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Devonian Proetidae (Trilobita) from the Ardennes Massif (Belgium, N France) and the Eifel Hills (W Germany)

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Devonian Proetidae (Trilobita) from the Ardennes Massif (Belgium, N France) and the Eifel Hills (W Germany) BULLETIN DE L’INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE SCIENCES DE LA TERRE, 78: 9-29,2008 BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN AARDWETENSCHAPPEN, 78: 9-29, 2008 Devonian Proetidae (Trilobita) from the Ardennes Massif (Belgium, N France) and the Eifel Hills (W Germany) by Allart P. VAN VIERSEN «fe Harald PRESCHER Viersen, A.P. van & Prescher, H., 2008 - Devonian Proetidae Introduction (Trilobita) from the Ardennes Massif (Belgium, N France) and the Eifel Hills (W Germany). In: Steurbaut, E., Jagt, J.W.M. & Jagt- The family Proetidae (proetids) encompasses a large Yazykova, E.A. (Editors), Annie V. Dhondt Memorial Volume. group of trilobites in the Devonian of the Ardennes. Bulletin de l ’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 78: 9-29, 5 figs, 5 pis, Brussels, October 31, Early Devonian trilobite faunas in this region are low in 2008-ISSN 0374-6291. diversity, and dominated by the families Acastidae and Homalonotidae. The appearance of Proetidae during the Middle Devonian here is relatively late when compared Abstract to abundant Early Devonian occurrences elsewhere, e.g. in Bohemia (S n a jd r , 1980) and Morocco (ALBERTI, New data are presented on trilobites of the family Proetidae 1969). In the Ardennes, the stratigraphically earliest (subfamilies Proetinae, Dechenellinae, Cornuproetinae) from the confirmed records of Proetidae come from the lower Devonian of the Ardennes Massif and the Eifel Hills (Rhenohercynian zone). Preliminary stratigraphie distributions of the Proetidae from Eifelian (Couvin and Jemelle Formations), which more the southern border of the Dinant Synclinorium are introduced. or less coincides with putative initial migration of Eight taxa are recorded, namely: Gerastos prox lessensis n. ssp., trilobites from Gondwana towards the northern margin Gerastos catervus (VAN VlERSEN, 2006), Gerastos sp. B, Dohrniella of the Rheic Ocean (S c h r a u t <fe Feist , 2004) and the tenuiornata n. sp., Dechenella daumeriesi n. sp., Dechenella sp. A, development of shallow-water carbonate deposition Cornuproetus cornutus ssp. 2, and DiademaproetusI sp. In addition, Dohrniella stumporum n. sp. andGerastos sp. A are described from in the Ardennes(C r ô n ier «fe v a n V ie r se n , 2007). the Eifel. The genus Rhenocynproetus BASSE, 2002 is regarded as a Claimed occurrences of proetids in the upper Emsian of junior subjective synonym ofGerastos G o ld fu ss, 1843. Belgium (e.g. M a il l ie u x , 1933; A s se l b e r g h s , 1946) have not been confirmed but are possible given that Keywords: Trilobites, Proetidae, Devonian, Ardennes, Eifel. they were recently reported from upper Emsian strata Résumé in adjacent regions (e.g. Proetinae and Cornuproetinae from Luxembourg by B a s s e , 2002 and B a sse «fe De nouvelles données sont présentées sur les trilobites de la famille F r a n k e , 2006). Throughout the Eifelian, family des Proetidae (sous-familles des Proetinae, des Dechenellinae et Proetidae is represented in the Ardennes by a substantial des Cornuproetinae) du Dévonien du Massif des Ardennes et des variety of species comprising at least six genera (Fig. collines de l’Eifel (zone rhéno-hercynienne). Une distribution stratigraphique préliminaire des Proetidae sur le bord méridional 1). Diversity of the family declined significantly in this du Synclinorium de Dinant est proposée. Huit taxa sont reconnus, region at around the Eifelian-Givetian boundary while à savoir Gerastos prox lessensis n. ssp., Gerastos catervus (VAN only Gerastos and Dechenella are known with certainty VlERSEN, 2006), Gerastos sp. B, Dohrniella tenuiornata n. sp., to continue into the Givetian. Dechenella daumeriesi n. sp., Dechenella sp. A, Cornuproetus cornutus ssp. 2 et DiademaproetusI sp. En complément, Dohrniella The proetid fauna of the Devonian of the Ardennes stumporum n. sp. et Gerastos sp. A sont décrites dans la région has been documented inadequately which is due mainly de l’Eifel. Le genre Rhenocynproetus BASSE, 2002 est considéré to a 60-year intermission of research on Devonian comme un synonyme plus récent de Gerastos G o ld fu ss, 1843. trilobites there. Although occurrences of proetids are frequently reported in the older literature (e.g. Mots-clefs: Trilobites, Proetidae, Dévonien, Ardennes, Eifel. M a il l ie u x , 1904, 1919, 1933, 1938; R ichter <fe 10 Allart P. VAN VIERSEN & Harald PRESCHER Netherlands, in 2004], Recent collecting by the authors at Devonian outcrops on the southern border of the ■c </> Dinant Synclinorium (Belgium, northern France) has yielded a wealth of new trilobite material which has O N enabled