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PROGRAMA FONDECYT

INFORME FINAL

ETAPA 2017

COMISIÓN NACIONAL DE INVESTIGACION CIENTÍFICA Y TECNOLÓGICA

VERSION OFICIAL NÂº 2

FECHA: 15/03/2018

Nº PROYECTO : 1150664 DURACIÓN : 3 años AÑO ETAPA : 2017 TÍTULO PROYECTO : MIOCENE MARINE DIVERSITY ALONG THE COAST OF CENTRAL TO SOUTHERN CHILE ACROSS MULTIPLE TAXA

DISCIPLINA PRINCIPAL : PALEONTOLOGIA GRUPO DE ESTUDIO : CS. DE LA TIERRA INVESTIGADOR(A) RESPONSABLE : SVEN NIKOLAUS NIELSEN DIRECCIÓN : CAMPUS ISLA TEJA S/N, EDIFICIO PUGÍN COMUNA : VALDIVIA CIUDAD : Valdivia REGIÓN : XIV REGION

FONDO NACIONAL DE DESARROLLO CIENTIFICO Y TECNOLOGICO (FONDECYT) Moneda 1375, Santiago de Chile - casilla 297-V, Santiago 21 Telefono: 2435 4350 FAX 2365 4435 Email: [email protected] INFORME FINAL PROYECTO FONDECYT REGULAR

MODIFICACIONES ACADÉMICAS

El informe no presenta modificaciones académicas. PROJECT RESULTS:

Describe the results of your research in reference to its original and/or modified Project objectives.

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Results The main objective of this project was to better know and understand early Miocene latitudinal gradients in Chile. For this, several fieldwork campaigns were conducted in the areas of Navidad, Arauco, Valdivia, Chiloé, Chonos islands, and Taitao peninsula to improve sampling intensity in the previously known regions and extend sampling towards the south. Several molluscan (and other) taxa new to science were discovered and geographic ranges of known species can be extended. The collections of Chilean fossils housed at the Paleontological Research Institution in Ithaca, USA, were visited and, as far as possible in only one week, revised and documented. A number of rare and undescribed taxa were identified and will be included in monographs on the respective molluscan groups. A few highlights for the areas:

Navidad: Most new localities reproduced the typical well-known mollusk fauna. However, especially a lithologically unusual locality near Maitenlahue yielded undescribed taxa of mollusks (e.g. Trivia) and small decapod crabs. Previously not recorded brachiopods were collected from Punta Perro. Arauco: A 3-dimensionally preserved fish skull was found at Ranquil and described in the master thesis of J. Pérez as belonging to the genus Hypoplectrodes. This genus occurs today mostly off Australia and New Zealand and has one species off Chilean tropical oceanic islands (thesis J. Pérez attached). Our fossil represents the first fossil record for the genus and adds to the Miocene Chile- New Zealand connection previously reported for mollusks. Valdivia: Little information exists about fossil invertebrates of the Santo Domingo Formation. An undergraduate thesis in biology will review and extend the knowledge of this unit starting this semester. At least one new bivalve species (Panopea) is present in our collection. Chiloé: New collections extend the range of several taxa from Navidad down south. Also it is now possible to include the enigmatic locality Puñihuil in the Lacuy Formation, due to the findings of gastropods, a coral and a brachiopod known from Navidad or equivalent units. Two geology undergraduate theses on fauna and stratigraphy are starting this semester. Fieldwork included detailed stratigraphic logging of the ~100m vertical coastal bluff section at Cucao. Chonos islands: Unusual findings include decapod crabs and new echinoderms. A master thesis on the foraminifera, clarifying age and bathymetry, will be finished during 2018 (not listed because not inscribed during 2017). Taitao peninsula: Taxa found which are new to science include mollusks, decapod crabs, and echinoderms. We were able to limit the age of the poorly known islands in the center of Tres Montes gulf to the Miocene (submitted master thesis Robles attached). Additionally, we found previously unreported localities that are significantly younger in age, likely Pliocene-Pleistocene, which extend the fossil record of this age by 2°lat to the south.

