification used in the Red Data Book of (BINoT et al., 1998) and therefore with the criteria and categories developed by the IUCN-The World Conserva­ tion Union (column RL-C in Table 1). Introduction - EW: Extinct in the wild. Species not located in the wild / during the last fifty years. For the spiders of Flanders these During recent years Red lists of particular taxonomic groups are usually species which were only mentioned for that became more and more indispensable tools in nature conserva­ region by Becker and for which we have no reason to doubt tion research. This also holds true for Flanders. As well as for the correctness of the determination by that author. the development of a modem nature conservation policy as - CR: Critical. These are species that became ve1y rare in our for their use in site assessment studies such lists summaris­ region due to a drastic reduction of their habitats. These ing the present status of each involved species can be very species will go extinct if the causal factors for that habitat useful instruments (ALDERWE1RELDT & MAELFAJT, 1992; MAEL­ loss continue operating. FAIT, 1993, 1994, 1995; MAES et a!., 1995 ; VERHEY EN et a/., - EN: Endangered. Species, that are rare becaH~e of extensive 1994). deterioration and destruction of their habitats. The study of spiders in Belgium (and Flanders) began by - VU: Vulnerable. Species, that became quite uncommon be­ the end of the nineteenth century with the very luxurious cause of habitat loss during the past decades. books treating the occurrence of these in Belgium by - RG: Rare geographically restricted species. These are spe­ BECKER (1882, 1896). During the fifties of the twentieth cen­ cies for which Flanders lays at the northern (N), sou them (S) tmy faunistic studies were resumed by J. KEKENBOSCH (Royal or western (W) limit of their geographical range. Belgian Institute of Natural Sciences). It was only from the - IN: Indeterminate. Species assumed to be threatened but seventies onwards however that a large variety of habitats where there is not enough information to decide which of distributed over the whole of the country were sampled. This the three above-mentioned categories is appropriate was mostly done by scientists associated with the LaboratOiy - IK: Insufficiently known. Species, that are suspected but not of Ecology (University of Gent) and, later on, with the Royal definitely known to belong to any of the above categories, Institute of Natural Sciences and with the Institute of because of lack of information. Also the species mentioned to Nature Conservation. Amateur arachnologists also contribu­ occur in Flanders doubtfully. ted a lot to that inventory work. Most of these faunistic studi es were were published in the newsletter of the Belgian Arachno- In Table 1, the habitat for each species of the categories EW, 132 J.-P. MAELFAIT, L. BAERT, M. JANSSEN & M. ALDERWEIRELDT I I

Table I. - Spider species occurring in Flanders (FI) with indi­ FAMILY, SPECIES FL RL-C HABITAT cation of their status and habitat choice (explanation: see text). Dysdera erythrina (WALCKENAER, 1802) FL EN Fddd FAMILY, SPECIES FL RL-C HABITAT Harpactea hombergi (Scorou, 1763) FL EN Fddd Atypidae Segestriidae Atypus affinis EICHWALD,1830 FL vu Godts Segestria bavarica C.L. KocH, 1813 FL Eresidae Segestria f/orentina (Rossi, 1790) FL Eresus cinnaberinus (OuVJER, 1789) FL IK y Segestria senoculata (LiNNAEUS, 1758) FL Eresus sandafiatus (MARTINI & Gom, 1778) FL EW Hdbs Scytodidae Scytodes thoracica LATREILUE, 1801 FL claustrarius (HAHN, 1831) Pholcidae (KEYSERUNG, 1863) Pholcus phalangioides (FUESSUN, 1775) FL Amaurobius fenestra/is (STROEM, 1768) FL Psilochorus simoni (BERLAND, 1911) FL (WALCKENAER, 1825) FL Gnaphosidae Amaurobius simi/is (BLACKWALL, 1815) FL Cal/ilepis nocturna (LiNNA£us, 1758) FL RG N Dictynidae Drassodes cupreus (BLACKWALL, 1831) FL A/tel/a Iucida (SIMON, 1871) FL RG N Drassodes hypocrita (SIMON, 1878) Argenna patula (SIMON, 1871) FL CR Sa Drassodes lapidosus (WALCKENAER, 1802) FL Argenna subnigra (O.P. -CAMBRIDGE , 1861) FL EN Go db Drassodes pubescens (THORELL, 1856) FL EN Godt Cicurina cicur (FABRICIUS, 1793) FL Gnaphosa leporina (L. KocH, 1866) FL EN Hw Dictyna arundinacea (LINNAEUS, 1758) FL Gnaphosa lucifuga (WALCKENAER, 1802) FL CR Godb Dictyna civica (LucAS, 1819) FL RG N Gnaphosa lugubris (C.L. KocH, 1839) Dictyna latens (FABRICIUS, 1775) FL EN Godd Gnaphosa nigerrima L. KocH, 1877 FL CR Hws Dictyna major MENGE, 1869 FL RG s Gnaphosa opaca HERMAN, 1879 FL RG N Dictyna pusi//a THORELL, 1856 FL EN Godd Haplodrassus dalrnatensis (L. KocH, 1866) FL EN Godt Dictyna uncinata THORELL, 1856 FL Haplodrassus kulczynskii LOH¥J1NDER, 1912 Lathys humilis (BLACKWALL, 1855) FL Haplodrassus moderatus (KuLCZVNSKJ, 1897) Lathys pula (O.P-CAMBRIDGE, 1863) FL RG N Haplodrassus signifer (C.L. KocH, 1839) FL Mastigusa arietina (THORELL, 1871) FL EN Godta Haplodrassus silvestris (BLACKWALL, 1833) FL EN Fddd Mastigusa macrophtha/mus (KuLCZVNSKI, 1896) Haplodrassus umbratilis (L. KocH, 1866) FL CR Godts Nigma flavescens (WALCKENAER, 1825) FL Micaria dives (LuCAS, 1816) FL CR Godta Nigma walckenaeri (ROEWER, 1951) FL Mic9Fia formicaria (SuNDEVALL, 1832) Micaria fulgens (WALCKENAER, 1802) FL EN Godta quadriguttata (HAHN, 1883) FL CR Fddvs Micaria guttulata (C.L. KocH, 1839) Uloboridae Mica ria pulicaria (SuNDEVALL, 1831) FL Hyptiotes paradoxus (C.L. KocH, 1831) FL Micaria romana L. KocH, 1866 FL RG N Uloborus plumipes LucAS, 1816 FL Micaria silesiaca L. KocH, 1875 FL RG N Uloborus walckenaerius LATREILLE, 1806 Micaria subopaca WESTRING, 1861 FL "' Oecobiidae Phaeocedus braccatus (L. KocH, 1866) FL EN Hdb Oecobius navus BLACKWALL, 1859 FL Poecilochroa conspicua (L. KocH, 1867) FL RG N Zodariidae Poecilochroa variana (C.L. KocH, 1839) Zodarion gal/icum (SIMON, 1872) FL IK y Scotophaeus blackwalli (THORELL, 1873) FL Zodarion itaficum (CANESTRINI, 1868) Scotophaeus scutulatus (L. KocH, 1866) FL Zodarion rubidum (SIMON, 191 1) FL RG N Zelote vi/licus (THORELL, 1875) Oonopidae Zelotes aeneus (SIMON, 1878) FL RG N Oonops domesticus DED ALMAS, 1916 FL Zelotes apricorum (L. KocH, 1876) FL IN X Oonops pu/cher TEMPLETON, 1835 FL Zelotes electus (C.L. KocH, 1839) FL vu Godt Dysderidae Zelotes erebeus (THORELL, 1871) Dysdera crocota C.L. KocH, 1838 FL Zelotes latreillei (SIMON, 1878) FL A Red list for the spiders of Flanders II 133

