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Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
Native Plants for Wildlife Habitat and Conservation Landscaping Chesapeake Bay Watershed Acknowledgments
U.S. Fish & Wildlife Service Native Plants for Wildlife Habitat and Conservation Landscaping Chesapeake Bay Watershed Acknowledgments Contributors: Printing was made possible through the generous funding from Adkins Arboretum; Baltimore County Department of Environmental Protection and Resource Management; Chesapeake Bay Trust; Irvine Natural Science Center; Maryland Native Plant Society; National Fish and Wildlife Foundation; The Nature Conservancy, Maryland-DC Chapter; U.S. Department of Agriculture, Natural Resource Conservation Service, Cape May Plant Materials Center; and U.S. Fish and Wildlife Service, Chesapeake Bay Field Office. Reviewers: species included in this guide were reviewed by the following authorities regarding native range, appropriateness for use in individual states, and availability in the nursery trade: Rodney Bartgis, The Nature Conservancy, West Virginia. Ashton Berdine, The Nature Conservancy, West Virginia. Chris Firestone, Bureau of Forestry, Pennsylvania Department of Conservation and Natural Resources. Chris Frye, State Botanist, Wildlife and Heritage Service, Maryland Department of Natural Resources. Mike Hollins, Sylva Native Nursery & Seed Co. William A. McAvoy, Delaware Natural Heritage Program, Delaware Department of Natural Resources and Environmental Control. Mary Pat Rowan, Landscape Architect, Maryland Native Plant Society. Rod Simmons, Maryland Native Plant Society. Alison Sterling, Wildlife Resources Section, West Virginia Department of Natural Resources. Troy Weldy, Associate Botanist, New York Natural Heritage Program, New York State Department of Environmental Conservation. Graphic Design and Layout: Laurie Hewitt, U.S. Fish and Wildlife Service, Chesapeake Bay Field Office. Special thanks to: Volunteer Carole Jelich; Christopher F. Miller, Regional Plant Materials Specialist, Natural Resource Conservation Service; and R. Harrison Weigand, Maryland Department of Natural Resources, Maryland Wildlife and Heritage Division for assistance throughout this project. -
An Encyclopedia of Shade Perennials This Page Intentionally Left Blank an Encyclopedia of Shade Perennials
An Encyclopedia of Shade Perennials This page intentionally left blank An Encyclopedia of Shade Perennials W. George Schmid Timber Press Portland • Cambridge All photographs are by the author unless otherwise noted. Copyright © 2002 by W. George Schmid. All rights reserved. Published in 2002 by Timber Press, Inc. Timber Press The Haseltine Building 2 Station Road 133 S.W. Second Avenue, Suite 450 Swavesey Portland, Oregon 97204, U.S.A. Cambridge CB4 5QJ, U.K. ISBN 0-88192-549-7 Printed in Hong Kong Library of Congress Cataloging-in-Publication Data Schmid, Wolfram George. An encyclopedia of shade perennials / W. George Schmid. p. cm. ISBN 0-88192-549-7 1. Perennials—Encyclopedias. 2. Shade-tolerant plants—Encyclopedias. I. Title. SB434 .S297 2002 635.9′32′03—dc21 2002020456 I dedicate this book to the greatest treasure in my life, my family: Hildegarde, my wife, friend, and supporter for over half a century, and my children, Michael, Henry, Hildegarde, Wilhelmina, and Siegfried, who with their mates have given us ten grandchildren whose eyes not only see but also appreciate nature’s riches. Their combined love and encouragement made this book possible. This page intentionally left blank Contents Foreword by Allan M. Armitage 9 Acknowledgments 10 Part 1. The Shady Garden 11 1. A Personal Outlook 13 2. Fated Shade 17 3. Practical Thoughts 27 4. Plants Assigned 45 Part 2. Perennials for the Shady Garden A–Z 55 Plant Sources 339 U.S. Department of Agriculture Hardiness Zone Map 342 Index of Plant Names 343 Color photographs follow page 176 7 This page intentionally left blank Foreword As I read George Schmid’s book, I am reminded that all gardeners are kindred in spirit and that— regardless of their roots or knowledge—the gardening they do and the gardens they create are always personal. -
