Marine Flooding Event in Continental Triassic Facies Identified by A
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O'keefe, F. R. 2006
12 Neoteny and the Plesiomorphic Condition of the Plesiosaur Basicranium F. Robin O’Keefe Introduction Historically, the systematics of the Plesiosauria (Reptilia, Sauroptery- gia) were based largely on postcranial characters (Persson, 1963; Brown, 1981). Several factors account for this bias: plesiosaur skulls tend to be del- icate and are often crushed even when preserved, postcranial elements are relatively common and cranial elements are not, lack of knowledge about the relationships of stem-group sauropterygians, and lack of knowledge of plesiosaur cranial anatomy itself. However, recent detailed examinations of plesiosaur cranial anatomy have identified many characters of use in plesiosaur systematics (Brown, 1993; Cruickshank, 1994; Storrs & Taylor, 1996; Storrs, 1997; Carpenter, 1997; Evans, 1999; O’Keefe, 2001, 2004), and the systematics of the group have changed markedly in response (Carpenter, 1997; O’Keefe, 2001, 2004). The work of Rieppel and others has clarified the anatomy and relationships of stem-group sauropterygians (Storrs, 1991; see Rieppel, 2000, for review). This work has laid the anatomic and phylogenetic foundations for a better understanding of ple- siosaur cranial anatomy. The purpose of this paper is to describe the condition of the braincase in stratigraphically early and morphologically primitive plesiosaurs. In- formation on the braincase of plesiomorphic taxa is important because it establishes the polarity of characters occurring in more derived ple- siosaurs. This paper begins with a short review of braincase anatomy in stem-group sauropterygians. Data on braincase morphology of the ple- siomorphic plesiosaur genera Thalassiodracon and Eurycleidus are then presented and interpreted via comparison with other plesiosaurs, stem- group sauropterygians, and stem diapsids (Araeoscelis). -
A New Species of the Sauropsid Reptile Nothosaurus from the Lower Muschelkalk of the Western Germanic Basin, Winterswijk, the Netherlands
A new species of the sauropsid reptile Nothosaurus from the Lower Muschelkalk of the western Germanic Basin, Winterswijk, The Netherlands NICOLE KLEIN and PAUL C.H. ALBERS Klein, N. and Albers, P.C.H. 2009. A new species of the sauropsid reptile Nothosaurus from the Lower Muschelkalk of the western Germanic Basin, Winterswijk, The Netherlands. Acta Palaeontologica Polonica 54 (4): 589–598. doi:10.4202/ app.2008.0083 A nothosaur skull recently discovered from the Lower Muschelkalk (early Anisian) locality of Winterswijk, The Nether− lands, represents at only 46 mm in length the smallest nothosaur skull known today. It resembles largely the skull mor− phology of Nothosaurus marchicus. Differences concern beside the size, the straight rectangular and relative broad parietals, the short posterior extent of the maxilla, the skull proportions, and the overall low number of maxillary teeth. In spite of its small size, the skull can not unequivocally be interpreted as juvenile. It shows fused premaxillae, nasals, frontals, and parietals, a nearly co−ossified jugal, and fully developed braincase elements, such as a basisphenoid and mas− sive epipterygoids. Adding the specimen to an existing phylogenetic analysis shows that it should be assigned to a new species, Nothosaurus winkelhorsti sp. nov., at least until its juvenile status can be unequivocally verified. Nothosaurus winkelhorsti sp. nov. represents, together with Nothosaurus juvenilis, the most basal nothosaur, so far. Key words: Sauropterygia, Nothosaurus, ontogeny, Anisian, The Netherlands. Nicole Klein [nklein@uni−bonn.de], Steinmann−Institut für Geologie, Mineralogie und Paläontologie, Universtät Bonn, Nußallee 8, 53115 Bonn, Germany; Paul C.H. Albers [[email protected]], Naturalis, Nationaal Natuurhistorisch Museum, Darwinweg 2, 2333 CR Leiden, The Netherlands. -
