The Transylvanian Dinosaur Museum Project
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A Model Based on Curvatures of Extant Avian Ungual Bones
Inferring lifestyle for Aves and Theropoda: A model based on curvatures of extant avian ungual bones A thesis submitted to the University of Manchester for the degree of Master of Science by Research in the Faculty of Science & Engineering 2019 Savannah E. Cobb School of Earth and Environmental Sciences Contents List of Figures.........................................................................................................................4-5 List of Tables..............................................................................................................................6 List of Abbreviations..............................................................................................................7-8 Abstract......................................................................................................................................9 Declaration...............................................................................................................................10 Copyright Statement...............................................................................................................11 Acknowledgements..................................................................................................................12 1 Literature Review........................................................................................................13 1.1 Avians, avialans, and theropod dinosaurs..........................................................13 1.2 Comparative study and claws............................................................................18 -
An Aberrant Island-Dwelling Theropod Dinosaur from the Late Cretaceous
An aberrant island-dwelling theropod dinosaur SEE COMMENTARY from the Late Cretaceous of Romania Zoltán Csikia,1, Mátyás Vremirb, Stephen L. Brusattec,d, and Mark A. Norellc,d aLaboratory of Paleontology, Department of Geology and Geophysics, University of Bucharest, 1 N. Bălcescu Boulevard, Bucharest 010041, Romania; bDepartment of Natural Sciences, Transylvanian Museum Society (EME), 2-4 Napoca Street, Cluj-Napoca 400009, Romania; cDivision of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024; and dDepartment of Earth and Environmental Sciences, Columbia University, New York, NY 10025 Edited* by Paul E. Olsen, Columbia University, Palisades, NY, and approved July 8, 2010 (received for review May 19, 2010) Islands are noted for the occurrence of aberrant, endemic, and large carnivores on the European paleoislands. It exhibits a suite dwarfed taxa (the “island effect”). Late Cretaceous vertebrate of peculiar features, most notably an enlarged pedal digit I, which assemblages of Romania and elsewhere in Europe are classic exam- along with digit II is modified for hyperextension. Nearly 20 ad- ples of island faunas in the fossil record, and are characterized by ditional autapomorphies also characterize this taxon, many of dwarfed herbivorous dinosaurs and other endemic taxa that are which relate to extensive fusion of elements in the hand and distal noticeably primitive relative to their mainland contemporaries. hindlimb and enlarged pelvic musculature. This unique dromaeo- Fossils of -
A Fast-Growing Basal Troodontid (Dinosauria: Theropoda) from The
www.nature.com/scientificreports OPEN A fast‑growing basal troodontid (Dinosauria: Theropoda) from the latest Cretaceous of Europe Albert G. Sellés1,2*, Bernat Vila1,2, Stephen L. Brusatte3, Philip J. Currie4 & Àngel Galobart1,2 A characteristic fauna of dinosaurs and other vertebrates inhabited the end‑Cretaceous European archipelago, some of which were dwarves or had other unusual features likely related to their insular habitats. Little is known, however, about the contemporary theropod dinosaurs, as they are represented mostly by teeth or other fragmentary fossils. A new isolated theropod metatarsal II, from the latest Maastrichtian of Spain (within 200,000 years of the mass extinction) may represent a jinfengopterygine troodontid, the frst reported from Europe. Comparisons with other theropods and phylogenetic analyses reveal an autapomorphic foramen that distinguishes it from all other troodontids, supporting