Addenda, corrigenda et emendanda

C.G.G.J. van Steenis c.s.

At times colleagueshave asked me whether my effort to collect the Addenda, Corrigendaet Emendanda was worthwhile.

The main purpose is to keep readers up to date with the of Malesia in onework and keep them aware of additions, name changes, etc.

also for They are important as a source -geographical purposes, to correct names of useful plants, etc.

Another facet of keeping up with the records is that they reflect the degree of completeness of collections at the time of the original revision, and form a certain check on the degree of exploration.

In an overall review of the 'Floristic inventory of the Tropics: Where do we stand?' PRANCE has made use of the Addenda in comparing the state of explorationin the neotropics with that of Africa and Malesia (Ann.

Mo. Bot. Gard. 64, 1977, 657-685, especially p. 671). He found the number of addenda and novelties much larger in the neotropicsthan in Malesia, obviously due to a lower, and especially less varied exploration (collec-

This tends conviction ting density). comparison to support my that the bulk of the Malesian has become gradually represented in the herbarium.

It that the careful record of the Addenda and was pleasant to experience keeping on serves good purposes should therefore be continued.

Printing errors have only been corrected if they might give rise to confusion.

Volume and page number are separated by a colon. Page numbers provided with either a or b denote the left and right columns of a page respectively.

Alismataceae Araliaceae

5: 319, Caldesia parnassifolia (BASSI ex L.) PARL. 9: 39a Osmoxylon lineare (MERR.) PHILIPSON.

In central 320a Malesia a very rare plant (see map in Fl. Substitute in description after 'the

Males. I, 5: 322, fig. 3). In New Guinea it branch 4-5 mm long': bearing an umbel of

was only known from Cape Vogel Peninsu- c. 12 sterile, globose, bacciform flowers, c.

la, but it is now also collected in West New 8 mm 0 when dry.

Guinea: Star Mts, Sibil Valley, 1200-1300 Distr. Add: Negros Occidental, (PAN-

m (KALKMAN4188) and in Papua New Gui- CHO 1845).

nea: Kubor Ra., Nona Minj Divide, 1920 m Ecol. Low bushy shrub along river-

(VINK 16512); Morobe Distr., 15 miles west banks. Flowers orange; fruit dark purple or

of Lae, 150 m (HARTLEY 9778) and near black. Cultivated (recently) as an ornamen-

Mumeng, 950 m (W. Moi 166). In the last- tal for its dark green and shiny foliage.

named collection the inflorescence consists Vern. Miagos

of only one whorl of 3 flowers and a ter- 9: 67 Arthrophyllum stonei Aii-I.an Lim, Mai. minal flower. For. 43 (1980) 263, f. 1; Stone, Fed. Mus.

J. n.s. 26(1) (1981) 71, f. ii.

Anacardiaceae (DING HOU) Distr. Malesia: Malaya: Pahang-Selan-

gor (Stone 12358, 13754, 14140, KLU

8: 483 Spondias pinnata (L. /.) KURZ. 27353, 30006, 30007).

KOSTERMANS (Quart. J. Taiwan Mus. 34, Ecol. Montane forest, 1000-1800 m,

1981, 108-111) suggested that what was sometimes dwarfed in elfin forest on sum- named Spondias pinnata from Malesia mit ridges.

(Malaya toNew Guinea) would differ from Notes. Said to be intermediate between

the typical,S. pinnatafrom India and repre- A. montanum and A. alternifolium, dis-

sent an undescribed species, S. malayana tinct from the latter by the purple fruit, de-

KOSTERMANS. gree of branching of the inflorescence, and

When the he could the thin-coriaceous leaflets. writing paper only re- narrow,

ly on his memory of former experience and A. montanum and A. alternifoliumwere

ononly three specimens from Malaya. I ob- alreadydistinguishedby Philepson(F1. Ma-

served that in one of them the leaf differ- les. I, 9: 55) by weak characters and the ad-

ence mentioned in his key does not fit. His dition of a third one, with intermediate leaf

experience can only relate to the size of characters is therefore hardly welcome; the

trees and bears no testimony on the flower differentiating characters are also slight.

details mentioned in the and their Mr. Lim should have clear for key geo- given a key

graphical variability, which is in this genus the three species instead of this haphazard

difficult tojudgebecause of the cultivation, description.

domestication, and running wild of Prof. Philipson (inlitt.) finds ‘A. stonei’

cultivated trees. For the present his represents rather larger than usual speci-

delimitation does not seem convincing, es- mens of A. alternifolium.

pecially as there are also discrepanciesin his 9: 103 Replace: 16. ELEUTHEROCOCCUS

key and descriptions. MAXIM. M6m. Ac. Sc. St. Petersb. Sav. Etr.

553 554 Flora Malesiana [ser. I, vol. 9 3

9 (1859) 132; S. Y. Hu, J. Arn. Arb. 6 is established. The procedure is established

(1980) 108. Panax subg. Acanthopanax onuninjuredrootlets. Naturally it does not

DECNE& Rev. Hort. PLANCH. IV, 3 (1854) exclude the possibility that infection can al-

105. Acanthopanax (DF.CNE& PLANCH.) so proceed on injured roots or even stem

H. WITTE, Ann. Hort. Bot. 4 (1861) 89; bases of host plants. MIQ. Ann. Mus. Bot. Lugd.-Bat. 1 (1863)

10; PHILIPSON, Fl. Males. I, 9 (1979) 103. Bignoniaceae

Note. UnfortunatelyAcanthopanax has

The Malesian 8: 137FTbancana STEEN. to give way. two species Deplanchea (SCHEFFER)

should be called: Add to Distr.: Central Celebes (Baolu, near

I. Eleutherococcus trifoliatus (L.) S. Y. Palopo; Usu near Malili; near Matana

Hu, J. Arn. Arb. 61 (1980) 110. Acan- lake).

Add to thopanax trifoliatus (L.) Voss. Vilmor. Notes: The Celebes specimens were

Blumengartn. 1 (1894) 406; MERR. Philip. poor and originally assigned to D. glabra

J. Sc. 1 (1906) Suppl. 217; PHILIPSON, Fl. (STEEN.) STEEN. With abundant recent ade-

Males. 9 103. material I, (1979) quate from localities nearby, they

2. Eleutherococcus malayanus (M. R. HEN- appear to belong to D. bancana.

DERSON) STONE, Mai. For. 43 (1980) 395. 8: 138 a Change in Fig. 11, map: the localities from

Acanthopanax malayanus M. R. HENDER- Celebes of Deplanchea glabrabelong to D.

SON, Gard. Bull. S. S. 7 (1933) 105, pi. 22; bancana.

PHILIPSON, Fl. Males. I, 9 (1979) 103. 8: 141 Deplanchea glabra (STEEN.)aSTEEN.

Delete Celebes from the distribution.

Balanophoraceae 8: 148ft 3. Stereospermum colais (HAM. ex DILLW.)

MABBERLEY, Taxon 27 (1978; publ. 1979)

797 7: BalanophoraelongataABL. 553. Bignoniacolais HAM.ex DILLW. Re-

Add to literature: B. C. STONE, Mai. Nat. view of the references to the Hortus Mala-

RHEE- J. 33 (1979) 129, fig.; Fed. Mus. J. n.s. 26 baricus etc. (1839) 28, based onpadri

72. Malab. (1) (1981) B. papuana (non Schltr) DE, Hort. 6, t. 26. S. personatum

Soepadmo, Nature Malaysiana 3 (1) (1978) (HASSK.) CHATTERJEE; STEEN. Fl. Males. I,

24, with col. illus. — B. hansenii HAMBALI, 8 (1977) 148.

Reinwardtia 9 (1980) 425. Nomencl.: On the strength of HAMIL-

7: 798 a Add to Distr.: Malaya: Selangor/Pahang TON'S opinionthat this was different from

G. Ulu in dwarf forest chelonoides L. border, Kali, on Bignonia /., DILLWUN pro-

ridge at 1500-1700 m, on Pentaphylax posed a new name, which antedates that of

auryoides. HASSKARL.

Add to Note: According to Dr. B. HANSEN 8: 159a,Fernandoa macroloba (MIQ.) STEEN.

(in litt.) the new record from Malaya be- 153ft The mention of this N. Sumatran endemic

longs undoubtedly to B. elongata to which tree (as ‘Heterophragma macrolobium’=

it keys out without difficulty in the key in Haplophragma macrolobum (MIQ.)

Fl. Males. I, 7: 793. STEEN.) by FLENLEY & RICHARDS (eds.),

7: 802i> Balanophora abbreviata BL. The Krakatao Centenary Expedition, in

Add to Distr.: New Hebrides: Mallicolo I. Dept. Geogr., Univ. Hull, Misc. ser. 25

(N. HALLE RSNH 6351), parasitic onFicus. (1982) 16, 27,48, 166 is derived from a mis-

Cf. Halle, Adansonia 17 (1978) 260. identification; the specimens belong to the

Add to Note: Mr. G.D. Arekal & G.R. common Radermachera glandulosa (BL.) Shivamurthy (Phytomorph. 26, 1976, MIQ.

135-138, 6 fig.) have, for the first time, 8: 160, These two pages have unfortunately be-

been able to examine, in situ - since all for- -161 come transposed.

mer experimental infections have failed - 8: 177 Pandorea pandoranaa(ANDR.) STEEN.

the germination of the seed of B. ab- Add to Distr.: Central Celebes (VAN BAL-

achievement with and breviata, a magnificent GOOY 3915), change Fig. 38, map, ac- these utterly minute seeds. Afterthe break- cordingly.

ing of the seed coat the endosperm cells

produce 4-8 narrow tubular extensions Burmanniaceae

coated with a sticky substance adhering to the fine rootlets for anchorage. Then the 4: 20, Gymnosiphon BL.

hardly differentiated embryo produces 1-4 593 a Add to Distr.: Lesser Sunda Islands: Flores

tubular processes to form the primary haus- (SCHMUTZ 4802, prob. G. aphyllus BL.).

torium, penetrating into the rootlet and The genus is not yet reported from the Mo-

making contact with the vascular tissue of luccas and the Philippines.

the rootlet of the host; a nodule develops, 4: 21 Thismia GRIFFITH.

throwing off the seed coat, and parasitism Add the synonym: i Geomitra BECC. Malesia 1982] Addenda, corrigenda et emendanda 555

1 (1877) 250. Distr. Malesia: Malay Peninsula (?Pe-

Add Note. 15. Thismia clavigera seems to rak, G. Hiau above Taiping; Langkawi Is.),

be better accommodated in Thismia sect. N. Sumatra (Gajo Lands), Borneo (Sara-

Sarcosiphon (BL.) JONKER, from the three wak: Lundu Distr.).

species ofwhich it is distinct by the free cla- 4: 25 Delete 4. Geomitra etc.

vate inner tepals.

4: 22 Insert in the key after the second lead of 1: (LEENHOUTS)

la. Innerperianth lobes connate at the tips,

5: formingan erect mitre with large holes. 222 Dacryodes rugosaA(BL.) H. J. LAM.

Continue to 12. Add to Distr.: Nicobar Is., Katchal I. (var.

la. Inner perianth lobes connate at the tips, rugosa).

forming a mitre with large holes, the 5: 228b Dacryodes longifolia (King) H. J. Lam.

lobes continued above the mitre as 3 Add to literature: Kochummen in Whit-

free, almost erect, filiform, clavate seg- more, Tree PI. Malaya 1 (1972) 141.

ments 15. T. clavigera Add to description: Tree up to 40 m with

4: 25 Insert after 14. Thismia crocea: buttresses up to 3.5 m (cf. KOCHUMMEN, 15. Thismia clavigera (BECC.) F.V.M. Vict. 1972).

Nat. (1890) 235; Pap. & Proc. R. Soc. Add to Distr.: Throughout the Malay Pen-

Tasm. for 1890 (1891) 235; STONE, Blumea insula (Kochummen, 1972); var. longifolia 26 (1980) 420, pi. Geomitra clavigera also in Borneo.

BECC. Malesia 1 (1877) 250, t. 10, f. 1; Jon- Insert: Ecol. Usually on coastal hills or

ker, Fl. Males. I, 4 (1948) 25. Fig. 1. hills which were coastal in the Pleistocene

300 (Kochummen, 1972), up to m. Fr. April.

5: 232 Santiriaa grandiflora Kalkman.