us to recognise a number of new proetid species and subspecies, some of which are described herein. In timorensis addition, one new species is recorded from Eifelian Terres d'Haurs strata in the Eifel. Trois-Fontaines hemiansatus Hanonet Locations and stratigraphy ensensis La Lomme Ardennes Massif (southern border of Dinant Synclinorium) kockelianus Loc002, Resteigne Abandoned quarry at Resteigne, Belgium. Trilobites come from the basal part of the Hanonet Formation, just below the Eifelian-Givetian boundary (Polygnathus ensensis conodont australis Jemelle Zone). This locality was recently discussed and indicated on a map by van Viersen (2007b) who recorded several trilobite species from it and provided references to additional literature sources. Trilobites from the uppermost Eifelian at this locality include; Calycoscutellum goolaertsi VAN costatus VlERSEN, 2007b, Nyterops hollandi v a n VlERSEN, 2007b, Dechenella daumeriesi n. sp., Dohrniella tenuiornata n. sp., Gerastosprox lessensis n. ssp., Hypsipariops? sp., Cyphaspis sp. [= Otarioninae gen. & sp. indet. o f VAN VlERSEN (2007b)], and Astycoryphe sp. Accompanying macrofauna Couvin partitus (det. H.P.): Brachiopods: abundant occurrences oGypidula f abunda STRUVE ssp. and Invertrypa cf. kelusiana (STRUVE); Primipilaria pripilaroides STRUVE, Productella subaculeata Eau Noire (MURCHISON), Desquamatia (Synatrypa) cf. subzonata patulus (BlERNAT), and Atryparia cf. instita COPPER. Gastropods: St.-Joseph "Laxonema" sp., Omphalocirrus sp., and Bellerophon serotinus Hlerges sp. Cephalopods, as well as crinoid ossicles and crowns belonging to Abbreviatocrinites BOHATŸ are common. Fig. 1 — Generalised overview of stratigraphie units on Loc004, Jemelle the southern border of the Dinant Synclinorium Embankment south of the Jemelle railway station, Belgium (Belgium, northern France) (after B u l t y n c k (see M agrean & van Viersen, 2005). Trilobite specimens & DEJONGHE, 2001, fig. 5). Indications of old come from argillaceous limestones belonging to the Jemelle (Belgian) stratigraphie divisions are auxiliary. Formation Polygnathus( partitus - ensensis conodont zones), Continuous lines indicate known stratigraphie Chavées Member; middle Eifelian. The rich macrofauna is ranges of proetid trilobite genera; broken lines highly diverse in trilobites which are represented here by at are tentative ranges. For occurrences in other least 21 species: Koneprusia maillieuxi M a g r e a n , 2006, formations and further explanation on these Kettneraspis bayarti VAN VlERSEN, 2007a, Radiaspis cf. stratigraphie ranges, see under Systematic comes (B a s s e , 1998), Ceratatges cf. armatus (GOLDFUSS, palaeontology. 1839), Eifliarges sp. cf. Eifliarges sp. o f BASSE in BASSE & MÜLLER (2004) [= Eifliarges caudimurus (sic) o fMAGREAN (2006)], Cornuproetus cornutus ssp. 2 (see below), Gerastos sp., Dohrniella sp. 1 o f VAN VlERSEN (2006b), R ichter , 1918; v a n Tu ijn , 1927; A sse l b e r g h s , Tropidocoryphe sp. cf. Tropidocoryphe n. sp. G o f BASSE 1946), examination of some of the original specimens (2002), Astycoryphe senckenbergiana RICHTER & RICHTER, by one of us (A.V.) led us to the conclusion that genera 1919, Cyphaspis sp. B and Cyphaspis? sp. C o fVAN VlERSEN and species have not been properly discriminated [e.g. & PRESCHER (2007), Cyphaspis cf. hoepfneri (KOWALSKI, the van Tuijn Collection (v a n Tu ijn , 1927) from the 1990), Phaeops cf. imitator STRUVE, 1970, Pedinopariops cf. Ferrières area (Fig. 2), Natuurmuseum Nijmegen, the brongniarti (STEININGER, 1831), Pedinopariops sp., Geesops Devonian Proetidae from the Ardennes and the Eifel 11 Generalised geolo- gical map o f the Ardennes (after Bu l t y n c k & D e jo n g h e , 2001, Namur .Sambra fig. 1). ¡harleroi Meuse LOC004 Ü3C002 f l post-Palaeozoic □ Carboniferous Rocroi Massif P I Middle and Upper Devonian I Lower Devonian I Silurian, Ordovician. Cambrian sp., Asteropyge cf. comes B a s s e , 2003, Calycoscutellum and Crônier & van Viersen(2008). sp., Scutellum sp., and Scabriscutellum canaliculatum (GOLDFUSS, 1843) [= Scabriscutellum vogeli A r c h in a l , Eifel 1994 fide B a s s e in Basse & M üller (2004)]. GerolsteinOl, Lissingen Temporary site (1998 to 1999) near Lissingen, by the road Loc005, Wellin between Gerolstein and Miillenhorn; coordinates are r: Quarry “Les Limites”, just north of Wellin, Belgium. (25)45440, h: (55)65280 (Blatt Gerolstein, topographical Trilobites come from the X FormationPolygnathus ( ensensis map 1:25,000). Specimens come from the lower part of the - Ihemiansatus conodont zones); upper Eifelian to ?lower Bildstock Member (Polygnathus costatus conodont Zone), Givetian. This formation comprises crinoidal and reefal Ahrdorf Formation; middle Eifelian. Trilobites include limestones, and has yielded rich coral and stromatoporoid Dohrniella stumporum n. sp., Gerastos sp. A (see below), faunas. According to
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