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Zelotes longipes (L. KocH, 1866) FL vu Godt Phrurolithus festivus (C.L. KocH, 1835) FL Zelotes lutetianus (L. KocH, 1866) FL EN Gowt Phrurolithus minimus C.L. KocH, 1839 FL CR Godta Zelotes pedestris (C.L. KocH, 1839 FL E N Godt Scotina celans (BLACKWALL, 1811) FL RG N Zelotes petrensis (C.L. KocH, 1839) FL vu Godt Scotina gracilipes (BLACKWALL, 1859) FL CR Hdb Zelotes praeficus (L. KocH, 1866) FL CR Hdb Zoridae Zelotes pseudoclivicolus GRIMM, 1982 lora nemoralis (BLACKWALL, 1861) FL CR Fddvs Zelotes pumilus (C.L. KocH, 1839) lora silvestris KuLCZVNSKI, 1897 FL EN Hdb Zelotes pusil/us (C.L. KocH, 1833) FL lora spinimana (SUNDEVALL, 1833) FL Zelotes subterraneus (C.L. KocH, 1833) FL Anyphaenidae Clubionidae Anyphaena accentuata (WALCKENAER, 1802) FL Cheiracanthium erraticum (WALCKENAER, 1802) FL Eusparassidae Cheiracanthium montanum (L. KocH, 1877) FL RG N Micrommata virescens (CLERCK, 1757) FL CR Fdwv Cheiracanthium pennyi O.P. -CAMBRIDGE, 1873 FL RG N Thomisidae Cheiracanthium punctorium (VILLERS,1789) FL EW Godds Coriarachne depressa (C.L. KocH, 1837) FL RG s Cheiracanthium virescens (SuNDEVALL, 1833) FL EN Godt Diaea dorsata (FABRICIUS, 1777) FL Clubiona brevipes BLACKWALL, 1811 FL Heriaeus hirtus (LATREILLE, 1819) FL RG N Clubiona caerulescens L. KocH, 1867 FL EN Fddo Misumena vatia (CLERCK, 1757) FL vu God! Clubiona compta C.L. KocH, 1839 FL Misumenops tricuspidatus (FABRICIUS, 1775) FL RG N Clubiona corticalis (WALCKENAER, 1802) FL Ozyptila atomaria (PANZER, 181 0) FL EN Godt Clubiona diversa D.P.-CAMBRIDGE, 1862 FL Ozyptila blackwalli SIMON, 1875 Clubiona frisia WUNDERLICH & SCHIT, 1995 FL vu Godt Ozyptila brevipes (HAHN, 1831) FL EN Gowt Clubiona frutetorum L. KocH, 1867 FL Ozypti/a nigrita (THORELL, 1875) FL CR Godt Clubiona genevensis L. KocH, 1867 FL CR Godr Ozyptila praticola (C.L. KocH, 1837) FL Clubiona germanica THORELL, 1872 FL Ozypti/a pul/ata (THORELL, 1875) Clubiona juvenis SIMON, 1878 FL CR Mrr Ozyptila rauda SIMON, 1875 Clubiona lutescens WESTRING, 1851 FL Ozyptila sanctuaria (D.P.-CAMBRIDGE, 1871) FL EN Godt Clubiona neglecta D.P.-CAMBRIDGE , 1862 FL Ozyptila scabricula (WESTRING, 1851) FL EN Hdb Clubiona pallidula (CLERCK, 1757) FL Ozyptila simplex (O.P. -CAMBRIDGE, 1862) FL Clubiona phragmitis C.L. KocH, 181 3 FL Ozyptila trux (BLACKWALL, 1816) FL Clubiona pseudoneglecta WuNDERLICH, 1994 FL IN X Pisgus truncatus (PALLAS, 1772) FL CR Fddo Clubiona reclusa O.P.- CAMBRIDGE, 1863 FL Synaema globosum (FABRICIUS, 1775) FL RG N Clubiona stagnati/is KULCZVNSKI, 1897 FL Thomisus onus/us WALCKENAER, 1806 FL RG N Clubiona subsultans THORELL, 1875 FL RG s Tmarus piger (WALCKENAER, 1802) FL RG N Clubiona subtilis L. KocH, 1867 FL Xysticus acerbus THORELL, 1872 FL CR Hdb Clubiona terrestris WESTRING, 1862 FL Xysticus audax (SCHRANK, 1803) FL Clubiona trivia/is C.L. KocH, 1811 FL vu Godt Xysticus bifasciatus C.L. KocH, 1837 FL CR God! Liocranidae Xysticus cristatus (CLERCK, 1757) FL Agraecina lineata (SIMON, 1878) FL RG N Xysticus erraticus (BLACKWALL, 1831) FL EN Godt Agraecina striata (KuLCZVNSKI, 1882) Xysticus ferrugineus MENGE, 1875 FL RG N Agroeca brunnea (BLACKWALL, 1833) FL Xysticus kempelini THORELL, 1872 FL RG N Agroeca cuprea MENGE, 1873 FL EN Godt Xysticus kochi THORELL, 1872 FL Agroeca dentigera KuLCZYNSKI, 1913 FL CR Hws Xysticus lanio C.L. KocH, 1821 FL Agroeca inopina O.P. -CAMBRIDGE, 1886 FL IK y Xysticus linea/us (WESTRING , 19 51) FL E W Godd Agroeca lusatica (L. KocH, 1875) FL vu Godd Xysticus luctator L. KocH, 1870 Agroeca proxima (D.P.-CAMBRIDGE, 1871) FL Xysticus luctuosus (BLACKWALL, 1836) FL CR Fddv Apostenus fuscus WESTRING, 1851 FL EN Fddd Xysticus ninnii THORELL, 1872 FL EN Godb Liocranum rupicola (WALCKENAER, 1825) FL Xysticus robustus (HAHN, 1832) .. 134 J.-P. MAELFAIT, L. BAERT, M. JANSSEN & M. ALDERWEIRELDT

