Cave Hill RNA Flora Shawnee National Forest, Illinois

Surveyor: Steven Olson 1986-87. Appendix Item IV of the Cave Hill Research Natural Area, Shawnee National Forest Establishment Record 1990. Unpublished report on file at Vienna Administrative Office. Updated by Sue Lietz, Northern Research Station and Elizabeth Shimp, Shawnee National Forest December 2014.

Appendix Item IV. List of plants collected in the Cave Hill Research Natural Area and vicinity.

Nomenclature follows USDA, NRCS PLANTS database [accessed December 2014].

R = collected in RNA * = Saline County Record

Scientific name Common name Former Scientific name Trees (Dicots) Acer rubrum Red maple Acer saccharinum Silver maple R Acer saccharum Sugar maple R Amelanchier arborea Common serviceberry R Asimina triloba Pawpaw Carpinus caroliniana American hornbeam R Carya glabra Pignut hickory * Carya laciniosa Shellbark hickory R Carya ovalis Red hickory R Carya ovata Shagbark hickory R Carya texana Black hickory Celtis laevigata Sugarberry R Celtis occidentalis Common hackberry R Cercis canadensis Eastern redbud R Cornus drummondii Roughleaf dogwood R Cornus florida Flowering dogwood Crataegus crus-galli Cockspur hawthorn * Crataegus engelmannii Englemann’s hawthorn * Crataegus mollis Downy hawthorn R Diospyros virginiana Common persimmon R * Elaeagnus umbellata Autumn olive Fraxinus americana White ash Fraxinus pennsylvanica Green ash R Gleditsia triacanthos Honeylocust Ilex decidua Possumhaw Juglans nigra Black walnut R Juniperus virginiana Eastern redcedar Liquidambar styraciflua Sweetgum R * Liriodendron tulipifera Tuliptree

1 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Trees (Dicots) R Morus rubra Red mulberry R Nyssa sylvatica Blackgum R Ostrya virginiana Hophornbeam * Pinus echinata Shortleaf pine * Pinus strobus Eastern white pine Platanus occidentalis American sycamore Populus deltoides Eastern cottonwood R Prunus serotina Black cherry R Quercus alba White oak R Quercus coccinea Scarlet oak Quercus falcata Southern red oak * Quercus lyrata Overcup oak Quercus macrocarpa Bur oak R Quercus marilandica Blackjack oak * Quercus michauxii Swamp chestnut oak R Quercus montana Chestnut oak Quercus prinus * Quercus pagoda Cherrybark oak Quercus palustris Pin oak R Quercus prinoides Dwarf chinquapin oak R Quercus rubra Northern red oak Quercus shumardii Shumard's oak R Quercus stellata Post oak R Quercus velutina Black oak R Rhus copallinum Winged sumac Rhus copallina R Rhus glabra Smooth sumac Robinia pseudoacacia Black locust Salix nigra Black willow R Sassafras albidum Sassafras R Staphylea trifolia American bladdernut Staphylea trifoliata Taxodium distichum Bald cypress R Ulmus alata Winged elm Ulmus americana American elm Ulmus rubra Slippery elm

Shrubs (Dicots) * Amorpha fruticosa False indigo bush Cephalanthus occidentalis Common buttonbush Corylus americana American hazelnut * Dirca palustris Eastern leatherwood

2 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Shrubs (Dicots) Forestiera acuminata Eastern swampprivet R Hydrangea arborescens Wild hydrangea R Hypericum hypericoides St. Andrew's cross Hypericum stragulum R * Lindera benzoin Northern spicebush R Rosa carolina Carolina rose * Rosa multiflora Multiflora rose R Rosa setigera Climbing rose * Rubus flagellaris Northern dewberry Rubus enslenii Rubus pensilvanicus Pennsylvania dewberry R Symphoricarpos orbiculatus Coralberry R Vaccinium arboreum Farkleberry R Vaccinium pallidum Blue Ridge blueberry