Add to description: Branchlets up to 1.75

cm thick. Leaflets finally glabrous beneath. Fruits nearly transversely obovate, 2 by

1.25 by 1.5 cm, stigma more than 90° ex-

centric.

Add to Distr.: N. Borneo (Sipitang Distr.,

Mendalong, Lumaku For. Res.).

5: 251 Add in the key to the species of ,

after the second lead of 6:

6b. Stipules up to 1.5, rarely 2.5 cm long,

not distinctly exceeding the terminal

bud. Branchlets solid. Stamens 6.

7a. Branchlets 1.5-3 cm 0; pith thick and

soft.

7A. Petiole terete with the stipules in-

serted at the base. Fruit glabrous

56. C. batjanense

7A. Petiole sharp-angular with the sti-

pules inserted on the angles up to 5

cm from the base. Fruit prickly-

hairy 43. C. hirsutum

5: 275 Canarium grandifoliuma(Ridl.) H. J. Lam.

Add to Distr.: According to Kochummen

(in Whitmore, Tree Fl. Malaya 1, 1972,

129) also in Borneo (Brunei).

5: 2766 Canarium decumanum GAERTN.

Add to description: Leaflets of the upper

pairs sometimes on the basiscopic side de-

current till the rachis. Fruits up to 12 cm

long.

Add to Distr.: Admiralty Is. (Manus I.).

Add to Ecol.: sago swamp.

5: 283b Canarium fusco-calycinum RIDL.

Add Tree 21 to description: up to m high

and 60 cm 0. Infructescences c. 15-30 cm

Fig. 1. Thismia clavigera (BECC.) F.V.M., x2 (after long with few short oblique-erectbranches

BECCARI). and with few fruits; calyx funnel-shaped, 3 556 Flora Malesiana [ser. I, vol. 9

trigonous, c. 1 cm diam. Fruits (immature) and W. marginata.

narrowly oblong,taperingto both ends, tri- Distr. Malesia: Papua New Guinea:

in 3.5 1.25 bank from Iswan gonous section, by cm, glabrous; along leading swamp to in section Koma Creek Fl the kernel with flat sides and (Mt Victoria), c. 2650 m.

slightly rounded angles, the lids with a very fr. May.

faint median rib, lids c. 1.5 mm thick. Seeds Note. According to the author closely

2, the sterile cell moderately reduced. related to the glabrous New Zealand species

Add to Ecol.: On clay, up to 340 m. Fl. W. albomarginata HOOK./.

Aug. 6: 122, P. VAN ROYEN has provided a new key to

5: 290 Canariuma rigidum (Bl.) Miq. 928 the New Guinea species of Lobelia. Cf.

Add to Distr.: Moluccas (Obi I.). Bot. J. Linn. Soc. 77 (1978) 120.

5: 2906, Canarium cestracion Leenh. 6: 136, Add to the species of Lobelia:

291 a Add to description: Twigs up to 11 mm 0. 928 Lobelia victoriensis v. Royen, Bot. J.

Stipules linear, taperingto an acute point, Linn. Soc. 77 (1978) 118, f. 1.

1 4 by mm. Leaves up to 8-jugate; leaflets Stemless dwarf, 7-20 mm high, with a from 2.5 broad cm on; acumen sometimes single central flower in a rosette of ovate

long and slender; nerves up to 17 pairs. leaves. Corolla isomerous, without a dorsal

Fruits up to 3cm high and 1.75 cm broad. slit, lobes 5, equal, filiform, long.

Add to Distr.: Milne Bay Distr. Distr. Malesia: Papua New Guinea: Is-

5: 55. 296, Insert after Canarium pimela: wan swamp on Mt Victoria, 2660 m alt. Fl.

6: 928 56. Canarium batjanense Leenh. Blumea Jr. May. 27(1981)211.

Distr. Malesia: Moluccas: Batjan I. (Mt Caprifoliaceae

Sibela).

Ecol. Canopy or subcanopy tree in pri- 4: 175 Add to the family description: Leaves very

mary forest, 250-1050 m. Fl. fr. Oct. rarely scattered or in pseudowhorls (sub-

Note. The present species seems to be fam. Alseuosmoideae). allied with C. acutifolium MERR., C. balsa- 4: 176 Insert in the key after the second lead of 1:

miferum WILLD., and C. oleosum Engl., la. Leaves scattered or in pseudowhorls

which is also well in accordance with its S. Periomphale

geographicalposition (cf. Leenhouts, Blu- la. Leaves decussate. Continue to 2.

mea 9, 1959, 317, f. 9). It seems nearest to 4: 191 Sambucus javanica BL.a

C. balsamiferum, which differs by the ab- Add to literature: STEEN. Blumea 24 (1978)

of sence stipules, the far stronger reduced 479.

pistil in the male flower, and the fruit cells 4: 192 a Add to Distr.: West New Guinea: Arfak

BW that are all three well developed. The only Mts, Mt Lensemoi, 1850 m, 12657;

moderately reduced pistillode is well in ac- Wissel Lake region, EYMA 4618; Papua

cordance with C. oleosum, different in New Guinea: Morobe Distr., Sattelberg,

both other points mentioned under C. bal- Timbe R., 1500 m, CLEMENS 7793.

and in the Add fill of samiferum, however, moreover to Notes: These records the gap

much smaller, nearly ovoid fruits. Vegeta- the generic range in East Malesia, two spe-

tively, C. acutifolium and C. batjanenseare cies being recorded from Australia, which

hardly separable but for the longer and are discussed in VAN STEENIS, I.E.

more persistent stipules of the former; C. 4: 194 Insert after 4. Carlemannia:

acutifolium differs, however, in the strong- 5. PERIOMPHALE

ly reduced pistillode and in the smaller and BAILL.BUII. Mens. Soc. Linn. Paris 1 (1888)

more globular fruits. 731; STEEN. Blumea 24 (1978) 480. Pa-

7: 822 Dacryodes multijugaaLEENH. Add: chydiscus GILC, & SCHLTR, Bot. Jahrb. 39

Distr. South Sumatra (FORBES3O73, L), (1906) 270. Memecylanthes GILG &

fruits only. Hitherto only known from one SCHLTR, I.e. 269.

collection in Malaya. Shrublets. Leaves scattered or in pseudo-

whorls, entire or with a few faint teeth to-

Campanulaceae wards the apex. Flowers actinomorphic,

5-merous, fascicled or solitary axillary. 6: 118 Insert after 4. Wahlenbergia marginata: Ovary inferior, 2-celled, with rather few

5. Wahlenbergia papuana v. ROYEN, Bot. (1 -4) ovules on the septum. Corolla gamo-

J. Linn. Soc. 77 (1978) 121, f. 2. phyllous, somewhat barrel-shaped. Sta-

Branched dwarf herb, prostrate. Twigs, mens 5, free.

leaves and calyx hairy. Leaves at end of Distr. New Caledonia (4 spp.), Papua

twiglets, narrow-oblanceolate, 8-10 mm New Guinea (1 sp.).

long. Filaments glabrous, not widened at Notes. This genus belongs to the sub- base, thus different from both W. confusa family Alseuosmoideae, best known from 1982] Addenda, corrigenda et emendanda 557

the small Alseuosmia A. CUNN. from genus Chenopodiaceae

New Zealand.

There is no unanimity of opinion about 4: 104 Arthrocnemum MOQ.

its taxonomic place. Sometimes it is treated After a long and careful study of Australian

as a separate family, but mostly it is assign- Salicornieae P. G. WILSON has concluded

ed an isolated place in Caprifoliaceae, to that the concept of the genus Arthrocne-

I New which agree. This is another Caledo- mum is polymorphous, and that the Male-

nian turned in New Guinea. sian attributed it genus now up species to (A. indicum) A third, yet undescribed monotypic genus should belong to a new genus, Halosarcia,

of the subfamily occurs in Queensland(van differing from Arthrocnemum in sclereids

Steenis, I.e.). and the adaxial stamen which he finds of fundamental importance; he suggested that

1. the Periomphalepapuana Steen. Blumea 24 two genera are not closely related.

(1978)481. Within Halosarcia indica WILSON distin-

75 Glabrous, epiphytic shrublet, c. cm; guishes four subspecies, two of which are

stem and branches very slender. Leaves endemic to Australia and two others occur

scattered and in pseudowhorls, lanceola- also outside Australia, the type ssp. indica

te-oblong, 15-25 by 6-8 mm, cuneate at also in India and Tanzania, and a new sub-

base, acuteat apex, margin entire or mostly species in South Malesia.

1 in with or 2 short gland-tippedteeth; nerves As ssp. indica might also be found Ma-

extract WILSON'S 2-3 pairs; very erect; venation impressed lesia, I key.

above. Petiole 3-4 mm. Flowers solitary;

Halosarcia pedicels pink, 1-2 mm. Calyx lobes 5, indica (WILLD.) P. G. WILSON,

0.5 — thickish, blunt-deltoid, 1.25 by mm. Nuytsia 3 (1980) 63. Arthrocnemum in-

Corolla narrowly barrel-shaped, 6 mm dicum (WnxD.) Moq.: Backer, Fl. Males.

long, pinkish light green; lobes 1 mm, ca- I, 4 (1949) 104.

al- runculate inside apex. Stamens 5, free,

ternipetalous. Ovules 4-5 in each cell, at- KEY TO THE SUBSPECIES

tached to the septum, flattish. Style colum-

nar, as long as the corolla; stigma globular, 1. Plant decumbent or prostrate. Articles

rugose. of branches corky with age, ± trun-

Distr. Malesia: Papua New Guinea: cate, entire ssp. indica

West Sepik Distr., Telefomin Subdistr., 1. Plant ± erect. Cortical tissue of articles

3000 ± ci- m alt., in Podocarpus-Phyllocladus shrivelling with age, often lobed,

woodland with Gahnia tussocks dominat- liolate ssp. leiostachya ing the undergrowth (LAE 670687).

ssp. leiostachva (BTH.) P. G. WILSON,

Celastraceae (DING HOU) Nuytsia 3 (1980) 66. — Arthrocnemum ci-

liolatum BUNGE ex UNG.-STERNB. Versuch

6: 233 Celastrus hindsii BTH. Cf. STONE, Mai. Syst. Salicorneen 69 (1866); UNG.-STERNB. For. 43 (1980) 244. D. Atti Congr. Int. Bot. Firenze 1874

This species was not treated in the paper, (1876) 283, based on specimens from Java

but its occurrence in Malaya is mentioned and Lesser Sunda Is.; A. J. SCOTT, Bot. J.

twice in the summary. The specimen in Linn. Soc. 75 (1977) 370. — Salicornia leio-

— Ar- question (STONE 14039) was misidentified stachya BTH. Fl. Austr. 5 (1870) 203.

and the C. belongs to common monosper- throcnemum leiostachya (BTH.) PAULSEN, moides LOES. Dansk Bot. Ark. 2 (8) (1918) 61.

6: 243 Xylonymus KALKMAN. Distr. Australia, South Malesia: north-

Change in description: Flowers 4-5-mer- coast of Java, incl. Madura and Kangean

ous. Is., Lesser Sunda Is. (Sumba, Sumbawa,

6: 245 Xylonymus versteeghii KALKMAN. Timor).

Add to Distr.: West New Guinea: Dalman,

Nabire, KANEHIRA & HATUSIMA 12241; (Leenhouts) Darmi Distr., BW 9317. Moluccas: Obi 1.,

5: DE VOCEL 4347. 495 Anatomy. W. C. Dickison (J. Elisha

6: 420 Salacia kalahiensis KORTH. Mitchell Sc. Soc. 87, 1971,77-86; ibid. 88,

Hitherto known from the Philippines, Bor- 1972, 120-136; ibid. 89, 1973, 121-138)

neo and Java. Now also from the Lesser concluded that Connaraceae are distinctly

His Sunda Is.: W. Flores (Paku, 400 m, allied with Leguminosae and Rosaceae.

SCHMUTZ4S23; vern.: wase mantur). conclusions regarding relationships within

the family deserve a more critical attitude,

however, as at that time the author was in- 558 Flora Malesiana [ser. I, vol. 9 3

sufficiently aware of the difficulties of in- TRIMEN Handb. FL. Ceyl. 3 (1895) 221; terpreting , especially regarding AUSTIN, POWELL & NICOLSON, Brittonia 30

synonymy; he was too much inclined to (1978) 196. — Adamboe RHEEDE, Hort.

of herbarium Mai. 11 take identifications speci- (1692) 115, t. 56 (lectotype). — I.

mens for granted. illustris (CLARKE) PRAIN, Beng. PI. 2 (1903)

Palynology. W. C. DICKISON, Pollen et 735; OOSTSTR. FL. Males. I, 4 (1953) 485.