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Xysticus sabulosus {HAHN, 1832) FL EN Go db Leptorchestes berolinensis (C.L. KocH, 1816) Xysticus striatipes (L. KocH, 1870) Marpissa muscosa (CLERCK, 1757) FL Xysticus ulmi (HAHN, 1832) FL Marpissa nivoyi (LucAS, 1816) FL EN Godt Philodromidae Marpissa pomatia (WALCKENAER, 1802) FL CR Mrr Philodromus albidus KuLCZVNSKJ, 1911 FL EN Fddv Marpissa radiata (GRUBE, 1859) FL vu Mrr Philodromus aureo/us (CLERCK, 1757) FL Myrmarachne formicaria (OEGEER, 1778) FL Philodromus buxi SIMON, 1881 FL RG N Neon rayi (SIMON, 1875) Philodromus cespitum (WALCKENAER, 1802) FL Neon reticulatus (BLACKWALL, 1853) FL Philodromus co/linus C.L. KocH, 1835 FL Neon valentulus FALCONER, 1912 FL CR Hws Philodromus dispar WALCKENAER, 1825 FL Pel/enes tripunctatus (WALCKENAER, 1802) FL EN Go db Philodromus emarginatus (SCHRANK, 1803) FL IN X Phlegra fasciata (HAHN, 1826) FL vu Godb Phi/odromus fa/lax SUNDEVALL, 1833 FL CR Godtr Pseudicius encarpatus (WALCKENAER, 1803) FL IK y Philodromus histrio (LATREILLE, 1819) FL vu Hdb Sa/ficus cingulatus (PANZER, 1797) FL Philodromus margaritatus (CLERCK, 1757) FL CR Fdwl Salticus scenicus (CLERCK, 1757) FL Philodromus poecilus (fH ORELL, 1872) Salticus zebraneus (C.L. KocH, 1837) FL Philodromus praedatus D.P.-CAMBRIDGE, 1871 FL EN Fddv Sitticus caricis (WESTRING, 1861) FL EN Mo Philodromus rufus WALCKENAER, 1825 FL RG N Sitticus distinguendus (SIMON, 1868) FL EN Godb Thanatus arenarius THORELL, 1872 Sitticus floricola (C.L. KocH, 1837) FL vu Mo Thanatus formicinus (CLERCK, 1757) FL IN X Sitticus pubescens (FABRICIUS, 1775) FL Thanatus striatus C.L. KocH, 1815 FL vu Godt Sitticus saltator D.P.-CAMBRIDGE, 1868 FL EN Godb Tibellus maritimus (MENGE, 1875) FL vu Gowt Synageles hilarulus (C.L. KocH, 1816) FL RG N Tibellus ob/ongus (WALCKENAER, 1802) FL vu Gowt Synageles venator (LucAS, 1836) FL Salticidae Dxyopidae Aeluril/us v-insignitus (CLERCK, 1757) FL vu Godd Oxyopes heterophthalmus LATREILLE, 1801 FL EW Hdbs Asia nellu s festivus (C.L. KocH, 1831) FL IK y Oxyopes lineatus LATREIILLE, 1806 FL RG N Ballus chalybeius WALCKENAR, 1802 FL Oxyopes ramosus (PANZER, 1801) FL vu Hd Bianor aurocinctus (DHLERT, 1865) FL EN Go db Lycosidae Dendryphantes nidicolens (WALCKENAER, 1802) Alopecosa accentuata (LATREILLE, 1817) Dendryphantes rudis (SuNDEVALL, 1833) FL AlopeJosa barbipes (SUNDEVALL, 1833) FL vu Go db Euophrys aequipes (D.P. -CAMBRIDGE, 1871) FL vu Godbs ~ttfPecosa cuneata (CLERCK, 1757) FL vu Godb Euophrys aperta MILLER, 1971 A/opecosa cursor (HAHN, 1831) FL EW Hdb Euophrys erratica (WALCKENAER, 1825) FL Alopecosa fabrilis (CLERCK, 1757) FL EN Go db Euophrys frontalis (WALCKENAER, 1802) FL Alopecosa inquilina (CLERCK, 1757) Euophrys lanigera (SIMON, 1871) FL Alopecosa pulverulenta (CLERCK, 1757) FL Euophrys petrensis C.L. KocH, 1837 FL EN Go db Alopecosa striatipes (C.L. KocH, 1837) Evarcha arcuata (CLERCK, 1757) FL Alopecosa trabalis (CLERCK, 1758) FL 'cR Fddvs Evarcha fa/cata (CLERCK, 1757) FL Arctosa cinerea (FABRICIUS, 1777) FL CR Rg Evarcha jucunda (LucAs, 1818) FL IK y Arctosa figurata (SIMON, 1876) FL RG N Evarcha laetabunda (C.L. KocH, 1818) FL EN Hdb Arctosa leopardus (SUNDEVALL, 1833) FL vu Gowt Hasarius adansoni (SAVIGNY & AuDOUIN, 1827) FL Arctosa perita (LATREILLE, 1799) FL EN Go db Heliophanus aeneus (HAHN, 1831) FL EW Godbs Aulonia albimana (WALCKENAER, 1805) FL EW Godr Heliophanus auratus C.L. KocH, 1835 FL EN Mrr Hygro/ycosa rubrofasciata (DHLERT, 1865) FL EN Fdmot Heliophanus cupreus (WALCKENAER, 1802) FL Pardosa agrestis (WESTRING, 1861) FL EN Rs Heliophanus dampfi ScHENKEL, 1923 Pardosa agricola (f HORELL, 1856) FL CR Rg Heliophanus dubius C.L. KocH, 1831 FL RG s Pardosa a/acris (C.L. KocH, 1833) Heliophanus f/avipes (HAHN, 1832) FL Pardosa amentata (CLERCK, 1757) FL Heliophanus tribulosus SIMON, 1868 FL RG N Pardosa bifasciata (C.L. KocH. 1831) A Red list for the spiders of Flanders 11 135