Contributions in BIOLOGY and GEOLOGY
MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - --- -
Sauropterygia I Placodontia, Pachypleurosauria, Nothosauroidea, Pistosauroidea
Teil 12A / Part 12A Sauropterygia I Placodontia, Pachypleurosauria, Nothosauroidea, Pistosauroidea by O. RlEPPEL With 80 Figures Verlag Dr. Friedrich Pfeil • Munchen 2000 ISBN 3-931516-78-4 Contents Foreword (P. WELLNHOFER) V Acknowledgements VI Institutional Acronyms VII Figure Abbreviations VIII Introduction: History of the Concept of Sauropterygia 1 Phylogenetic Relationships of Stem-Group Sauropterygia 4 Stratigraphic and Geographic Distribution of Stem-Group Sauropterygia 6 General Skeletal Anatomy of Stem-Group Sauropterygia 10 Systematic Review 16 Superorder Sauropterygia OWEN, 1860 16 Order Placodontia COPE, 1871 16 Suborder Placodontoidea COPE, 1871 16 Family Paraplacodontidae PEYER & KUHN-SCHNYDER, 1955 17 Anatomy of Paraplacodontidae 17 Genus Paraplacodus PEYER, 1931 18 Family Placodontidae COPE, 1871 19 Anatomy of Placodontidae 19 Genus Placodus AGASSIZ, 1933 21 Suborder Cyamodontoidea NOPCSA, 1923 23 Anatomy of Cyamodontoidea 23 Interrelationships of Cyamodontoidea 25 Superfamily Cyamodontida NOPCSA, 1923 25 Family Henodontidae F. v. HUENE, 1948 26 Genus Henodus F. v. HUENE, 1936 26 Family Cyamodontidae NOPCSA, 1923 27 Genus Cyamodus MEYER, 1863 27 Superfamily Placochelyida ROMER, 1956 32 Family Macroplacidae nov. fam 32 Genus Macroplacus SCHUBERT-KLEMPNAUER, 1975 32 Family Protenodontosauridae nov. fam 33 Genus Protenodontosaurus PINNA, 1990 33 Family Placochelyidae ROMER, 1956 34 Genus Placochelys JAEKEL, 1902 34 Genus Psephoderma MEYER, 1858 36 Genus Psephosaurus E. FRAAS, 1896 38 Cyamodontoidea indet 39 IX Order Eosauropterygia -
Tesis Doctoral 2018
TESIS DOCTORAL 2018 HISTORIA EVOLUTIVA DE SIMOSAURIDAE (SAUROPTERYGIA). CONTEXTO SISTEMÁTICO Y BIOGEOGRÁFICO DE LOS REPTILES MARINOS DEL TRIÁSICO DE LA PENÍNSULA IBÉRICA CARLOS DE MIGUEL CHAVES PROGRAMA DE DOCTORADO EN CIENCIAS FRANCISCO ORTEGA COLOMA ADÁN PÉREZ GARCÍA RESUMEN Los sauropterigios fueron un exitoso grupo de reptiles marinos que vivió durante el Mesozoico, apareciendo en el Triásico Inferior y desapareciendo a finales del Cretácico Superior. Este grupo alcanzó su máxima disparidad conocida durante el Triásico Medio e inicios del Triásico Superior, diversificándose en numerosos grupos con distintos modos de vida y adaptaciones tróficas. El registro fósil de este grupo durante el Triásico es bien conocido a nivel global, habiéndose hallado abundantes restos en Norteamérica, Europa, el norte de África, Oriente Próximo y China. A pesar del relativamente abundante registro de sauropterigios triásicos ibéricos, los restos encontrados son, por lo general, elementos aislados y poco informativos a nivel sistemático en comparación con los de otros países europeos como Alemania, Francia o Italia. En la presente tesis doctoral se realiza una puesta al día sobre el registro ibérico triásico de Sauropterygia, con especial énfasis en el clado Simosauridae, cuyo registro ibérico permanecía hasta ahora inédito. Además de la revisión de ejemplares de sauropterigios previamente conocidos, se estudian numerosos ejemplares inéditos. De esta manera, se evalúan hipótesis previas sobre la diversidad peninsular de este clado y se reconocen tanto formas definidas en otras regiones europeas y de Oriente Próximo, pero hasta ahora no identificadas en la península ibérica, como nuevos taxones. La definición de nuevas formas y el incremento de la información sobre otras previamente conocidas permiten la propuesta de hipótesis filogenéticas y la redefinición de varios taxones. -