Spores 21 (1979) 31-79. In this extensive Note. Apparently Adamboe RHEEDE

survey of the pollen of the Connaraceae, does not belong to Stictocardia - as VAN

the author avoided the mistakes made in his OOSTSTROOM hesitantly assumed - and be-

earlier anatomical papers. This makes his comes the type through the new typifica-

The conclusions of greater importance. tax- tion. The name change is a nuisance, but

onomic opinions brought forward in the has to be accepted.

Flora Malesiana revision ofthis family are 4: 4856 Change 37. Ipomoeacrassicaulis etc. as fol-

to a high degreeconfirmed. lows:

5; 504a MERR. Agelaea trinervis (LLANOS) 37. Ipomoea carnea J acq. ssp. fistulosa Add Distr.: to Hainan (c/. ANON., Fl. Hai- (Mart, ex Choisy, in DC.) D. Austin,

nan 3, 1974, 113). Taxon26(1977)237. — I. fistulosa MART.

5: 507ft Roureopsis asplenifolia SCHELLENB. ex CHOISY in DC. Prod. 9 (Jan. 1845) 349.

Add to Distr.: Borneo. — Batatas crassicaulis BTH. Voy. Sulph. 5

5: 520a Rourea prainiana Talbot. (June 1845) 134. — I. crassicaulis (BTH.) B.

Add to Distr.: Northern Thailand (cf. Vi- L. ROBINSON, Proc. Am. Ac. Sc. 51 (1916)

dai, Fl. Thailand 2, 1972, 124). 530; OOSTSTR. Fl. Males. I, 4 (1953) 485. 5: 5246 Ellipanthus beccarii PIERRE. Note. AUSTIN has convincingly shown

Add to Distr.: Sumatra (var. beccarii). that the differences between I. carneaand I.

5: 526 Correct the key to the species of Connarus fistulosa do not merit specific distinction.

as follows: As the two forms, which are distinct by mi-

second lead of 18: endocarp sparse- nor vegetative characters, grow in South

ly to densely non-glandularpubescent, America largely geographicallyisolated (al-

couplet 19: delete 'Sepals blunt' versus 'Se- lopatric), a subspecific rank (as a race)

pals acute'. seems the best disposition.

5: 528 Insert after 2. Connarus euphlebius: 4: 487a Change 40. Ipomoea tuba etc. as follows:

2a. Connarus impressinervis STONE, Mai. 40. Ipomoea macrantha R. & S. Syst. Veg.

For. 43 (1980) 255, fig. 4 (1819)451; POWELL, NICOLSON& AUSTIN,

Differs from C. euphlebiusMerr. as fol- Brittonia 30 (1978) 201. — I. tuba

lows: Branches and leaves glabrous. Petio- (SCHLECHTEND.) G. DON, Gen. Syst. 4

4 lules 0.8 cm. Nerves 5-6(-8) pairs. Petals (1838) 271; OOSTSTR. Fl. Males. I, (1953)

(under the fruit) outside appressed short- 487.

hairy. Beak of fruit faint, lateral, slightly Note. An unfortunate but unavoidable

below the apex; pericarp outside glabrous, name change of a pantropical species.

inside with a few scattered hairs.

Distr. Malesia: Borneo (Sarawak; S Crassulaceae (H. OHBA, Tokyo)

14731, S 18903, S 24232, S 24715, S 28417).

Note. in 1962 studied the Already I type 4: 197 Insert the key and replace the species de- L. material onwhich this new species is based scription under the genus Sedum by the

and found that it is allied to C. euphlebius, following new treatment:

but differed in several points and seemed to

represent a new species. I pointed out the KEY TO THE SPECIES

differences cited above (in sched.). Because

floral characters are important in the 1. Sepals free.

genus, I refrained from describing it, all 2. Flowers 4-merous, sepals narrowly ob-

material hitherto known being in fruit. long-spathulate. Styles very short (less

5: 5386 L. ssp. malayensis than 0.3 mm). Flowering stems simple.

LEENH. Cauline leaves spathulate

Add Distr.: Thailand 1. S. australe to SE. (cf. VlDAL, Fl. erythrospermum ssp. Thailand 2, 1972, 129). 2. Flowers 5-merous, sepals spathulate(to

oblong). Styles long (0.9-1.2 mm).

Convolvulaceae Flowering stems tri- (rarely bi-)furcate.

Cauline leaves spathulate to broadly

4: 485a Change 36. Ipomoea illustris etc. as fol- obovate 2. S. formosanum

lows: 1. Sepals connate for c. 0.5 mm. Flowers

36. Ipomoea campanulata LINNE, Sp. PI. 5-merous, sepals linear-lanceolate to

(1753) 160; MOON, Cat. PI. Ceyl. (1824) 14; very narrowly oblong. Style long (c. 1.2 1982] Addenda, corrigenda et emendanda 559

short mm). Flowering stems with a ster- than the petals, erect at anthesis; filaments

ile shoot or simple. Cauline leaves oblan- filiform, 3-4 mm, oppositipetalousonesc.

1 ceolate to narrowly obovate mm from the base connate with the petal.

3. S. Nectar-scales 0.8 parvisepalum ssp. philippinense flat, narrowly oblong, by 0.3 3-4 mm. Gynoecium mm long, the

ovaries for 1. Scdiim erythrospermum Hayata, Ic. PI. basally c. 0.5 mm connate, ven- Formos. 3 (1913) 110. trally gibbose, dorsally round, suberect, 1.5

ssp. australe Merr. H. Ohba, J. Jap. Bot. mm wide at the middle,style very short, less

52 (1977) 322. — S. australe MERR. GOV. than 0.3 mm. Ovules c. 20 in each locule.

Lab. Publ. Philip. 29 (1905) 16, non ROSE Distr. Malesia: Philippines (Luzon,

(1903); Philip. J. Sc. 5 (1910) Bot. 350, Benguet Prov.: Mt Santo Tomas, Elmer

= En. 2 — s. 6568 PNH The p.p.; Philip. (1923) 217, p.p. 114365). ssp. erythrosper- R. T. Formosa. ambiflorum Clausen, Cact. Succ. J. mum occurs in

18 (1946) 58; Backer, Fl. Males. 1,4 (1951) E c o 1. On ledgesand gravelly hillside near

— 2000-2200 197, versim., p.p. Fig. 2a-d. the summit of mountain, m.

Note. Ssp. erythrospermum differs

5-merous from ssp. australe in having flo- lin- wers, narrowly oblong-oblanceolateor

ear sepals, subulate or narrowly lanceolate

petals, and usually trifurcate flowering

stems.

2. Sedum formosanum N. E. BROWN,

Gard. Chron. n.s. 24 (1885) 134; FORBES& HEMSLEY, J. Linn. Soc. 26 (1888) 285; HA-

YATA, Ic. PI. Formos. 2 (1912) 12; PRAE-

GER, Not. R. Bot. Gard. Edinb. 13 (1921)

83; J. R. Hort. Soc. 46 (1921) 295; R.-HA-

MET, Candollea4 (1929) 32; BERGERUI E. & Fig. 2. Sedum erythrospermum HAYATA ssp. austrate P. Nat. Pfl. Fam. ed. 2, 18a (1930) 460;

H. OHBA. a. Petal with stamen, b. sepal, c. nectar FROD. Act. Hort. Gothob. 6 (1931) app. 97,

scale, d. ovaries. — S. parvisepalum YAMAMOTO ssp. f. 778-784, t. 62; OHWI, Fl. Jap. Engl. ed. philippinenseH. OHBA. e. Petal with stamen, f. calyx (1965) 497; HATUS. Mem. Fac. Agr. Kago-

lobe, g. nectar scale, h. ovaries. All x 5 (a-d ELMER shimaUniv. 5 (1966) 31; Fl. Ryukyus (1971)

6568, e-h MUNI 5644). 301; MORANin Walker, Fl. Okinawa (1976)

508; H. OHBA, J. Jap. Bot. 52 (1977) 322.

S. mariae R.-Hamet in Fedde, Rep. 8

Succulent, glabrous, perennial herb. (1910) 143.

Roots thin. Flowering stems annual, 5-10 Succulent, glabrous, perennial herb, up

cm long, 1.5-2 mm 0, simple, erect from to 25 cm high. Flowering stems erect from

nearly decumbent base, smooth. Leaves al- a creeping or procumbent base, once or

ternate (rarely opposite or ternate), remote- twice tri- (rarely bi-)furcate,terete, smooth.

ly arranged, sessile, shortly spurred (the Leaves alternate, remotely arranged, ses-

with round-truncate 0.5 less than spur a apex, c. mm sile, very shortly spurred (the spur 1 long),entire, spathulate, apex round or ob- mm), entire, spathulate to broadly obo-

tuse, base long attenuate, 7—18(—27) by vate, apex round, base long-attenuate,

2-5(-ll) mm, thick-herbaceous, flat, 1.5-3.5 by 0.6-1.6 cm, fleshy, bright green

smooth, spreading or ascending, fnflores- above, paler below, smooth. Inflorescences

cences terminal, a ternate bracteate cyme. terminal, a ternate, bracteate, loosely

Bracts leafy, spathulate to obovate. many-flowered cyme. Bracts leafy. Flowers

Flowers mostly 4-merous, erect, sessile. 5-merous, sessile, 6-9 mm at anthesis. Sep-

Sepals free, narrowly oblong-spathulate, als free, very shortly spurred (c. 0.3 mm),

base round-truncate spurred (the spur with entire, spathulate or rarely oblong, apex

apex, 0.2 mm long), apex round to nearly round, somewhat broadening towards the

truncate, somewhat broadening towards base, nearly equal in size, 2.8-3.7 by

the base, entire, somewhat unequal in size, 0.8-1.3 mm, fleshy, ascending at anthesis.

for 2-4 by 0.8-1 mm, more or less fleshy, Petals bright yellow, basally connate c.

0.5 green, smooth, suberect throughout, per- mm, lanceolate to oblong-lanceolate,

sistent at fructescence. Petals bright yellow, acute, 5.2-6.2 by 1.3-1.6 mm, nearly erect

connate c. 0.3 mm from the base, lanceo- or ascending at anthesis. Stamens 10, shor-

4 late, acuminate, c. mm by 1.2 mm, widely ter than the petal, nearly erect at anthesis;

spreading at anthesis. Stamens 8, shorter filaments filiform, c. 4.5 mm, the oppositi- 560 Flora Malesiana [ser. I, vol. 93

for 0.8 connate with 1.2 petalous ones c. mm apex from the middle, style c. mm, slen-

the petal, anthers oblong, c. 0.4 mm long, der, stigma not papillate. Ovules (20-) 24 deep yellow before dehiscence. Nectar- (-34) in each locule. Follicles brownish,

4-5 scales broadly oblong-obovate to broadly by c. 2.5 mm, carpels widely spreading.

oblong, c. 0.5 by c. 0.4 mm, creamy white, Seeds brownish red, oblong-cylindrical, 0.7

flattish. Gynoecium 5.6-6.2 mm long, the mm long, apex round, testa very minutely

ovaries for c. 1.5 mm connate, ventrally puncticulate.

gibbose just above the ventral connection, Distr. Malesia: Philippines (Luzon,

dorsally round, erect throughout, c. 1.6 Benguet Prov.: BS 4279, 4451, 5373, 5644,

mm wide at the middle part, abruptly nar- Merrill 4861, van Steenis 17950, Wil-

rowing near the apex; style 0.9-1.2 mm, liams 117). The ssp. parvisepalum occurs in

slender, tapering towards the apex; stigma Formosa.

lowly papillate, yellowish. Ovules 20-24 in Ecol. On rocks or boulders along steep

each locule. Follicles 6-7 mm long, whit- slopes, c. 300-1700 m.

ish, obovate, carpels not spreading. Seeds Note. Ssp. philippinense is distinguish-

0.7 ed from the Formosan oblong, c. mm long, testa brown, very ssp. parvisepalum

minutely puncticulate throughout. by the narrowly obovate to oblanceolate

i and shorter and D s t r. Formosa Japan(Ryukyu and leaves, flowering stems, longer Kyushu), in Malesia: N. Philippines (Batan petals. Ssp. parvisepalum has linear-oblan-

I.: Hatusima& Sato28624). ceolate leaves, flowering stems up to 25 cm

Ecol. On rocks at the shore, rarely in- long, and petals attaining 7.5 mm long.

land. Fl. June-August. Ssp. philippinensediffers from the Philip-

S. australe the pine erythrospermum ssp. by 3. Sedum parvisepalum Yamamoto, Suppl. 5-merous flowers, the ovaries with long

Ic. PI. Formos. 2 (1926) 22, f. 14. style, the broadly connate, linear-lanceo-

ssp. philippinen.se H. OHBA, J. Jap. Bot. 52 late sepals, and the narrowly obovate to

(1977) 323. — S. australe (non MERR.) oblanceolate cauline leaves. STEEN. Bull. Jard. Bot. Btzg III, 13 (1934)

195.— S. ambiflorum(non R. T. CLAUSEN) Cyperaceae

4 BACKER, Fl. Males. I, (1951) 197, p.p.-

Fig. 2e-h. 7: 468 Add to Fig. 10, map of Mapania: Lesser

Succulent, glabrous, perennial herb, up Sunda Islands: Flores 4-

-- ■ . .- - - to 15 cm high. Flowering stems ascending 7:471a Mapania macrocephala. (GAUDICH.) K.

or erect from creeping, sometimes branch- SCH.

ed base, with a short sterile shoot or simple, Add to Distr.: Lesser Sunda Is. (W. Flores:

terete, smooth. Leaves alternate, remotely Paku, Wae Meleng, 700 m, damp stream-

arranged, sessile, very shortly spurred (c. bed in rain-forest, SCHMUTZ 4913).