its identifcation as a new genus and species, Tamarro insperatus. Bone histology shows that it was an actively growing subadult when it died but may have had a growth pattern in which it grew rapidly in early ontogeny and attained a subadult size quickly. We hypothesize that it could have migrated from Asia to reach the Ibero‑Armorican island no later than Cenomanian or during the Maastrichtian dispersal events. During the latest Cretaceous (ca. 77–66 million years ago) in the run-up to the end-Cretaceous mass extinc- tion, Europe was a series of islands populated by diverse and distinctive communities of dinosaurs and other vertebrates. Many of these animals exhibited peculiar features that may have been generated by lack of space and resources in their insular habitats. -
Some Maastrichtian Vertebrates from Fluvial Channel Fill Deposits at Pui (Hațeg Basin)
Muzeul Olteniei Craiova. Oltenia. Studii şi comunicări. Ştiinţele Naturii. Tom. 31, No. 2/2015 ISSN 1454-6914 SOME MAASTRICHTIAN VERTEBRATES FROM FLUVIAL CHANNEL FILL DEPOSITS AT PUI (HAȚEG BASIN) SOLOMON Alexandru, CODREA Vlad Abstract. Latest Cretaceous deposits are cropping out in various localities of the Hațeg basin (Romania). Among these localities Pui is of peculiar interest, being the southeastern most one where Maastrichtian fluvial deposits are exposed. These terrestrial deposits are represented mainly by red beds, which yielded since the end of the 19th century, rich vertebrate assemblages. From a channel fill block discovered ex situ, a diverse fossil vertebrate assemblage was recovered (turtles, crocodilians, pterosaurs, and various herbivore and carnivore dinosaurs). This study is focused on the fossil taxa collected from this block and their fossilization processes. Keywords: latest Cretaceous, fluvial deposits, vertebrates, Hațeg basin, Romania. Rezumat. Câteva vertebrate maastrichtiene din depozite de canal fluvial de la Pui (Bazinul Hațeg). Depozite cretacic terminale aflorează în varii localități din Bazinul Hațeg (România). Dintre acestea, Pui este localizată în extremitatea sud-estică a bazinului, unde apar la zi depozite fluviale maastrichtiene. Aceste depozite sunt dominate de red beds, din care au fost colectate, încă de la finele secolului XIX, bogate asociații de vertebrate fosile. Dintr-un bloc cu umplutură de canal descoperit ex situ a fost extrasă o asociație diversă de vertebrate fosile (țestoase, crocodili, pterosauri și variați dinozauri erbivori și carnivori). Asociația de fosile din acest bloc și procesele de fosilizare evidențiate sunt descrise în acest studiu. Cuvinte cheie: Cretacic terminal, depozite fluviale, vertebrate, Bazinul Hațeg, România. INTRODUCTION In latest Cretaceous, an emerged land occurred in the actual Transylvania named the “Hațeg Island”. -
Body-Size Evolution in the Dinosauria
8 Body-Size Evolution in the Dinosauria Matthew T. Carrano Introduction The evolution of body size and its influence on organismal biology have received scientific attention since the earliest decades of evolutionary study (e.g., Cope, 1887, 1896; Thompson, 1917). Both paleontologists and neontologists have attempted to determine correlations between body size and numerous aspects of life history, with the ultimate goal of docu- menting both the predictive and causal connections involved (LaBarbera, 1986, 1989). These studies have generated an appreciation for the thor- oughgoing interrelationships between body size and nearly every sig- nificant facet of organismal biology, including metabolism (Lindstedt & Calder, 1981; Schmidt-Nielsen, 1984; McNab, 1989), population ecology (Damuth, 1981; Juanes, 1986; Gittleman & Purvis, 1998), locomotion (Mc- Mahon, 1975; Biewener, 1989; Alexander, 1996), and reproduction (Alex- ander, 1996). An enduring focus of these studies has been Cope’s Rule, the notion that body size tends to increase over time within lineages (Kurtén, 1953; Stanley, 1973; Polly, 1998). Such an observation has been made regarding many different clades but has been examined specifically in only a few (MacFadden, 1986; Arnold et al., 1995; Jablonski, 1996, 1997; Trammer & Kaim, 1997, 1999; Alroy, 1998). The discordant results of such analyses have underscored two points: (1) Cope’s Rule does not apply universally to all groups; and (2) even when present, size increases in different clades may reflect very different underlying processes. Thus, the question, “does Cope’s Rule exist?” is better parsed into two questions: “to which groups does Cope’s Rule apply?” and “what process is responsible for it in each?” Several recent works (McShea, 1994, 2000; Jablonski, 1997; Alroy, 1998, 2000a, 2000b) have begun to address these more specific questions, attempting to quantify patterns of body-size evolution in a phylogenetic (rather than strictly temporal) context, as well as developing methods for interpreting the resultant patterns. -