Diversity and Evolution of Monocots
Lilioids - petaloid monocots 4 main groups: Diversity and Evolution • Acorales - sister to all monocots • Alismatids of Monocots – inc. Aroids - jack in the pulpit ! • Lilioids (lilies, orchids, yams) – grade, non-monophyletic – petaloid . orchids and palms . ! • Commelinoids – Arecales – palms – Commelinales – spiderwort – Zingiberales –banana – Poales – pineapple – grasses & sedges Lilioids - petaloid monocots Asparagales: *Orchidaceae - orchids • finish the Asparagales by 1. Terrestrial/epiphytes: plants looking at the largest family - typically not aquatic the orchids 2. Geophytes: herbaceous above ground with below ground modified perennial stems: bulbs, corms, rhizomes, tubers 3. Tepals: showy perianth in 2 series of 3 each; usually all petaloid, or outer series not green and sepal-like & with no bracts 1 *Orchidaceae - orchids *Orchidaceae - orchids The family is diverse with about 880 genera and over 22,000 All orchids have a protocorm - a feature restricted to the species, mainly of the tropics family. Orchids are • structure formed after germination and before the mycotrophic (= fungi development of the seedling plant dependent) lilioids; • has no radicle but instead mycotrophic tissue some are obligate mycotrophs Cypripedium acaule Corallorhiza striata Stemless lady-slipper Striped coral root Dactylorhiza majalis protocorm *Orchidaceae - orchids *Orchidaceae - orchids Cosmopolitan, but the majority of species are found in the Survive in these epiphytic and other harsh environments via tropics and subtropics, ranging from sea -
Wildflowers and Ferns Along the Acton Arboretum Wildflower Trail and in Other Gardens FERNS (Including Those Occurring Naturally
Wildflowers and Ferns Along the Acton Arboretum Wildflower Trail and In Other Gardens Updated to June 9, 2018 by Bruce Carley FERNS (including those occurring naturally along the trail and both boardwalks) Royal fern (Osmunda regalis): occasional along south boardwalk, at edge of hosta garden, and elsewhere at Arboretum Cinnamon fern (Osmunda cinnamomea): naturally occurring in quantity along south boardwalk Interrupted fern (Osmunda claytoniana): naturally occurring in quantity along south boardwalk Maidenhair fern (Adiantum pedatum): several healthy clumps along boardwalk and trail, a few in other Arboretum gardens Common polypody (Polypodium virginianum): 1 small clump near north boardwalk Hayscented fern (Dennstaedtia punctilobula): aggressive species; naturally occurring along north boardwalk Bracken fern (Pteridium aquilinum): occasional along wildflower trail; common elsewhere at Arboretum Broad beech fern (Phegopteris hexagonoptera): up to a few near north boardwalk; also in rhododendron and hosta gardens New York fern (Thelypteris noveboracensis): naturally occurring and abundant along wildflower trail * Ostrich fern (Matteuccia pensylvanica): well-established along many parts of wildflower trail; fiddleheads edible Sensitive fern (Onoclea sensibilis): naturally occurring and abundant along south boardwalk Lady fern (Athyrium filix-foemina): moderately present along wildflower trail and south boardwalk Common woodfern (Dryopteris spinulosa): 1 patch of 4 plants along south boardwalk; occasional elsewhere at Arboretum Marginal -
Vegetation Community Monitoring at Congaree National Park: 2014 Data Summary