0.5 mm), entire, narrowly obovate to ob- Add to Notes: The species was known from

lanceolate, apex obtuse, base attenuate, the S. Philippines and N.-Central Moluc-

0.9-1.7 by 4-6 mm, thick-herbaceous, cas eastwards; Flores is an importantexten-

smooth. Inflorescences a terminal, ternate sion westwards. The nuts are on the small

cyme, densely 30-50-flowered, sparsely side, some 4-4.5 by 3 mm and pearl grey.

bracteate. Bracts leafy. Flowers 5-merous, 7: 474a Mapania cuspidata (MIQ.) UITTIEN.

sessile, c. 10 mm wide at anthesis. Sepals Add to Distr.: Lesser Sunda Is. (W. Flores:

1 basally for c. 0.5 mm connate, linear-lan- Paku, 500 m, plant m, locally gregarious,

ceolate in to very narrowly oblong, entire, rain-forest, SCHMUTZ 4816).

in 2.5-3.5 0.5-0.7 Add Notes: The widest distributed slightly unequal size, by to spe-

mm, apex round, slightly broadening to- cies ofthe genus, of which KERN(F1. Males.

wards the base, base spurless, fleshy, I, 7, 1974, 466) expressly stated that it lack-

ed in Sunda the ascending or spreading through anthesis. the Lesser Islands, gap now

Petals bright yellow, basally for c. 0.5 mm being filled.

acumi- 522a chinensis KERN. connate, lanceolate, apex acute to 7: Lipocarpha (OSB.)

nate-acute, 5-6.5 by 0.8-1 mm, spreading Add to Distr.: Lesser Sunda Is. (Flores:

at anthesis. Stamens shorter than the VELDKAMP 7102).

petals, filaments 3.5-3.7 mm, the oppositi- 7: 567a Fimbristylis eragrostis (NEES) HANCE.

petalous onesfor c. 1 mm connate with the Add to literature: VELDKAMP, Reinwardtia

petals, anthers ovate, c. 0.6 mm long, red- 10 (1982) 26.

dish before dehiscence. Nectar-scales Add to Distr.: Australia: Northern Terri-

P. broadly oblong to square, c. 0.5 by 0.3 mm. tory (Arnhem Land: K. LATZ2836).

Gynoecium 4-5 mm long, the ovaries for 5: 567b Fimbristylis fusca (NEES) CLARKE.

0.7 mm connate, ventrally gibbose, dorsally Add to literature: VELDKAMP, Reinwardtia

1 round, c. mm wide, tapering towards the 10 (1982) 26. 1982] Addenda, corrigenda et emendanda 561

Add to Distr.: Northern Terri- in several Australia, bryony occurs species (see p. 263, tory (Katherine Gorge: DUNLOP 3733). A 270).

new record for Australia. 9: 257 Line 13 from top: change SASAHUUO SASA-

7: 6056 Cyperus esculentus LINNE. KI.

Add to literature: EVERAARTS, Weeds Vege- Paragraph 'Morphology', line 2, first tables Java (1981) 79. KERN knew this wide- word: change cylindrical into circular.

ly distributed species only from a single 9: 267 Paragraph 3, line 4, add after 'species lev-

'suspected' collection. This suspicion is el.': Dipterocarpoideae differ from other

now removed by Mr. A. P. EVERAARTS in subfamilies in the presence of resin canals

his study on weeds of Java. He collected and in their multiseriate rays.

this species in fields (with vegetables) near 9: 269 Paragraph 2, last line, add between the

Lembang and Pengalengan in West Java brackets: SOMEGO, 1978.

and on Mt Tengger (Tosari, Ngadisari) in 9: 272 Line 16 from top: change ‘hypochroa’into East Java at 1300-1400 m. hypochra.

7: 640 Cyperus compactus aRETZ. Line 15 from bottom: change DC. into

Add to literature: K. L. WILSON, Telopea 1 KURZ).

(1980) 462. Line 4 from bottom: change DYER into

Add to Distr.: Australia: Northern Terri- KING.

tory. 9: 273 Line 4 from top: change MIQ. into BL. 7: 686 Oreobolus kükenthaliiASTEEN. Line 7 from bottom: omit ‘Balanocarpus’.

Add to literature: STEEN. Reinwardtia 10 9: 276 Line 5 from bottom: change'flora' into flo- (1982) 26. ral.

7: 687 a Add to Distr.: NW. Borneo: Sarawak (4th 9: 279 Paragraph 2, lines 8& 9, change sentence

Div., N. side ofMt Murud), and dot this lo- into: The mature fruit of Pakaraimaea is

cality in Fig. 85. small, capsular, but the method of germi-

Add to Ecol.: Forming dense tufts in wet nation is unknown.

rock holes, 2100 m, BURTT& MARTIN 5482. 9: 284 Substitute paragraph 5 by: A picture thus

Add Note: Hitherto only known from N. emerges of the Asiatic subfamily originat-

Sumatra and Malaya; Mt Kinabalu is the ing in Central Gondwanaland and migrat-

westernmost locality of another species, O. ing eastwards. It appears likely that the

ambiguus KUK & STEEN. family originated in the seasonal tropics;

9: 149 Carex breviscapaaCLARKE. subsequent immigration and rapid diversi-

Add to Distr.: Central Celebes (Mt Lokila- fication in the oceanic climates of S.W.

laki: W. MEIJER 9876). Ceylon, West Malesia and New Guinea be-

Add to Ecol.: 1700-2200 m alt. ing accompanied by a reduction in stamen

9: 164 Curex oedorrhamphaaNELMES. number and pollen production, and tomen-

Add to Distr.: Central Celebes (Mt Lokila- tum; and the evolution of a thin-walled and

laki: W. MEUER 9869). poorly dispersed fruit.

9: 290 Line 6 from bottom: change 'sort' into (ASHTON) short.

9: 293 Line 5 from top: replace ’D. apterus’ by D.

16 9: 239 Line from top: Cotylelobium has 5 spp., validus.

not 6. Paragraph 5, line 3: change 'but' into by.

" 9: 242 Paragraph 5: replace 2nd and 3rd sentence 9: 304a Dipterocarpus baudii KORTH.

It by: is not impossible that they were de- Line 11 from top: change ‘D. elongatus

rived from the Indian subcontinent, as ac- KORTH.' by D. validus BL.

cording to geophysical theory this rafted 9: 310 DipterocarpusAkunstleri KING.

block ofland had by the Eocene joinedthe Line 16 from top: change ‘D. elongatus

Asian plate, while, moreover, Dipterocar- KORTH.' by D. validus BL.

pus could have migratedthrough S.W. Asia 9: 339 aln caption Fig. 35 replace 'place' by tree,

before it became arid in the Miocene. and add: The collector MUJAH indicates

9: 244 Paragraph 2, lines 10-12: ssp. philippinen- scale.

sis also in S.E. Borneo; the doubtful sterile 9: 341 Cotylelobium PIERRE.

specimens belong to ssp. philippinensis. Paragraph 2, after 'Distr.', change 6 into 5.

9: 246 Line 16 from top: change SASAHinto SASA- 9: 342b Cotylelobium melanoxylon (HOOK. /.)

KI. PIERRE.

9: 246 Change the last two lines from bottom into: Delete the Note.

In and its intensity would tend to inhibit vec- 9: 344 caption Fig. 41: b belongs to Vatica um-

tor numbers from reaching adequate levels bonata (HOOK./.) BURCK ssp. acrocarpa

for effective pollination. CHAN (1980) (SLOOT.) ASHTON, d-e to ssp. umbonata.

found most species tohave high self-incom- 9: 355 a Vatica sarawakensis HEIM.

11 patibility, but apomictic adventive em- Line from top: change ‘V. oblongifolia’ 562 Flora Malesiana [ser. I, vol. 9 3

into V. sarawakensis. origin.

9: 365 a Vatica mangachapoi BLCOissp. mangacha- 9: 5436 In caption Fig. 113, line 2, add after 'Bru-

poi. nei': The collector is LADI ANAK BIKAS.

Under'Ecol.' line 3: 'confirmed' in- 547 DYER change 9: parvifolia assp. parvifolia.

to confined. In 'Vern.' line 2, change ‘bung’into bunga. 9: 4046 Hopea latifolia SYM.

In 'Vern.' change ‘jongkang’ into jang- Droseraceae

kang.

9: 4056 Hopea pierrei HANCE. 4: 377, B. J. CONN (Brunonia 3, 1980, 209-216, 2

Add to Distr.: Sumatra (N.W. coast). 5: 557, fig.) gave a review of Drosera L. in New

9: 424 Hopea plagata (BLCO)aVIDAL. 6: 943 Guinea, where all Malesian species occur.

Delete the Note. He gave a new key, cited new records, and

9: In Note 429 under Subsection Pierrea, line 1, added a 7th species, which was hitherto on-

change 'The New Guinea species' into ly known from lowland savannahs in

Some New Guinea species. Northern Australia.

1 9: 4366 Hopea siranda MIQ. under 'Excluded', Drosera banksii R.BR, ex DC. Prod.

change '4B9' into 491. (1824) 319; B. J. CONN, Brunonia 3 (1980)

9:437 Shorea ROXB. ex GAERTN./. 209, f. 2A.

Under 'Uses', line 4, change 'timber ven- Similar to D. peltata J. E. SMITH, but dif-

eer' into veneer timber. fering by absence of bracts under the flo-

and of 9: 4486 Shorea guiso (BLCO) BL., in synonymy: wers presence stipules.

Lines 1 & 2 from top: change'263' into 45. Distr. Northern Australia; in Malesia:

9: 4596 Shorea superba SYM. Papua New Guinea (W. Div.: near More-

Under 'Distr.', line 2, read: Sabah, Tidung, head, JOHNS 2201; Wassi Kussa R., HGF

Berau; Sampit, sterile coll.). 38747).

9: 4606 Shorea astylosa Foxw. Ecol. Under seasonal climatic condi-

Delete the third paragraph of the Notes. tions in open Banksia dentata savannahs,

9: 472 Line 2 from top in text: change 'sepalled' with Melaleuca,Acacia, Eucalyptus, at

into sepals. c. 20 m altitude.

9: 4836 Shorea kudatensis WOOD ex MEIJER.

In Distr. change 'Kilias' into Klias. Ericaceae

9: 487 Shorea dealbata Foxw.a

Under 'Vern.' change ‘bunbong’into bum- 6: 668 Add the Rhododendron species 298-301;

bong. see below.

6: 9: 4916 Shorea assamica DYER ssp. globifera 483, 298. Rhododendron capellae P. Kores, (RIDL.) SYM. 493 Blumea 24 (1978) 181, f. 1.

Under 'Distr.' delete line 4 and read: Ben- Distr. Malesia: Papua New Guinea, W.

coolen. Sepik, Mt Capella(Star Mts), 3800 m (LAE

9: 4996 Shorea albida Sym. 68056).

