'Ja,{\oU 'ectur.rllz IInJ ur ueq,!\relncupeds eru suorlzpdodpue umlJnqaluotl3 eql dn eqeur 1eqt ereue8 eql Jo srea.og F;:noloJ tsotu pue 1se3ru1eqt s?q snua8 eql spled 'slueurBas Uos o8rel pu€ unlqluzd,{q drruq,(1pse xf,pc A1 Jo ecuur€edd?er{l Suhr8 peqolrq,tlaleep '?lpour,ul?1s qedeseql ou pu?suaulsls l'eJfed 96 Suprequ1e.reue8 esoql uroJJ sJeJJIp { ?qolpro?luat "Jsa(f pus e8pnu olulh.tDe wn?rnqawoq) 01 pe&]p.t,{lesopsr puu (986I s33ug ? uosur{of '616I uosuqol ? s33ug) ecueqp wnlcnqawoqJ eql Jo reqluelue^Dcuqsp frerr. e sm[uoalld

'snue8eql u1 pespEocar mou 'suone.qsn 'suopdpcsap sercedslq8re aql ro; {e1u puusdeur uoqnqluslp l aslcuocsepplo.rd.redud qq; 'pesluEocu Euleqsercedsqns ,t eu u puz selceds,reu:no; ur polFsoJolurl prrus/,s oJruJal d elulsurerol 'snueE paaueql pel?llsuoruep e^Eq lu€qcJ€I,^l C'N r0r^\uowJoq€lloo ul .{nJed eqlJo selp$s lueceu

'pelsleJ{lesolc lou Flq€qordore tuql sexelduocsolceds luoroJJ;p ,{-reluro;; selcedsgo pespdruoc are pue seuro.rpu,{s uopeullod luueggp .{lepuessesdeq;ed erz suopcas '(s!.nryrunpad 'd 'srypq 'd 'snclJuornD eseql p:fe snpuruorqnrd 4 Sulsudruoc,r\ou) papl Tpun suerrjglsaql qll^{ sfl{,uDafdouow txespuv (snso)rura^'d pw snlolt{'d'srlguouluatdas 'd 's!Jow!l 'd Sursuduioc,/t\ou) pap D sueuI€tsqlr^{ l'3ant wou snpuoaydngsE) snryuoa\d tces'suolceso,{l teselp otul snua8eql pepvlp (t061)a4un) ,{ur4uou,{sol'srsaJtrrr^ d Joeuo pecnpeJ uruqluag lnq ',,slsuelp.qsnverold,, u1 snueE eqtr go luaulBaJl (r981) s,urErtlueg otr .ropd peqpcsap uaeq peq selceds mod €l1e4snvuelse N uJeqlnosol cFuepuesqmls snoJeJIJoUJo snueEgerus e \ snqJuoalq

uo!lcnpoJluI

'pe]€lsuleJ sl enntrd 'peqF 'dsqrs snsoJllua^ d pu€ sepere (tnr4?rc;1sual11td sl.nlncunpad7) selcedsqns.reu euopue '4 'tnq?nll ',{:eq31a; ([nq?re>I slpuotttuardas d ptre frarlSo>Isflpltluorqnr sry1aq4, s r!truuno 'pesmSocer snqtuoall4)saoeds /llLeu mog sercedsqnso,,!\l s?q qJrq,r Jo euo eje sercedstq31g pe,neFe: '?ll€.nsnv ' sl uelsed! uJeqlnosol clrrJepuesqnqs snoJeJIJoU;o snue8gerus e lTqz'I sm{uoa\d '(ZO\d I9-Lt'.0)91 otsl(nN (e?eJ?u,{nl)snqJuDafd snue8 aql Jo ,rar^erv ,{req3}e) IO

lJBJlsqY

S909€ll?Jlsnv uIalsaA'oorauuelA'19 xog Od'alueJ qrr?aseueJrlpI/A lua(ua8eu?I,\lpup'I pu? uory^resuoCJo tuauruedeo

,ftaq8lax 'f 'C

(auaee.r,(tr41)snqtuoald snue8eql Jo ,leeper V

LE (ZOOO\9- LEtrJ)9| D!sltnN Nr)fria Vol. 15, No. I (2002)

Materials and methods

Herbariumcollections ftom PERTH and most major Australianherbaria were examinedduring the course of the study. Type collections and/or photographsof type collections were examined ftom relevantherbaria. All measurementswere taken from dry pressedmaterial supplemented by observations of living plants in the field and cultivation.

The conseryationcodes given in this paper arc thoseused by the Departmentof Conseryationand Land Management.An explanationof thesecodes is given in eachissue ofN&)tuia.

Taxonomic treatm€nt

Pileanthus Labill., Nov. Holl. Pl. Sp. 2, 1I , t. 149 (Feb, 1801).- PileqnthusLabrIl. sect.Pileanthus las sect.Eupileanthus nom. illeg. azT. von Post& O. Kuntze,Lex. Gen.Phan.438 (Dec. 1903)1.Type: P ILe ant hus limacis Labill.

Pileanthus sect.MonopiLeanthus Kuntze lfl T. von Post & O. Kuntze, Lex. Gen. Phan.438 (Dec. 1903). Type: PiLeanthuspeduncularis Endl.

