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Axioidea of the World and a Reconsideration of The Axioidea of the World and a Reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida) CRUSTACEANA MONOGRAPHS constitutes a series of books on carcinology in its widest sense. Contri- butions are handled by the Series Editor(s) and may be submitted through the office of KONINKLIJKE BRILL Academic Publishers N.V., P.O. Box 9000, NL-2300 PA Leiden, The Netherlands. Series Editors for the current volume: J.C. VON VAUPEL KLEIN, P.O. Box 30, NL-3720 AA Bilthoven, The Netherlands; e-mail: [email protected] and C.H.J.M. FRANSEN, c/o National Museum of Natural History, P.O. Box 9517, NL-2300 RA Leiden, The Netherlands; e-mail: [email protected] Editorial Committee: N.L. BRUCE, Wellington, New Zealand; Mrs. M. CHARMANTIER-DAURES, Mont- pellier, France; Mrs. D. DEFAYE, Paris, France; H. DIRCKSEN, Stockholm, Sweden; J. FOREST,Paris, France; R.C. GUIA¸SU, Toronto, Ontario, Canada; R.G. HARTNOLL,PortSt.Mary,IsleofMan; E. MACPHERSON, Blanes, Spain; P.K.L. NG, Singapore, Rep. of Singapore; H.-K. SCHMINKE,Old- enburg, Germany; F.R. SCHRAM, Langley, WA, U.S.A.; G. VA N D E R VELDE, Nijmegen, Netherlands; H.P. WAGNER, Leiden, Netherlands; D.I. WILLIAMSON, Port Erin, Isle of Man. Published in this series: CRM 001 - Stephan G. Bullard Larvae of anomuran and brachyuran crabs of North Carolina CRM 002 - Spyros Sfenthourakis et al. The biology of terrestrial isopods, V CRM 003 - Tomislav Karanovic Subterranean Copepoda from arid Western Australia CRM 004 - Katsushi Sakai Callianassoidea of the world (Decapoda, Thalassinidea) CRM 005 - Kim Larsen Deep-sea Tanaidacea from the Gulf of Mexico CRM 006 - Katsushi Sakai Upogebiidae of the world (Decapoda, Thalassinidea) CRM 007 - Ivana Karanovic Candoninae (Ostracoda) from the Pilbara region in Western Australia CRM 008 - Frank D. Ferrari & Hans-Uwe Dahms Post-embryonic development of the Copepoda CRM 009 - Tomislav Karanovic Marine interstitial Poecilostomatoida and Cyclopoida (Copepoda) of Australia CRM 010 - Carrie E. Schweitzer et al. Systematic list of fossil decapod crustacean species CRM 011 - Pedro Castro et al. (eds.) Studies on Brachyura: a homage to Danièle Guinot CRM 012 - Patricio R. De los Ríos-Escalante Crustacean zooplankton communities in Chilean inland waters CRM 013 - Katsushi Sakai Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida) In preparation (provisional titles): CRM 014 - Charles H.J.M. Fransen et al. (eds.) Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume CRM 015 - Akira Asakura et al. (eds.) New frontiers in crustacean biology – Proceedings of the TCS Summer Meeting, Tokyo, 20-24 September 2009 CRM 016 - Danielle Defaye et al. (eds.) Studies on freshwater Copepoda: a volume in honour of Bernard Dussart CRM 00x - Chang-tai Shih Marine Calanoida of the China seas Author’s address: Prof. Dr. K. SAKAI, 1570 Nishiyama, Kamihachiman-cho, Tokushima 770-8041, Japan; e-mail: [email protected] Manuscript first received 9 October 2008; final version accepted 5 April 2010. Cover: Gyanacaris hirsutimana (Boesch & Smalley, 1972), male; see p. 122, fig. 22A. Axioidea of the World and a Reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida) By Katsushi Sakai CRUSTACEANA MONOGRAPHS,13 LEIDEN • BOSTON This book is printed on acid-free paper. Library of Congress Cataloging-in-Publication Data The Library of Congress Cataloging-in-Publication Data is available from the Publisher. ISBN: 978 90 04 17086 5 © 2011 by Koninklijke Brill NV, Leiden, The Netherlands. Koninklijke Brill NV incorporates the imprints BRILL, Hotei Publishing, IDC Publishers, Martinus Nijhoff Publishers and VSP. All rights reserved. No part of this publication may be reproduced, translated, stored in a retrieval system, or transmitted in any form or by any means, electronic, mechanical, photocopying, recording or otherwise, without prior written permission from the publisher. Authorization to photocopy items for internal or personal use is granted by Koninklijke Brill NV provided that the appropriate fees are paid directly to The Copyright Clearance Center, 222 Rosewood Drive, Suite 910, Danvers, MA 01923, USA. Fees are subject to change. PRINTED IN THE NETHERLANDS CONTENTS Preface..................................................................... xi Introduction ................................................................ 1 Infraorderclassification...................................................... 9 Superfamily Axioidea Huxley, 1879 . ......................................... 13 Family Axiidae Huxley, 1879 . ............................................ 15 Genus Acanthaxius Sakai & De Saint Laurent, 1989 . ................. 