The Dinosaur Field Guide Supplement
The Dinosaur Field Guide Supplement September 2010 – December 2014 By, Zachary Perry (ZoPteryx) Page 1 Disclaimer: This supplement is intended to be a companion for Gregory S. Paul’s impressive work The Princeton Field Guide to Dinosaurs, and as such, exhibits some similarities in format, text, and taxonomy. This was done solely for reasons of aesthetics and consistency between his book and this supplement. The text and art are not necessarily reflections of the ideals and/or theories of Gregory S. Paul. The author of this supplement was limited to using information that was freely available from public sources, and so more information may be known about a given species then is written or illustrated here. Should this information become freely available, it will be included in future supplements. For genera that have been split from preexisting genera, or when new information about a genus has been discovered, only minimal text is included along with the page number of the corresponding entry in The Princeton Field Guide to Dinosaurs. Genera described solely from inadequate remains (teeth, claws, bone fragments, etc.) are not included, unless the remains are highly distinct and cannot clearly be placed into any other known genera; this includes some genera that were not included in Gregory S. Paul’s work, despite being discovered prior to its publication. All artists are given full credit for their work in the form of their last name, or lacking this, their username, below their work. Modifications have been made to some skeletal restorations for aesthetic reasons, but none affecting the skeleton itself. -
An Aberrant Island-Dwelling Theropod Dinosaur from the Late Cretaceous of Romania
An aberrant island-dwelling theropod dinosaur from the Late Cretaceous of Romania Zoltán Csikia,1, Mátyás Vremirb, Stephen L. Brusattec,d, and Mark A. Norellc,d aLaboratory of Paleontology, Department of Geology and Geophysics, University of Bucharest, 1 N. Bălcescu Boulevard, Bucharest 010041, Romania; bDepartment of Natural Sciences, Transylvanian Museum Society (EME), 2-4 Napoca Street, Cluj-Napoca 400009, Romania; cDivision of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024; and dDepartment of Earth and Environmental Sciences, Columbia University, New York, NY 10025 Edited* by Paul E. Olsen, Columbia University, Palisades, NY, and approved July 8, 2010 (received for review May 19, 2010) Islands are noted for the occurrence of aberrant, endemic, and large carnivores on the European paleoislands. It exhibits a suite dwarfed taxa (the “island effect”). Late Cretaceous vertebrate of peculiar features, most notably an enlarged pedal digit I, which assemblages of Romania and elsewhere in Europe are classic exam- along with digit II is modified for hyperextension. Nearly 20 ad- ples of island faunas in the fossil record, and are characterized by ditional autapomorphies also characterize this taxon, many of dwarfed herbivorous dinosaurs and other endemic taxa that are which relate to extensive fusion of elements in the hand and distal noticeably primitive relative to their mainland contemporaries. hindlimb and enlarged pelvic musculature. This unique dromaeo- Fossils of the predators -
Postcranial Skeletal Anatomy of the Holotype and Referred