Srrrbs to 3 m tall. BranchLetstercte,brown, with oil glandsepidermis becoming grey andshedding, with persistentstipular outgrowths often presentin the leaf axils. Leaves opposite and decussate, sometimesin denseopposite clusters, sessile, linear to clavate,with prominentoil glands,margins entire to laciniate.Floral leavessimilar to vegetativeones except in P. Limacis. Inflorescenceof solitaxy flowers in upper axils. Bracteoles2, opposite,united at baseinto a cup, enclosingthe bud, scarious, usually promin€ntly umbonat€,often viscid with a white oily exudate,free portions above the cup caducousl cup turbinate, around the hypanthium. Flowers regrlar, bisexual, large. Hypanthium campanulate,pilose outside, adnate to ovary for most of its length. Sepals5, erectto spreading,deeply bilobed giving the appearanceof separatelobes, usually sparselypilose. Perals 5, soft, exceedingthe calyx, often brightly coloured,margins usually shortly dentate.Androecium of 20 stamensin a single row, with onestamen opposite each sepal andpetal and the rest alternating betweenthe sepals andpetals, united at the base, antipetalinestamens olten distinctly longer than antisepalineones, the anthers sometimesalso distinctly different in size; free filaments slenderbut dilated at the base;anther cells parallel, opening longitudinally, either attachedcontiguous to the thickenedend of the filament or separatelyattached to the branchesof a forked filament. Oval') l-celled, with 4 or 8 ovules in 2 rows on an erectfree basalplacenta. Sr1,/e simple, filifom, terete;stigma small, terminal.Frzit usually a 1- seedednut formed from the hardenedbase of the hypanthium,dispersed as a unit with the dded upper hypanthiumand sepalsattached.

Chromosomenumber, n = ll hasbeen recorded for four species.There areno recordsof polyploidy or dysploidy in the genus(Rye 1979,.

Distribution and habitat.^fhe genusPileanllras consists of eight speciesendemic to southemWestem Australia. Members are found on sandy soils in the Avon Wheatbelt and Geraldton Saldplain Biogeographicregionsofsouth WestemAustaliaandextend into theadj acent Camarvon Biogeographic region of arid Western Australia. There is a concentration of species on the northem margins of the Geraldton Sandplains. This is a very different pattem to that shown by the related g enera Chamelaucium '61.61 '9i61 'oulxv rec 9'.I']ensettfillo ASStU{ Z :(HJUEd) r auplD) V) JeC :(HJUgd) 868i llupDlg g Al11y61 dr5 y7'sffur:dg cuql:yl1ya1snv NUAISEI& pautwoxasuautcads paltalag

(C V I e.rniu) 'snonJrdi^uoJur'esoqo18 '8uot rru8rls :snorq?l; ruor g t a|(,?S t talr,1o Suol urul I J stuaul?lrl eurledasque:3uol ruur Z g l stueGrBlrJourlqeduuE :stuorutlrJ poqollqqlrr wn))orpry etBtuq)-eterurJEl 'esuel 'epr,tt '3uo1 'Surddppelo urF:tllu orLu71 6 Lurugt ZI slriad esnlqoxedr'snouuqueur 'eprstno '?lr]rurJPl'?lruts 'rpr^\ '8uol sur8Jeur'eprsursnoJqrJB osolcl-,{Ills rum t-[ urru 9-t'eleJuqtur '3uo1 'etr:lnuudrucr,{lpeoJq sTrrla5 esolrd ruur g g LuliqtltDdiH 3uo1ruu 7 1 oqrun luefund c qtr,tr 'peuop-purl.Q'u,rorq '3uo1 ruur g g suoruodaeil:3uo1 tuur g g'peqollqdno:r1Dd aloapD.tg snoJqul8 'uorlJcs^-sso.ro 'peltop-puulF'opr,m ur rrldrllool-rulnxjr)'Fuol ruur 0z L al)unpad alni)1]xsdn urur l ,:r '3uolrutu(02)tI 'osEq 'Ir:rour'porolsnll /(9) tr]pouallBLl lnq snoJtcnbu] 1 ,(lesueplou .r;7ir):r7tueulqc]r]1r? 'pe18ur 'posmJ'lueuruoJd ,no1eqpaiurn lsoLup t sereue.t.trceplrel :spur[8 Surppeqspun ,{er8 'spue18 'u,roiq 'sneJqrl8 Surnroccq Jropcsru.r lr:luorsr:rJJo qtr^\ s;,t1.tuo-rt1Sttnotr q8rq ur 1 o1rini45

'(-lat{') il)rri).,Zt ou'lloi qt9fuo trunre I ' 'lrr '.rc^ 's8u I EIiI-0gil I lr.rtsnV urctsclA rU rlt[us^^\o.r]V rlcu udS llrs u.rnqloJ| ,, )-SrrudSauinoqloJ 'rri^rehl edo.rclc"-oueJu arlruulcl ur,, 'xliv_ (LS8l) !f :l log')ot uLltj / snlloJllusnrlluBalld'I