Problems The main obstacle for the execution of the project was that PI Nielsen had to step up as director of the newly created Institute of Earth Sciences at Universidad Austral de Chile (document attached). This was not foreseen and occurred simultaneous to the arrival of funds for the projects first year. As a consequence, the time dedicated to the project could not be met as promised and more than a quarter of the funds for the first year had to be returned to Fondecyt. While fieldwork was conducted and most samples are processed, classification of new taxa and writing of manuscripts are behind schedule. Start and development of the project were also hindered by the late arrival of funds, especially in the first year (July!), which effectively eliminated the envisaged work during winter holidays 2015. This resulted in postponing the trip to revise collections in the USA to the following year, disturbing the following plans. Both proposed international collaborators dropped out from the project as detailed below, so that the hypothesis on water temperatures could not be addressed and the part on foraminifera was severely crippled. Several students doing their theses within the frame of this project were hired before finishing. While this is certainly good for them, it delays progress of important parts of the project.

Specific goals I) To establish and compare latitudinal diversity gradients of mayor taxonomic groups from the Chilean Miocene. There are few definite results yet for this goal. For foraminifera and ostracods, this is partly due to the unexpected early (forced) retirement of international collaborator Finger, which also resulted in him dropping out of this project. However, a master thesis on the latitudinal gradient of foraminifera (Chávez) will be finished during 2018. Chávez is now teaching paleontology at Universidad Central in Santiago, which clearly slowed down progress of her thesis. Work on decapod crustaceans is advanced for the Chonos islands including some other localities like Mocha island. Seven new species will be described from relatively few specimens (22), significantly increasing the overall diversity of Chilean Miocene decapods. Work on classification of fish otoliths along the gradient, not originally considered in the proposal, has begun in collaboration with German specialist Werner Schwarzhans. The number of new mollusk taxa discovered during this project seems to be higher for the northernmost area (i.e., Navidad), but geographic ranges can now be extended for taxa formerly limited to single areas (e.g. Mitridae formerly only reported from Chiloé were collected in areas both to the north and south). Results of analysis of spatiotemporal biodiversity pattern of chondrichthyan fishes were done by co-PI Rivadeneira and his (former) student Villafaña (attached publication Villafaña & Rivadeneira in press), leading to the conclusion that multiple environmental exogenous factors interacted with taxon traits during the Neogene, creating a mosaic of biogeographic dynamics. To understand changes in faunal and functional diversity (see also point IV below) for the Miocene, we first needed to get a better grip on more (geologically) recent changes. An analysis of changes in gastropods was published for the Pliocene-Quaternary by the two PIs of this project (attachment Rivadeneira & Nielsen 2017).

II) To establish geographic patterns of different mayor and minor taxonomic groups from the Chilean Miocene (Naticidae, Muricidae, Protobranchia, Pectinidae). Work on the bivalve group Protobranchia (families Nuculidae and Nuculanidae) is conducted with master student L. Pérez and preliminary results were presented at several conferences (see there). After starting his thesis, Pérez was hired as curator of fossil invertebrates at Chiles National Museum of Natural History in Santiago, which clearly slowed down progress. Manuscripts on Naticidae, Lucinoidea, and Mitridae by PI Nielsen are advanced, while progress on Muricidae and Pectinidae is little. Understanding the driving factors for the evolution of latitudinal gradients is fundamental for our research and is treated in the attached publications by Rivadeneira et al. (2015) and Spano et al. (2016). with participation of co-PI Rivadeneira.

Abstract Lucinoidea (Nielsen, Reich, Kiel): The Cenozoic species of Lucinidae, Ungulinidae and Thyasiridae from Chile are revised. Types and new material of the 13 species described by Philippi in 1887 are discussed and figured. Three new Neogene species are described: Lucinoma aaa sp. nov., Thyasira bbb sp. nov., and Thyasira ccc sp. nov.

Abstract Naticidae (Nielsen): A total of twelve species of Naticidae are described from early Miocene deposits of coastal Chile. Four species are newly described and new combinations are proposed for six oothers. Several available names are synonymized. The naticid species represent all three currently recognized subfamilies and belong in seven genera. Aaa gen. nov. is erected for Natica striolata Sowerby, 1846. Newly described species are Bulbus bbb sp. nov., Euspira ccc sp. nov., and Falsilunatia ddd sp. nov.