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Pardosa hortensis (THORELL, 1872) FL RG N Tegenaria parietina (FouRcRov, 1785) FL Pardosa lugubris (WALCKENAER, 1802) FL vu Fddv Tegenaria picta SIMON, 1870 FL Pardosa monticola (CLERCK, 1757) FL EN Godg Tegenaria saeva BlACKWALL, 1811 FL Pardosa nigriceps (THORELL, 1856) FL Tegenaria silvestris L. KocH, 1872 FL vu Fddd Pardosa oreophila SIMON, 1937 Textrix caudata L. KocH, 1872 FL Pardosa paludicala (CLERCK, 1757) FL EW Mo Textrix denticu/ata (OLMER, 1789) FL Pardosa pa/ustris {lJNNAEus, 1758) FL Hahniidae Pardosa prativaga (L. KocH, 1870) FL vu Me Antis tea e/egans (BlACKWALL, 1811) FL Pardasa proxima (C.L. KocH, 1817) FL RG N Hahnia candida SiMON, 1875 FL RG N Pardosa pullata (CLERCK, 1757) FL Hahnia helveola SIMON, 1875 FL vu Fddd Pardasa purbeckensis F.O.P. -CAMBRIDGE, 1895 FL CR Sa Hahnia montana (BlACKWALL, 1811) FL Pardosa sa/tans TiPFER-HOFMANN &V oN HELVERSEN, 1990 FL vu Fddv Hahnia nava (BlACKWALL, 1811) FL EN Godr Pardasa sphagnicala (F. DAHL, 1908) FL CR Mo Hahnia ononidum SIMON, 1875 Pirata hygraphi/us THORELL , 1872 FL Hahnia pusilla C.L. KocH, 1811 FL IN X Pirata knorri (ScoPou, 1763) FL EW Rg Mimetidae Pirata latitans (BlACKWALL, 1811) FL Era aphana (WALCKENAER, 1802) FL RG N Pirata piraticus (CLERCK, 1757) FL Era cambridgei KULCZVNSKI , 1911 FL Pirata piscatorius (CLERCK , 1757) FL EN Mo Era furcata (VILLERS 1789) FL Pirata tenuitarsis SIMON, 1876 FL CR Mo Era tuberculata (DEGEER, 1778) FL vu Godd Pirat a u/iginosus (THORELL, 1872) FL CR Hws Theridiidae Tricca lutetiana (S IMON, 1876) Achaearanea lunata (CLERCK, 1757) FL Trochosa rabusta (SIMON, 1876) FL CR God I Achaearanea riparia (BlACKWALL , 183 1) FL vu Godba Trochosa ruricala (DEGEER, 1778) FL Achaearanea simulans (THORELL, 1875) FL Trochasa spinipalpis (F.O.P.-CAMBRIDGE, 1895) FL vu Gowt Achaearanea tepidariorum (C.L. KocH, 1811) FL Trochosa terrico/a THORELL, 1856 FL Achaearanea veruculata (URQUHART, 1885) FL Xero/ycosa miniata (C .L. KocH, 1831) FL EN Godb Anelosimus au/icus (C.L. KocH, 1838) FL RG N Xero/ycosa nemaralis (WESTRING, 1861) FL vu Fddv Ane/osimus pu/chellus (WALCKENAER, 1802) FL RG N Pisauridae Anelasimus vittatus (C.L. KocH, 1836) FL Dalamedes fimbriatus (CLERCK, 1757) FL CR Mo Carniel/a brigna/ii THALER & STEINBERGER, 1988 Dalamedes plantarius (CLERCK, 1757) FL CR Wed Crustlj}ina guttata (WiDER, 1831) FL vu Godt Pisaura mirabilis (CLERCK , 1757) FL CruStulina sticta (O .P.-CAMBRI OGE , 1861) FL CR Mo Cybaeidae Dipaena caracina (C.L. KocH, 1811) Argyraneta aquatica (CLERCK, 1757) FL vu Wes Dipoena erythropus (SIMON, 1881) Cybaeus angustarium L. KocH, 1868 FL IK y Dipoena inomata (D .P.-CAMBRIDGE, 1861) FL CR Hd Agelenidae Dipoena melanogaster (C.L. KocH, 181 5) FL EN Godd C.L. KocH, 1811 FL IN X Dipaena prana (MENGE, 1868) FL EN Godd Agelena gracilens "' Agelena labyrinthica (CLERCK, 1757) FL Dipoena tristis (HAHN, 1833) FL CR Godd Caelotes atropos (WALCKENAER, 1825) FL IK y Enaplagnata pal/ens BlACKWALL, 1831 FL Caelates inermis (L. KocH, 1855) FL RG N Enaplagnatha latimana HIPPA & OKSAlA, 1982 FL Caelates terrestris (Wi DER, 1831) FL vu Fddd Enap/agnatha mardax {TH ORELL, 1875) FL vu Godd Cryphaeca silvicala (C.L. KocH, 1831) FL EN Fddd Enoplagnatha aelandica {THORELL, 1875) FL EN Godd Histapona tarpida (C.L. KocH, 1831 ) FL RG N Enaplagnatha avata (CLERCK, 1757) FL Tegenaria agrestis (WALCKENAER, 1802) FL Enaplognatha tecta (KEYSERUNG, 1881) FL RG N Tegenaria africa C.L. KocH, 181 3 FL Enap/agnatha tharacica (HAHN , 1833) FL Tegenaria damestica (CLERCK, 1757) FL Episinus angulatus (BlACKWALL, 1836) FL Tegenaria ferruginea (PANZER, 1801) FL Episinus truncatus LATREILLE, 1809 FL CR Godd Tegenaria pagana C.L. KocH, 1811 FL IK y Euryopis flavomacu/ata (C.L. KocH, 1836) FL vu Godr 136 J.-P. MAELFAIT, L. BAERT, M. JANSSEN & M. ALDERWEIRELDT

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Euryopis /aeta (WESTRING, 1861) Metellina merianae (ScoPou, 1773) FL Pho/comma gibbum (WESTRING , 1851 ) FL vu Godr Metellina segmentata (CLERCK, 1757) FL Robertus arundineti (D.P.-CAMBRIDGE, 1871) FL EN Gowr Zygiel/a africa (C.L. KocH, 181 5) FL Robertus kuehnae BAUCHHENS & UHLENHAUT, 1993 FL IN X Zygiel/a stroemi (THORELL, 1870) FL IN X Robertus lividus (BLACKWALL, 1836) FL Zygiella x-notata (CLERCK, 1757) FL Robertus neg/ectus (D.P.-CAMBRIDGE, 1871) FL vu Fdwv Araneidae Robertus scoticus JACKSON, 1911 FL RG s Acu/epeira ceropegius (WALCKENAER, 1802) Steatoda albomaculata (DE GEER, 1778) FL vu Hd Agalenatea redii (ScoPou, 1763) FL vu Godd Steatoda bipunctata {lJNNAEus, 1758) FL Araneus a/sine (WALCKENAER, 1802) FL vu Fdwv Steatoda grossa (C.L. KocH, 1838) FL Araneus angulatus CLERCK, 1757 FL EN Fddv Steatoda phalerata (PANZER, 1801) FL vu Hd Araneus diadematus CLERCK, 1757 FL Steatoda triangu/osa (WALCKENAER, 1802) FL Araneus marmoreus CLERCK, 1757 FL Theonoe minutissima (O.P.- CAMBRIDGE, 1879) FL vu Hws Araneus quadratus CLERCK, 1757 FL bellicosum (SIMON, 1873) Araniella a/pica (L. KocH, 1869) Theridion bimaculatum {LlNNAEus, 1767) FL Araniella cucurbitina (CLERCK, 1757) FL Theridion blackwalli D.P.-CAMBRIDGE, 1870 FL Araniel/a displicata (HENTZ, 1817) FL Theridion familiare D.P.-CAMBRIDGE, 1871 FL Araniella insconspicua (SIMON, 1871) Theridion hannoniae DENIS, 1911 FL Araniella opistographa (KuLCZVNSKI, 1905) FL Theridion hemerobium SIMON, 19 11 FL Araniella proxima (KuLCZVNSKI, 1885) Theridion impressum L. KocH, 1881 FL Argiope bruennichi (ScoPou, 1772) FL RG N Theridion instabile O.P. -CAMBRIDGE, 1871 FL EN Me Ate a sturmi (HAHN, 1831) FL Theridion melanurum HAHN, 1831 FL Atea triguttata (FABRICIUS, 1757) FL Theridion mystaceum L. KocH, 1870 FL Cercidia prominens (WESTRING, 1851) FL Theridion pictum (WALCKENAER , 1802) FL Cyclosa conica (PALLAS, 1772) FL Theridion pinastri L. KocH, 1872 FL Cyclosa oculata (WALCKENAER, 1802) FL RG N Theridion simile C.L. KocH, 1836 FL Gibbaranea bituberculata (WALCKENAER, 1802) FL IN X Theridion sisyphium (CLERCK, 1757) FL Gibbaranea gibbosa (WALCKENAER, 1802) FL Theridion tinctum (WALCKENAER, 1802) FL Hypsosinga albovittata (WESTRING, 1851) FL vu Godd Theridion uh/ighi MARTIN, 1975 FL CR Hdb Hypsosinga heri (HAHN, 1831) FL EN Gowr Theridion varians HAHN, 1833 FL HypstJsinga pygmaea (SuNDEVALL, 1831) FL EN Hw Nesticicdae Hypsosinga sanguinea (CL KocH, 181 1) FL EN Hd Nesticus cel/ulanus (CLERCK, 1757) FL Larinioides cornutus (CLERCK, 1757) FL Tetragnathidae Larinioides patagiatus (CLERCK, 1757) FL Pachygnatha clercki SuNDEVALL, 1823 FL Larinioides sclopetarius (CLERCK, 1757) FL Pachygnatha degeeri SUNDEVALL, 1830 FL Mangora acalypha (WALCKENAER, 1802) FL Pachygnatha /isteri SUNDEVALL, 1830 FL vu Fdmo Neoscona adianta (WALCKENAER, 1802) FL "' Tetragnatha dearmata THORELL, 1873 FL IK y Nuctenea umbratica (CLERCK, 1757) FL Tetragnatha extensa {LlNNAEus, 1758) FL Singa hamata (CLERCK, 1757) FL EN Gowr Tetragnatha montana SIMON, 1871 FL Singa nitidula C.L. KOCH, 1815 Tetragnatha nigrita LENDL, 1886 FL Zilla diodia (WALCKENAER, 1802) FL Tetragnatha obtusa C.L. KocH, 1837 FL Anapidae Tetragnatha pinicola L. KocH, 1870 FL Comaroma simoni BERTKAU, 1889 Tetragnatha reimoseri (RoscA, 1939) FL CR Mr Theridiosomatidae Tetragnatha striata L. KocH, 1862 FL EN Mr Theridiosoma gemmosum (L. KocH, 1877) FL EN Fdmo Metidae Erigoninae Meta menardi (LATREILLE, 1801) Acartauchenius scurrilis (D.P. -CAMBRIDGE, 1872) FL EN Godta Metellina mengei (BLACKWALL, 1869) FL Araeoncus crassiceps (WESTRING , 1861) FL EN Hws A Red list for the s piders of Flanders I I 13 7