'd 't- 'elrq,r sgleuoglualdas 3uo1nrur t t slulad opr,\\[rur t 0 r sa^rc'l , slJerull'd't Sur';urur y ., 'c.rlLroi .L po,ro1 ytrrcl qlrm olrqm ro lurd oluclslrlod opr,r ruor I ,) sc^l?c-l '1urclolrcl .ro elrqm srenolg 'I snsoJluJa^'d'z lLrnlqoqLLrl c1ocl:rurg ""uo1 urLU I I il slulcd pc.rs.rc,\\olc 9 snlloJllg'd'l " " " " polurocloqu.rn oloatrnig ';uol rulu ql-Zl sllrlod osr.roJs.ra,{\olC g 'ltcl lo osr.roJs_ro^\oljl I 'slucLu?ll.l pol.roluo sllooioqluv I snolluuJnc'd'S snorcry1;-e1{tg oiullo s.rc,ro1g g 'd ', " " " " " " " " " "pe^inJur snprlruoJqnJ sialllur|aurltcleJ pal s.ra^\olJ t snllaq'd'9 l)era s-reqluneurFdal lurcl sJe^\olJ t 'oi^uq ]c ,(rrrq 31,{15 per ro lurcl sre,rolC I t scln^o llcrclsrl[]lou so^ua'l z 'd '8 'slelqcul)lq slreln)unpad 8 seln^O eplslloqs uo PaJal.-nli)so^llc'l Z 'slueun:111pedrqs-qnlJuosnon8rlroJ sllel.reqluv I

snqtuaaFd ro snraodsol ,{cX

'(266l '(.:^n!4DllP uotsnoH) r-u urls poqolrclsDq Lptqt J) sorJaclseuo ur petueuncop uaeq sl?rl uorluurllod eeq crlr:reds frc,y "ttqttutap4 ur uedduq1ou saop srqJ uedo s.re,rol]eqt c.rqaq e1{1scql1o pue eql tr: rolucscld uclpd r otuo 1c1clo.rp,{1ro ur ur uclpd rroqt1o uorlrsodapeql se^lolur DlpJo)utaA pu12Dlutt\\JD(J'LLtnDnDpLLpLl) rrauei pelrJe,reql ur euro,rpu,{suo4lrurllod eq]- {3olo1(l uoltD lllod

.pnq je^\oll aql Jo selrnlrr:q Surgrlesop aql ol 3ur..r:a1er's..re,lroLJ - sorylu, pur drr - r'oa7ir1'1ool9c\t utoJf [Soloutl]g

'llaqlceq^\uo^V eql ur selueorourlu qlr,{\:'urrldpu?S oJucred"-g cql uo pue sureldpueguolplucg uJcqlnosoql Jo EoJeJnonsc-1 ttrAI aqt uo csre,rrp-sereedsere qartql\olLut .to1 p\e

'{r.q;r.) (au3]IlIK) rr'trlrrirJ snu:t :qt to ,rar^ V fC Nr)tsia Vol. 15,No. I (2002)

',,$ffiffiI EhD

J

ifi

Figure1.A-D. Pikanthusrtldolius. A - leafand flower, B - bmct€oles,C - stamens,D-style; E J. P lir??acis.E-leaf andflower,F-leaf{iom below,G-leafftomside, H -b&cteoles,I- stamens,J-style;K-N. P.septe tio alis.K-flowerinJg branclilet,L - bracteoles,M stamens,N - style:O-S . P.1)emicosus.O-leaves andflower, P- leaf,Q-bracteoles, R- stamens, S - style. Drawn ftom CA. Garhrer 5 Jar..1931 (A-D), S.J. Claymorc& A.S. Westn264 (E-I), C.A. Gardner3184 (K-N) andA.S.Georqe 16949(O-S). Scalebals for bmcteoles= 5 mm, for flowers = 10 mm. 'seroods srqtJo s3^[el urojrtrj eqt ol 8uu]e]er '(+uel)Lulltlof pue (pB?'Ilt) run+Jl4te'I aqt .u].otgt SoytuttlE

's{JEd leuortENe^rJ puEselreser oJnlru lq8re t"-re1tr:o1 pepro.e6 pe,^lesuo:l11e,t put: pte.rdseptl6 sntotsuollzMasuo)

'f;unue1 ol lequreceq fluwru 3uue,rolg pott;;d Suuanolg

(VZ einSu) qperl re.\o aellrhl ,u7r?lpol '\B"r.I 'pul?s sntd,llnnl puBpljIlpoo^\ ^\ol lr?slil?r6r DuDu looLlDlqrrB, ur JDuellrIlo pupsuo s,l^oJD 'urFurD 'ere8uoq lo quou ol qtnospue ,raue8ur4 o1puelur uoo^\taqsinlllo Itl?./DL!pLtDuolinquit^lg

'(HIdAd) '9961 ',ncuc8urlrJ LtlDWU V ,t0y g :(HJUAd) t8SI ptr{1y 1ptuat131a;1t D'Lg6l ur,lZ.-'ur8urg1og'e,ueseX 1le/\\ staltrlrg:(HJUA& I99Z ullJiD VI

..\ns0jll1.u'|'./g:.st1Du|)t'l,Dkl)|'..]|)|||,'.|t)t)ul]].]..|lr]1]]]l!!]UD) SV

I LOqN I f rT In€ Aluo{.s tturi{une \r

quorPlDr.C i\\ la .\t: h\ f h\N

(

"! r2

(.?.rrllN) s,?r1rlr,rr|dsnui; ,qt lo $.r^er V {r3qir.) I D M/)/rid Vol. 15.No. I (2002)

Notes.Houstoo ( 1992) documentedthe poll ination of this speciesby the monolecticbee, Eun gLossa semaphore.Further studieson the pollination of the gerus are underway.This specieshas the largest flowers of the genus.