Abstract Mitridae (Nielsen): The two Miocene mitrid species described by Philippi in 1887 are revised. Mitra martini Philippi, 1887 is a junior homonym and is renamed and reclassified as Aaa bbb nom. nov. while Mitra chiloensis Philippi, 1887 is confirmed in Imbricaria Schumacher, 1817. Contrary to published statements, the original type material of both species is still available. The range of Aaa bbb nom. nov. is extended north to Mocha island and south to the Chonos Archipelago, while the record of I. chiloensis is extended north towards the Navidad Formation in central Chile and south to the Chonos Archipelago. Aaa bbb nom. nov. is the first record of that genus from South America.

III) To relate geographic patterns to environmental factors or ecological traits. We conducted a preliminary analysis for the area of Chiloé, comparing functional richness (FRic) versus species richness (SRic) of early Miocene and Plio-Pleistocene age. The curves of FRic versus SRic during the Miocene and Plio-Pleistocene look very similar. Each point represents one collection site and the colors indicate to what geological time they belong; red for the warmer Miocene, blue for the colder Plio-Pleistocene.

Figure 1: Functional Richness (FRic) versus Species Richness (SRic). Each Point represents one collection site.

There is a clearly visible positive trend (Fig.2) showing a higher FD with a higher amount of species. The graphic shows also a higher SRic in the Miocene compared to the Plio-Pleistocene about the double amount of species in one site of the Miocene. There are a total of seven points for the Miocene, with 35 different species and 13 different functional groups. For the Plio- Pleistocene there are six points with a total of 17 different species and 11 different functional groups.

Figure 2: Curve fitted to Michaelis-Menten Model for Miocene (RED) and Pliocene/Pleistocene (BLUE). Dashed lines - interval of confidence. Solid lines – curve of best fit.

The fitted curve for the Miocene (RED) shows a non-linear linear progression. All the points are within the calculated interval of confidence (dashed lines) with more point below the curve of best fit (solid line). This indicates an assemblage with low functional diversity (FD) and more similar functional trait values than expected by chance. In comparison, the fitted curve for the Pliocene/Pleistocene shows a more linear progression. Most points are within the line of confidence, except one. This indicates a nonstandard distribution for the Plio-Pleistocene. Additionally, three points above and one point below the curve of best fit are directly on the lines of the interval of confidence. Here the majority of points are above the curve of best fit. This is pointing to an assemblage with high FD at a given level of SRic meaning it has more dispersed functional traits than expected by chance. Changes in FD are greater than expected if extinction and colonization were random. So comparing the mean functional originality of species that are colonizing and species that are going extinct, indicate colonizers to have more unique traits. If assemblages loose and gain the same number of species, FD tends to increase. In this case this is not absolutely clear. There is a decrease in number of species from the Miocene (35 species) to the Plio-Pleistocene (17 species). With that there is a decrease in FD as well, from 13 to 11 different unique functional groups. The point is the amount of decrease. The difference in SRic is far greater than the difference between FRic in the Miocene and Plio-Pleistocene. This still could mean an increase in FD from the Miocene to the Plio-Pleistocene. Interesting as well is the difference indicated by the fact of more points being above or below the curve of best fit. Even though there is a higher number of species as well as functional groups, the fitted curve indicates a higher similarity of functional traits, whereas the fitted curve for the Plio-Pleistocene indicates more dispersed functional traits, as stated above. This underlines an increase in FD from the Miocene to the Plio-Pleistocene as well. Here it would be interesting to analyze the composition of functional traits and compare, which functional groups are lost and compare the composition of the species from both geological timescales as well. It would be useful to compare migrations, extinctions and survivors from the change from the Miocene with its warm waters and from the Plio-Pleistocene with its cold waters, like was done by Rivadeneira & Nielsen (attached publication). So it is clear that the FD in the fossil gastropods and bivalves changes from the Miocene to the Plio-Pleistocene. The greater dissimilarity in species functional traits during the Plio-Pleistocene indicates a possible change due to higher stress, for example competition. Even though it is at this time not possible to identify mechanisms from this pattern, which are responsible for the change, it is possible to say, that a type of environmental filtering occurs. It could indicate that species are particularly adapted to their environment or habitat.