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Araeoncus humilis (BLACKWALL, 1811 ) FL Gongylidium rufipes (SUNDEVALL, 1829) FL Baryphyma duffeyi (MILUDGE, 1951) FL CR Sa Hy/yphantes graminicola (SUNDEVALL, 1829) FL Baryphyma maritimum (CROCKER &P ARKER, 19 70) FL EN Godtr Hylyphantes nigritus (SIMON, 1881) FL RG N Baryphyma pratense (BLACKWALL, 1861) FL vu Gowr Hypomma bituberculatum (WIDER, 1831) FL Baryphyma trifrons (D.P.-C AMBRIDGE , 1863) FL CR Hdb Hypomma cornutum (BLACKWALL, 1833) FL Ceratine/la brevipes (WESTRING, 1851) FL Hypomma fulvum BosENBERG, 1902 FL vu Mrr Ceratine/la brevis (WiDER, 1831) FL Hypselistes jacksoni (D.P.-C AMBRIDGE, 1902 ) FL EN Hws Ceratinella scabrosa (D.P.-CAMBRIDGE, 1871) FL Jacksonella falconeri (JACKSON , 1908 ) Ceratinopsis romana (D.P.-CAMBRIDGE, 1872) FL EN Godt Lessertia dentic/1elis (SIMON, 1881) FL Ceratinopsis stativa (SIMON, 1881) FL RG N Lophomma punctatum (BLACKWALL, 1811 ) FL Cinetata gradata (SIMON, 1881) Maso gallicus SIMON, 1891 FL EN Gowr Cnephalocotes obscurus (BLACKWALL, 1831) FL Maso sundevalli (WESTRING, 1851) FL Dicymbium brevisetosum LocKET, 1962 FL Metopobactrus prominulus (D.P.-C AMBRIDGE, 1872) FL vu Godb Dicymbium nigrum (BLACKWALL, 1831) FL Micrargus apertus (D.P.-CAMBRIDGE, 1870) FL IN X Dicymbium tibiale (BLACKWALL, 1836) FL Micrargus herbigradus (BLACKWALL, 1851 ) FL Dip/acephalus caecus DENIS, 1952 Micrargus laudatus (D.P. -CAMBRIDGE, 1881 ) Dip/acephalus crista/us (BLACKWALL, 1833) FL Micrargus subaequalis (WESTRING, 1851) FL Dip/acephalus latifrons (D.P.-CAMBRIDGE, 1863) FL Microctenonyx subitaneus (D.P. -CAMBRIDGE, 1875) FL Dip/acephalus lusiscus (SIMON, 1872) Milleriana inerrans (D.P.-CAMBRIDGE, 1881) FL Dip/acephalus permixtus (D.P. -CAMBRIDGE, 1871) FL Minicia margine/fa (WiDER , 1831) Dip/acephalus picinus (BLACKWALL, 1811 ) FL Minyriolus pusillus (WiDER, 1831) FL Dismodicus bifrons (BLACKWALL, 1811) FL Mioxena blanda (SIMON, 1884) FL RG N Dismodicus elevatus (C.L. KocH, 1838) FL RG s Moebelia penicillata (WESTRING, 1851) FL Entelecara acuminata (WIDER, 1831) FL Monocephalus castaneipes (SIMON. 1881) FL RG s Entelecara congenera (D.P.-C AMBRIDGE, 1879) FL RG s Monocephalus fuscipes (BLACKWALL, 1836) FL Entelecara errata D.P.-CAMBR IDGE, 1913 FL R G N Nematogmus sanguinolentus (WALCKENAER, 1837) Entelecara erythropus (WESTRING, 1851) FL Notioscopus sarcina/us (D.P.-CAMBRIDGE, 1872) FL CR Hws Entelecara flavipes (BLACKWALL, 1831) FL IN X Oedothorax agrestis (BLACKWALL, 1853) FL Entelecara media KuLClYNSKI, 1887 FL RG s Oedothorax apicatus (BLACKWALL, 1850) FL Entelecara omissa D.P.- CAMBRIDGE, 1902 FL CR Gowt Oedgthorax fuscus (BLACKWALL, 1831 ) FL Erigone arctica (WHITE, 1852) FL rledothorax gibbosus (BLACKWALL, 1811 ) FL vu Fdmo Erigone atra (BLACKWALL, 1811) FL Oedothorax retusus (WESTRING, 1851) FL Erigone dentipalpis (WIDER, 1831) FL Panamamops sulcifro ns (WIDER, 1831) Erigone longipalpis (SuNDEVALL, 1830) FL Parapelecopsis nemoralis (BLACKWALL, 1841) FL EN Godtr Erigone promiscua (D.P.-CAMBRIDGE, 1872) FL CR Godb Pelecopsis elongata (WiDER , 1831) FL IK y (WIDER, 1831) Erigonella hiemalis (BLACKWALL, 1811) FL RG s Pelecopsis para/lela FL ., Erigonella ignobilis (D.P. -CAMBRIDGE, 1871) FL EN Gowt Pelecopsis radicicola (L. KocH, 1875) FL IN X Erigonop/us g/obipes (C.L. KocH, 1872) Peponocranium ludicrum (D.P. -CAMBRIDGE, 1861) FL Erigonoplus justus (D.P. -CAMBRIDGE, 1875) Peponocranium orbiculatum (D.P.- CAMBRIDGE, 1882) G/yphesis cottonae (LA ToucHE, 19 15) FL CR Hws Pocadicnemis juncea LocKET & MILUDGE, 1953 FL G/yphesis servulus (SIMON, 1881) FL EN Fdmot Pocadicnemis pumila (BLACKWALL, 1811) FL Gnathonarium dentatum (WiDER, 1831) FL Prinerigone vagans AuDOUIN, 1826 FL Gonatium rubellum (BLACKWALL, 181·1) FL Pseudocarorita thaleri (SAARISTO, 1971) FL RG s Gonatium rubens (BLACKWALL, 1833) FL Pseudomaro aenigmaticus DENIS, 1966 FL RG N Gongylidiellum latebricola (D.P.- CAMBRIDGE, 1871) FL Saloca diceros (D.P. -CAMBRIDGE, 1871) FL vu Fclwo Gongylidiellum murcidum SiMON, 1881 FL CR Mrr Satilatlas britteni (JACKSON, 1913) FL CR Hw Gongy/idiellum vivum (D.P. -CAMBRIDGE, 1875) FL Savignia frontata BLACKWALL, 1833 FL vu Gowr 138 J.-P. MAELFAIT, L. BAERT, M. JANSSEN & M. ALDERWEIRELDT I'

