Chapter 1. Introduction
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A New Middle Cambrian Trilobite with a Specialized Cephalon from Shandong Province, North China
Editors' choice A new middle Cambrian trilobite with a specialized cephalon from Shandong Province, North China ZHIXIN SUN, HAN ZENG, and FANGCHEN ZHAO Sun, Z., Zeng, H., and Zhao, F. 2020. A new middle Cambrian trilobite with a specialized cephalon from Shandong Province, North China. Acta Palaeontologica Polonica 65 (4): 709–718. Trilobites achieved their maximum generic diversity in the Cambrian, but the peak of morphological disparity of their cranidia occurred in the Middle to Late Ordovician. Early to middle Cambrian trilobites with a specialized cephalon are rare, especially among the ptychoparioids, a group of libristomates featuring the so-called “generalized” bauplan. Here we describe an unusual ptychopariid trilobite Phantaspis auritus gen. et sp. nov. from the middle Cambrian (Miaolingian, Wuliuan) Mantou Formation in the Shandong Province, North China. This new taxon is characterized by a cephalon with an extended anterior area of double-lobate shape resembling a pair of rabbit ears in later ontogenetic stages; a unique type of cephalic specialization that has not been reported from other trilobites. Such a peculiar cephalon as in Phantaspis provides new insights into the variations of cephalic morphology in middle Cambrian trilobites, and may represent a heuristic example of ecological specialization to predation or an improved discoidal enrollment. Key words: Trilobita, Ptychopariida, ontogeny, specialization, Miaolingian, Paleozoic, Longgang, Asia. Zhixin Sun [[email protected]], Han Zeng [[email protected]], and Fangchen Zhao [[email protected]] (cor responding author), State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palae ontology and Center for Excellence in Life and Palaeoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; University of Chinese Academy of Sciences, Beijing 100049, China. -
Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
Copertina Guida Ai TRILOBITI V3 Esterno
Enrico Bonino nato in provincia di Bergamo nel 1966, Enrico si è laureato in Geologia presso il Dipartimento di Scienze della Terra dell'Università di Genova. Attualmente risiede in Belgio dove svolge attività come specialista nel settore dei Sistemi di Informazione Geografica e analisi di immagini digitali. Curatore scientifico del Museo Back to the Past, ha pubblicato numerosi volumi di paleontologia in lingua italiana e inglese, collaborando inoltre all’elaborazione di testi e pubblicazioni scientifiche a livello nazonale e internazionale. Oltre alla passione per questa classe di artropodi, i suoi interessi sono orientati alle forme di vita vissute nel Precambriano, stromatoliti, e fossilizzazioni tipo konservat-lagerstätte. Carlo Kier nato a Milano nel 1961, Carlo si è laureato in Legge, ed è attualmente presidente della catena di alberghi Azul Hotel. Risiede a Cancun, Messico, dove si dedica ad attività legate all'ambiente marino. All'età di 16 anni, ha iniziato una lunga collaborazione con il Museo di Storia Naturale di Milano, ed è a partire dal 1970 che prese inizio la vera passione per i trilobiti, dando avvio a quella che oggi è diventata una delle collezioni paleontologiche più importanti al mondo. La sua instancabile attività di ricerca sul terreno in varie parti del globo e la collaborazione con professionisti del settore, ha permesso la descrizione di nuove specie di trilobiti ed artropodi. Una forte determinazione e la costruzione di un nuovo complesso alberghiero (AZUL Sensatori) hanno infine concretizzzato la realizzazione -
Malformed Agnostids from the Middle Cambrian Jince Formation of the Pøíbram-Jince Basin, Czech Republic