Preliminary results also exist for ecological trade gradients, but analyses were not done yet in sufficient detail:

Figure 3: Trophic type of mollusk species for three time slices across latitudinal gradients, early Miocene, Plio-Pleistocene, and Extant. A) Bivalvia absolute numbers. B) Gastropoda absolute numbers. C) Bivalvia percentage. D) Gastropoda percentage. Nav-Navidad Fm (~33°S), Ran- Ranquil Fm (~37°S), Lac-Lacuy Fm (~42°S), Cho-Chono islands (~45°S), Coq-Coquimbo Fm (~30°S), Tub-Tubul Fm (~37°S), Guf-Guafo island (~43°S), Gum-Guamblín island (~44°S).

Figure 4: Mode of life of mollusk species for three time slices across latitudinal gradients, early Miocene, Plio-Pleistocene, and Extant. A) Bivalvia absolute numbers. B) Gastropoda absolute numbers. C) Bivalvia percentage. D) Gastropoda percentage. Nav-Navidad Fm (~33°S), Ran- Ranquil Fm (~37°S), Lac-Lacuy Fm (~42°S), Cho-Chono islands (~45°S), Coq-Coquimbo Fm (~30°S), Tub-Tubul Fm (~37°S), Guf-Guafo island (~43°S), Gum-Guamblín island (~44°S).

IV) To increase the taxonomic knowledge of Miocene faunas from Chile. While the taxonomic diversity can be clearly increased for most newly sampled localities, no additional records of modern genera were identified for the early Miocene. Classification of taxa from most new localities, however, is still under way, especially for the Taitao peninsula area. From the new (supposedly) Pliocene localities at Taitao, new fossil records for modern genera include the gastropod Argobuccinum and the bivalve Ensis. Abstracts shown for goal II show some of the finished identifications. On a more geological and paleobiogeographic topic, PI Nielsen participated in collaborative work reviewing the Oligo-Miocene transgression (attached book chapter by Encinas et al. in press, http://www.springer.com/gb/book/9783319677736) that formed the geological units yielding the fossils under study, namely the Navidad Fm and its southern equivalents.

V) To determine Miocene water temperatures along the central and southern Chilean coast with stable oxygen isotopes and Mg/Ca ratios. This hypothesis had to be abandoned due to the fact that the international collaborator, M. Regenberg, left academia and had to be replaced as an international collaborator. However, this topic is important for ongoing work, so we need to find other possibilities to tackle it in the future.

Impact Involving several students of the recently founded (2014) first Paleontology Master program in Chile helped to finance their studies. Recruitment in professional jobs prior to finishing their theses is problematic for getting results ‘in time’ but shows the need for professional paleontologists in Chile. While education is not the aim of a Fondecyt project, this fact is considered here as a positive impact of the project on Chilean society. Two museum curators and two university lecturers do or have done their master thesis within this project. Topic related to the project, but not directly treating results, were transmitted to the criminal police Valdivia and a group of school children (see outreach document in attachments).

COOPERACIÓN INTERNACIONAL

Nº Proyecto: 1150664 Nombre Colaborador (a) Extranjero (a): STEFFEN KIEL Afiliación Institucional Actual: SWEDISH MUSEUM OF NATURAL HISTORY Fechas de estadía Desde :10/12/2017 Hasta :17/12/2017

Describa las actividades realizadas y resultados obtenidos. Destaque su contribución al logro de los objetivos del proyecto. Si es pertinente, indique las publicaciones conjuntas generadas, haciendo referencia a lo informado en la etapa Productos. Agregue en la etapa anexos la información necesaria. Project Fondecyt 1150664 originally had two international collaborators, Drs. Kenneth Finger (Foraminifera) and Marcus Regenberg (paleoceanography). As indicated in the final report, Finger was retired due to 'strategic reasons' of his institution and Regenberg left academia because his contract was not renewed, so both were not available as planned. However, Finger is still helping with foraminifera identification. Due to these problems, Dr. Steffen Kiel was invited as international collaborator. PI Nielsen and Kiel have worked together in the past on Chilean latitudinal gradients (doi:10.1130/G31282.1) and Kiel is now collaborating on latitudinal gradients as well as specific taxonomic groups. Kiel visited Chile for one week during December 2017. Fieldwork was done with Nielsen in the Navidad Fm, central Chile, where especially the methane seep reported in the recent literature was visited. Kiel gave a public talk at Universidad Austral de Chile, which was attended by approximately 40 persons (see announcement in annexes). Apart from discussions and planning with Nielsen, a working group meeting was held to better connect students with Kiel and which ended with a Chilean barbecue. Most advanced at the moment is the work on Chilean Neogene Lucinoidea (Bivalvia) which needs preparation of map and plates, but the text is almost ready for submittal: The Cenozoic species of Lucinidae, Ungulinidae and Thyasiridae from Chile are revised. Types and new material of the 13 species described by Philippi in 1887 are discussed and figured. Three new Neogene species are described: Lucinoma aaa sp. nov., Thyasira bbb sp. nov., and Thyasira ccc sp. nov.