See fhe notes given under Pileanthus peduncularis regarding the application of the name Chame Lquc ium dil atat un.

2. Pileanthus vernicosus F. Muell.. Fragm. Phyt. Austral. l, 225 (Dec. 1859). Type citation: "Ad flumen Murchison.Oldtleld et Walcott." [MurchisonRiver, WesternAustralia], OldJiekl (.s1,n: MELI).

Upright slenderslzrab to 1 m tall. Yotutg,brtncltes brown, oil glandsproninent; leaf decurencies obvious.with redtriangular-shaped stipulcs in axils.Zeaues clustered on shortopposite shoots, linear- triquetrous,3-5(9) mrnJong, c. I mm wide,oil glandsprominent, margins and midrib with prominent tooth-likc hirirs,apcx runcate to acute.Fbrul let|es similar to vegetativeleaves bul not as chlstcrcd. Peduncl.esleotier,l - I 5 rnm long,with prominentoil glands.B rac:teo[e pair: cupc.3 rnm long,l-2 mm wide: freeportions 3 5 mm long, with midrib not prominent,mar-gins entjrc, umbo not obviousbut apex trreathickened with proninent oil glancls.Pedicel 2 3 mm long. Htlttutthium ca]npanulate,3 4 mm long,c. 3 mrndiarr., si)ky-pilosc, adnate portion 2-2.5 mm long.Se2als oblong, c. 3 mm krng,r.. 2 nrrn wide,with scattercdhails becoming silky-pilose towards base, apex obtuse. P?/,rlJ 8-ll mrnlong,7 9 mm widc, rcd wilh bascpinkish, margins Iaciniate, apcx bliLn'rt. Androet:iunt: antipctaline l'ilamcnts conspicuouslybilobed , l- I .5 rrrr long;antisepaline filaments less prorn inently bilobed, less than I nrm long.Ovules 4. S4,/c:2-3 mrn long,glabrous; stigma globose, inconspicuous. (Figure lO-S)

SeLede(L.\pecimens extLrrincrl. WESTERN AUSTRALIA: CapeLesueur, Percn Station, 9 Nov. l9ll2, R.J.Cranlielel252l (PERTH); Cooloornia Nature Rcserve, l9 Sep.1979, S.D. Hol?er 1385(PERTH); 40 knr N of Gerrldtoron NorthWest Coastal Highway, K/i Kenneully4707 (PERTH); 392 mile peg 185krrl N of Geraldton,l8 Sep. 1974,11.L. PoweU 74140 (PERTH)I Kalbarri National Park, R.C l/erzr I (PERTH).

Distribution urullutlrir./I. Occurs between Geraldton and PeronPeninsula, Shark Blly. Grows on rcclor yellow sanclin Banks ia st:eptruush.l rbl,,tnal, Trec heath,constal heath and Acacia shrubland. (Figure 2B)

Fbwering periocl. Betweenmicl Septembcrantl early Novernbcr.

ChrcmosomenLulbet. n=l1(Rye 1979),voucher B.L. Powell11140lasPileantltu.t sp. 1).

Etyrnobgy. From thc Latin ye,?i(o.r.l'r aboundiDg in varnish,plobably rcfcrring to the shinyoily coating on the youngfloral buds.

Conservqtionstatus. Restricted in occulrencebut wcll conservedin Kalbarri National Park,Zuytdorp National Park and proposedreselves within the Shark Bay Wolld Hedtage area.

Noles. Closely relatedto PileLmthusJ'ililb lir.r, a northemleplacement for that species.A singlerecord notesplant size as 2.5 m high by 2.5 m across. pep^\oJcr'rnra7 spulz18Suppeqs pue fer8 Sunuoreq'pe op-puEI8'u,$oJq-per ,{le.rrJ ro scr?uarn.ep 'lT4 1ua1,{q uepprq saqcuotq 3uruttr epr,r ru I ot ruJ 0B ot qnrlls alu..rtso:d;o Supre;ds ,to1

'(iIC) IprlpJlsnv uJatsald '3 . ur^\nee'I-ur^grrol ur,. ird,{., (908I qac)6tT 1'T I d5 14 1pg .,to1'1'llrqu'IslrBrullsnqluEalld't

'sa1nd4s Sur,requr suouroedsleqlo eqt uro.gsregrp gy9.ia3rlpg T o trql etoN se^ealunD eql pul] sluled Su rddul.re^o uou qlr,r sJa,{\oUctrq,,d:al1nrus etp u t Suueglptnq s?J?ri.,l?/drot pat€ter'{losolJ lzloN

'snueBerp go requeu lsoru uieqtJouaqt sr s[lJ uroqlnu lol plo,\\ ulluJ B ruo.Upe,rucp st lcqltde eql- iSopa!\iil