Malformed agnostids from the Middle Cambrian Jince Formation of the Pøíbram-Jince Basin, Czech Republic OLDØICH FATKA, MICHAL SZABAD & PETR BUDIL Two agnostids from Cambrian of the Barrandian area bear different types of skeletal malformations. The tiny pathologi- cal exoskeleton of Hypagnostus parvifrons (Linnarsson, 1869) has asymmetrically developed pygidial axis, while the posterior pygidial rim in the larger Phalagnostus prantli Šnajdr, 1957 has an irregular outline. • Key words: agnostids, Middle Cambrian, Jince Formation, Příbram-Jince Basin, Barrandian area, Czech Republic. FATKA, O., SZABAD,M.&BUDIL, P. 2009. Malformed agnostids from the Middle Cambrian Jince Formation of the Příbram-Jince Basin, Czech Republic. Bulletin of Geosciences 84(1), 121–126 (2 figures). Czech Geological Survey, Prague. ISSN 1214-1119. Manuscript received November 11, 2008; accepted in revised form January 9, 2009; published online January 23, 2009; issued March 31, 2009. Oldřich Fatka, Department of Geology and Palaeontology, Faculty of Science, Charles University, Albertov 6, Praha 2, CZ -128 43, Czech Republic; [email protected] • Michal Szabad, Obránců míru 75, 261 02 Příbram VII, Czech Re- public • Petr Budil, Czech Geological Survey, Klárov 3, Praha 1, CZ -118 21, Czech Republic; [email protected] Numerous examples of exoskeletal abnormalities have discussed by Babcock and Peng (2001). Öpik (1967) de- been described in various polymerid trilobites (e.g., Owen scribed and figured one pathological pygidium of Glyp- 1985, Babcock 1993, Whittington 1997), including para- tagnostus stolidotus Öpik, 1961 with hypertrophic devel- doxidid trilobites from the Cambrian Příbram-Jince Basin opment of the left side of the pygidium. of the Barrandian area (Šnajdr 1978). -
1500 Peng.Vp
Intraspecific variation and taphonomic alteration in the Cambrian (Furongian) agnostoid Lotagnostus americanus: new information from China SHANCHI PENG, LOREN E. BABCOCK, XUEJIAN ZHU, PER AHLBERG, FREDRIK TERFELT & TAO DAI The concept of the agnostoid arthropod species Lotagnostus americanus (Billings, 1860), which has been reported from numerous localities in the upper Furongian Series (Cambrian) of Laurentia, Gondwana, Baltica, Avalonia, and Siberia, is reviewed with emphasis on morphologic and taphonomic information afforded by large collections from Hunan in South China, Xinjiang in Northwest China, and Zhejiang in Southeast China. Comparisons are made with type and topotype material from Quebec, Canada, as well as material from elsewhere in Canada, the USA, the United Kingdom, Sweden, Russia, and Kazakhstan. The new information clarifies the limits of morphologic variability in L. americanus owing to ontogenetic changes and variation within holaspides, including inferred microevolution. It also provides details on apparent variation of taphonomic origin. The Chinese collections demonstrate a moderately wide variation in L. americanus, indicating that arguments favoring restriction of Lotagnostus species to narrowly defined, geographi- cally restricted forms are unwarranted. Species described as L. trisectus (Salter, 1864), L. asiaticus Troedsson, 1937, and L. punctatus Lu, 1964, for example, fall within the range of variation observed in L. americanus, and are regarded as ju- nior synonyms. The effaced form Lotagnostus obscurus Palmer, 1955 is removed from synonymy with L. americanus.A review of the stratigraphic distribution of L. americanus as construed here shows that the earliest occurrences of the spe- cies in all regions of the world are nearly synchronous. • Key words: Cambrian, Furongian, agnostoid, Lotagnostus americanus, China, Quebec. -
The Evolution of Trilobite Body Patterning
ANRV309-EA35-14 ARI 20 March 2007 15:54 The Evolution of Trilobite Body Patterning Nigel C. Hughes Department of Earth Sciences, University of California, Riverside, California 92521; email: [email protected] Annu. Rev. Earth Planet. Sci. 2007. 