PRODUCTOS

ARTÍCULOS Para trabajos en Prensa/ Aceptados/Enviados adjunte copia de carta de aceptación o de recepción.

Nº : 1 Autor (a)(es/as) : Rivadeneira, MM; Nielsen, SN Nombre Completo de la Revista : PLoS ONE Título (Idioma original) : Diversification dynamics, species sorting, and changes in the functional diversity of marine benthic gastropods during the Pliocene-Quaternary at temperate western South America Indexación : WoS ISSN : Año : 2017 Vol. : 12 Nº : 10 Páginas : e0187140 Estado de la publicación a la fecha : Publicada Otras Fuentes de financiamiento, si las hay : Fondecyt 1140841, INACH G_05-11

Envía documento en papel : no Archivo(s) Asociado(s) al artículo : Rivadeneira_Nielsen2017Pliocene_QuaternaryGastropodaChile.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_articulos/descarga/24292360/1150664/2017/105608/1/

Nº : 2 Autor (a)(es/as) : Spano, CA; Hernández, CE; Rivadeneira, MM Nombre Completo de la Revista : Ecography Título (Idioma original) : Evolutionary dispersal drives the latitudinal diversity gradient of stony corals Indexación : WoS ISSN : Año : 2016 Vol. : 39 Nº : Páginas : 836-843 Estado de la publicación a la fecha : Publicada Otras Fuentes de financiamiento, si las hay : Fondecyt 1140692, 1130868, 1140841

Envía documento en papel : no Archivo(s) Asociado(s) al artículo : SpanoEtAl2016StonyCoralsDiversityGradient.PDF https://servicios.conicyt.cl/sial/index.php/investigador/f4_articulos/descarga/24292360/1150664/2017/105609/1/

Nº : 3 Autor (a)(es/as) : Rivadeneira, MM.; Alballay, AH.; Villafaña, JA.; Raimondi, PT.; Blanchette, CA.; Fenberg, PB. Nombre Completo de la Revista : Global Ecology and Biogeography Título (Idioma original) : Geographic patterns of diversification and the latitudinal gradient of richness of rocky intertidal gastropods: the ?into the tropical museum? hypothesis Indexación : WoS ISSN : Año : 2015 Vol. : 24 Nº : Páginas : 1149-1158 Estado de la publicación a la fecha : Publicada Otras Fuentes de financiamiento, si las hay : Fondecyt 1110582, 1140841 The David and Lucile Packard Foundation, The Gordon and BettyMoore Foundation, Bureau of Ocean Energy Management, State of California, The Monitoring Enterprise, The National Park Service, The National Marine Sanctuary Program, and SCCWRP.

Envía documento en papel : no Archivo(s) Asociado(s) al artículo : Rivadeneira_et_al_2015_GEB_online.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_articulos/descarga/24292360/1150664/2017/109146/1/

Nº : 4 Autor (a)(es/as) : Villafaña, JA.; Rivadeneira, MM. Nombre Completo de la Revista : Paleobiology Título (Idioma original) : The modulating role of traits on the biogeographic dynamics of chondrichthyans from the Neogene to the present Indexación : WoS ISSN : Año : Vol. : Nº : Páginas : Estado de la publicación a la fecha : En Prensa Otras Fuentes de financiamiento, si las hay : Fondecyt 1140841

Envía documento en papel : no Archivo(s) Asociado(s) al artículo : Villafana___Rivadeneira_2018__in_press_.PDF https://servicios.conicyt.cl/sial/index.php/investigador/f4_articulos/descarga/24292360/1150664/2017/109147/1/