Vines (Dicots and Monocots) R Ampelopsis cordata Heartleaf peppervine Amphicarpaea bracteata American hogpeanut Amphicarpa bracteata * Apios americana Groundnut Calystegia sepium Hedge false bindweed R Campsis radicans Trumpet creeper * Celastrus orbiculatus Oriental bittersweet R * Celastrus scandens American bittersweet Cocculus carolinus Carolina coralbead R Dioscorea villosa Wild yam R Lonicera japonica Japanese honeysuckle * Matelea obliqua Climbing milkvine Menispermum canadense Common moonseed R Parthenocissus quinquefolia Virginia creeper R Passiflora lutea Yellow passionflower Potentilla simplex Common cinquefoil * Sicyos angulatus Oneseed bur cucumber R Smilax glauca Cat greenbrier * Smilax pulverulenta Downy carrionflower Smilax tamnoides Bristly greenbrier Smilax hispida * Trachelospermum difforme Climbing dogbane R Vitis aestivalis Summer grape * Vitis cinerea Graybark grape

Other Dicots (forb/herb/subshrub) R Acalypha gracilens Slender threeseed mercury

3 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) R Acalypha rhomboidea Common threeseed mercury R Achillea millefolium Common yarrow R Agalinis tenuifolia Slenderleaf false foxglove Agalinis besseyana R Ageratina altissima White snakeroot Eupatorium rugosum R Agrimonia parviflora Harvestlice R Agrimonia pubescens Soft agrimony R Agrimonia rostellata Beaked agrimony Amaranthus retroflexus Redroot amaranth R Ambrosia artemisiifolia Annual ragweed Ambrosia bidentata Lanceleaf ragweed R Ambrosia trifida Great ragweed * Ammannia coccinea Valley redstem Amsonia tabernaemontana Eastern bluestar R Anemone virginiana Tall thimbleweed R Antennaria plantaginifolia Woman’s tobacco R Apocynum cannabinum Indianhemp R Aquilegia canadensis Red columbine R Arabis canadensis Sicklepod Arabis laevigata Smooth rockcress R Aristolochia serpentaria Virginia snakeroot R Arnoglossum atriplicifolium Pale Indian plantain Cacalia atriplicifolia Asclepias hirtella Green milkweed Asclepias incarnata Swamp milkweed * Asclepias perennis Aquatic milkweed Asclepias quadrifolia Fourleaf milkweed Asclepias syriaca Common milkweed Asclepias tuberosa Butterfly milkweed R Asclepias variegata Redring milkweed Asclepias verticillata Whorled milkweed * Asclepias viridiflora Green comet milkweed R Aureolaria flava Smooth yellow false foxglove Baptisia alba Largeleaf wild indigo Baptisia lactea Bidens aristosa Bearded beggarticks * Bidens coronata Crowned beggarticks R Boehmeria cylindrica Smallspike false nettle Boltonia asteroides White doll’s daisy Campanulastrum americanum American bellflower Campanula americana R Cardamine concatenata Cutleaf toothwort Dentaria laciniata Cardamine hirsuta Hairy bittercress

4 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) R Cardamine parviflora Sand bittercress R Chamaecrista fasciculata Partridge pea Cassia fasciculata R Chamaecrista nictitans Sensitive partridge pea Cassia nictitans Chamaesyce maculata Spotted sandmat Chamaesyce supina Chenopodium album Lambsquarters Cichorium intybus Chicory Broadleaf enchanter's R Circaea lutetiana nightshade R Cirsium carolinianum Soft thistle R Cirsium discolor Field thistle R Claytonia virginica Virginia springbeauty R Clitoria mariana Atlantic pigeonwings R Conyza canadensis Canadian horseweed Coreopsis tripteris Tall tickseed R Corydalis flavula Yellow fumewort Croton capitatus Hogwort R Croton michauxii Michaux's croton Crotonopsis elliptica Croton monanthogynus Prairie tea R Cryptotaenia candensis Canadian honewort Cunila origanoides Common dittany Cuphea viscosissima Blue waxweed * Cuscuta gronovii Scaldweed Cuscuta pentagona Fiveangled dodder Cuscuta campestris * Cuscuta polygonorum Smartweed dodder Cynanchum laeve Honeyvine Cynoglossum virginianum Wild comfrey R Dalea candida White prairie clover Datura stramonium Jimsonweed Daucus carota Queen Anne’s lace Desmanthus illinoensis Illinois bundleflower R * Desmodium canescens Hoary ticktrefoil Desmodium ciliare Hairy small-leaf ticktrefoil R Desmodium glutinosum Pointedleaf ticktrefoil R Desmodium nudiflorum Nakedflower ticktrefoil R Desmodium paniculatum Panicledleaf ticktrefoil Dianthus armeria Deptford pink R Dicentra cucullaria Dutchman’s breeches R Diodia teres Poorjoe R Dodecatheon meadia Pride of Ohio