19 Genus Albatrossaxius gen.nov........................................ 29 Genus Alienaxiopsis gen.nov......................................... 32 Genus Allaxiopsis gen.nov........................................... 36 Genus Allaxius Sakai & De Saint Laurent, 1989 ........................ 40 Genus Amakusaxius gen.nov. ........................................ 44 Genus Anophthalmaxius De Man, 1905 ............................... 45 Genus Australocaris Poore & Collins, 2009 . .......................... 48 Genus Axiopsis Borradaile, 1903 ..................................... 50 Genus Axiorygma Kensley & Simmons, 1988 .......................... 63 Genus Axius Leach, 1815 . .......................................... 64 Genus Balssaxius gen.nov. .......................................... 73 Genus Boasaxius gen.nov............................................ 76 Genus Bouvieraxius Sakai & De Saint Laurent, 1989 ................... 79 Genus Bruceaxius gen.nov........................................... 82 Genus Calaxius Sakai & De Saint Laurent, 1989 ....................... 84 Genus Calocarides Wollebæk, 1908 .................................. 86 Genus Coelhocalaxius gen.nov....................................... 98 Genus Colemanaxius gen.nov........................................ 103 Genus Dorphinaxius Sakai & De Saint Laurent, 1989 . ................. 106 Genus Eutrichocheles Wood-Mason, 1876 . ............................ 109 Genus Guyanacaris gen.nov. ........................................ 119 Genus Heterocalaxius gen.nov. ...................................... 123 Genus Leonardsaxius gen.nov........................................ 124 Genus Levantocaris Galil & Clark, 1993 . ............................ 129 Genus Litoraxius Komai & Tachikawa, 2007 ........................... 130 Genus Manaxius Kensley, 2003 (sensu nov.) . .......................... 131 Genus Marianaxius Kensley, 1996 . ................................. 142 Genus Michelaxiopsis Poore & Collins, 2009 . ......................... 143 Genus Newzealandaxius gen.nov. .................................... 148 Genus Oxyrhynchaxius Parisi, 1917 ................................... 150 Genus Parascytoleptus Sakai & De Saint Laurent, 1989 ................. 152 Genus Paratrichocheles gen.nov...................................... 155 Genus Paraxiopsis De Man, 1905 . .................................. 156 Genus Paraxius Bate, 1888 . ........................................ 164 vi CRM 013 – Katsushi Sakai Genus Pilbaraxius Poore & Collins, 2009 ............................. 167 Genus Pillsburyaxius gen.nov........................................ 169 Genus Planaxius Komai & Tachikawa, 2008 ........................... 171 Genus Platyaxiopsis gen.nov......................................... 172 Genus Platyaxius Sakai, 1994 ........................................ 174 Genus Pseudoaxiopsis gen.nov....................................... 177 Genus Ralumcaris gen.nov........................................... 182 Genus Scytoleptus Gerstaecker, 1856 . ................................. 185 Genus Spongiaxius Sakai & De Saint Laurent, 1989 . ................... 193 Family Callianideidae Kossmann, 1880 . ................................... 197 Genus Callianidea H. Milne Edwards, 1837 . ......................... 198 Genus Paracallianidea Sakai, 1992 . ................................. 205 Family Calocarididae Ortmann, 1891 . ..................................... 209 Genus Ambiaxius Sakai & De Saint Laurent, 1989 . ................... 211 Genus Briancaris Sakai & Ohta, 2005 . .............................. 213 Genus Calastacus Faxon, 1893 . ..................................... 214 Genus Calaxiopsis Sakai & De Saint Laurent, 1989 . .................... 223 Genus Calocaris Bell, 1846 . ........................................ 228 Genus Calocarisopsis gen.nov........................................ 242 Genus Eucalastacus Sakai, 1992 . ..................................... 244 Genus Lophaxiopsis gen.nov......................................... 245 Genus Lophaxius Kensley, 1989 ...................................... 246 Genus Nipponcalaxiopsis gen.nov. ................................... 248 Genus Paracalocaris Sakai, 1991 . ................................... 250 Family Coralaxiidae Sakai & De Saint Laurent, 1989 ......................... 253 Genus Coralaxius Kensley & Gore, 1981 . ............................ 254 Genus Meinertaxius gen.nov......................................... 260 Family Eiconaxiidae Sakai & Ohta, 2005 . ................................... 267 Genus Eiconaxius Bate, 1888 . ..................................... 268 FamilyEiconaxiopsididaefam.nov. ........................................ 289 Genus Eiconaxiopsis gen.nov. ....................................... 290 Family Meticonaxiidae Sakai, 1992 . .....................................
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