Under 'Ecol.' line 10: change '6.5' into 65. Note. Keys out to aff. R. vinkii SLEUM.,

Ditto line 21: add after 'community': 3. but its affinity is obviously with R. pullea-

9: 503 Under Subsection Smithiana, line 3, delete num KOORD.

'inflaked'. 6: 4906 Rhododendron scortechinii K. &G.

9: 5046 Shorea smithiana SYM. Add to Distr.: Borneo: Sarawak (Mt Mulu,

In 'Vern.' line 3, change ‘belong’ into be- enroute from 4th camp to summit, on west-

long. ern ridge), 2000-2400 m (Hotta 14907).

9: 5156 Shorea platyclados SLOOT. ex Foxw. 6: 550 299. Rhododendron roseiflorum P. F. Ste-

Delete at the end of the Note: 'not by lateral vens, Adansonia II, 18 (1978) 55, 1 fig. plagiotropic shoots'. Distr. Malesia: West New Guinea: Mt

9: 518 a In captionFig. 102, line 2, read: (Sar)awak, Carstensz, near mining on S. slope,

Semengoh Arboretum, with collector SAL- 2100-2700 m (Raynai 17580, 17672).

LEHstanding beside it (Photogr. SMYTHIES). Note. Keys out near R. ruttenii J.J.S.

9: 524 Shorea macrophyllaa(DE VRIESE) ASHTON. 6: S80, 300. Rhododendron burttii P. J. B. In 'Uses', line 1, read for Illippe: Illipe. 582 Woods, Not. R. Bot. Gard. Edinb. 37

9: 529 In Note under Subsection Auriculatae, end (1978) 157, f. 1 a-d.

of line 1, read: three others. Distr. Malesia: Borneo: Sarawak (Mt

9: 5416 Shorea platycarpa HEIM. Murud), 1500 m.

Note, line 2, read: lowland, dry land for- Note. Belongs to ser. Buxifolia; culti-

ests. vated at Edinburgh. No affinity given but

9: 5426 Shorea curtisii DYER ex KINO ssp. curtisii. keys out to R. frey-wysslingii J.J.S. from

Note, first sentence, after 'Selangor;', read: N. Sumatra.

6: 568 301. Rhododendron rubineiflorum CRA- subspecies grandis may have the same 1982] Addenda, corrigenda et emendanda 563

ven, Not. R. Bot. Gard. Edinb. 38 (1980) probably closest to D. ingens (SLEUM.) STE-

141, f. 1. VENS. Distr. Malesia: Papua New Guinea.

E c o 1. Alpineshrubberies, 2650-3400 m. Fagaceae

Note. Closely allied to R. anagalliflo-

rum WERNH. 7: 277 Nothofagus BL.

6: 646b Rhododendron nervulosum SLEUM. the of the var. Unfortunately typification genus exuberans SLEUM. Nothofagus by N. betuloides (MlRB.)

Add to literature: P. J. B. WOODS, Not. R. Oerst. (Taxon 7, 1958, 145) has been over-

Bot. Gard. Edinb. 37 (1978) 159, f. 1 e-i. looked, and consequently some names of

Note. Additional remarks; attention is infrageneric taxa need correction.

drawn towards similarity with R. steno- 7: 278 Changes in the key at the bottom of the

phyllum HOOK./. page:

6: 657 a Correction as follows: Line 1, replace '1. Sect. Nothofagus'by: 1.

277. Rhododendron irroratum FRANCH. Sect. Calucechinus (Hombr. & J acq.)

ssp. kontumense (SLEUM.) CHAMBERLAIN, Krasser, and omit this name from line 2. Not. R. Bot. Gard. Edinb. 37 (1978) 117. Line 6, replace 'la. Subsect. Antarcticae

R. atjehense SLEUM. FL. Males. I, 6 (1966) Steen.' by: la. Subsect. Calucechinus, and

657, f. 47 & 48. add the former name to its synonymy.

Note. R. irroratum is a Chinese and In- Line 9, replace '2. Sect. Calusparassus dochinese species. (Hombr. & Jacq.) Krasser' by: 2. Sect.

6: 878, Add the following species: Nothofagus, and add the former name to

altiterrae VELDK. 111 242. Vaccinium Blumea its synonymy.

25 (1979) 479. Line 13, replace '2a. Subsect. Quadriparti-

Distr. Malesia: Papua New Guinea tae Steen.' by: 2a. Subsect. Nothofagus,

(Western Highlands: Mts Burgers and Ke- and add the former name to its synonymy. gum). 7: 280 Replace '1. Section Calusparassus (Hombr.

Ecol. Edges of Drimys/Rapanea coppi- & Jacq.) Krasser' by: 1. Section Nothofa-

ces, 3400-3675 m. gus, and add the former name to its syno-

Note. Allied to V. oranjense J.J.S. nymy.

6: 8926, New reduction proposed: 7: 398 Trigonobalanus FORMAN.

8956 Dimorphanthera amblyornidis (Becc.) Through the paper by D. H. MAI on the

F.v.M. var. steinii Stevens, J. Arn. Arb. Tertiary fossils of the genus in the Eocene

58 (1977) 439. — D. steinii Sleum. Fl. Ma- in Europe (Jahrb. Geol. 3, 1970, 381-409)

les. I, 6 (1967) 895. it has become clear that the two living spe-

6: Reduction cies of the in West Malesia and Thai- 895, proposed to variety: genus

8926 Dimorphantheraapoana (Merr.) Schltr. land are relicts of a former much larger

var. mindanaensis (Merr.) Stevens, J. range. T. doichangensis, hitherto only

Am. Arb. 58 (1977)440.—D. mindanaensis known from Thailand, has now also been

Merr.; Sleum. Fl. Males. I, 6 (1967) 892. recorded from Yunnan (Hsu, WANG, WU&

6: 914 Add the following three Dimorphanthera LI, Acta Bot. Yunnan. 3, 1981, 213).

species: This conclusion is now unexpectedly con-

71. Dimorphantheranapuensis P. F. STE- firmed by the find of a third living species

VENS, J. Am. Arb. 58 (1977) 441. in the mountains of the National Park of

Distr. Malesia: West New Guinea (Ba- Colombia (NW. South America). G. LOZA-

liera; Bele R.; Lake Habbema). NO-C, J. I. HERNANDES-C& J. E. HENAO-S

Ecol. Sprawling or scandent shrub in published this as T. excelsa nov. sp. (Calde-

valley forest, 2000-2350 m. sia 12 (n. 60), 1979, recvd June 1980,

D. Note. Specimens were confused with 517-537, 3 pi.). Itgrows in mountain forest

wrightiana(KOORD.) J.J.S. Not assigned to at 1550-1800 m, as a large tree, 20-40 m

a section. high. Its leaves are scattered, as in the Thai-

72. Dimorphanthera wisselensis P. F. STE- land-Yunnan species. It is clear that the ge-

VENS, J. Am. Arb. 58 (1977) 442. nus once had an ancient, large I aurasian

Distr. Malesia: West New Guinea (Wis- range. It should also occur in the fossil state

sel Lakes); one collection, formerly con- in the southern U.S.A.

fused with D. wrightiana (Koord.) J.J.S. 73. Dimorphanthera albida P.F. Stevens, Flacourtiaceae

J. Arn. Arb. 58 (1977) 437.

Distr. Malesia: Papua New Guinea, 5: 466 Ryparosa javanica (BL.) KURZ.

Morobe Distr., Edie Creek, 2700 m, van Add to Distr.: Lesser Sunda Is. (Flores: Pa-

der Kloet 35875, in fern heath;//. Aug. ku, 500 m, SCHMUTZ 4818).

Note. Belongs to sect. Pachyanthae, 5: 51 Homalium JACQ. 564 Flora Malesiana [ser. I, vol. 9 3

CRAVEN (Brunonia 2, 1979, 107-124, 9 single specimen. From the key given it ap-

has of the Pa- that differences fig.) given a new treatment pears single vegetative are

puasian species by proposing 8 new species, frequently used for their delimitation and

by which the number of 2 distinguished by almost all are compared with the widely dis-

in- variable H. Future SLEUMER(F1. Males. I, 5, 1954, 51) has tributed, very foetidum.

creased to 10, apart from the 11th, H. ta- collections will show whether they will

tambense SLEUM. from the Solomons which stand the test of time. CRAVEN'S key and

is included here. This is surprising to me, diagnoses are extracted here:

the more so as 7 of them are based on a

KEY TO THE PAPUASIAN SPECIES

1. Stamens always solitary before each petal.

2. Flowers sessile or subsessile, in simple spikes or racemes.

3. Leaf-apex long-attenuate, acute; leaf margin entire or obscurely distantly crenate. Papua New Guinea

(Morobe Distr.; 1 old coll., not seen) 1. H. acutissimum GILO

3. Leaf-apex shortly acuminate, obtuse; leaf margin distinctly crenate. — Differs from H. foetidum by

sessile flowers and solitary stamens. Papua New Guinea (Western Distr.; 1 coll.)7.H. reductum CRAVEN

2. Flowers distinctly pedicellate, in racemes or panicles.

4. Petiole c. 10 mm long. Leaf margin strongly crenate. — Differs from H. acutissimum by distinctly

crenate leaves and pedicelled flowers in panicles. Papua New Guinea (Madang Distr. and Bagabag I.;

2 coll.); New Ireland (1 coll.) 2. H. bismarckense CRAVEN

4. Petiole c. 5-6 mm long. Leaf marginweakly crenate. — Differs from H. acutissimum by elliptic acumi-

nate leaves and distinctly pedicelled flowers. West New Guinea (Vogelkop; 1 coll.)

H. caput-avis CRAVEN

1. 2 Stamens in fascicles of (1) or more before each petal.

5. Stamens constantly in pairs, onebehind the other. Papua New Guinea and Bismarcks, many collections

5. H. foetidum (ROXB.) BTH.

5. Stamens generallyin fascicles of 3 or more.

6. Leaves broadly ovate to subcordate. Stamens in fascicles of5-8. — Differs from H. foetidumby broad

5 1 subcordate leaves and stamens in fascicles of or more flowers. Solomons (Guadalcanal; coll.)

9. H. subcordatum CRAVEN

6. Leaves elliptic to lanceolate. Stamens in fascicles of not more than 5.

— H. 7. Perianth segments sparsely hairy, at least the sepals glandularon the margins. Differs from foetidum

by glandular perianth. Papua New Guinea (Milne Bay Distr.; 1 coll.) ... 8. H. streimannii CRAVEN

7. Perianth segments densely hairy and non-glandular.

8. Leaf marginweakly crenate to entire. — Possibly allied toH. subcordatum but different in elliptic nar-

rower leaves and stamens usually in fascicles of 3-4. Solomons (Santa Isabel I.; 1 coll.)

10. H. tatambense SLEUM.

8. Leaf margin distinctly crenate.

9. Leaves velutinous. — Differs from H. foetidum by the velutinous leaves and stamens mostly in fasci- cles of 3-4. Papua New Guinea (Central Distr.; 1 coll.) 11. H. velutinum CRAVEN

9. Leaves glabrous.

10. Leaf margin entire in the basal 1/3-1/2. Branchlets glabrescent. — Differs from H. foetidum by:

petals and sepals similar, stamens mostly in fascicles of 3. Normanby I. (1 coll.)

4. H. dentrecasteauxense CRAVEN

10. Leaf margin wholly crenate. Branchlets glabrous. — Differs from H. foetidum by larger flowers

and stamens in fascicles of 3-5. Papua New Guinea (Milne Bay Distr.; 1 coll.)

6. H. maneauenseCRAVEN

Flagellariaceae moner form with small, globularfruits clo-

fruit ser set, by spaced, larger, ovate-acute 4: 2496 Hanguana malayana (JACK) MERR. up to 1.5-2 by 1.25 cm. SHAW added that

Add to Distr.: N. Australia: Northern Ter- the small-fruited form does not occur in

ritory, cf. AIRY SHAW, Kew Bull. 33 (1978) this area.