5 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) R Draba brachycarpa Shortpod draba Eclipta prostrata False daisy R * Enemion biternatum eastern false rue anemone Isopyrum biternatum R Erechtites hieraciifolius American burnweed R Erigeron annuus Eastern daisy fleabane R * Erigeron philadelphicus Philadelphia fleabane Erigeron strigosus Prairie fleabane Eupatorium coelestinum Blue mistflower R Eupatorium serotinum Lateflowering thoroughwort R Euphorbia corollata Flowering spruge Euphorbia dentata Toothed spruge Poinsettia dentata * Euphorbia spathulata Warty spruge Euphorbia obtusata Euthamia graminifolia Flat-top goldenrod Eutrochium fistulosum Trumpetweed Eupatorium fistulosum R * Galactia regularis Eastern milkpea R Galium circaezans Licorice bedstraw Galium concinnum Shining bedstraw R Galium pilosum Hairy bedstraw * Galium trifidum Threepetal bedstraw R Geum canadense White avens R Geum vernum Spring avens R Gillenia stipulata American ipecac Porteranthus stipulatus * Glycine max Soybean Gnaphalium Gnaphalium purpurea Spoonleaf purple everlasting purpureum Hedeoma pulegioides American false pennyroyal Helenium flexuosum Purplehead sneezeweed * Helianthus annuus Common sunflower * Helianthus decapetalus Thinleaf sunflower R Helianthus divaricatus Woodland sunflower R Helianthus microcephalus Small woodland sunflower R Heliopsis helianthoides Smooth oxeye R Heuchera americana American alumroot Hibiscus lasiocarpos Rosemallow Hibiscus lasiocarpus R Hieracium gronovii Queendevil R Hieracium scabrum Rough hawkweed R Hordeum pusillum Little barley R Houstonia longifolia Longleaf summer bluet Hedyotis longifolia R Houstonia pusilla Tiny bluet Hedyotis pusilla

6 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) R Hylotelephium telephioides Allegheny stonecrop Sedum telephioides R Hypericum gentianoides Orangegrass Hypericum mutilum Dwarf St. Johnswort Impatiens capensis Jewelweed Ipomoea hederacea Ivyleaf morning-glory Ipomoea lacunosa Whitestar Ipomoea pandurata Man of the earth * Iva annua Annual marsh elder R Krigia biflora twoflower dwarfdandelion Kummerowia stipulacea Korean clover Kummerowia striata Japanese clover R Lactuca canadensis Canada lettuce * Lactuca floridana Woodland lettuce Lactuca serriola Prickly lettuce R Lechea tenuifolia Narrowleaf pinweed R Lepidium virginicum Virginia pepperweed R * Lespedeza cuneata Sericea lespedeza R Lespedeza hirta Hairy lespedeza Lespedeza procumbens Trailing lespedeza R Lespedeza repens Creeping lespedeza R Lespedeza virginica Slender lespedeza * Liatris aspera Tall blazing star R Liatris spicata Dense blazing star R Liatris squarrosa Scaly blazing star * Lithospermum canescens Hoary puccoon R Lobelia inflata Indian-tobacco Lobelia spicata Palespike lobelia R Ludwigia alternifolia Seedbox Lycopus americanus American water horehound Lycopus rubellus Taperleaf water horehound Lysimachia ciliata Fringed loosestrife R Lysimachia lanceolata Lanceleaf loosestrife Lythrum alatum Winged lythrum Matricaria R * Matricaria discoidea Disc mayweed matricarioides Medicago lupulina Black medic Melitotus officinalis Sweetclover includes M. alba Mimulus alatus Sharpwing monkeyflower R Monarda bradburiana Eastern beebalm