Sauropterygia, Placodontia) from the Late Triassic of Schesaplana Mountain, Graubu¨Nden, Switzerland
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by RERO DOC Digital Library Swiss J Geosci (2014) 107:349–357 DOI 10.1007/s00015-014-0173-9 New specimen of Psephoderma alpinum (Sauropterygia, Placodontia) from the Late Triassic of Schesaplana Mountain, Graubu¨nden, Switzerland James M. Neenan • Torsten M. Scheyer Received: 22 October 2014 / Accepted: 22 October 2014 / Published online: 20 November 2014 Ó Swiss Geological Society 2014 Abstract Psephoderma alpinum is an armoured, duro- Abbreviations phagous placodont known from the alpine Late Triassic. IVPP Institute of Vertebrate Paleontology and Here we present a new, well-preserved isolated skull dis- Paleoanthropology, China covered in the Alplihorn Member (Late Norian–Early MB.R. Museum fu¨r Naturkunde der Humboldt Rhaetian) of the Ko¨ssen Formation, Schesaplana Mountain, Universita¨t, Berlin, Germany which straddles the Swiss/Austrian border. Micro-com- MSNM Museo Civico di Storia Naturale di Milano, Italy puted tomographic (lCT) scanning was used to create an PIMUZ Palaeontological Institute and Museum, accurate osteological reconstruction of the specimen, the University of Zurich, Switzerland first time this has been conducted for Psephoderma.We thus clarify disputed anatomical features from previous descriptions, such as a lack of a lacrimal and a pineal foramen that is enclosed by the parietal. We also present 1 Introduction the first description based on lCT data of the lateral braincase wall, sphenoid region and some cranial nerve Psephoderma alpinum Meyer 1858 is a highly-nested, canals for Psephoderma, with the location of the hypo- armoured placodont best known from the Norian and physeal seat, cerebral carotid foramina, dorsum sellae, Rhaetian (Late Triassic) of the Alps (e.g., Rieppel 2000, prootic foramen, lacrimal foramen, as well as all dental 2001). -
O'keefe, F. R. and M. T. Carrano. 2005
Paleobiology, 31(4), 2005, pp. 656±675 Correlated trends in the evolution of the plesiosaur locomotor system F. Robin O'Keefe and Matthew T. Carrano Abstract.ÐThis paper investigates trends in the evolution of body size and shape in the Plesiosau- ria, a diverse clade of Mesozoic marine reptiles. Using measures from well-preserved plesiosaur specimens, we document and interpret evolutionary patterns in relative head size, body size, and locomotor variables. Size increase is a signi®cant trend in the clade as a whole, and in constituent clades. The trend in relative head size is of variance increase; observed head sizes are both smaller and larger than ancestral values. In the locomotor system, changes in propodial and girdle pro- portions appear concomitant with body size increase and are interpreted as allometric responses to the physical constraints of large body size. Other trends in the locomotor system are signi®cantly correlated with both body size and relative head size. These locomotor trends evolved convergently in several clades of plesiosaurs, and may have had an ecomorphological basis, although data are lacking to constrain speculation on this point. The evolution of the locomotor system in plesiosaurs sheds new light on the response of aquatic tetrapods to the physical constraints of foraging at large body size. F.Robin O'Keefe. Department of Anatomy, NYCOM II, Room 326, New York College of Osteopathic Med- icine, New York Institute of Technology, Old Westbury, New York 11568. E-mail: [email protected] Matthew Carrano. Department of Paleobiology, Post Of®ce Box 37012, MRC 121, Smithsonian Institution, Washington, D.C. -
A Cladistic Analysis and Taxonomic Revision of the Plesiosauria (Reptilia: Sauropterygia) F
Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences 12-2001 A Cladistic Analysis and Taxonomic Revision of the Plesiosauria (Reptilia: Sauropterygia) F. Robin O’Keefe Marshall University, [email protected] Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Aquaculture and Fisheries Commons, and the Other Animal Sciences Commons Recommended Citation Frank Robin O’Keefe (2001). A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia). ). Acta Zoologica Fennica 213: 1-63. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. Acta Zool. Fennica 213: 1–63 ISBN 951-9481-58-3 ISSN 0001-7299 Helsinki 11 December 2001 © Finnish Zoological and Botanical Publishing Board 2001 A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia) Frank Robin O’Keefe Department of Anatomy, New York College of Osteopathic Medicine, Old Westbury, New York 11568, U.S.A Received 13 February 2001, accepted 17 September 2001 O’Keefe F. R. 2001: A cladistic analysis and taxonomic revision of the Plesio- sauria (Reptilia: Sauropterygia). — Acta Zool. Fennica 213: 1–63. The Plesiosauria (Reptilia: Sauropterygia) is a group of Mesozoic marine reptiles known from abundant material, with specimens described from all continents. The group originated very near the Triassic–Jurassic boundary and persisted to the end- Cretaceous mass extinction. This study describes the results of a specimen-based cladistic study of the Plesiosauria, based on examination of 34 taxa scored for 166 morphological characters. -
Evolution Et Extinction Des Reptiles Marins Au Cours Du Mesozoique
EVOLUTION ET EXTINCTION DES REPTILES MARINS AU COURS DU MESOZOIQUE par Nathalie BARDET * SOMMAIRE Page Résumé, Abstract . 178 Introduction ..................................................................... 179 Matériel et méthode . 181 La notion de reptile marin . 181 Etude systématique . 182 Etude stratigraphique. 183 Méthodes d'analyse. 183 Systématique et phylogénie. 184 Le registre fossile des reptiles marins . 184 Affinités et phylogénie des reptiles marins. 186 Analyses taxinomique et stratigraphique. 187 Testudines (Chelonia) . 187 Squamata, Lacertilia . 191 Squamata, Serpentes. 193 Crocodylia ............................................................... 194 Thalattosauria . 195 Hupehsuchia . 196 Helveticosauroidea . 197 Pachypleurosauroidea . 197 Sauropterygia .... 198 Placodontia. 198 * Laboratoire de Paléontologie des Vertébrés, URA 1761 du CNRS, Université Pierre et Marie Curie, Case 106,4 Place Jussieu, 75252 Paris cédex 05, France. Mots-clés: Reptiles marins, Tortues, Lézards, Serpents, Crocodiles, Thalattosaures, Hupehsuchiens, Helveticosaures, Pachypleurosaures, Nothosaures, Placodontes, Plésiosaures, Ichthyosaures, Mésozoïque, Evolution, Extinction, Assemblages et Renouvellements fauniques. Key-words: Marine Reptiles, Turtles, Lizards, Snakes, Crocodiles, Thalattosaurs, Hupehsuchians, Helveticosaurs, Pachypleurosaurs, Nothosaurs, Placodonts, Plesiosaurs, Ichthyosaurs, Mesozoic, Evolution, Extinction, Faunal Assemblages and Turnovers. Palaeovertebrata. Montpellier. 24 (3-4): 177-283, 13 fig. (Reçu le 4 Juillet 1994, -
Now, on to Mesozoic Marine Reptiles
Now, on to Mesozoic Marine Reptiles NOT DINOSAURS! • They are reptiles, but some have adopted different skull fenestration • “Euryapsid” and “Anapsid” conditions are likely modified Diapsids • First reptiles returned to the sea in the Permian (Mesosaurus) How they are related: Sauropterygia Mosasauridae Thalattosuchia Dinosauria Phytosauria Lepidosauromorpha Crurotarsi Prolacertiformes Archosauromorpha Thalattosauriformes? Ichthyosauromorpha Sauria Claudiosaurus Neodiapsida Morphology • 4 basic body plans (Baupläne): (a) Thunniform advanced ichthyosaur (b) Long neck/small head plesiosaur (elasmosaur) (c) Short neck/big head pliosaur (d) Undulatory mosasaur (and basal ichthyosaur) • + “functional group 3” after Robert L. Carroll (“swimming lizards”) Morphological Trends • Limbs become rigid, often with hyperphalangy (many phalanges) • Polydactyly in ichthyosaurs Morphological Trends • (A) Merriamia (basal ichthyosaur) manus • (B) Opthalmosaurus (Jurassic ichthyosaur) manus • (C) Elasmosaurus (Cretaceous plesiosaur) manus • (D) Nothosaur pes • (E) Mosasaur pes Morphological Trends • Thoracic stiffening a usual trend • Lateral flexion directed posteriorly, or propulsion moved paraxially Carrier’s Constraint • Because of their sprawling gait, *GASP!