OTRAS PUBLICACIONES / PRODUCTOS

Nº : 1 Autor (a)(es/as) : Encinas, A; Folguera, A; Nielsen, SN; Ramos, V et al. Título (Idioma original) : The Late Oligocene-Early Miocene marine transgression of Patagonia Tipo de publicación o producto : Capítulo de Libro ISBN : 331967773X Editor (es) (Libro o Capitulo de libros) : Folguera, A; et al. Nombre de la editorial /Organización : Springer País : ALEMANIA Ciudad : Heidelberg Fecha : Febrero - 2018 Año : 2018 Vol. : Nº : Páginas : Otras Fuentes de financiamiento, si las hay : Fondecyt 1010691, 3060051, 11080115, 1090165, 1110914, 1151146 Universidad Nacional de Río Negro project PI 40-B-159 ANPCyT projects PICT 2010?2051, PICT 2014-2240

Envía documento en papel : no Archivo(s) Asociado(s) al artículo : Transgresiones.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_otras_publicaciones/descarga/24292360/1150664/2017/30906/1/

Book_Cover.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_otras_publicaciones/descarga/24292360/1150664/2017/30906/2/

CONGRESOS

Nº : 1 Autor (a)(es/as) : Robles, X.; Nielsen, SN. Título (Idioma original) : Antecedentes preliminares de la bioestratigrafía y paleobatimetría a partir de foraminífero neógenos, Golfo Tres Montes, Región de Aysén Nombre del Congreso : V Simposio de Paleontología en Chile País : CHILE Ciudad : Concepción Fecha Inicio : 07/11/2016 Fecha Término : 11/11/2016 Nombre Publicación : Libro de Resúmenes, V Simposio de Paleontología en Chile Año : 2016 Vol. : Nº : Páginas : 155 Envía documento en papel : no Archivo Asociado : Abs_2016_Robles_Nielsen_SPC.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170727/1/

Nº : 2 Autor (a)(es/as) : Bustos, B.; Nielsen, SN.; Rivadeneira, MM. Título (Idioma original) : Variación geográfica en las interacciones bióticas a lo largo del Plio-Pleistoceno en Chile centro-sur Nombre del Congreso : V Simposio de Paleontología en Chile País : CHILE Ciudad : Concepción Fecha Inicio : 07/11/2016 Fecha Término : 11/11/2016 Nombre Publicación : Libro de Resúmenes, V Simposio de Paleontología en Chile Año : 2016 Vol. : Nº : Páginas : 76-78 Envía documento en papel : no Archivo Asociado : Abs_2016_BustosEtAl_SPC.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170728/1/

Nº : 3 Autor (a)(es/as) : Nielsen, SN. Título (Idioma original) : New paleontological records from the Neogene of southern Chilean fjords: from Guafo via Los Chonos to Taitao Nombre del Congreso : V Simposio de Paleontología en Chile País : CHILE Ciudad : Concepción Fecha Inicio : 07/11/2016 Fecha Término : 11/11/2016 Nombre Publicación : Libro de Resúmenes, V Simposio de Paleontología en Chile Año : 2016 Vol. : Nº : Páginas : 103 Envía documento en papel : no Archivo Asociado : Abs_2016_Nielsen_SPC.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170750/1/

Nº : 4 Autor (a)(es/as) : Pérez Barria, L.; Nielsen, S. Título (Idioma original) : El género Limopsis Sassi, 1827 (Bivalvia: Arcida) en el Mioceno de Chile Nombre del Congreso : V Simposio de Paleontología en Chile País : CHILE Ciudad : Concepción Fecha Inicio : 07/11/2016 Fecha Término : 11/11/2016 Nombre Publicación : Libro de Resúmenes, V Simposio de Paleontología en Chile Año : 2016 Vol. : Nº : Páginas : 153 Envía documento en papel : no Archivo Asociado : Abs_2016_Perez_Nielsen_SPC_Limopsis.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170752/1/

Nº : 5 Autor (a)(es/as) : Pérez Barria, L.; Nielsen, S. Título (Idioma original) : La subclase Protobranchia (Mollusca: Bivalvia) en el Mioceno marino de la costa Pacífica Chilena: problemáticas y desafíos actuales Nombre del Congreso : V Simposio de Paleontología en Chile País : CHILE Ciudad : Concepción Fecha Inicio : 07/11/2016 Fecha Término : 11/11/2016 Nombre Publicación : Libro de Resúmenes, V Simposio de Paleontología en Chile Año : 2016 Vol. : Nº : Páginas : 104-106 Envía documento en papel : no Archivo Asociado : Abs_2016_Perez_Nielsen_SPC_Protobranchia.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170775/1/