7 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) R Monarda fistulosa Wild bergamot Myosotis verna Spring forget-me-not Myosurus minimus Tiny mousetail Nuphar lutea Yellow pond-lily Oenothera biennis Common evening primrose R Oenothera linifolia Threadleaf evening primrose R Opuntia humifusa Devil’s tongue R Orbexilum pedunculatum Sampson’s snakeroot Psoralea psoralioides R Orobanche uniflora Oneflowered broomrape R Oxalis stricta Common yellow woodsorrel R Oxalis violacea Violet woodsorrel * Packera aurea Golden ragwort Senecio aureus * Packera glabella Butterweed Senecio glabellus R Parietaria pensylvanica Pennsylvania pellitory R Paronychia fastigiata Hairy forked nailwort R Parthenium integrifolium Wild quinine * Penstemon alluviorum Lowland beardtongue R Penstemon pallidus Pale beardtongue Penthorum sedoides Ditch stonecrop R Phlox divaricata Wild blue phlox Phlox pilosa Downy phlox R Phryma leptostachya American lopseed Phyla lanceolata Lanceleaf fogfruit * Physalis lanceolata Sword groundcherry * Physalis virginiana Virginia groundcherry R Phytolacca americana American pokeweed R Pilea pumila Canadian clearweed R * Plantago cordata Heartleaf plantain Plantago lanceolata Narrowleaf plantain Plantago rugelii Blackseed plantain R Plantago virginica Virginia plantain Podophyllum peltatum Mayapple Polemonium reptans Greek valerian Polygonatum R Polygonatum biflorum Smooth Solomon's seal commutatum Polygonum punctatum Dotted smartweed R Polytaenia nuttallii Nuttall’s prairie parsley * Prenanthes crepidinea Nodding rattlesnake root R Prunella vulgaris Common selfheal

8 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) R Pycnanthemum tenuifolium Narrowleaf mountainmint Pyrrhopappus carolinianus Carolina desert-chicory R Ranunculus abortivus Littleleaf buttercup R Ranunculus fascicularis Early buttercup R Ranunculus hispidus Bristly buttercup R Ranunculus micranthus Rock buttercup * Ratibida pinnata Pinnate prairie coneflower R Rudbeckia hirta Blackeyed Susan Rudbeckia subtomentosa Sweet coneflower R Ruellia humilis Fringeleaf wild petunia Ruellia strepens Limestone wild petunia Rumex crispus Curly dock Sabatia angularis Rosepink Salvia lyrata Lyreleaf sage Samolus valerandi Seaside brookweed R Sanguinaria canadensis Bloodroot R Sanicula canadensis Canadian blacksnakeroot Saururus cernuus Lizard’s tail R * Scutellaria incana Hoary skullcap Scutellaria nervosa Veiny skullcap * Scutellaria parvula Small skullcap Scutellaria australis R Sedum pulchellum Widowscross Sida spinosa Prickly fanpetals R Silene stellata Widowsfrill Silphium integrifolium Wholeleaf rosinweed Solanum carolinense Carolina horsenettle R * Solanum ptycanthum West Indian nightshade R Solidago canadensis Canada goldenrod R Solidago juncea Early goldenrod R Solidago nemoralis Gray goldenrod R Solidago ulmifolia Elmleaf goldenrod Spigelia marilandica Woodland pinkroot Stachys tenuifolia Smooth hedgenettle R Stylosanthes biflora Sidebeak pencilflower R Symphyotrichum turbinellum Smooth violet prairie aster Aster turbinellus R Taenidia integerrima Yellow pimpernel R Taraxacum officinale Common dandelion R Tephrosia virginiana Virginia tephrosia Teucrium canadense Canada germander