4. The species H. malayana, as conceived

Note. The specimens belong to var. by BACKER (Fl. Males. 1,4,1951,249)is ad-

anthelminthica (BL.) BAKH./. mittedly variable, BACKER mentioning the

4: 249 Hanguana major SHAW, Kew Bull. 35 fruit also to attain 2 cm. Extensive field ob-

(1981) 819, 1 fig. servation would be needed to check the

This proposed new species, confined to racial and population variability in detail

the Kinabalu differs from the the entire area, com- over range. 1982] Addenda, corrigenda et emendanda 565

Geraniaceae to their satisfaction, but further considera-

tion has induced them 'to regard the taxa as

4: 445 Geranium LINNE. species'. These considerations are, amongst

The number of Malesian species, according others, that there may be chromosome

to CAROLIN (Fl. Males. I, 6, 1964, 445) 3 taxa, that most taxa are represented by

and one variety, has unexpectedly been in- more than one collection, and that they ap-

creased by VELDKAMP & MOERMAN (Blumea pear to be homogeneous, althoughin sever-

24, 1978, 463 —477) to 15, including 9 new al cases occurring on more than one moun-

species from New Guinea and 1 from SW. tain. In the key the exact description ofthe

Celebes. In their introduction the authors leaves plays an important role. Future re- state that the evaluation of taxonomic sta- search, especially karyologic, must show

tus ofthese taxa has posed a bit of a prob- whether the specific status of the new spe-

lem, which they have not been able to solve cies can be maintained.

KEY TO THE SPECIES

1. Leaves (sub)compound, middle 'leaflet' free for at least 0.9th of its length.

2. 'Leaflets' repeatedly2-3-partitewith ± linear-lanceolate,overlapping segments; the middle 0.2-0.3 mm

wide at base. Papua New Guinea (Mt Suckling; 4 coll.) 9. G. leptodactylon VELDK.

2. 'Leaflets' 3-lobed to -fid, segments broader, not overlapping, the middle 0.3-1 mm wide at base. Papua

New Guinea (Star, Saruwaket, Owen Stanley Mts; 12 coll.) 13. G. subcompositum VELDK.

its 1. Leaves palmatifid to -partite, middle segment free for at most 0.9th of length, usually much less.

3. Lower leaf surface very densely grey to silvery hairy, sometimes brown when dried, the indument obscur-

ing the venation. West New Guinea (Carstensz to Star Mts; 15 coll.) 10. G. monticola RIDL.

3. Lower leaf surface variously strigose, the indument not obscuring the venation, often making it more

prominent, instead.

4. Middle leaf segment entire, the laterals sometimes with a lobe, rarelythe middle segments of some leaves

2- or 3-lobed, leaves then glabrous on the upper surface, 5-partite, and petals pink.

5. Upper surface of the leaves strigose-setose. Peduncle at anthesis already 18-23 mm long. Papua New Guinea (Mt Saruwaket; 1 coll.) 6. G. editum VELDK.

5. Upper surface of the leaves glabrous. Peduncle in fruit 0-11 mm.

6. Leaves 7-fid, middle segment free for 0.5-0.6th of its length,0.8-2 mm wide at the base of its free

part. Papua New Guinea (Mts Giluwe, Wilhelm, Bangeta; 10 coll.) 7. G. hyperacrion VELDK.

6. Leaves 5-partite, middle segment free for 0.8-0.9th of its length, 0.5-0.7 mm wide at the base of its

free part. West New Guinea (Mt Wilhelmina; 3 coll.) 15. G. wilhelminae VELDK.

4. Middle leaf segment lobed, in some upper or reduced leaves occasionally entire and rarely also glabrous

on the upper surface, then 5 —7-fid or petals purple.

7. Upper leaf surface glabrous or sparsely and patchily long-strigose.

8. Petals 9-16 mm long, purple (? always), 1.7-2.1 times as long as the 5.5-7.6 mm long sepals. West

New Guinea (Carstensz, Star, Piora Mts; 9 coll.) 12. G. papuanum RIDL.

8. Petals 4.5-6 mm long, white to pink, 1.2-1.4 times as long as the 3.5-4.8 mm long sepals.

9. Upper leaf surface patchily long-strigose. West New Guinea (Mt Wilhelmina; 1 coll.)

8. G. lacustre VELDK.

9. Upper leaf surface glabrous or with some long hairs near the margin. Papua New Guinea (Wharton

Ra.; 6 coll.) 14. G. whartonianum VELDK.

7. Upper leaf surface evenly strigulose to strigose.

10. Leaf blade fairly large, usually over 15 by 25 mm.

11. Inflorescences strictly 1-flowered. Peduncle 25-41 mm. Sepals 5-6 mm long, in fruit 7.5-9 mm.

Petals 5.5-7.5 mm long. Java (Mt Merbabu eastward to Mt Tengger; 13 coll.) 2. G. ardjunense Z. & M.

11. Inflorescences usually 2-flowered. Pedicels 9-16 mm. Sepals 3.5-5 mm long, in fruit 3.8—7.5 mm.

Petals 3.5-6 mm long. New Zealand, Tasmania, East Australia, Timor, East Java (Mt Tengger); 12

coll 3. G. homeanum TURCZ.

10. Leaf blade fairly small, usually less than 15 by 25 mm.

Middle fruit 12. leaf segment pinnately lobed. Peduncle in 55-150 mm. Ceylon, Nilghiris, Himalayas

(Afghanistan to W. China), N. Sumatra (one coll.) 1. G. nepalense SWEET

12. Middle leaf segment 3-lobed, its outer lobes rarely with a lateral tooth. Peduncle in fruit 0-45 mm.

13. Leaf blades 7-fid to -partite. Peduncle in fruit 0-3 mm long. Papua New Guinea (Star, Kinkain,

Wilhelm and Saruwaket Mts; 11 coll.) 5. G. balgooyi VELDK.

13. Leaf blades 5-partite. Peduncle in fruit 6-45 mm.

14. Inflorescences 1- or 2-flowered. Peduncle in fruit 6-20 mm, pedicel then 4-15 mm. Petals red-

purple. SW. Celebes (Mt Bonthain; 2 coll.) 4. G. frigidurbis MOERMAN 566 Flora Malesiana [ser. I, vol. 9 3

Inflorescences 14. strictly 1-flowered. Peduncle in fruit 19-45 mm, pedicel then 16-31 mm. Petals

white to pinkish. Papua New Guinea (Sugarloaf, Giluwe, Saruwaket to Dayman Mts; 24 coll.)

11. G. niuginiense VELDK.

Goodeniaceae lu), 3300-3760 m alt.

Veldkamp (Reinwardtia 10, 1982, 25)

5: 339o, 1. Scaevola sericea VAHL, Symb. Bot. 2 noted that the specimens differ in many de-

6: 951O — (1791) 37. S. taccada (GAERTN.) ROXB. tails from the common form. I still consider

Hort. 15. Kew Bull. Beng. (1814) JEFFREY, it an alien brought by tourists.

34 (1980) 543. 4: 214 Add the following species:

For extremely formal meticulous-no- 5. Juncus nupela Veldk. Blumea 24 (1977)

menclatural reasons JEFFREY has advanced 415.

that S. should sericea be the proper combi- Distr. Malesia: Central New Guinea

nation. (Star Mts, Tel Basin, 3000 m, Veldkamp 6369).

Hydrocharitaceae Note. The author finds this to be allied

to the northern hemisphere J. balticus

5: 396o, Change„Limnobium1. stoloniferum(laevi- WILLD. and tabulated differences with three

7: 828b gata) into: other species, including J. inflexus L. No

& B. 1. Hydromystria laevigata (H. ex developed ovules could be found.

WILLD.) DIAZ-MIRANDA& PHILCOX, Bot. J. 4: 214 Luzula DC.

Linn. Soc. 83 (1981) 321, 6 fig. R. BROWN and BENTHAM assumed the Aus-

The generic name Hydromystria G. tralasian specimens of Luzula to belong to

Prim. Fl. 152 MEYER, Esseq. (1818) ante- a broad concept of L. campestris. In this

dates the synonym Limnobium RICH, ex they were followed by BUCHENAU in his

STEUD. (1841). monograph, except that the latter distin-

guishedthem to represent an outlyingvarie-

Icacinaceae ty, var. australasica. MERRILL (1922) ac-

commodated the Philippinespecimens also

7: 15 Gonocaryum MIQ. in L. campestris sensu lato. BACKER (Fl. Add to Distr.: Lesser Sunda Is. (West Males. 1,4, 1951,215) identified the Philip-

G. Flores), probably macrophyllum (BL.) pine and Papuan specimens as belonging to SLEUM. the variety.

7: 55 Replace the name Nothapodytesfoetidaby: The precise naming of the Australasian

1. Nothapodytes nimmoniana (J. GRAH.) specimens of this affinity complex from

MABBERLEY in K.S. Manilal (ed.), Bot. Australia, Tasmania and New Zealand has

Hist. Hort. Mai. (1980) 88; Taxon 29 (1980) given rise to a confused nomenclature and

606. — N. foetida (WIGHT) SLEUM.; Fl. species distinction which M. E. JANSENhas

Males. I, 7 (1971) 55. — Premna nimmo- tried to solve (Blumea 24, 1978, 527-532, 1

niana J. GRAH. Cat. PI. Bombay (1839) fig.). For Malesia he distinguishedtwo new

155. endemic species, L. philippinensis M. E.

The antedates that of Stemo- and basionym JANSEN L. papuana M. E. JANSEN. In

WIGHT six nurus foetidus by years. the key he did not include how they are dis-

7: 56 Stemonurus Bl. tinguished from L. campestris (L.) DC. It

Add to Distr.: Lesser Sunda Is. (Flores; must be left to a future monographic study

near Paku, 900 m, SCHMUTZ 3371, speci- of the genus to see whether these two taxa

men in fr.; identified by SLEUMER). can be upheld at specific rank.

This collection fills in the nicely a gap

genericrange and shows again that the Les- Labiatae

ser Sunda Islands flora fits in with the rest

of Malesia. 8: 3386 Leucas lavandulifolia J. E. SM. Change 7: 73 Sarcostigma W. & A. into:

Add to Distr.: Lesser Sunda Islands (Flo- 3. Leucas linifolia (ROTH) SPRENO. Syst.

res). Veg. ed. 15, 2 (1825) 743; BTH. Lab. Gen.

7: 15b Sarcostigma paniculata PIERRE. Sp. (1834) 617, (1835) 744; FOSBERG& SA-

Add toDistr.: Lesser Sunda Is. (Flores; Pa- CHET, Smithson. Contr. Bot. 47 (1981) 25.

ku, 500 m, SCHMUTZ 4818a). — Phlomis linifolia ROTH, NOV. PI. Sp.

Ind. Or. (1821) 260. — L. lavandulaefolia

Juncaceae J. Sm. in Rees, Cycl. (1812) Leu 20, nom.

Meg. (superfl.); Keng, Fl. Males. 1,8 (1978)

4: 2136 Juncus bufonius L. 338.

Note. Add to Distr.: Borneo (Sabah: Mt Kinaba- Smith's name is illegitimateas a 1982] Addenda, corrigenda et emendanda 567

superfluous name since he cited Leonurus comm., 1980).

Leonurus indicus L. as a synonym. indicus 6: 308A Fagraea fragrans Roxb.

can no longer be transferred to Leucas, be- Add to Distr.: Ceylon, the whole of

R.BR, SW. Guinea. causeofL. indica ex SPRENG., based Indo-China, New

on Phlomis indicus L. 6: 320a Insert the following species:

8: 340b Leucas flaccida R.BR. Change into: 8a. Fagraea graciliflora Leenh. Blumea 27

6. Leucas decemdentata (WILLD.) J.SM. in (1981)209.

Rees, Cycl. (1812) LEU 20; DRAKE, 111. Fl. Distr. Malesia: Central Celebes (Mt Ro-

Baloooy Ind. Mar. Pac. (1890) 263; FOSBERG& SA- roka Timbu; van 3247, deVogel

CHET, Smithson. Contr. Bot. 47 (1981) 25. 5390).

— Stachys decemdentata Sol. ex Forst./. Ecol. Montane ridge forest dominated

Prod. (1768) 91, nomen. — Phlomis de- by conifers and Fagaceae, at c. 2100 m. Fl.

cemdentata WILLD. Sp. PI. 3 (1800) 124. — May.

L. flaccida R.BR. Prod. (1819) 505; KENG, Note. Though the texture of the flowers

Fl. Males. I, 8 (1978) 340. — L. stachyoides and the distinctly exserted stamens and

SPRENG. Syst. Veg. ed. 15, 2 (1825) 743. style remind of the situation in sect. Cyrto-

8: 361 Satureia gracilis (BTH.) LOES. phyllum, the new species will have to be in-

Add to Distr.: Borneo: Sabah (Headquar- cluded in sect. Fagraea which is the most

ters Taman Negara Sabah, c. 1560 m alt., primitive in the genus (cf. Punt & Leen-

R. H. WILLEMSE 605, in disturbed places). houts, Grana Palynol. 7, 1967, 510-515).

It is closest allied to F. tubulosa BL. which

Liliaceae is found in Sumatra and Malaya.