'SSAJOSO}ICJnll]N roru.rog pul] elloo'SIn1d 'eiutg 1nuoqe51 e8url ,{peuua;1 edu3 urtuosoJdpc^.rcsuoc 11e,tr ptn pneldseptl6 \^nlDtt^LtottD^.o''uo)

'| '(6t6 st)Duttl J-lt1ds |nLllw)alld rlt ) gZOtL lpao.I Z 6rrcqcno^ | o^U) | l=Lt .1)qLtttllt )tLtosot.ttor4J

'Jaqolog ol yn3lny ltotL;tdiiuuatutlg

(VZ oin8rC) seunppms pcr uo lloc-IJIJ clu'l 'o.aunu -K)eprs u.rcls^ull cql Lro ,{Pouua) oql pul?ciuPl cch:3Lloc,tloq sjt]ooo ,l/n/tltltl l|uD uoltnqutstQ

(HJdsd) 'Lro^r?u.nc,lo 'I.nd gzotLll)t14od7f1'!16I qcc i I N LLrIcot xc stur) 1rPon?^rllnr:(HJUad) Izt I I ' 'aiuru 'LpLUrpur?S ,iaqwp y y 2,91118nV IZ ,{pcuuo) rt\.to /AN ul qt :(HJUEd)tlll )'i.kDD S'V '0961 flnV It'qtt()Lu.c't.loSlu)t ZI I salrlull :(HJUgd)fl\t.laup.|DD V)'Zt6t ""rry97'1o06 '8gfrI 3uruur16.ncu'uorlDlsrloo8odA:(HJUA471y9talutp-g1'g dasZ {r,\\q8rHlulsr?oJlsc^A t0roN uo osnoqproucltl pr.rJugto iNSiA nrl Zi :VI'IVUJSnV NUAISEL^'pdulLuD|a su)Lut.)a.ls Pal)aPS

(N-) | 'olnurLu 'fluol '3uo1 'paqoltq oJnFrC)csoqolS DLuirls:snolqcln- tutug ot ,/.!5 y sa1r,rp ururg 1ot {ldoop lou slueurulrlaurludosrluc :3uol LLrur t ol'peqoJIq slueutelr.l oLlluochlu? rr,/r1Jrr-l/rry oll!uop-eltruIcnl 'e1r 'epr,ll 'fluol '3u 'cprstno xadu q,n ururE 3 urury-g rddupe,to 1ou sprrd elDluepsurBicut crlsJILI'cpr.{\ 'olnsrtq-c1tq,n'3uo1 urug r'3uo1mur€ J'snoartl.mqcsl?rrl;rg spuul8 lto tpt,{\ nru El g-9 611.rcdaa.tl 'epr,r 'pcuop-Pultli urur; ot'3uc1 urtu g ol urnttlltuttltll Suol ortu I g 0 oclurnJcJIUoJ luc8und u qtrm 'u,trorq'3uol ,{puourruojd mruL ol suoluodcaJ}:3uoIurLu j7oldrJ :JtD.lalozt:rD.tg spurl31to lueututord '3uol 'spurlS 'ePL\\ qll,r ruu (El)lI gft) )pu pad osnlqoxcdt lro pasn: lueutuo.rdqlr,\\ tultl t 'l '3uo1 'osuq 'Jccurl uru I t "-prp,{\otpeueuuu lnq sno;lanbu r'rnra? seqJuDlqleplo uo ]ueuturojd 'suorsnJlxe '.llet 'r7r,riys rclnpuelSqtr,n u,uorq clncls aLlJLtDrg epti \rr I ot uI g l o1 ucdo Sutpecld5

'(tzttoEzo'892n0920 "lEW'SNVJ HJdgd ..^r t8LZt09Z0HJdAd iolotl) 'Prlt?rlsrlv ']:rloo8oy'A lglt rauplDu \/'J'zE6l lsn3nv8z uletssiN'1oo43n[rur16 lEou .rr./,t

'prrl 'n8uol u[u t-Z urur t-t 1]lp]uqoll-uou 'p8uol '€IffeurT cluted rtel u[u t 0 Luur8-l trlenbu] €IIoC sntel!u t ta sntlEtu I BJrrJ snsnltaselnrC

',rori ds',{;eq8rey qpuol.rlualdes snqluealld'€

(.i.rnrll,,i)rr?n/rrlrd snua6 eqr Jo .^\ar^ V naq8ra) rD ,V lrsi,4vol. 15,No. I (2002) at endsof branches,5-8 mm long, c. I mnl wide, coveredby oil glands,adaxial surface flat, adaxial curved, margins hyaline with small tooth like projections,apex obttse. Peduncle4 8(15) mm long. Bracteok pair: cup 2 3 mm long; free portions,1-6 mm long, 4-5 mm wide, brown, with conspicuous oil glands, soon caducous,with a more or less globular umbo c. 1 mm lor'g. Hypanthium broadly campanulate,4-5 mm long, 4-5 mm wide, silky-pilose,gland-dotted. Sepals deeply divided, lobes 3- 5 mm long, 4-5 mm wide, slightly pilose, apexobtuse, dentate. Petals overlapping,c. 8 mm long, c. 6 mm wide, white with cente flushed pink to red, or pale pink, margins laciniate, apex rounded. Androecium: antipetalinestamens bilobed, to 3 mm long; antisepalinestamens not deeply bilobed, c. 2 mm long. Ovuks 4. Style c.5 mm long, glabrous;stigma minute, globular. (Figure lE-J)

Selectedspecimens exanrined. WESTERN AUSTRALIA: Blowholes, QuobbaStation, 18 July 1964, J.S.Beard 3480 (PERTH); Waroora Station,near homestead, Sep. 1962,J.C. Malone s.n. (PERTH); Dorre Island,Disaster Cove, 15 July 1959,R.D. Royce5876 (PERTH); N end of Bemier Island,5 Oct. 1947,D.L. Serventys.n. (PERTH).