9 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other Dicots (forb/herb/subshrub) * Thalictrum dioicum Early meadow-rue R Thalictrum thalictroides Rue anemone Trifolium hybridum Alsike clover Trifolium pratense Red clover Trifolium repens White clover R Triodanis perfoliata Clasping Venus' looking-glass Valerianella radiata Beaked cornsalad Verbascum blattaria Moth mullein R Verbascum thapsus Common mullein Verbena hastata Swamp verbena Verbena stricta Hoary verbena Verbena urticifolia White vervain R Verbesina helianthoides Gravelweed R * Vernonia baldwini Baldwin's ironweed Veronica arvensis Corn speedwell R Viola bicolor Field pansy Viola rafinesquii R * Viola nephrophylla Northern bog violet Viola pratincola R Viola pedata Birdfoot violet R Viola sororia Common blue violet R Viola triloba Three-lobe violet * Xanthium strumarium Rough cocklebur * Zea mays Corn Zizia aurea Golden zizia

Grasses (Monocots) R Agrostis gigantea Redtop Agrostis alba Agrostis hyemalis Winter bentgrass R * Agrostis perennans Upland bentgrass Agrostis scabra Rough bentgrass R Andropogon gerardii Big bluestem * Andropogon ternarius Splitbeard bluestem Andropogon virginicus broomsedge bluestem Aristida oligantha Prairie threeawn R Aristida purpurascens Arrowfeather threeawn R Brachyelytrum erectum Bearded shorthusk R Bromus pubescens Hairy woodland brome R Bromus racemosus Bald brome Chasmanthium latifolium Indian woodoats Cinna arundinacea Sweet woodreed

10 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Grasses (Monocots) Dactylis glomerata Orchardgrass R Danthonia spicata Poverty oatgrass R Dichanthelium acuminatum Tapered rosette grass R Dichanthelium boscii Bosc’s panicgrass Dichanthelium clandestinum Deertongue * Dichanthelium commutatum Variable panicgrass R Dichanthelium dichotomum Cypress panicgrass R Dichanthelium linearifolium Slimleaf panicgrass R * Dichanthelium malacophyllum Softleaf rosette grass Dichanthelium R Dichanthelium sphaerocarpon Roundseed panicgrass polyanthes R Dichanthelium sphaerocarpon Roundseed panicgrass R * Digitaria filiformis Slender crabgrass Digitaria ischaemum Smooth crabgrass * Digitaria villosa Shaggy crabgrass * Echinochloa crus-galli Barnyardgrass Echinochloa muricata Rough barnyardgrass * Eleocharis acicularis Needle spikerush * Eleocharis obtusa Blunt spikerush Eleusine indica Indian goosegrass R Elymus canadensis Canada wildrye R Elymus hystrix Eastern bottlebrush grass R Elymus villosus Hairy wildrye R Elymus virginicus Virginia wildrye R Eragrostis capillaris Lace grass Eragrostis pectinacea Tufted lovegrass R Eragrostis spectabilis Purple lovegrass R * Festuca paradoxa Clustered fescue R Glyceria striata Fowl mannagrass R Koeleria macrantha Prairie Junegrass Leersia lenticularis Catchfly grass Leersia virginica White grass R Melica nitens Threeflower melicgrass R Muhlenbergia sobolifera Rock muhly R Panicum anceps Beaked panicgrass Panicum dichotomiflorum Fall panicgrass Phleum pratense Timothy R Poa annua Annual bluegrass R Poa compressa Canada bluegrass