35:401–34 Key Words First published online as a Review in Advance on Trilobita, trilobitomorph, segmentation, Cambrian, Ordovician, January 29, 2007 diversification, body plan The Annual Review of Earth and Planetary Sciences is online at earth.annualreviews.org Abstract This article’s doi: The good fossil record of trilobite exoskeletal anatomy and on- 10.1146/annurev.earth.35.031306.140258 togeny, coupled with information on their nonbiomineralized tis- Copyright c 2007 by Annual Reviews. sues, permits analysis of how the trilobite body was organized and All rights reserved developed, and the various evolutionary modifications of such pat- 0084-6597/07/0530-0401$20.00 terning within the group. In several respects trilobite development and form appears comparable with that which may have charac- terized the ancestor of most or all euarthropods, giving studies of trilobite body organization special relevance in the light of recent advances in the understanding of arthropod evolution and devel- opment. The Cambrian diversification of trilobites displayed mod- Annu. Rev. Earth Planet. Sci. 2007.35:401-434. Downloaded from arjournals.annualreviews.org ifications in the patterning of the trunk region comparable with by UNIVERSITY OF CALIFORNIA - RIVERSIDE LIBRARY on 05/02/07. For personal use only. those seen among the closest relatives of Trilobita. In contrast, the Ordovician diversification of trilobites, although contributing greatly to the overall diversity within the clade, did so within a nar- rower range of trunk conditions. -
Dinosaur Provincial Park, Alberta
For Potuakhali University - http://www.pstu.ac.bd/ Fossil insect from Green River Formation Eocene Green River Shale, Wasatch County, Utah, USA http://www.ucmp.berkeley.edu/tertiary/eoc/greenriver.h tml https://en.wikipedia.org/wiki/Green_River_Formation Along the White River south of Bonanza, Utah. The low light colored outcrops near the river on the right are the Green River Formation Ammonite (Hematite replacement) Era – Cretaceous From – Morocco Triassic Scute Locality- Saint john, Arizona, USA (Blue-green Algae/ cyanobacterial colony) Stromatolite from Morocco Billion Years – 600 Million Years Old. Pre-Cambrian Locality – Near Atlas Mountain, Ourzarzate Morocco https://www.geocaching.com/geocache/GC2NKN4_moroccan- stromatolites http://epod.usra.edu/blog/2006/02/morocco- stromatolites.html Mosasaur Tooth, Cretaceous, Morocco . Triassic/Permian reptile bone ? Need better picture Ancient Mollusk, The Orthoceras 400 MYO, Devonian Morocco http://www.fossils-facts-and-finds.com/orthoceras.html https://en.wikipedia.org/wiki/Orthoceras Echinoid fossil Jurassic, Morocco Baltic Amber (Please confirm) 44 MYO - Eocene epoch Baltic amber (succinite) From-Poland (collected from Baltic Sea shores) Dinosaur Bone and black iron stone from Dinosaur Provincial park, Canada. Cretaceous ~65 MYO (reference Pictures below) Silurian Horn Coral – Rugos Kentucky, USA Jelly fish Carboniferous Era, Mazon Creek , USA (reference Pictures below) Carboniferous Trilobite, 300 Million Years=৩০ ক ো綿 বছর পুরোন ো Era – Carboniferous Locality – Bangor Limestone Formation, Northern Alabama, USA http://mrdata.usgs.gov/geology/state/sgmc- unit.php?unit=TNMbh%3B3 https://books.google.ca/books?id=zcdUAAAAYAAJ&pg=SL8- PA7&lpg=SL8- PA7&dq=Bangor+Limestone+Formation&source=bl&ots=JlCU5 _bKBc&sig=wCZ87cTc8hWHZHzXfSsteqG5azA&hl=en&sa=X&ve d=0ahUKEwi- lqL7_qnKAhULjZQKHQgzBQMQ6AEIOjAF#v=onepage&q=Bang or%20Limestone%20Formation&f=false LAVA STONE - SNOWFLAKE OBSIDIAN Could not recognize from picture the left one and the one under the paper & the one under the bigger ammonite in Ziplock. -
Ptychopariid Trilobites in the Middle Cambrian of Central Bohemia (Taxonomy, Biostratigraphy, Synecology)
Ptychopariid trilobites in the Middle Cambrian of Central Bohemia (taxonomy, biostratigraphy, synecology) VRATISLAV KORDULE A revision of ptychopariid trilobites from the Middle Cambrian of central Bohemia is presented. With a few exceptions, they were previously referred only to Ptychoparia striata. Three genera are recently distinguished: Ptychoparia Hawle & Corda, 1847, Ptychoparioides Růžička, 1940, and Mikaparia gen. nov. Seven species are described: three are revised, four are new, and two is left in open nomenclature; their stratigraphical ranges and significance are discussed. A new stratigraphical subdivision of the Middle Cambrian of the