Taxonomy of Hymenoxys Subgenus Picradenia and a Conspectus of the Subgenera of Hymenoxys (Asteraceae: Helenieae: Tetraneurinae)
NUMBER 4 BIERNER: REVISION OF HYMENOXYS SUBGENUS PICRADENIA 37 TAXONOMY OF HYMENOXYS SUBGENUS PICRADENIA AND A CONSPECTUS OF THE SUBGENERA OF HYMENOXYS (ASTERACEAE: HELENIEAE: TETRANEURINAE) Mark W. Bierner Section of Integrative Biology and Plant Resources Center, The University of Texas, Austin, Texas 78712 Abstract: Hymenoxys (Helenieae, Tetraneurinae) is composed of 28 taxa distributed in 8 subgenera: 1) subg. Hymenoxys, 2) subg. Dugaldia, 3) subg. Macdougalia, 4) subg. Phileozera, 5) subg. Picradenia, 6) subg. Picradeniella, 7) subg. Plummera, and 8) subg. Rydbergia. Eleven taxa comprise Hymenoxys subg. Picradenia. Resumen: Hymenoxys (Helenieae, Tetraneurinae) contiene 28 taxones en 8 subge neros: 1) subg. Hymenoxys, 2) subg. Dugaldia, 3) subg. Macdougalia, 4) subg. Phi leozera, 5) subg. Picradenia, 6) subg. Picradeniella, 7) subg. Plummera, and 8) subg. Rydbergia. El subgenero Hymenoxys subg. Picradenia contiene 11 taxones. Keywords: Asteraceae, Helenieae, Tetraneurinae, Hymenoxys, taxonomy. Hymenoxys Cass. has been consigned to (1994) returned Hymenoxys to tribe Hele various tribes and subtribes of Asteraceae, nieae, placing it with Helenium, Gaillardia, beginning with its placement by Cassini and other genera in subtribe Gaillardiinae. (1828) in tribe Heliantheae (subtribe) He Most recently, Baldwin and Wessa (2000), lenieae (which also included Helenium L. using analyses of nuclear 18S-26S rDNA se and Gaillardia Foug.). Lessing (1832) and quences of the internal transcribed spacer Candolle (1836) placed it in tribe Senecio region, classified Hymenoxys (sensu Bierner, neae (as Senecionideae) subtribe Helenieae. 1994, and this paper) along with Amblyo Bentham (1873) recognized the Helenieae lepis DC., Baileya Harv. & A. Gray, Psilos (as Helenioideae) as a tribe, placing Hyme trophe DC., and Tetraneuris Greene in tribe noxys (along with Helenium and Gaillardia) Helenieae subtribe "Riddelliinae" (an ille in subtribe Euhelenieae.
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