Distribution and habitat. Occurs in near coastalsituations and offshore islands from Shark Bay to Waroora Station,on coastaldunes usually over limestone.(Figure 2A)

Flowering peiod. July to October,depending on rainfall.

Consenaion sraus. Widespreadand not endangered,

Etymology.From limacis Greekfor slug, owing to the shortfat leaveshaving a resemblanceto a slug.

Notes.Although often noted as having white flowers, this speciesnormally has pale pink flowers or white flowem suffusedpink ]lnlrke P. septenftionaliswhere they are invariably white. Pileanthus limacis andP. septentionalis occsr entirely within the arid CamarvonBasin, one along the coastand the other inland.

5. Pileanthus auranticus Keighery, .rp.nov.

Futex erectus,2.5 m altus.Differt a Pileanthobello fo\is 10-17 mm versus4-12 mm longis.Flores auranticis,umbonibus curvatis longis pungentibus.

Typus: 30.1 km west of North West CoastalHighway on Vermin Proof Fence,27"15'S, 114"19'E, WesternAustralia, 21 October 1995,M.N. Lyons 2319 (iolo: PERTH; isor CANB, K, MEL).

Upright woody sftrab to 2 m tall, denselybranched at base.Young branches with raisedoil glands and occasionalhairs; leaf decunencesprominent, becoming grey and shedding.Zeaves not densely clustered,linear, 10-17 mm long, c. 1 mm wide, with raisedoil glands,upper surfacedeeply grooved, undersurfaceconvex, apex actte. Peduncle filiform 15-20 mm long, with prominent oil glands. Bracteolepair: cup 3-4 mm long; free portions 9-11 mm long, greenishyellow, with prominent oil glands,with a curvedpungent umbo 3-4 mmlong. Hypanthiun narrow-campanulate,5 6 nrm long, 3 4 nrm diam.,ftee pafi c.2mmlong. Sepalsdeeply divided, lobes3-4 mm long, 4-5 mm wide, slightly pilose,apex obtuse, dentate. Petdls overlapping, c. 8 mmlong, c. 6 mm wide, orange,margins laciniate, apex rounded.Androeciwn with entire filaments; antipetalinestamens c. 2 mm long; antisepaline stamens0.5-1 mm long. Ovules4. S4,lec. 5 mm long, glabrous;stigma minute. (Figure 3A-E) 'ruru 0I = $eAog roJ'urul S = seloap?rqrcJ s.rBqolers (I-d) 196I'rc] ttunH uIaO S )) 08€6ar{oy O y'(H) 6829tlaupo)'V)'C't'D' 920n to ag Ot'g(g v) \BEZsuoq NtI ruolJu^\urc 'sueuels ... -O'Ie,1dou - 'sueu?N S'seloolcprq-- d'Jeel pup se^eel- d snpllaotqud J-d:el,ts O N'saloalr?Jq - IN 'ro^\ou 'sueurqs 're^\ou pueJeel- X srrrlfiru pad dsqls suolnxmpad d O-) :el,&s- I - I'seloep?lq - H SeeI- C pue '!n 'E aq nqtuoald l g'.el.{ts. g'sueulPls O'seloepprq C'J3el g're^]\oupup sa^eq V sficltuDntod A-V Ifl [ls ll I W I il \mAnd Eu"Vq]" Hj\1 l,/'n a wf{ IA\ /\U MA rffi

I fi t1 i1

(eEe.Erf.t,) snqtmafd snue8 eql Jo ^\er ar V',ftoq8ra){ ID Nr)trld Vol. 15,No. 1 (2002)

Other specimens examfued. WESTERN AUSTRALIA: near northern boundarJ of Murchison House Stationon Vemin Proof Fence,29 Oct. 1986,A.S. George16848 (PERTH); 32.3km W of North West CoastalHighway on Vemin Proof Fence,21 Ocl 1995,M.N. Zlons 2380 (PERTH); 32.6 km west of North West CoastalHighway towardsZuytdorp Cliffs, I Nov. 1985,P. Roberts755 (PERTH).

Distribution and habitat, Found on the north-west comer of the Eurady sand sheet,in undulating sandplaindorninatedby Actinostrobus arerxarlus shrubland over low heath.In this areaall collections arefrom interdunalsites on yellow sandover yellow sandyclays. One collection hasbeen recorded on red sandover limestonein low mixed heathcloser to the coast.(Figure 4A)

Flowering period. Recordedflowering in October.