Skryje-Týřovice area is suggested, including three assemblage zones and three barren zones. Four substrate and bathymetrically controlled trilobite associations are recognized in the Skryje-Týřovice area. • Key words: Bohemia, Middle Cambrian, trilobites, new taxa, biostratigraphy. KORDULE, V. 2006. Ptychopariid trilobites in the Middle Cambrian of Central Bohemia (taxonomy, biostratigraphy, synecology). Bulletin of Geosciences 81(4), 277–304 (13 figures). Czech Geological Survey, Prague. ISSN 1214-1119. Manuscript received February 17, 2005; accepted in revised form December 4, 2006; issued December 31, 2006. Vratislav Kordule, Dlouhá 104, 261 01 Příbram III, Czech Republic; [email protected] Representatives of the genera Ptychoparia Hawle & Corda to study the materials in Vokáč’s private collection, the 1847, Ptychoparioides Růžička, 1940, and Mikaparia gen. specimens described and figured by Vokáč (1997) can be nov. are significant components of the trilobite fauna of identified, and their taxonomic position is revised. the Middle Cambrian marine deposits in the Jince and Following the previous stratigraphical models (their Skryje-Týřovice areas. Specimens referred to these taxa summary is given in Havlíček 1998 and Fatka et al. -
GSSP) of the Drumian Stage (Cambrian) in the Drum Mountains, Utah, USA
Articles 8585 by Loren E. Babcock1, Richard A. Robison2, Margaret N. Rees3, Shanchi Peng4, and Matthew R. Saltzman1 The Global boundary Stratotype Section and Point (GSSP) of the Drumian Stage (Cambrian) in the Drum Mountains, Utah, USA 1 School of Earth Sciences, The Ohio State University, 125 South Oval Mall, Columbus, OH 43210, USA. Email: [email protected] and [email protected] 2 Department of Geology, University of Kansas, Lawrence, KS 66045, USA. Email: [email protected] 3 Department of Geoscience, University of Nevada, Las Vegas, Las Vegas, NV 89145, USA. Email: [email protected] 4 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. Email: [email protected] The Global boundary Stratotype Section and Point correlated with precision through all major Cambrian regions. (GSSP) for the base of the Drumian Stage (Cambrian Among the methods that should be considered in the selection of a GSSP (Remane et al., 1996), biostratigraphic, chemostratigraphic, Series 3) is defined at the base of a limestone (cal- paleogeographic, facies-relationship, and sequence-stratigraphic cisiltite) layer 62 m above the base of the Wheeler For- information is available (e.g., Randolph, 1973; White, 1973; McGee, mation in the Stratotype Ridge section, Drum Moun- 1978; Dommer, 1980; Grannis, 1982; Robison, 1982, 1999; Rowell et al. 1982; Rees 1986; Langenburg et al., 2002a, 2002b; Babcock et tains, Utah, USA. The GSSP level contains the lowest al., 2004; Zhu et al., 2006); that information is summarized here. occurrence of the cosmopolitan agnostoid trilobite Pty- Voting members of the International Subcommission on Cam- chagnostus atavus (base of the P. -
Early and Middle Cambrian Trilobites from Antarctica
Early and Middle Cambrian Trilobites From Antarctica GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Early and Middle Cambrian Trilobites From Antarctica By ALLISON R. PALMER and COLIN G. GATEHOUSE CONTRIBUTIONS TO THE GEOLOGY OF ANTARCTICA GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Bio stratigraphy and regional significance of nine trilobite faunules from Antarctic outcrops and moraines; 28 species representing 21 genera are described UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1972 UNITED STATES DEPARTMENT OF THE INTERIOR ROGERS C. B. MORTON, Secretary GEOLOGICAL SURVEY V. E. McKelvey, Director Library of Congress catalog-card No. 73-190734 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price 70 cents (paper cover) Stock Number 2401-2071 CONTENTS Page Page Abstract_ _ ________________________ Dl Physical stratigraphy______________________________ D6 I&troduction. _______________________ 1 Regional correlation within Antarctica ________________ 7 Biostratigraphy _____________________ 3 Systematic paleontology._____-_______-____-_-_-----_ 9 Early Cambrian faunules.________ 4 Summary of classification of Antarctic Early and Australaspis magnus faunule_ 4 Chorbusulina wilkesi faunule _ _ 5 Middle Cambrian trilobites. ___________________ 9 Chorbusulina subdita faunule _ _ 5 Agnostida__ _ _________-____-_--____-----__---_ 9 Early Middle Cambrian f aunules __ 5 Redlichiida. __-_--------------------------_---- 12 Xystridura mutilinia faunule- _ 5 Corynexochida._________--________-_-_---_----_ -