National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science Vegetation Community Monitoring at Congaree National Park 2014 Data Summary Natural Resource Data Series NPS/SECN/NRDS—2016/1016 ON THIS PAGE Tiny, bright yellow blossoms of Hypoxis hirsuta grace the forest floor at Congaree National Park. Photograph courtesy of Sarah C. Heath, Southeast Coast Network. ON THE COVER Spiraling compound leaf of green dragon (Arisaema dracontium) at Congaree National Park. Photograph courtesy of Sarah C. Heath, Southeast Coast Network Vegetation Community Monitoring at Congaree National Park 2014 Data Summary Natural Resource Data Series NPS/SECN/NRDS—2016/1016 Sarah Corbett Heath1 and Michael W. Byrne2 1National Park Service Southeast Coast Inventory and Monitoring Network Cumberland Island National Seashore 101 Wheeler Street Saint Marys, GA 31558 2National Park Service Southeast Coast Inventory and Monitoring Network 135 Phoenix Drive Athens, GA 30605 May 2016 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado, publishes a range of reports that address natural resource topics. These reports are of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Data Series is intended for the timely release of basic data sets and data summaries. Care has been taken to assure accuracy of raw data values, but a thorough analysis and interpretation of the data has not been completed. -
Phylogenetic Relationships of Monocots Based on the Highly Informative Plastid Gene Ndhf Thomas J
Aliso: A Journal of Systematic and Evolutionary Botany Volume 22 | Issue 1 Article 4 2006 Phylogenetic Relationships of Monocots Based on the Highly Informative Plastid Gene ndhF Thomas J. Givnish University of Wisconsin-Madison J. Chris Pires University of Wisconsin-Madison; University of Missouri Sean W. Graham University of British Columbia Marc A. McPherson University of Alberta; Duke University Linda M. Prince Rancho Santa Ana Botanic Gardens See next page for additional authors Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Givnish, Thomas J.; Pires, J. Chris; Graham, Sean W.; McPherson, Marc A.; Prince, Linda M.; Patterson, Thomas B.; Rai, Hardeep S.; Roalson, Eric H.; Evans, Timothy M.; Hahn, William J.; Millam, Kendra C.; Meerow, Alan W.; Molvray, Mia; Kores, Paul J.; O'Brien, Heath W.; Hall, Jocelyn C.; Kress, W. John; and Sytsma, Kenneth J. (2006) "Phylogenetic Relationships of Monocots Based on the Highly Informative Plastid Gene ndhF," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 22: Iss. 1, Article 4. Available at: http://scholarship.claremont.edu/aliso/vol22/iss1/4 Phylogenetic Relationships of Monocots Based on the Highly Informative Plastid Gene ndhF Authors Thomas J. Givnish, J. Chris Pires, Sean W. Graham, Marc A. McPherson, Linda M. Prince, Thomas B. Patterson, Hardeep S. Rai, Eric H. Roalson, Timothy M. Evans, William J. Hahn, Kendra C. Millam, Alan W. Meerow, Mia Molvray, Paul J. Kores, Heath W. O'Brien, Jocelyn C. Hall, W. John Kress, and Kenneth J. Sytsma This article is available in Aliso: A Journal of Systematic and Evolutionary Botany: http://scholarship.claremont.edu/aliso/vol22/iss1/ 4 Aliso 22, pp. -
Spiranthes Bightensis (Orchidaceae), a New and Rare Cryptic Hybrid Species Endemic to the U
Phytotaxa 498 (3): 159–176 ISSN 1179-3155 (print edition) https://www.mapress.com/j/pt/ PHYTOTAXA Copyright © 2021 Magnolia Press Article ISSN 1179-3163 (online edition) https://doi.org/10.11646/phytotaxa.498.3.2 Spiranthes bightensis (Orchidaceae), a New and Rare Cryptic Hybrid Species Endemic to the U. S. Mid-Atlantic Coast MATTHEW C. PACE New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10348, U. S. A. [email protected]; https://orcid.org/0000-0002-2333-1924 Abstract Recognizing species diversity is challenging in genera that display interspecific similarity and intraspecific variation; hy- bridization and the evolution of cryptic hybrid species amplifies these challenges. Recent molecular and morphological research focused on the systematics of Spiranthes (Orchidaceae) support hybrid speciation as an important driver of species diversity, particularly within the S. cernua species complex. Working under an integrated history-bound phylogenetic spe- cies concept, new molecular and morphometric data provide evidence for a new and rare cryptic hybrid species resulting from the ancient hybridization of S. cernua × S. odorata, here described as S. bightensis. Although S. bightensis is regionally sympatric with S. cernua it does not co-occur with that species, and it is allopatric with respect to S. odorata. Endemic to a narrow region extending from the Delmarva Peninsula to Long Island, New York, this new species occurs in the shadow of the Northeast megalopolis and appears to have undergone a major population decline over the last 200 years. By recognizing this distinct evolutionary lineage as a new species, this research is the first step towards developing conservation protocols for this rare species and highlights the importance of the North American Geologic Coastal Plain for biodiversity conserva- tion and evolution. -