* reptiles cannot breathe and run at the same time Ahhh… • Same applies to marine reptiles with lateral flexion (they breathe air!) *GASP!* • Solved by moving propulsion posteriorly, stiffening thorax, or moving limbs independently of spine Triassic Seas • Pangea beginning to break up • Marine -
Sepkoski, J.J. 1992. Compendium of Fossil Marine Animal Families
MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. Department of the Geophysical Sciences University of Chicago Chicago, Illinois 60637 Milwaukee Public Museum Contributions in Biology and Geology Rodney Watkins, Editor (Reviewer for this paper was P.M. Sheehan) This publication is priced at $25.00 and may be obtained by writing to the Museum Gift Shop, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Orders must also include $3.00 for shipping and handling ($4.00 for foreign destinations) and must be accompanied by money order or check drawn on U.S. bank. Money orders or checks should be made payable to the Milwaukee Public Museum. Wisconsin residents please add 5% sales tax. In addition, a diskette in ASCII format (DOS) containing the data in this publication is priced at $25.00. Diskettes should be ordered from the Geology Section, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Specify 3Y. inch or 5Y. inch diskette size when ordering. Checks or money orders for diskettes should be made payable to "GeologySection, Milwaukee Public Museum," and fees for shipping and handling included as stated above. Profits support the research effort of the GeologySection. ISBN 0-89326-168-8 ©1992Milwaukee Public Museum Sponsored by Milwaukee County Contents Abstract ....... 1 Introduction.. ... 2 Stratigraphic codes. 8 The Compendium 14 Actinopoda. -
Novel Anatomy of Cryptoclidid Plesiosaurs with Comments on Axial Locomotion
NOVEL ANATOMY OF CRYPTOCLIDID PLESIOSAURS WITH COMMENTS ON AXIAL LOCOMOTION. A Thesis submitted to the Graduate College of Marshall University In partial fulfillment of The requirements for the degree of Master of Science In Biological Science by Benjamin C. Wilhelm Approved by Dr. F. Robin O’Keefe, Ph.D., Advisor, Committee Chairperson Dr. Brian L. Antonsen, Ph.D. Dr. Victor Fet, Ph.D. Dr. Suzanne G. Strait, Ph.D. Marshall University May 2010 Acknowledgements This thesis was greatly improved by the comments of my advisor Dr. F. R. O’Keefe and committee members Dr. B. Antonsen, Dr. V. Fet, and Dr. S. Strait. Description of the specimens presented in chapter 2 would not have been possible without the skilled preparation of J. P. Cavigelli. S. Chapman, H. Ketchum , and A. Milner arranged access to specimens at the BMNH. Chapter 2 was also improved by the comments of Dr. R. Schmeisser and an anonymous reviewer. This research was supported by a grant from the National Geographic Society (CRE 7627-04) and Marshall Foundation funds to Dr. F. R. O’Keefe and MURC student travel grants to B. C. Wilhelm. Many thanks are also due to A. Spriggs, my parents, George and Kris, and my brother Greg for all the love and support during the course of my Master’s studies. ii Table of Contents Acknowledgements…………………………………………………………………….……..ii Table of contents…………………………………………………………………………..iii-iv List of Figures, Tables, and Graphs………………………………………………………..v Abstract………………………………………………………………………………………...vi Chapter 1- Introduction to plesiosaurs………………………………………………...1-14