ConservationJtaluJ. ConservationCodes for Westem Australian Flora: Priority One. Has a very restrictedrange and populationsare entirely on pastorallands in Murchison HouseStation. It is highly likely the speciesalso occu$ in the adjoining Shark Bay World Heritagearea.

Etymology.From the Greek a ralx/ic&sfor the orangeflowers of this species.

Norer,Closely relatedtoPileanthus bellas, differing in the longer slenderfloral leaves,curyed pungent umbo and orangeflowers. The two speciesdo not appearto occur togetheror overlap in distdbution. Pileanthusbellus occurs on the tall yellow sanddunes of the Eurady Dune System(as doesanother endemicof thesedunes, Vetlicordia coolomia) which occvs to the eastof the rangeof P. auranticus.

6, Pileanthus bellus Keighery, sp. rou.

Frutex erectus,3 m altus. Foliis 4-12 mm longis. Differt a PiLeanthoaurantico fToibvs rcseis, umbonibusbrevibus pungentibus.

Typas. nearnumber one tank lNoth West CoastalHighway, just north of MurchisonRiver crossing], WesternAustralia, 2 November 1965,C.A. Gardner 16289(holo; PERTH 04365550;lsor CANB, K, MEL).

Erect shrub to 3 m tall. Youngbranches brown, with prominent oil glands;leaf decunenciesnot prominent,becoming grey and shedding,with large stipular outgrowthsin the leaf axils. Leayesnot denselyclustered, linear, triquetous, 4-12 mm long, c. I mm wide, glabrous,with prominentoil glands, adaxial surfacegrooved, abaxial convex, margin with scatteredhairs but not hyaline, apex obtuseor acl:te.Peduncle slender, 10-15 mm long. Bracteolepdir cup nafrowly turbinate,2-3 mm long; free ponionsH mm long,brown, margins minutely denticulate, with a pungentconicalumbo c, I mm long. Hyp&nthiumturbiLate,3-5mm long,3-4 mm wide, silky-pilose,gland-dotted, free part 1-1.5 mmlong. Pedicel24 mmlong Sepalsbilobed,24 mm long, 3-4 mm wide, shortlypilose, apex obtuse, margin minutely toothed,fused for c. 1 mmatbase,Petals 7-8 mm long, c. 8 mmwide, pink, baseerose,margin e\l]le. Androecium with entire filaments; antipetalinestamens c, 2 mm long; antisepalinestamens c. 1 mm long. Ovules4. SDle 4 7 mn long, ciliate at base;stigma minute. (Figure 3F-J)

Selectedspecimens erazrlned. WESTERN AUSTRALIA: l0 km NW of Nofth West CoastalHighway along ffack to Z bend, Kalbarri National Park, 30 Oct. 1996, B. & D. Bellairs 4026 (PERTH); 15 km =-

s|p4tuo.tqu d l:V rr,rrli?rd dsqns suDtlnunpad d puq. st.tDttliurp.d dsgrls tuDt]i.|npn d g:.r,llrq ./ pue f tniuonD tnltrunalt.J V sdnu uoDnqulsr(I i 3rn3rC

I I \ r+/ ri I Arcr*es i tt a \ t a

a. a

-LS:M H.LNOS 3H1 -\ vr'tvd-LsnvNaltstl x\N l \ 'Ez-t- .-t- l--t-. )) ttl V \.i,,,, -- l---l--/ | t ,v) I t (

(.'r.3nr,{J^t)sr{nrrrlrd snu.; .ql lo ^{.rri.r V ,tr.qira) f C 48 Nryhil7 Vol. 15.No. i (2002)

N of Murchison River Crossingon North West CoastalHighway, 12 Dec. 1984,H. Demarz 10489 (PERTH); 410 nile peg on Nofih West CoastalHighway [157 km N of Geraldton],30 Oct. 1963,n Lulffitz 2848 (PERTH).

Distribution and habitat. Mainly confined to the Eurardy sanddunes, where it occurson tall yellow dunes dominated by Actinostrobus arenarius shrubland,but also a disiunct southem record from shallow grey sandsover sandstoneemergent from low heath.(Figure 4A.)

Flowering period. October to December.

Conservqtionstcttus. Conservation Codes for Westem Austalian Flora: Priority Three. Currently of restricted distribution, but present in Kalbarri National Park and adjacentuncleared pastoral and unallocatedCrown larrds.Probably not at dsk, but requiring further survey.

Etlmology. Frcm the Greek,bellus for beautiful,refeffing to this specieswhen in flower it is a stunning sight.

Noles.A putativehybrid(A.5. George16858) between Pileanthus beLlus and P. peduncularishasbeen collectedat 45.8 km north of GalenaBddge, Murchison River on the North West CoastalHighway. This, if conect, is the only record of naturalhybridisation in the genus.

7. Pileanthus rubronitidus Keighery, sp. nov.

PileanthofiLifolio habitu et floribus apparentersimilis, sed diffefi flodbus rubds, petalis7-8 mm versus l2-15 mm longis, staminibusnon bilobis.

Typrs; 4 km west-north-westofYerina Spring,28'04'S, 114'19'E, WesternAustralia, I October1979, J. Taylor 1041,M.D. Crisp & R. Jackson(holo: PERTH 02503921;lso; BISH, CANB).