Nº : 6 Autor (a)(es/as) : Nielsen, SN. Título (Idioma original) : Gradientes latitudinales en moluscos marinos de Chile desde el Mioceno Nombre del Congreso : 11° Congreso de la Asociación Paleontológica Argentina País : ARGENTINA Ciudad : General Roca Fecha Inicio : 17/10/2016 Fecha Término : 21/10/2016 Nombre Publicación : 11° Congreso de la Asociación Paleontológica Argentina Año : 2016 Vol. : Nº : Páginas : 70 Envía documento en papel : no Archivo Asociado : 2016_CAPA.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170779/1/ Nº : 7 Autor (a)(es/as) : Pérez, JR.; Moreno, K.; Nielsen, SN.; Arratia, G. Título (Idioma original) : Descripción osteológica y taxonomía de un posible pez serránido (Actinopterygii, Perciformes) del Mioceno Temprano de la Provincia de Arauco, Chile Nombre del Congreso : 11° Congreso de la Asociación Paleontológica Argentina País : ARGENTINA Ciudad : General Roca Fecha Inicio : 17/10/2016 Fecha Término : 21/10/2016 Nombre Publicación : 11° Congreso de la Asociación Paleontológica Argentina Año : 2016 Vol. : Nº : Páginas : 78-79 Envía documento en papel : no Archivo Asociado : 2016_CAPA1.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170780/1/

Nº : 8 Autor (a)(es/as) : Pérez Barría, LP:; Nielsen, SN. Título (Idioma original) : El género Limopsis Sassi, 1827 (Bivalvia: Arcida) en el Mioceno de Chile Nombre del Congreso : 11° Congreso de la Asociación Paleontológica Argentina País : ARGENTINA Ciudad : General Roca Fecha Inicio : 17/10/2016 Fecha Término : 21/10/2016 Nombre Publicación : 11° Congreso de la Asociación Paleontológica Argentina Año : 2016 Vol. : Nº : Páginas : 79-80 Envía documento en papel : no Archivo Asociado : 2016_CAPA2.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170781/1/

Nº : 9 Autor (a)(es/as) : Pérez Barría, L:; Nielsen, SN. Título (Idioma original) : Los Protobranchia (Nuculoidea, Nuculanoidea y Solemyoidea) del Mioceno de Chile: Problemáticas y desafíos actuales Nombre del Congreso : 2. Congreso Argentino de Malacología País : ARGENTINA Ciudad : Mendoza Fecha Inicio : 10/08/2016 Fecha Término : 12/08/2016 Nombre Publicación : Libro de Resúmenes, Segundo Congreso Argentino de Malacología Año : 2016 Vol. : Nº : Páginas : 13 Envía documento en papel : no Archivo Asociado : Abs_2016_Perez_Nielsen_CAM.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170789/1/

Nº : 10 Autor (a)(es/as) : Nielsen, SN.; Rivadeneira, MM. Título (Idioma original) : La evolución de gradientes latitudinales en moluscos marinos de Chile desde el Mioceno Nombre del Congreso : X Congreso Latinoamericano de Malacología País : URUGUAY Ciudad : Piriápolis Fecha Inicio : 01/10/2017 Fecha Término : 06/10/2017 Nombre Publicación : X Congreso Latinoamericano de Malacología, Libro de Resúmenes Año : 2017 Vol. : Nº : Páginas : 170 Envía documento en papel : no Archivo Asociado : Abs_2017_CLAMA_Gradientes_Latitudinales.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170797/1/

Nº : 11 Autor (a)(es/as) : Pérez, JR.; Moreno, K.; Nielsen, SN.; Arratia, G. Título (Idioma original) : A serranid fish (Actinopterygii, Perciformes) from the Early Miocene of the Province of Arauco, Chile Nombre del Congreso : Society of Vertebrate Paleontology 77th Annual Meeting País : CANADA Ciudad : Calgary Fecha Inicio : 23/08/2017 Fecha Término : 26/08/2017 Nombre Publicación : Society of Vertebrate Paleontology 77th Annual Meeting, Meeting Program and Abstracts Año : 2017 Vol. : Nº : Páginas : 176 Envía documento en papel : no Archivo Asociado : Abs_PerezEtAl_SVP_Serranid.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_congresos/descarga/24292360/1150664/2017/170806/1/