11 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Grasses (Monocots) Poa pratensis Kentucky bluegrass * Poa sylvestris Woodland bluegrass * Schedonorus arundinaceus Tall fescue Festuca arundinacea R Schizachyrium scoparium Little bluestem R Setaria faberi Japanese bristlegrass Setaria pumila Yellow foxtail Setaria glauca Setaria viridis Green bristlegrass * Sorghum bicolor Sorghum Sorghum halepense Johnsongrass Sphenopholis obtusata Prairie wedgescale R Sporobolus heterolepis prairie dropseed R Tridens flavus Purpletop tridens R Vulpia octoflora Sixweeks fescue

Sedges & Rushes (Monocots) R Carex albicans Whitetinge sedge Carex artitecta * Carex albolutescens Greenwhite sedge R Carex annectens Yellowfruit sedge * Carex bebbii Bebb’s sedge Carex blanda Eastern woodland sedge * Carex brevior Shortbeak sedge R Carex bushii Bush’s sedge R Carex caroliniana Carolina sedge Carex digitalis Slender woodland sedge * Carex flaccosperma Thinfruit sedge * Carex frankii Frank’s sedge R Carex glaucodea Blue sedge * Carex granularis Limestone meadow sedge * Carex gravida Heavy sedge * Carex grayi Gray’s sedge R Carex hirsutella Fuzzy wuzzy sedge R * Carex jamesii James’ sedge Carex lurida Shallow sedge R Carex muehlenbergii Muhlenberg’s sedge R * Carex normalis Greater straw sedge Carex retroflexa Reflexed sedge R * Carex rosea Rosy sedge * Carex tenera Quill sedge R * Carex torta Twisted sedge

12 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Sedges & Rushes (Monocots) Carex tribuloides Blunt broom sedge R Cyperus echinatus Globe flatsedge Cyperus ovularis Cyperus erythrorhizos Redroot flatsedge R Cyperus lupulinus Great Plains flatsedge Cyperus filiculmis Cyperus strigosus Strawcolored flatsedge Juncus biflorus Bog rush Juncus effusus Common rush Juncus interior Inland rush R Juncus marginatus Grassleaf rush R Juncus tenuis Poverty rush R Luzula multiflora Common woodrush Scirpus pendulus Rufous bulrush

Other herbs (Monocots) Alisma plantago-aquatica European water plantain R Allium canadense Meadow garlic Allium vineale Wild garlic R Arisaema dracontium Green dragon R Arisaema triphyllum Jack in the pulpit R Commelina communis Asiatic dayflower Commelina virginica Virginia dayflower R Corallorhiza wisteriana Spring coralroot Erythronium americanum Dogtooth violet Hymenocallis * Hymenocallis occidentalis Northern spiderlily caroliniana R Hypoxis hirsuta Common goldstar Lilium michiganense Michigan lily Feathery false lily of the Smilacina racemosa R Maianthemum racemosum valley R Manfreda virginica False aloe Najas minor Brittle waternymph R Nothoscordum bivalve Crowpoison Potamogeton diversifolius Waterthread pondweed Sagittaria calycina Hooded arrowhead Sagittaria latifolia Broadleaf arrowhead * Sisyrinchium albidum White blue-eyed grass R Sisyrinchium angustifolium Narrowleaf blue-eyed grass Spiranthes cernua Nodding lady’s tresses R Tradescantia virginiana Virginia spiderwort

13 Plant List Cave Hill RNA, Shawnee National Forest, IL Cave Hill RNA Flora Shawnee National Forest, Illinois

Scientific name Common name Former Scientific name Other herbs (Monocots) R Trillium recurvatum Bloody butcher Uvularia grandiflora Largeflower bellwort

Ferns R Asplenium bradleyi Bradley’s spleenwort R Asplenium pinnatifidum Lobed spleenwort R Asplenium platyneuron Ebony spleenwort Botrychium dissectum Cutleaf grapefern R Botrychium virginianum Rattlesnake fern R Cheilanthes lanosa Hairy lipfern Cystopteris protrusa Lowland bladderfern R Dryopteris marginalis Marginal woodfern R Polystichum acrostichoides Christmas fern R Woodsia obtusa Bluntlobe cliff fern

14 Plant List Cave Hill RNA, Shawnee National Forest, IL