Spiranthinae, Cranichideae, Orchidoideae, Orchidaceae)
LEONARDO RAMOS SEIXAS GUIMARÃES Filogenia e citotaxonomia do clado Stenorrhynchos (Spiranthinae, Cranichideae, Orchidoideae, Orchidaceae) Tese apresentada ao Instituto de Botânica da Secretaria do Meio Ambiente, como parte dos requisitos exigidos para obtenção do título de DOUTOR em BIODIVERSIDADE VEGETAL E MEIO AMBIENTE, na área de concentração de Plantas Vasculares. SÃO PAULO 2014 LEONARDO RAMOS SEIXAS GUIMARÃES Filogenia e citotaxonomia do clado Stenorrhynchos (Spiranthinae, Cranichideae, Orchidoideae, Orchidaceae) Tese apresentada ao Instituto de Botânica da Secretaria do Meio Ambiente, como parte dos requisitos exigidos para obtenção do título de DOUTOR em BIODIVERSIDADE VEGETAL E MEIO AMBIENTE, na área de concentração de Plantas Vasculares. ORIENTADOR: DR. FÁBIO DE BARROS (INSTITUTO DE BOTÂNICA, SÃO PAULO, BRASIL) COORIENTADOR: DR. GERARDO A. SALAZAR (INSTITUTO DE BIOLOGÍA, UNIVERSIDAD NACIONAL AUTÓNOMA DE MÉXICO, D.F., MÉXICO) Ficha Catalográfica elaborada pelo NÚCLEO DE BIBLIOTECA E MEMÓRIA Guimarães, Leonardo Ramos Seixas G963fi Filogenia e citotaxonomia do clado Stenorrhynchos (Spiranthinae, Cranichideae, Orchidoideae, Orchidaceae) / Leonardo Ramos Seixas Guimarães -- São Paulo, 2014. 101 p. il. Tese (Doutorado) -- Instituto de Botânica da Secretaria de Estado do Meio Ambiente, 2014 Bibliografia. 1. Orchidaceae. 2. Citogenética. 3. Sistemática molecular. I. Título CDU: 582.594.2 Data de defesa: 9 Junho 2014 BANCA EXAMINADORA Prof. Dr. Fábio de Barros (IBt, São Paulo-SP) Prof. Dr. Rodolfo Aniceto Solano Gómez (CIIDIR Unidad Oaxaca, -
Tresses, Spiranthes Casei Var. Casei (Orchidaceae), in Ottawa, Ontario
Phenology, Population Dynamics, and Flowering Dynamics of Case’s Ladies’-tresses, Spiranthes casei var. casei (Orchidaceae), in Ottawa, Ontario JOYCE M. R EDDOCH and ALLAN H. R EDDOCH 548 Rivershore Crescent, Gloucester, Ontario K1J 7Y7 Canada; e-mail: [email protected] Reddoch, Joyce M., and Allan H. Reddoch. 2008. Phenology, population dynamics, and flowering dynamics of Case’s Ladies’- tresses, Spiranthes casei var. casei (Orchidaceae), in Ottawa, Ontario. Canadian Field-Naturalist 123(1): 19-31. We describe a seven-year study (2002–2008) of a population of Case’s Ladies’-tresses ( Spiranthes casei var. casei ) in the western Greenbelt in Ottawa, Ontario, that had been extant since at least 1972. We also record a temporary colonization in the eastern Greenbelt for 11 years (1998–2008). The seven life history stages identified were seed, juvenile, immature, flowering, vegetative, offshoot (ramet), and non-emergent. Mature plants present in the first two years of the study had mean half lives of 9.4 years. Flowering and seed production were limited by damp, cloudy weather when fall rosettes devel - op, by gastropod herbivory in early summer, by drought in mid-summer, and by deer herbivory in late summer. Given the resulting wide fluctuations in the fractions of plants flowering, as well as the lack of visibility of vegetative plants, this study demonstrates the limitations of annual censuses of flowering stems for determining the actual sizes of populations and for detecting long-term population trends. Key Words: Case’s Ladies’-tresses, Spiranthes casei , Brown-lipped Snail, Cepaea nemoralis , Banded Wood Snail, Stetho - baris ovata , White-tailed Deer , Odocoileus virginianus, flowering dynamics, gastropod, herbivory, long-term study, population dynamics, Ottawa, Ontario. -