(Upper Cambrian, Paibian) Trilobite Faunule in the Central Conasauga River Valley, North Georgia, Usa
Schwimmer.fm Page 31 Monday, June 18, 2012 11:54 AM SOUTHEASTERN GEOLOGY V. 49, No. 1, June 2012, p. 31-41 AN APHELASPIS ZONE (UPPER CAMBRIAN, PAIBIAN) TRILOBITE FAUNULE IN THE CENTRAL CONASAUGA RIVER VALLEY, NORTH GEORGIA, USA DAVID R. SCHWIMMER1 WILLIAM M. MONTANTE2 1Department of Chemistry & Geology Columbus State University, 4225 University Avenue, Columbus, Georgia 31907, USA <[email protected]> 2Marsh & McLennan, Inc., 3560 Lenox Road, Suite 2400, Atlanta, Georgia 30326, USA <[email protected]> ABSTRACT shelf-to-basin break, which is interpreted to be east of the Alabama Promontory and in Middle and Upper Cambrian strata the Tennessee Embayment. The preserva- (Cambrian Series 3 and Furongian) in the tion of abundant aphelaspine specimens by southernmost Appalachians (Tennessee to bioimmuration events may have been the re- Alabama) comprise the Conasauga Forma- sult of mudflows down the shelf-to-basin tion or Group. Heretofore, the youngest re- slope. ported Conasauga beds in the Valley and Ridge Province of Georgia were of the late INTRODUCTION Middle Cambrian (Series 3: Drumian) Bo- laspidella Zone, located on the western state Trilobites and associated biota from Middle boundary in the valley of the Coosa River. Cambrian beds of the Conasauga Formation in Two new localities sited eastward in the Co- northwestern Georgia have been described by nasauga River Valley, yield a diagnostic suite Walcott, 1916a, 1916b; Butts, 1926; Resser, of trilobites from the Upper Cambrian 1938; Palmer, 1962; Schwimmer, 1989; Aphelaspis Zone. Very abundant, Schwimmer and Montante, 2007. These fossils polymeroid trilobites at the primary locality and deposits come from exposures within the are referable to Aphelaspis brachyphasis, valley of the Coosa River, in Floyd County, which is a species known previously in west- Georgia, and adjoining Cherokee County, Ala- ern North America. -
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900 By ALLISON R. PALMER A SHORTER CONTRIBUTION TO GENERAL GEOLOGY GEOLOGICAL SURVEY PROFESSIONAL PAPER 264-D Of the 2ty species described prior to I(?OO, 2/ are redescribed and 2C} refigured, and a new name is proposedfor I species UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1954 UNITED STATES DEPARTMENT OF THE INTERIOR Douglas McKay, Secretary GEOLOGICAL SURVEY W. E. Wrather, Director For sale by the Superintendent of Documents, U. S. Government Printing Office Washington 25, D. C. - Price $1 (paper cover) CONTENTS Page Abstract..__________________________________ 55 Introduction ________________________________ 55 Original and present taxonomic names of species. 57 Stratigraphic distribution of species ____________ 57 Collection localities._________________________ 58 Systematic descriptions.______________________ 59 Literature cited____________________________ 82 Index __-_-__-__---_--______________________ 85 ILLUSTRATIONS [Plates 13-17 follow page 86] PLATE 13. Agnostidae and Dolichometopidae 14. Dorypygidae 15. Oryctocephalidae, Dorypygidae, Zacanthoididae, and Ptychoparioidea 16. Ptychoparioidea 17. Ptychoparioidea FIGUBE 3. Index map showing collecting localities____________________________ . Page 56 in A SHORTER CONTRIBUTION TO GENERAL GEOLOGY AN APPRAISAL OF THE GREAT BASIN MIDDLE CAMBRIAN TRILOBITES DESCRIBED BEFORE 1900 By ALLISON R. PALMER ABSTRACT the species and changes in their generic assignments All 29 species of Middle Cambrian trilobites