Postcranial skeletal anatomy of the holotype and referred specimens of Buitreraptor gonzalezorum Makovicky, Apesteguı´a and Agnolı´n 2005 (Theropoda, Dromaeosauridae), from the Late Cretaceous of Patagonia Federico A. Gianechini1, Peter J. Makovicky2,*, Sebastia´n Apesteguı´a3,* and Ignacio Cerda4,* 1 Instituto Multidisciplinario de Investigaciones Biolo´gicas (IMIBIO-SL), CONICET— Universidad Nacional de San Luis, San Luis, Argentina 2 Section of Earth Sciences, Integrative Research Center, Field Museum of Natural History, Chicago, IL, USA 3 CONICET, Fundacio´n de Historia Natural ‘Fe´lix de Azara’, CEBBAD, Universidad Maimo´nides, Buenos Aires, Argentina 4 CONICET, Instituto de Investigacio´n en Paleobiologı´a y Geologı´a, Universidad Nacional de Rı´o Negro, General Roca, Rı´o Negro, Argentina * These authors contributed equally to this work. ABSTRACT Here we provide a detailed description of the postcranial skeleton of the holotype and referred specimens of Buitreraptor gonzalezorum. This taxon was recovered as an unenlagiine dromaeosaurid in several recent phylogenetic studies and is the best represented Gondwanan dromaeosaurid discovered to date. It was preliminarily described in a brief article, but a detailed account of its osteology is emerging in recent works. The holotype is the most complete specimen yet found, so an exhaustive description of it provides much valuable anatomical information. The Submitted 18 January 2018 holotype and referred specimens preserve the axial skeleton, pectoral and pelvic Accepted 9 March 2018 girdles, and both fore- and hindlimbs. Diagnostic postcranial characters of this taxon Published 26 March 2018 include: anterior cervical centra exceeding the posterior limit of neural arch; eighth Corresponding author and ninth cervical vertebral centra with lateroventral tubercles; pneumatic foramina Federico A. -
The Evolution of Sauropod Locomotion
eight The Evolution of Sauropod Locomotion MORPHOLOGICAL DIVERSITY OF A SECONDARILY QUADRUPEDAL RADIATION Matthew T. Carrano Sauropod dinosaur locomotion, fruitful but also have tended to become canal- like that of many extinct groups, has his- ized. In this regard, the words of paleontologist torically been interpreted in light of potential W. C. Coombs (1975:23) remain particularly apt, modern analogues. As these analogies—along as much for their still-relevant summary of the with our understanding of them—have shifted, status quo in sauropod locomotor research as perspectives on sauropod locomotion have fol- for their warning to future workers: lowed. Thus early paleontologists focused on the “whalelike” aspects of these presumably aquatic It is a subtle trap, the ease with which an entire reptilian suborder can have its habits and habi- taxa (e.g., Osborn 1898), reluctantly relinquish- tat preferences deduced by comparison not ing such ideas as further discoveries began to with all proboscideans, not with the family characterize sauropod anatomy as more terres- Elephantidae, not with a particular genus or trial. Although this debate continued for over a even a single species, but by comparison with century, the essentially terrestrial nature of certain populations of a single subspecies. Deciding that a particular modern animal is sauropod limb design was recognized by the most like sauropods is no guarantee of solving early 1900s (Hatcher 1903; Riggs 1903). Aside the problem of sauropod behavior. from a few poorly received attempts (e.g., Hay 1908; Tornier 1909), comparisons have usually Similarly, modern analogues play a limited been made between sauropods and terrestrial role in illuminating the evolution of sauropod mammals, rather than reptiles. -
The Tetrapod Fossil Record from the Uppermost
geosciences Review The Tetrapod Fossil Record from the Uppermost Maastrichtian of the Ibero-Armorican Island: An Integrative Review Based on the Outcrops of the Western Tremp Syncline (Aragón, Huesca Province, NE Spain) Manuel Pérez-Pueyo 1,* , Penélope Cruzado-Caballero 1,2,3,4 , Miguel Moreno-Azanza 1,5,6 , Bernat Vila 7, Diego Castanera 1,7 , José Manuel Gasca 1 , Eduardo Puértolas-Pascual 1,5,6, Beatriz Bádenas 1 and José Ignacio Canudo 1 1 Grupo Aragosaurus-IUCA, Facultad de Ciencias, Universidad de Zaragoza, C/Pedro Cerbuna, 12, 50009 Zaragoza, Aragón, Spain; [email protected] (P.C.-C.); [email protected] (M.M.-A.); [email protected] (D.C.); [email protected] (J.M.G.); [email protected] (E.P.