Breeding Systems in Spiranthes Magnicamporum Sheviak Christopher D
Eastern Illinois University The Keep Masters Theses Student Theses & Publications 1993 Breeding Systems in Spiranthes magnicamporum Sheviak Christopher D. Loyet This research is a product of the graduate program in Botany at Eastern Illinois University. Find out more about the program. Recommended Citation Loyet, Christopher D., "Breeding Systems in Spiranthes magnicamporum Sheviak" (1993). Masters Theses. 2179. https://thekeep.eiu.edu/theses/2179 This is brought to you for free and open access by the Student Theses & Publications at The Keep. It has been accepted for inclusion in Masters Theses by an authorized administrator of The Keep. For more information, please contact [email protected]. THESIS REPRODUCTION CERTIFICATE TO: Graduate Degree Candidates who have written formal theses. SUBJECT: Permission to reproduce theses. The University Library is receiving a number of requests from other institutions asking permission to reproduce dissertations for inclusion in their library holdings. Although no copyright laws are involved, we feel that professional courtesy demands that permission be obtained from the author before we allow theses to be copied. Please sign one of the following statements: . Booth Library 0£ Eastern Illinois University has my permission to lend my thesis to a reputable college or university for the purpose 0£ copying it for inclusion in that institution's library or research holdings. o~ lot /"\~ Date I respectfully request Booth Library 0£ Eastern Illinois University not allow my thesis be reproduced because --------------------------- Date Author m Breeding systems in Spiranthes magnicamporum Sheviak. (TITLE) BY Christopher D. Loyet THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF Masters of Science Major in Botany IN THE GRADUATE SCHOOL, EASTERN ILLINOIS UNIVERSITY CHARLESTON, ILLINOIS 1993 YEAR I HEREBY RECOMMEND THIS THESIS BE ACCEPTED AS FULFILLING THIS PART OF THE GRADUATE DEGREE CITED ABOVE ~.2<:;DAT f(fq3 TABLE OF CONTENTS ABSTRACT. -
Navasota Ladies' Tresses
Figure 1. Distribution of Spiranthes parksii populations. 236 NATIVEPLANTS | 13 | 3 | FALL 2012 BIOLOGY, ECOLOGY, AND CONSERVATION OF Navasota ladies’ tresses (Spiranthes parksii Correll) AN ENDANGERED TERRESTRIAL ORCHID OF TEXAS Carissa L Wonkka, William E Rogers, Fred E Smeins, J Ryan Hammons, Sarah J Haller, and Martha C Ariza ABSTRACT Navasota ladies’ tresses (Spiranthes parksii Correll [Orchidaceae]) is a federally and state-listed endangered orchid of east-central Texas. Habitat loss and degradation re- lated to urban and industrial development are major threats to S. parksii populations. To ensure recovery, a complete understanding of the population dynamics, ecology, and biology of an endangered species is necessary to foster effective conservation that is compatible with human population growth and continued development. Here we provide an overview of the known aspects of Spiranthes parksii ecology and biol- ogy and highlight factors with implications for species conservation. Our intention is to provide a framework for development of future S. parksii related studies and back- ground for those interested in S. parksii conservation and management. Wonkka CL, Rogers WE, Smeins FE, Hammons JR, Haller SJ, Ariza MC. 2012. Biology, ecology, and conservation of Navasota ladies’ tresses (Spiranthes parksii Correll), an endangered terrestrial orchid of Texas. Native Plants Journal 13(3):236–243. CONVERSIONS 1 mm = 0.04 in K E Y W O R D S 1 cm = 0.4 in Orchidaceae, natural history, habitat management 1 m = 3.3 ft (°C * 1.8) + 32 = °F NOMENCLATURE USDA NRCS (2012) 237 NATIVEPLANTS | 13 | 3 | FALL 2012 avasota ladies’ tresses (Spiranthes parksii Correll [Or- Spiranthes parksii is sympatric with several other species in chidaceae]) is a federally listed endangered species the genus Spiranthes throughout its range, including cernua (L.) Nendemic to east-central Texas (USFWS 1996).