In the key given in Fl. Males. I, 6 (1962)

9: 213 Thysanotus tuberosus R.BR. 302, the new species keys out under the first

In revision ofthe lead of 19 F. tubulosa BL. from arecent genus Thysanotus couplet as

N. H. BRITTAN(Brunonia4, 1981,67-181) which it can be distinguished as follows:

assigned the Papuan specimens to T. bank- 19a. Leaves about 2 times as long as wide.

sii R.BR., a name which since BENTHAM'S Inflorescences glomerulous, with

Flora was considered a synonym. In the key bracteoles 8. F. tubulosa

it appears that the minute differential char- 19a. Leaves 13.5-22 by 4-6.5 cm, 2.5-4

acters either do not hold or are inconsistent times as long as wide. Inflorescence a

with the descriptions. I cannot accept T. terminal 3-flowered cyme and in the

banksii as a distinct species. axil of the upper leaf-pair either a

9: 234a Astelia alpina R.Br. cyme or a solitary flower. Bracteoles

Add to Distr.: West Central Celebes (Lati- absent 8a. F. graciliflora

modjong Range; Mt Rante Mario, summit 6: 3206 Fagraea ridleyi K. & G.

zone, very local). Add to Notes: Though typical specimens of

Note. The collection was made by Dr. J. F. blumei and F. ridleyi are clearly distinct,

M. B. Smith in February 1981 and is a re- fruiting material is not always easy to

markable addition. name, especially in Borneo. As a whole the

number of flowers and fruits is smaller in

I.oganiaceae (Leenhouts) the present species, the pedicel and calyx in

fruit are less densely warty, the calyx is lar-

6: 293 Wood anatomy. See A. M. W. Menne- ger, the nerves are fewer in number and

oa in E. & P. Nat. Pfl. Fam. ed. 2, 28b I more prominent beneath, the axillary scales

(1980) 112-161. are less conspicuous, and the dried leaves

Palynology. See W. Punt in E. & P. are more yellow, in contrast to the

Nat. PH. Fam. ed. 2, 28b I (1980) 162-191. usually dark redbrown ones of F. blumei.

6: 294 Phytochemistry. See N. G. BissETin E. 6: 328a Fagraea auriculata Jack.

& P. Nat. Pfl. Fam. ed. 2, 28b I (1980) Add toDistr.: Great Nicobar (cf. Fl. Males.

211-237. Bull, no 29, 1976, 2547) and Celebes.

6: 295 Delimitation and subdivision. See 6: 3286 Add to Notes: van Baloooy 3492 from

A. J. M. Leeuwenberg et al. in E. & P. Central Celebes combines the long-petioled

Nat. Pfl. Fam. ed. 2, 28b I (1980) 1-255. leaves with small auricles of ssp. parviflora

Fosberg & Sachet (Smithson. Contr. with the large flowers of ssp. auriculata.

Bot. 45, 1980, 18-19) suggest the combina- The collection VAN BALGOOY 3678, also

tion of Loganiaceae-Potalieaewith Gentia- from Central Celebes, represents ssp. bor-

neensis. naceae-Tachiineae, preferably as part of

the Gentianaceae, but on rather vague and 6: 331a Fagraea resinosa LEENH.

superficial arguments. This seems at least Add to description: Tree, 7.5-9 m by 2-20

contrary to wood anatomical data (Menne- cm, or climber. Leaves 8-16 cm long,

I.e. and width from 3.5 onwards. Bracteoles 2 ga, 158) to palynology (PUNT, pers. cm 568 Flora Malesiana [ser. I, vol. 93

and 3 3-3.5 in I cm resp. Calyx cm high, Note. doubt whether this new species

4 fruit up to cm. will be tenable. Edit.

Add to Distr.: Sarawak (4th Div.), N. Bor-

neo (Mt Kinabalu; Sandakan Distr., Telu- Myoporaceae pid).

to to 1550 KRAENZL. Add Ecol.: Altitude up m. 4: 265 MyoporumpapuanumA

6: 336 Buddleja HOUST. ex LINNE. Add to Distr.: Lesser Sunda Is.: W. Timor

Add to literature: LEEUWENB. Meded. (Kie, near Amanubar, C. W. KOOY 1271, Landbouwhogeschool 79-6 (1979) 1-163. dated 7-2-1981).

6: 359O Strychnos axillaris COLEBR.

Add to Distr.: Ceylon. Pittosporaceae

6: 365, Neuburgiacorynocarpa (A. GRAY) LEENH.

959 In a detailed study of the specimens ofN. 6: 9626 Pittosporumpumilum SCHODDE.

in corynocarpa Papua B. J. CONN (Bruno- Add to literature: Steen. Blumea 24 (1978) 4 nia 2, 1979, 99-105, fig.) has come to the 482.

BE- conclusion that N. sarcantha (GILG & Add to description: Pedicels 6 mm. Flowers

NED.) LEENH. FL. Males. I, 6 (1962) 366 can- functionallyfemale, deeppurple. Sepals 5,

not well be distinguished at specific level. free, ovate-oblong, blunt, long-hairy, 5 by

Accordingly he reduced this to a variety of 2.5 mm. Petals 5(-6), cohering at base, li-

the former, var. sarcantha (Gilo& Bened.) gulate, glabrous, bent outwards, apex

B. J. Conn, which he keyed out (I.e. 104). rounded, c. 12 by 2.5 mm. Stamens re-

also described structure He the ofseedlings. duced, c. 3 mm. Ovary densely brown-

Distr. Add: Moluccas (Obi) and New hairy, ellipsoid, 4 by 2 mm; style glabrous

Ireland. 1 mm. Fruit red to deep-brown.

Note. An isotype of N. sarcantha is Add to Distr.: Papua New Guinea, Mt Ke-

found in B (LEDERMANN 13005) and re- nive (Nisbet), 2500 m, LAE 6501 \\fl. July.

places the neotype with which it completely

agrees. Proteaceae

6: 371 Geniostoma FORST.

Recently B. J. CONN (Blumea 26, 1980, 5: 1956 Macadamia hildebrandii STEEN. Add:

245-364, 29 fig.) made a world revision of Uses. This endemic tree of Celebes has

the genus resulting in the name change of been successfully planted as a fire-lane tree

one and the creation of a new species. He in the very large plantations of Pinus mer- also gave a new key to the species of Papua- kusii in the Aek na Uli area, on the north

sia (incl. Solomons). In using my key (Fl. flank of Toba Lake (N. Sumatra). Thirty

Males. I, 6, 1962, 371) the two can be ac- years old trees have the virtue of keeping commodated in couplet 3, second lead, af- dense foliage and branching from the base

ter: upwards.

3. Corolla 2-4 mm long.

4. Pistil hairy 4. G. antherotrichum Rhizophoraceae

4. Pistil glabrous 5. G. leenhoutsii

4. Geniostoma antherotrichum GILG & BE- 5: 429 The distinction of sterile material of Rhizo-

NED. Bot. Jahrb. 54 (1916) 158, f. 2; phora and Bruguiera may give difficulty.

LEENH. Fl. Males. I, 6 (1962) 371, as syn. Dr. DING HOU found an easy way for iden-

under G. rupestre;; B. J. CONN, Blumea 26 tification (Blumea 10, 1960, 628) by means

— In (1980) 317, f. 15. G. arfakense Kan. & of a hand-lens. Bruguiera the leaf-scars

Hat. Bot. Mag. Tokyo 56 (1942) 163, f. 7; show 3 distinct, usually horseshoe-shaped

Leenh. Fl. Males. I, 6 (1962) 373. bundles of leaf traces. In Rhizophora there

Note. This species has a variety: are several vascular bundles in two rows or

var. archboldianum (Merr.& Perry) B. J. a crescentic pattern. This was recently also

Conn, Blumea 26 (1980) 320, f. 15G. — G. advanced by Kenneally C.S. in their work

archboldianum Merr. & Perry, J. Arn. onAustralian mangroves (Nuytsia 2, 1978,

Arb. 23 (1942) 408. 178-180, 1 fig.).

Distinct by smaller leaves (2-6 by 1—2

A cm) and flowers solitary or in triads. few Styracaceae

collections intermediate with the type vari-

ety. 4: 53a crotonoides Clarke fraserensis Styrax ,. ssp.w

5. Geniostoma leenhoutsii B. J. Conn, Bin- (Putz & Ng) Steen., comb. Nov. S.

mea 26 (1980) 323, f. 17. fraserensis PUTZ& NG, Mai. For. 40 (1977)

Distr. Solomons (type), in Malesia: Pa- 249, f. 1; Tree Fl. Malaya 3 (1978) 263.

Guinea Distr.: pua New (Central Woitape); A small tree. Average mature leaves in 2 all specimens known. 6-10.5 by 2.5-5 cm; nerves 4-6 pairs. 1982] Addenda, corrigenda et emendanda 569

2 Fruits roundish, c. cm diam. 8: 267 Symplocos ophirensis CLARKE var. kalien-

Distr. Matesia: Malaya: Fraser's Hill, sis Stone, Mai. For. 43 (1980) 260, f. 6.

Distr. Malesia: G. at c. 1300 m. Malaya: Pahang,

Notes. I have carefully compared the Ulu Kali, Stone 13965.

descriptive data with those of S. crotonoi- Note. According to Stone closest to

in des, but find no essential differences, and var. densereticulata NOOT., differing a

also that the ones mentioned are slightly glabrous disk, branches, and fruit, style 5

overlapping. In the species the mature mm. From all other varieties different by a

leaves average 8.5-20 by 4-10 cm, nerves rounded leaf base and a convex petiole.

5-9 pairs. Fruit round to ovoid, tending to

have a smallish or indistinct tip at apex, Ulmaceae

1.7—3 cm diam. Lowland, 90-300 m.

Obviously a hill race, with smaller leaves. 8: 616 Celtis paniculata (ENDL.) PLANCH.

The Add shape of the fruit induced me to main- to synonymy: Strombosia philippinen-

tain it as a taxon. sis sensu Lam & Holth. Blumea 5 (1942)

4: 546 Styrax ridleyanum PERK. 178.

PUTZ & No (Tree Fl. Malaya 3, 1978, 264) Add to Distr.: Moluccas (Talaud Is.: Lam

have discarded this from Malaya and find 3175).

the specimen YEOP KEP 3639 best to place

under the allied .S. benzoin DRY AND. be- Umbelliferae

causeof the short inflorescence. I must ad-

mit that this is indeed no differentiating 4: 131 Add the following species:

character. I had no opportunityto re-study 5. Oreomyrrhis plicata Mathias & Con-

this specimen. stance, J. Am. Arb. 58 (1977) 190, f. 1-6.

However, another one, also with short - Fig. 3.

inflorescence, exactly tallies with the differ- Leaves tufted at apex of a thick, hardly

enceswith S. benzoin as given in the key of branched ligneous stem 5-15 cm long,

elaborate Jard. Bot. linear the my treatment (Bull. erect, to lanceolate, 1-pinnate,

Btzg. 111, 12, 1932,223): twigs dark brown- sheathingpetiole about as long as the blade;

ish; leaves with smaller stellate scales than blade 1-4 cm by 1-3 mm, plicate, seg-

in S. benzoin; buds rather broad; petals in- ments 5-11, linear, 2-5 mm long, erect,

all side appressed-pubescent over; connec- entire. Peduncles erect, solitary, exceeding

tive ditto; stigma 3-lobed, much wider than the leaves, 2.5-12 cm long. Flowers pedi-

the style. celled, in a head.

Distr. Malesia: Malaya: K. Trengganu, Distr. Malesia: Papua New Guinea (Mt

Bt Lah, off Sg. Nerus, near Kp. Merjor, Suckling, summit of Goe Dendeniwa:

Kiah Sinclair* SF40896 (dupl. in L). Veldkamp& Stevens5748), 3325-3625 m,

Note. I expect that KEP 3639 from Ku- rocky ridges.

antan will turn out to belong to S. ridleya- Note. Distinctly different from all other

num, as I studied this in 1932. species by the simply pinnate leaves, folded

lengthwise, with closely adpressed, linear,

Symplocaceae entire leaf segments.