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Silometopus ambiguus (D.P.-CM1BRJDGE, 1905) FL CR Sa Agyneta conigera (D.P.-CM1BRJDGE, 1863) FL Silometopus bonessi CASEMIR, 1970 FL CR Hd Agyneta decora (D.P.-CM1BRIDGE, 1870) FL Silometopus elegans (D.P.-CM1BRJDGE, 1905) FL vu Gowr Agyneta ramosa JACKSON, 1912 FL Silometopus incurvatus (D.P.-CAMBRIDGE, 1873) FL EN Godb Agyneta subtilis (D.P.-CM1BRIDGE, 1863) FL Silometopus reussi (THORELL, 1871) FL vu Gowr Allomengea scopigera (GRUBE, 1859) FL CR Sa Tapinocyba biscissa (D.P.-CM1BRIDGE, 1872) FL IN X Allomengea vidua (L. KocH, 1879) FL vu Gowr Tapinocyba insecta (L. KocH, 1869) FL Aphileta misera (D.P.-CM1BRIDGE, 1882) FL EN Hws Tapinocyba pal/ens (D.P.-CM1BRIDGE, 1872) FL IN X Asthenargus paganus (SiMON, 1881) FL IN X Tapinocyba praecox (D.P.-CM1BRIDGE, 1873) FL Bathyphantes approximatus (D.P.-CM1BRIDGE, 1871) FL Tapinocyboides pygmaea (MENGE, 1869) Bathyphantes gracilis (BLACKWALL, 181 1) FL Thyreostenius biovatus (D.P.-CM1BRIDGE, 1875) FL CR Godta Bathyphantes nigrinus (WESTRING, 1851) FL Thyreostenius parasiticus (WESTRING, 1851) FL Bathyphantes parvulus (WESTRING, 1851) FL Tiso vagans (BLACKWALL, 1831) FL Bathyphantes setiger F.D.P.-C M1BRIDGE, 1891 FL EN Hws Tmeticus affinis (BLACKWALL, 1811) FL vu Mr Bathyphantes simillimus (L. KocH, 1879) Trematocephalus cristatus (WiDER, 1831) FL vu Fddv Bathyphantes vittiger SIMON, 1881 Trichoncus hackmanni MILUDGE, 1956 FL CR Rg Bolyphantes alticeps (SuNDEVALL, 1832) Trichoncus saxicola (D.P.-CM1BRIDGE, 1861) FL IN X Bolyphantes luteolus (BLACKWALL, 1833) FL RG s Trichoptema cito (D.P.-CM1BRIDGE, 1872) FL vu Godb Carorita paludosa DuFFEY, 1971 FL EN Mrr Trichoptema thorelli (WESTRING, 1861) FL EN Gowr Centromerita bicolor (BLACKWALL, 1833) FL Troxochrus cirrifrons (D.P.-CM1BRIDGE, 1871) FL Centromerita concinna (THORELL, 1875) FL Troxochrus nasutus ScHENKEL, 1925 Centromerus aequalis (C.L. KocH, 1841) FL Troxochrus scabriculus (WESTRING, 1851) FL Centromerus arcanus (D.P.-CM1BR IDGE, 1873) Typhochrestus digitatus (D.P. -CM1BRIDGE, 1872) FL Centromerus capucinus (SiMON, 1881) FL RG N Typhochrestus simoni DELEssERT, 1907 Centromerus dilutus (D.P. -CAMBRIDGE, 1875) FL Walckenaeria acuminata BLACKWALL, 1833 FL Centromerus incilium (L. KocH, 1881) Walckenaeria alticeps (DENIS, 1952) FL CR Mo Centromerus leruthi FAGE, 1933 FL RG N Walckenaeria an fica (WiDER, 1831) FL Centromerus levitarsus (SIMON, 1881) FL CR Mo Walckenaeria atrotibialis D.P. -CM1BRIDGE, 1878 FL Centromerus pabulator (D.P .-CM1BRIDGE, 1875) FL RG w Walckenaeria capita (WESTRING, 1861) FL IN X Centromerus persimilis (D.P. -CM1BRIDGE, 191 2) FL IN X Walckenaeria comiculans (D.P.-CAMBRIDGE, 1875) FL vu Fdwo Centrgrnerus prudens (D.P.-CAMBRIDGE, 1873) FL Walckenaeria cucullata (C.L. KocH, 1836) FL Cenfromerus semiater (L. KocH, 1879) FL EN Hw Walckenaeria cuspidata (BLACKWALL, 1833) FL Centromerus serratus (D.P.-CM1BRIDGE, 1875) FL RG N Walckenaeria dysderoides (WIDER, 1831) FL Centromerus sylvaticus (BLACKWALL, 1811 ) FL Walckenaeria furcillata (MENGE, 1869) FL Diplostyla concolor (WiDER, 1831) FL Walckenaeria incisa (D.P. -CAMBRIDGE, 1871) FL Donacochara speciosa (THORELL, 1875) FL vu Gowr Walckenaeria kochi (D.P.-CM1BRIDGE, 1872) FL RG s Drapetisca socialis (SUNDEVALL, 1832) FL "' Walckenaeria mitrata (MENGE, 1868) FL EN Fddd Drepanotylus uncatus (D.P.-CM1BRIDGE, 1873) FL EN Mo Walckenaeria monoceros (WIDER, 1831) FL Floronia bucculenta (CLERCK, 1757) FL Walckenaeria nodosa D.P. -CM1BRIDGE, 1873 FL EN Gowr Frontinellina frutetorum (C.L. KocH, 1831) FL RG w Walckenaeria nudipalpis (WESTRING, 1851) FL Halorates distinctus (SiMON, 1879) FL EN Gowr Walckenaeria obtusa BLACKWALL, 1836 FL Haloratus reprobus (D.P.-CM1BRIDGE, 1879) FL CR Sa Walckenaeria stylifrons (D.P. -CM1BRIDGE, 1875) FL EN Godbr Helophora insignis (BLACKWALL, 1811) FL Walckenaeria unicomis D.P.-CM1BRIDGE, 1861 FL Hilaira excisa (D.P. -CM1BRIDGE, 1870) FL IN X Walckenaeria vigilax (BLACKWALL, 1833) FL Kaestneria dorsalis (WIDER, 1831) FL vu Godd Wiehlea calcarifera (SIMON, 1881) Kaestneria pullata (D.P.-CM1BRIDGE, 1863) FL Linyphiinae Labull a tlwracica (WIDER, 1831) FL IN X Agyneta cauta (D.P. -CM1BRIDGE, 1902) FL EN Fdwo Lepthyphantes alacris (BLACKWALL, 1853) A Red list for the spiders of Flanders 139