-P.); [email protected] (B.B.); [email protected] (J.I.C.) 2 Área de Paleontología, Departamento de Biología Animal, Edafología y Geología, Universidad de La Laguna, Citation: Pérez-Pueyo, M.; 38200 San Cristóbal de La Laguna, Santa Cruz de Tenerife, Spain 3 Cruzado-Caballero, P.; Instituto de Investigación en Paleobiología y Geología (IIPG), Universidad Nacional de Río Negro, Moreno-Azanza, M.; Vila, B.; 8500 Río Negro, Argentina 4 IIPG, UNRN, Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Castanera, D.; Gasca, J.M.; 2300 Buenos Aires, Argentina Puértolas-Pascual, E.; Bádenas, B.; 5 GEOBIOTEC, Department of Earth Sciences, NOVA School of Science and Technology, Campus de Caparica, Canudo, J.I. The Tetrapod Fossil 2829-516 Caparica, Portugal Record from the Uppermost 6 Espaço Nova Paleo, Museu de Lourinhã, Rua João Luis de Moura 95, 2530-158 Lourinhã, Portugal Maastrichtian of the Ibero-Armorican 7 Institut Català de Paleontologia Miquel Crusafont, Edifici Z, C/de les Columnes s/n, Campus de la Island: An Integrative Review Based Universitat Autònoma de Barcelona, Cerdanyola del Vallès, 08193 Barcelona, Spain; [email protected] on the Outcrops of the Western Tremp * Correspondence: [email protected] Syncline (Aragón, Huesca Province, NE Spain). -
An Infant Ornithopod Dinosaur Tibia from the Late Cretaceous of Sebeş, Romania
AN INFANT ORNITHOPOD DINOSAUR TIBIA FROM THE LATE CRETACEOUS OF SEBEŞ, ROMANIA Stephen L. BRUSATTE*, Mátyás VREMIR**, Akinobu WATANABE***, Zoltán CSIKI-SAVA****, Darren NAISH*****, Gareth DYKE******, Gregory M. ERICKSON*******, Mark A. NORELL******** Introduction Since 2010 the Transylvanian Museum Society (Cluj-Napoca, Romania), the “Ioan Raica” Municipal Museum (Sebeş, Romania), the University of Bucharest (Romania), the American Museum of Natural History (New York, USA), and the University of Southampton (UK) have collaborated on a project focused on the vertebrate paleontology and geology of the Late Cretaceous of the Sebeş region of Romania. The aim of this project is to better understand the peculiar dinosaur- bearing faunas of the European terminal Cretaceous, which included bizarre dwarfed and late-surviving relict species that inhabited an ancient island archipelago.1 The most notable result of our project thus far has been the discovery of the aberrant new dromaeosaurid theropod Balaur bondoc, a close relative of the iconic Central Asian Velociraptor mongoliensis. The type specimen of B. bondoc was discovered by M. Vremir, described by our joint Cluj-Bucharest-New York team in 2010,2 and later monographed by our group.3 Here we describe a fragmentary, but intriguing, new specimen collected during fieldwork in 2011: the tibia of a small ornithopod dinosaur that may have been less than one year old at the time of death. * Division of Paleontology, American Museum of Natural History, New York, NY, USA; and Department of Earth and Environmental Sciences, Columbia University, New York, NY, USA; e- mail: [email protected]. ** Department of Natural Sciences, Transylvanian Museum Society (EME), Cluj-Napoca, Romania; e- mail: [email protected]. -
The Dinosaurs of Eastern Europe
The dinosaurs of Eastern Europe Michael J. Benton (University of Bristol) Bright Horizons 10 Thursday, October 6th, 2011 Discovering dinosaurs n Now, we know dinosaurs from every continent on Earth, including Antarctica, but that has not always been the case… n Dinosaurs were first found close to where scientists lived – Europe and North America – and then further afield n The dinosaurs from Eastern Europe are a surprising lot – many of them were dwarfs, and they were studied by some rum characters, including a man who failed in his efforts to become King of Albania, but spied for both sides in WW1 n Let’s explore how palaeontologists put together their understanding of the diversity of dinosaurs worldwide… and explore what they tell us about ancient geography and island effects… Scrotum n The first published record of a dinosaur bone was in Robert Plot’s Natural History of Oxfordshire published in 1677 n He identified it as part of a thigh bone from just above the knee joint, and he thought it came from a giant man or woman n Named Scrotum humanum by Brookes in 1763 - (un)fortunately determined as a nomen oblitum - otherwise the first formal dinosaur name Gideon Mantell • Mantell (1790-1852) was a working country physician - struggled against prejudice from other palaeontologists who did not have to work for a living • Named Iguanodon in 1825 and Hylaeosaurus in 1833 Richard Owen • Owen (1804-1892) had to work for a living, but based in London and established a strong position fast as Professor of Anatomy at Royal College of Surgeons