TESIS/MEMORIAS

Nº : 1 Título de Tesis : Nuevo serránido fósil (Actinopterygii, Perciformes) proveniente del Mioceno temprano de la región de Arauco, Chile Nombre y Apellidos del(de la) Alumno(a) : José Rubén Pérez Marín Nombre y Apellidos del(de la) Tutor(a) : Karen Moreno Fuentealba; Sven N. Nielsen Título Grado : Magister Institución : Universidad Austral De Chile País : CHILE Ciudad : Valdivia Estado de Tesis : Terminada Fecha Inicio : 08/04/2016 Fecha Término : 18/10/2017 Envía documento en papel : no Archivo Asociado : Resumen___Abstract___Tesis_Jose_Perez.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_tesis_memorias/descarga/24292360/1150664/2017/81414/1/ CERTIFICADO_T__TULO.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_tesis_memorias/descarga/24292360/1150664/2017/81414/2/

Nº : 2 Título de Tesis : Bioestratigrafía y paleobatimetría de las unidades sedimentarias marinas neógenas del Golfo Tres Montes, Península Tres Montes, Región de Aysén Nombre y Apellidos del(de la) Alumno(a) : Ximena P. Robles Cornejo Nombre y Apellidos del(de la) Tutor(a) : Sven N. Nielsen Título Grado : Magister Institución : Universidad Austral De Chile País : CHILE Ciudad : Valdivia Estado de Tesis : En Ejecución Fecha Inicio : 18/12/2015 Fecha Término : Envía documento en papel : no Archivo Asociado : Tesis_Mpaleo_XRC.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_tesis_memorias/descarga/24292360/1150664/2017/83248/1/ CI_Envio_Tesis_XIMENA_ROBLES_CORNEJO1.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_tesis_memorias/descarga/24292360/1150664/2017/83248/2/

Nº : 3 Título de Tesis : Modelo bioestratigráfico y paleobatimétrico de los depósitos marinos de Isla Guafo (43°52' y 43°69'S), Chile Nombre y Apellidos del(de la) Alumno(a) : Flavia C. Velásquez Ruiz Nombre y Apellidos del(de la) Tutor(a) : Sven N. Nielsen Título Grado : Magister Institución : Universidad Austral De Chile País : CHILE Ciudad : Valdivia Estado de Tesis : En Ejecución Fecha Inicio : 26/09/2016 Fecha Término : Envía documento en papel : no Archivo Asociado : Certificado_tesis_Flavia.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_tesis_memorias/descarga/24292360/1150664/2017/83276/1/

Nº : 4 Título de Tesis : Moluscos fósiles neógenos del Golfo Tres Montes, Región de Aysén, Chile Nombre y Apellidos del(de la) Alumno(a) : Javiera Paz Bravo Fuentes Nombre y Apellidos del(de la) Tutor(a) : Sven N. Nielsen Título Grado : Pregrado Institución : Universidad Austral De Chile País : CHILE Ciudad : Valdivia Estado de Tesis : En Ejecución Fecha Inicio : 07/12/2017 Fecha Término : Envía documento en papel : no Archivo Asociado : constancia_Javiera_Bravo.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f4_tesis_memorias/descarga/24292360/1150664/2017/84267/1/

ANEXOS

Nº : 1 Archivo Asociado : Permiso_Oficial_CMN_Fondecyt_1140841.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f5_anexos/descarga/24292360/1150664/2017/82015/

Nº : 2 Archivo Asociado : PermisoCMN_Alfonso_Encinas.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f5_anexos/descarga/24292360/1150664/2017/82016/

Nº : 3 Archivo Asociado : ColaboracionInternacional1150664.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f5_anexos/descarga/24292360/1150664/2017/82658/

Nº : 4 Archivo Asociado : Informe_Resumen_Difusion_P1150664.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f5_anexos/descarga/24292360/1150664/2017/82961/

Nº : 5 Archivo Asociado : Resolucion2015_N__322_Director_ICT_Nielsen.pdf https://servicios.conicyt.cl/sial/index.php/investigador/f5_anexos/descarga/24292360/1150664/2017/83079/

A continuación se detallan los anexos físicos/papel que no se incluyen en el informe en formato PDF.