FAMILY, SPECIES FL RL-C HABITAT FAMILY, SPECIES FL RL-C HABITAT Lepthyphantes angulatus (O. P.-CAMBRIDGE, 1881) Meioneta saxatilis (BlACKWALL, 1811) FL Lepthyphantes angulipalpis (WESTRING, 1851) Meioneta simplicitarsis (SIMON, 1881) Lepthyphantes cristatus (MENGE, 1866) FL Microlinyphia impigra (O.P.-CAMBRIDGE, 1871) FL vu Mrr Lepthyphantes crucifer (MENGE, 1866) Microlinyphia pusil/a (SuNDEVALL, 1829) FL Lepthyphantes ericaeus (BlACKWALL, 1853) FL Microneta via ria (BlACKWALL, 1811) FL Lepthyphantes flavipes (BlACKWALL, 1851 ) FL Neriene c/athrata (SuNDEVALL, 1829) FL Lepthyphantes insignis O.P.-CAMBRIDGE, 1913 FL Neriene emphana (WALCKENAER, 1837) FL vu Fddv Lepthyphantes leprosus (OHLERT, 1865) FL Neriene furtiva (O.P.-CAMBRIDGE, 1871) FL EN Fddv Lepthyphantes lepthyphantiformis (STRAND, 1907) FL RG w Neriene hammeni 0/AN HELSDINGEN, 1963) FL EN Fddv Lepthyphantes mengei KULCZVNSKI, 1887 FL Neriene montana (CLERCK, 1757) FL Lepthyphantes minutus (BlACKWALL, 1833) FL Neriene peltata (WiDER, 1831) FL Lepthyphantes mughi (FICKERT, 1875) FL IK y Neriene radiata (WALCKENAER, 1811) FL CR Fddv Lepthyphantes obscurus (BlACKWALL, 1811) FL Oreonetides vaginatus (THORELL, 1872) Lepthyphantes pallidus (O.P.-CAMBRIDGE, 1871) FL Ostearius melanopygius (O .P. -CAMBRIDGE, 1879) FL Lepthyphantes Simoni CHYZER & KULCZVNSKI, 1891 Pityohyphantes phrygianus (C.L. KocH, 1836) Lepthyphantes tenebricola (WIDER, 1831) FL Poeciloneta globosa (WIDER, 1831) FL Lepthyphantes tenuis (BlACKWALL, 1852) FL Porrhomma campbelli F.O.P.-CAMBRIDGE, 1891 FL Lepthyphantes zimmermanni BERTKAU, 1890 FL Porrhomma convexum (WESTRING, 1861) FL Leptorhoptrum robustum (WESTRING, 1851) FL vu Rb Porrhomma egeria SIMON, 1881 FL Leptothrix hardyi (BlACKWALL, 1850) FL EN Gowd Porrhomma errans (BlACKWALL, 1811) FL Linyphia hortensis SUNDEVALL, 1829 FL Porrhomma microphthalmum (O.P.-CAMBRIDGE, 1871) FL Linyphia tenuipalpis SIMON, 1881 FL CR Hd Porrhomma myops SIMON, 1881 Linyphia triangularis (CLERCK, 1757) FL Porrhomma ob/itum (O.P.-CAMBRIDGE, 1870) FL Macrargus carpenteri (O.P.-CAMBRIDGE, 1891) FL RG s Porrhomma pallidum JACKSON, 1913 Macrargus rufus (WIDER, 1831) FL Porrhomma pygmaeum (BlACKWALL, 1831) FL Mara lehtineni SMRISTO, 1971 Porrhomma rosenhaueri (L. KocH, 1872) FL IN X Mara tepidus CASEMIR, 1963 Saaristoa abnormis (BlACKWALL, 1811) FL Mara minutus O.P.-CAMBRIDGE, 1906 FL IN X Saaristoa firma (O.P.-CAMBRIDGE, 1901) Mara sublestus FALCONER, 1915 FL EN Fdmo Sintula cornigera (BlACKWALL, 1856) FL IN X Mega/epthyphantes nebu/osus (SUNDEVALL, 1830) FL Stemonyphantes lineatus (LINNAEUS, 1758) FL Meioneta affinis (KULCZVNSKI, 1898) FL EN Godg Syedra gracilis (MENGE, 1866) FL IN X Meioneta fuscipa/pus (C.L. KocH, 1836) FL RG w Tal/usia experta (O.P.-CAMBRIDGE, 1871) FL Meioneta innotabilis (O.P.-CAMBRIDGE, 1863) FL Tapinopa longidens (WIDER, 1831) FL Meioneta mol/is (O.P.-CAMBRIDGE, 1871) FL Taranucnus setosus (O.P.-CAMBRIDGE, 1863) FL vu Hw Meioneta rurestris (C.L. KocH, 1836) FL Theonina cornix (SIMON, 1881)

CR, EN and VU the habitat is characterised by means of a - Fdmo: open Fdm code. For the RG-species it is indicated if they are at the - Fdmot: open Fdm with tussocks of sedges northern (N), southern (S) or western (W) limit of their geo­ - Fdw: wet deciduous forest graphical range. The following habitat types (and codes) were - Fdwl: Fdw rich in lichens distinguished: - Fdwo: open Fdw - Fdwv: verges of Fdw - Fd: deciduous forest - Go: oligotrophic grassland - Fdd: dry deciduous forest - God: dry oligotrophic grassland - Fddd: Fdd with large quantities of dead wood in the litter - Godb: God with patches of bare ground layer - Godba: God with patches of bare ground and ants - Fddo: open Fdd with well-developed undergrowth - Godbr: God with patches of barre ground on or close to first - Fddv: verges ofFdd dune ridge - Fddvs: southerly exposed verges ofFdd - Godbs: southerly exposed God with patches of bare ground - Fdm: marshy deciduous forest - Godd: God with dwarf shrubs 140 J.-P. MAELFAIT, L. BAERT, M. JANSSEN & M. ALDERWEIRELDT

EW

SA

IK (explanation: see text). Figure I. - Distribution of the Flemish spider species over Red list categories

Sa We

species (explanation: see text). Figure 2. - Habitat distribution of the extinct and threatened Flemish spider A Red list for the spiders of Flanders I I 14 1