4: 1316 Replace 2. Apiumtenuifolium by:

8. 239b Insert the following species: 2. Apium leptophyllum (PERS.) F.V.M. ex

5a. Symplocos columbuli Noot. Blumea 26 BTH. FL. Austr. 3 (1866) 372; BACK. & (1980) 417, fig. BAKH./. Fl. Java 2 (1965) 175. Cntdium

Distr. Malesia: N. Sumatra, Leuser Re- tenuifolium MOENCH. Meth. (1794) 98,

serve, in montane moss forest, 2100-2500 excl. syn. Pimpinelladioica L., nom. illeg.

m. - Pimpinella leptophylla PERS. Syn. 1

Note. In Nooteboom's key to fruiting (1805) 324. Helosciadium leptophyllum

material (Fl. Males. I, 8, 1977, 231) it (PERS.) DC. Mem. Soc. Phys. Geneve 4

comes out in two places because the leaves (1828) 493; Prod. 4 (1830) 105, cum syn.

are hairy underneath but finally glabrous, numer. Apium tenuifolium (MOENCH.)

viz. via lead 3a to couplet 8, from the two Thell. in Hegi, 111. Fl. Mitteleur. 5, 2

species of which it is distinguishedby: fruits (1926)1140,nom. illeg.; Buwalda, Fl. Ma-

narrow-ellipsoid, c. 15 mm long, with c. 10 les. I, 4(1949) 131.

conspicuous lengthwise ribs. Via lead 3b Distr. Southern hemisphere, intro-

one arrives at couplet 25, where the same duced in various countries: Europe, India,

characters single it out. Taiwan, Pacific Islands (Hawaii: Kauai; Fi-

Sofar only known in the fruiting state; ji, H. J. Lam 6827); in Malesia: Java(W.:

affinity probably with S. atjehensis NOOT. Lembang; E.: Pudjong, Tretes, Ever-

from the same area. AARTS, in litt.), Philippines (Luzon: Ba- 570 Flora Malesiana [ser. I, vol. 9 3

Fig. 3. Oreomyrrhis plicata MATHIAS & CONSTANCE. a. Habit, x 0.5, b. foliageleaf, nat. size, c. fruitingum- bel, x 2.5, d. petal, x 9, e. intact mature fruit, x 7, f. fruit transection, x 10 (Courtesy Journal Arnold Arbo- retum). 1982] Addenda, corrigenda et emendanda 571

guio,PNH 35028,35842), Papua New Gui- walls, 800-1500 m alt.

NGF nea (Western Highlands, Minj Distr.,

41801;Morobe Distr., Sattelberg, CLEMENS Xyridaceae (B. HANSEN, Copenhagen)

1720; Wau: NGF 29146; Goroka: MCKEE

1160). 4: 368 Replace the key tothe species ofXyris L. by

Ecol. Fields, wastelands, roadsides, old the following:

KEY TO THE SPECIES

1. Median bracts at apex emarginate, margin otherwise entire, grey field 2by 1.2 mm. Burma, Thailand,Viet-

nam X. lobbii RENDLE

1. Median bracts not emarginate, althoughsometimes irregularly torn, if emarginate then margins obviously

lacerate above.

2. Median bracts with upper margin lacerate.

3. Margin ofleaves formed by numerous retrorsely pointing, coalescent rows of cells, rough from small tu-

with bercles; scape compressed, usually two strong ridges 1. X. complanata

3. of leaves with low 2. X. bancana Margin not so; scape terete, ridges

2. Median bracts not lacerate at margin, rarely with a few dents.

with 4. Scape terete 6-15 longitudinalridges, leaves up to 8 mm wide with short transverse ribs connect-

ing the longitudinal ones 3. X. indica

4. Scape terete to compressed, without ridges or with 1-7 ridges, leaves never with transverse ribs.

5. Leaves 6-20 mm wide, inflorescence depressed hemispherical, wider than long;plant extremely robust

4. X. grandis

5. Leaves inflorescence wide. up to 6, rarely 7.5 mm wide, ellipsoid, globular or longer than

6. Median bracts without a greenish or greyish field below apex, at most with a ± conspicuous, narrow

ridge 5. X. capensis

6. Median bracts with a greyish or greenish field below apex.

7. Leaf sheath with margin ± finely ciliate below, oblong-triangulargreyish-brown field below apex of

median bracts 3-10 times longer than wide, not protruding upwards 6. X. wallichii

7. Leaf sheath not ciliate below, triangular greyish field below apex of median bracts at most 2.5 times

longer than wide.

8. Scape 4-angularwith papillatelongitudinalridges; leaves with papillatemargins, otherwise smooth

7. X. oligantha

8. Scape terete, without ridges.

9. Lateral sepals with smooth, entire crest.

10. Plants gracile, leaves at most 3 mm wide, often rough by protrudingcell walls (when dry), grey

field on median bracts ridge-like protruding towards apex of bract 8. X. pauciflora

10. Plants robust, leaves more than 3 mm wide, not rough, grey field on median bracts not protrud-

towards of bract Vietnam ing apex although convex. Burma, Thailand, Laos, Cambodia,

X. intersita MALME

Lateral 9. sepals with crest coarsely serrate, often by 1-3 dents only.

11. Plants robust, field on median bracts 1.8-2.6 by 1.4-2.6 mm, not protruding; with subglobose

underground tubers. Thailand, Laos, Cambodia X. tuberosa RIDL.

1 11. Plants gracile, grey field onmedian bracts -2.2 by 0.4-1.5 mm, ridge-likeprotrudingtowards

apex of bract; without tubers 8. X. pauciflora

4: 369 1. Xyris complanata R.BR. Prod. 1 (1810) 2. Xyris bancana Miq. 11. Ind. Bat., Suppl.

256; v. ROYEN, Fl. Males. I, 4 (1953) 369. (1861)608; v. Royen.F1. Males. 1,4(1953)

indica - X. auct., non LINNE: VAHL, 369, incl. var. lacerata v. ROYEN, I.e. 370.

Bot. 3 Symb. (1794) 7, p.p. quoad spec. -X. ridleyi Rendle, J. Bot. 37 (1899) 505;

— X. KOENIG. anceps auct., non LAMK: v. Royen, Blumea 7 (1953) 309, incl. var.

En. PI. 2 — VAHL, (1805) 205. X. elongata penicillata v. ROYEN, Fl. Males. I, 4 (1953)

RUDOE, Trans. Linn. Soc. 10(1811)289. — 370, 371. X. borneensis RENDLE, J. Bot.

X. walkeri KUNTH, En. PI. 4 (1843) 19. — 37 (1899) 506; v. ROYEN, Fl. Males. I, 4

X. malaccensis Steud. Syn. PI. Glum. 2 (1953) 370. X. chinensis MALME, Svensk (1855) 287. Bot. Tidskr. 21 (1927) 386. X. glaucella

Distr. Ceylon, India, China (Hainan), MALME, Bull. Jard. Bot. Btzg 111, 10 (1929)

Indochina, Thailand, throughout Malesia 388. X. subcomplanata MALME, Bull.

to Australia. Mus. Hist. Nat. Paris 11, 2 (1930) 685. X.

Ecol. Open, wet places on sandy soil, papuana v. ROYEN, Blumea 7 (1953) 307;

often among grasses and Sphagnum, from Fl. Males. I, 4 (1953) 371.

sea level to 1600 m. Distr. Thailand, Cambodia, Vietnam, 3 572 Flora Malesiana [ser. I, vol. 9

Fig. 4. Distribution of Xyris bancana MIQ. The dots provided with anoblique line represent localities above

1000 m altitude.

Hong Kong; in Malesia: Sumatra (Banka), v. ROYEN, Fl. Males. I, 4 (1953) 373. X. Malaya (Kedah Peak), Borneo, New Gui- robusta MART, in Wall. PI. As. Rar. 3

Aru Fl. Ind. nea (Misool I.; Is.: Trangan) as far east (1832) 30. X. calocephala MIQ.

as Sepik. Fig. 4. Bat. 3 (1857) 527. X. capito HANCE, J.

Ecol. On (temporarily) boggy or wet, Bot. 14 (1876) 262.

invariably acid, sandy soils (kerangas, Distr. Ceylon, India, Burma, China

sandstone, heath woodland), largely con- (Hainan), Thailand,throughoutMalesia to

fined to low altitudes below 100 m, but in Australia.

Indochina and Malaya at 1100-1300 m and Ecol. On open, swampy places and

in West New Guinea on the Star Mts at along or in ricefields in the lowland, rarely

1200-1300 m and on white sands in the Ba- up to 900 m.

liem Valley at 2000 m, nowhere at altitudes 4. Xyris grandis RIDLEY, J. Linn. Soc. Bot.

4 between. Fig. 4. 38 (1908) 332; v. ROYEN, Fl. Males. I,

X. 3. Xyris indica LinnS, Sp. PI. 1 (1753) 42; (1953) 372. chlorocephala v. ROYEN, 1982] Addenda, corrigenda et emendanda 573

Blumea 7 (1953) 308; Fl. Males. I, 4 (1953) Ecol. In wet, mossy places on rocks, 372. 850-1300 m.

Distr. Indochina, Thailand,inMalesia: 7. Xyris oligantha STEUD. Syn. PI. Glum. 2

Malay Sumatra. Peninsula, (1855)288. v. ROYEN.F1. Males. 1,4 (1954)

E c o In wet places, 599. — X. 1. among mosses over pauciflora auct., non WILLD.: R. 500-2200 rocks, m. BR. Prod. 1 (1810) 256, quoad descr.

5. Xyris capensis Thunb. Prod. Fl. Cap. D i s tr. Australia (Cape York Peninsu-

(1794) 12; NnssoN, Ofvers. Forh. Kongl. la); in Malesia: Aru Is. and New Guinea Svenska Vet.-Akad. 48 (1891) 155, incl. (Papua, Western Distr.).

var. nilagirensis (STEUD.) NILSSON et var. E c o In 1. wet, sandyplaces among sedges schoenoides (MART.) NILSSON; V. ROYEN, and grasses, up to 60 m. Fl. Males. 4 incl. I, (1953) 374, var. K. 8. Xyris pauciflora WILLD. Phytogr. 1 schoenoides MART, in Wall. PI. As. Rar. 3 (1794) 2; v. ROYEN, Fl. Males. I, 4 (1953) (1832) 30. X. nilagirensis STEUD. Syn. 371.— X.pauciflora var. oryzetorum MIQ.

PI. Glum. 2 X. — (1855)288. melanocepha- Fl. Ind. Bat. 3 (1857) 529. X. dajacensis la MIQ. Fl. Ind. Bat. 3 (1857) 528. X. su- v. ROYEN, Blumea 7 (1953) 308; Fl. Males.

matrana MALME, Bull. Jard. Bot. Btzg 111, I, 4 (1953) 372. — X. maritima KOYAMA, 10 391. (1929) X. novoguineensis Philip. J. Sc. 84 (1956) 367. HATUS. Tokyo Bot. Mag. 56 (1942) 422. Distr. Ceylon, India, Burma, Thailand, flabellata X. v. Royen, Blumea 7 (1953) Indochina, China, Hong Kong, Taiwan,

Fl. Males. 4 308; I, (1953) 375. throughout Malesia to Australia. Distr. S. America, S. Africa, India, Ecol. In open, wet places on sandy soil,

Thailand, Indochina, China, throughout from sea level up to 900 m. Malesia. The following species have to be Exclud- Ecol. In open, swampy places among ed:

sedges onSphagnum, 600-3300 m. 4: 3696 Xyris tuberosa RIDLEY, J. Fed. Mai. St.

6. Xyris wallichii KUNTH, En. PI. 4 (1843) Mus. 10 (1920) 122.

16. X. oreophila RIDL. J. Fed. Mai. St. D i s t r. So farrecorded from Laos, Cam- Mus. 7 (1916) 121; v. ROYEN, Fl. Males. I, bodia and Thailand (type, 'Mainland

4 (1953) 372. X. malmei V. ROYEN, Blu- shores of Takuapa') only.

mea 7 Fl. Males. (1953) 307; 1,4 (1953) 370. 4: 3716 Xyris lobbii RENDLE, J. Bot. 37 (1899) 506, Distr. India (Silhet, Khasya), Burma t. 403, f. 17-24.

(Moulmein), Vietnam (Chapa), Thailand; Distr. So far recorded from Burma

in Malesia: Malay Peninsula. (type), Thailand and Vietnam only.