- Godds: southerly exposed God with dwarf shrubs BAERT, L., 1996. Catalogus van de spinnen van Belgie. Deel - Godf: God rich in flowering plants XIV. Linyphiidae (Erigoninae). Studiedocumenten van he! - Godg: short-grazed God Koninklijk Belgisch liTstituut voor Natuunvetenschappen, 82: - Godl: lime-rich God 1-1 56. - Godr: God with rough vegetation BECKER, L., 1982. Les Arachnides de Belgique. Premiere partie. - Godt: God with grassy tussocks A nnales du Musee naturelle de Belgique, I 0: 1-246. tussocks and ants - Godta: God with grassy BECKE R, L., 1996. Les Arachnides de Belgique. Deuxieme et grassy tussocks on or close to first dune - Godtr: God with troisieme partie. Annates du Musee naturelle de Belgique, 12: ridge 1-378. - Godts: southerly exposed God with grassy tussocks UTTKE, H. & PRETSCHER, P., - Gow: wet oligotrophic grassland BI NOT, M., BLESS, R., BOYE, P., GR schlands, Bunde­ - Gowd: Gow with dwarf shrubs 1998. Rote Liste der gefuhrdeter Tiere Deut ad Godesberg. 434 pp. - Gowr: Gow with rough vegetation samt fiir Naturschutz, Bonn-B - Gowt: Gow with tussocks DE BLAUWE, R. & L. BAERT, 198 1. Catalogue des araignees de - Hd: dry heathland Belgique. Premiere partie. Famille des Agelenidae. Bulletin - Hdb: Hd with patches of bare ground de I 'fnstitut royal des Sciences naturelle de Belgique, 53 ( I): - Hdbs: southerly exposed Hd with patches of bare ground 1-37. - Hw: wet heathland JACOBS, J., 1993. Catalogus van de spinnen van Belgie. Dee! - Hws: Hw with Sphagnum carpets XII. Metidae. Studiedocumenten van het Koninklijk Belgisch ~ M: marshland Jnstituut voor Natuurwetenschappen, 7 1: 46-51. - Me: M with large sedges (Carex) JANSSEN, M. , 1993. Catalogus van de spinnen van Belgie. Dee! - Mo: oligotrophic marshland XIII. Thomisidae. Studiedocumenten van het Koninklijk Be!­ (Phragmitis) - Mr: M with reed vegetation gisch lnstituut voor Natuunvetenschappen, 71: 53-75. - Mrr: M with rough reed vegetation ERT. 1987. Catalogus van de spinnen van - R: riparian habitat JANSSEN, M. & L. BA edocumenten van het Ko­ - Rb: R with bare ground Belgie. Dee! IV. Salticidae. Studi Natuurwetenschappen, 43: - Rg: R with gravel llink!ijk Belgisch Instituut voor - Rs: sandy R 1-32. - Sa: saltmarsh MAELFAIT, J.-P., 1993. Rode lijsten: wat? hoe? waarom? Bulle­ - We: eutrophic stagnant or only slow-llowing water tin et Anna!les de Ia Societe royal beige d 'Entomologie, 129: - Wed: as We, covered with lloating water-plants 302-310. - Wes: as We, with well-developed submerged vegetation MAELFAIT, J-P., 1994. Rode lijsten van spi nnen, een nieuwe taakstelling voor ARABEL? Nieuwsbrief van de Belgische The habitat choice of the fN- and IK-species is not charac­ Arachnologische Vereniging, 9( I): 20-25. terised; the code X and Y is used respectively. MAELFAIT, J.-P., 1995. Recente ont\;vikkelingen in het projekt betreffende ongewervelde dieren van het Insti tuut voor Natuur­ behoud. Phegea, 23 (1): 59-64. Results and discussion MAES, D., J.-P. MAELFA IT & E. KUIJKEN, 1995. Rode lijsten: een onmisbaa r instrument in het m oderne Vlaamse nanmrbehoud. In Table I we have listed all the spider species mentioned as De Wie!p'aal, 61 (5): 149-156. occurring in Belgium. T hose occurring in Flanders are indi­ RANsY, M. & L. BAERT, 1985. Catalogus van de spinnen van cated (FL). In Figure 1 it can be seen that of the 604 Flem­ Belgie. Deel II. De Cribell atae. Studiedocumenten van het ish species only about half is safe or at low risk for the Koninklijk Be!gisch lnstituut voor Natuurwetenschappen, 26: moment (SA). The habitat distribution of the extinct (EW) 1-23. tened (CR, EN, VU) species is shown in Figure 2 . and threa RANsv, M. & L. BAERT, 1987a. Catalogue des araignees de fi gure we can d educe that the threatened part of From this Belgique. Troisieme partie. Les Araneidae. Documents de Tra­ biodiversity of Flanders occurs in oligotrophic the spider vail de I 'Jnstitut royal des Sciences naturel/es de Belgique, 36: heathland and several kinds grasslands, deciduous forests, 1-41. of wetland habitats. These are the habitat types prior atten­ ,,, des araignees de tion shou ld be directed t o by nature conservation policy ma­ RANSY, M. & L. BAERT, 1987b. Catalogue kers. Belgique. Cinquieme partie. Anyphaenidae, Argyronetidae, Atypidae, Dysderdidae, Mimetidae, Nesticidae, Oonopidae, Oxyopidae, Pholcidae, Pisauridae, Scytodidae, Segestriidae, Eusparassidae, Zodariidae et Zoridae. Documents de Travail References de I 'lnstitut royal des Sciences naturelles de Belgique, 46: 1-25. ALDERWE IRELDT, M. & J.-P. MAELFA IT, 1990. Catalogus van de RANSY, M. , KEKENBOSCH, J. & L. BAERT, 1987. Catalogue des spinnen van B elgie. Deel VII. Lycosidae. Studiedocumenten araignees de Belgique. Sixieme partie. Clubionidae et Liocra­ van het Koninklijk Belgisch lnstituut voor Natuurwetenschap­ nidae. Documents de Travail de I 'fnstitut royal des Sciences pen, 61: 1-92. naturelles d e Belgique, 67: 1-30. ALDERWEIRELDT, M. & J.-P. MAELFA IT, 1992. A proposal for the RANSY , M. & L. BA ERT. 199 1a. Catalogue des araignees de establishment of an red list of he Lycosid spiders in Flanders Belgique. Partie VIII. Gnaphosidae. Documents de Travail (Belgium). In: Van Goethem, J. & P. Grootae11 (Eds.), Proceed­ de I 'Institut royal des Sciences naturelles de Belgique, 64: ings of the 8th EIS meeting, Brussels, 1991 , 57-64. 7-30. 142 J.-P. MAELFAIT, L. BAERT, M. JANSSEN & M . ALDERWEIRELDT ''

RANSY, M. & L. BAERT, 1991b. Catalogue des araignees de Belgique. Partie X. Tetragnathidae. Documents de Tra vail de 1' Institut royal des Sciences naturelles de Belgique, 64: 43-52. SEGERS, H. & L. BAERT, 1991. Catalogus van de spinnen van Belgie. Dee! IX. Philodromidae. Studiedocumenten van het Koninklijk Belgisch Instituut voor Natuwwetenschappen, 64: 31 -42. VAN KEER, J. & H. VANUYTVEN , 1993. Catalogus van de spin­ nen van Belgie. Dee! XI. Theridiidae, Anapidae en Theridio­ somatidae. Studiedocumenten van het Koninklijk Belgisch In­ stituut voor Natuurwetenschappen, 71: 8-44. VERHEYEN, R.F., G. VAN SPAENDONCK, J.-P. MAELFAIT, M. HERMY, P. ULENAERS, K. ROMBOUTS & J. RUTTEN, 1994. Verlies van biodiversiteit.ln: A. Verbruggen (red.), Leren om te keren. Milieu- en natuurrapport Vlaanderen, Garant, Leuven/Apel­ doom: 497-520.

Jean-Pierre MAELFAIT Instituut voor Natuurbehoud Kliniekstraat 25 1070 Brussel

Leon BAERT Departement Entomologie Koninklijk Belgisch Instituut voor N atuurwetenscha ppen Vautierstraat 29 1000 Brussel

Marc JANSSEN Weg naar Ellikom 128 3670 Meeuwen-Gruitrode

Mark ALDERWEIRELDT Laboratorium voor Ecologie Ledeganckstraat 35 9000 Gent /

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