Erect shrub m 7 m tall. Youngbrancles brownish-red,with prominentoil glandsforming a sticky exudate;leaf decurrenciespromin€nt, persistent, becoming grey, with large red stipularoutgrowths in the leaf axils. Leaves not very clustered,linear, tdquetrous,5-13 mm long, less than 1 mm wide, glabrous, wlth prominent oil glands, adaxial surface grooved, abaxial convex, margin entire, apex obtuse or a.J$e.Peduncle slendet, (8)12-22 mm long, yellow-red. Bracteole pair: cup narrowly turbinate,2-3 mm long;freepoftions 4-6 mm long,brown, margins minutely denticulate, with aconical pungentumbo c. I mmlorg. Pedicelz4 mmlong. Hypanthiumturbinate, 3-5 mm long, 3-4 mm wide, silky-pilose,gland-dotted, ftee part 1-1.5 mmlong. Sepals5,2-4mm long,3-4 mm wide, the two lobes fused for c. I mm at base,sho ly pilose, apexobtuse, margin minutely toothed.Pel4ls 7-8 mm long, c. 8 mrn wide, red-orange,base erose, margtn ent;Lre.Androeciun with entirefilaments; stamens usually c. 1 mm long, (Mount Magnet collectionmuch longer). Ovules4. Style4-7 mm\ong, ciliate at base; stigma globose,minute. (Figure 3 P-T)

Selectedcollections examined.WESTERN AUSTRAIIA: Hutt River, W.H. Butler 198 (PERTH); Mount Magnet,2 Oct.1959,W.H.Butler s.n.(PERTH); cultivated at Kings Park,ex Murchison, l2 Feb. 1973,B.L. Powell13025 (KPBG); 25 km E of Kalbarri, 7 Nov. 1986,P. Robens 166 (PERTH). 'sa8uera8rul ernb e,leq I sercedsqnsqtoq puB sl;llLlluDalq.Jo sorrads petnqulsrll ,{lepr,\\lsoru 3q1sLsrqJ sntD$ rnlnQdsrn)

'(gt crndrC)NoU elnr8 ot u,{lrurtri trlo,qs,mf,f,O Ltounql.tlsle

'esoqo13'etu8t1s sdn)JaddoJ elnunu :xeclele Joled 'elclmd'srorqeli '3uo1 'fuol iururoceq u[rr 1-E ,lrls 8 srF1,1Ope: qsrlurd:oe1d-md LuorZ I sueon]ts :stlrolllulrleiDue qlr,r urilJao.tpuvetrruLcul sur8rru 'osuqqsppcr.ro oldrndu qtr,n.rotrroq8norql 'el 's.rrnl 'asolcl lSuulo'opr,rurru 9 y'3uo1nnu1,1 r'71l.r7 queu rS.ruru olulnrieqntqtr,l\ Jo pettop-puu13 'eprm '3uo1 '"ouo1 ,{11s urur7 .r rirury-g :rtr,noro fluo1qorlorla5 ruu7 I urd eltupl?'esoltd,{llts'€p1,1 LLnrrg g'3rro1uml g g'alulnuudrucruirrrrprird,(g Fuolruru g-7 7a,rtpa7SuolurLLr g 0 ,)oqlun lrJruo) 'ar 'luaLu 'spuu1"' '""uo1 ,{1puo.rc1u q1r,r rluesu r.a.mru uro.rdlclr cluprur po lueurtrorclq1r,n uru 6 g suoruod '8uol ee,rl:opr,\\urru t J nnu !-! dnJr-i ?/7 tl aloal.n.tg s1:tttllyiyoluoururrd tpr,tt3uo[utLLr 0Z-6'.ropuc[:- '3uo1 'rrlnJl^ru 'pcJotsnlr aJ.rrr7r.r7peir-tn,tr aslq s-ar\tlelaAllqeoael ueqt Jopr]oJcl ruru g ot torrJr,r/2.7/ 'spurlS l7-(//J ctnJrqns-() isnlqo xoch:"^.rrr11 o1r[-ql(x)l qlr,r uo]lour.Q.mrrr 1o lueuluordqtr,r'{rnq.ro ';-uo1 'snrxllnbr-rt'otr^ul. 's1xrt1s "'no.rtF13 urLuy-; io -nrorrrl olrsodch!oqs Lrollo.rcls-11lo .\.),r/2r7 slrxrl 'lueu '1Lrcu[uo.rd JLllu r srll.roJ.Qlr'ro.q ndrls 1ltrtrrs rprm rttrolclsc:ruc.unocp.lrcl :Surppcqs pul] {o.ri 3uru.rnl spuuloaIro'u^\o.rq r'.){.)l/r./q tr/?.4 ssol,(11unsn cpr,n ru g ,{q 11r:tur S l ol r/r./r/r'Strrpr:e.rcls ,no1

'lrlr.rls-11v 'Lll.rid.lo Lr.rJtsolA tsuo-Llt.rourLro.rl poc|.rasop :-iroocls lnq l)ir.lri)ods "tLjLut\.tcl x)rl:.xtt /, (!gflI)tof aOt:9 )stI/\ll).u){),11.1N lt)g f s,J ltx)H t!ttltrltlltl lt11t.)t1111)un)t.lJi,

'(Lrios ',^\ rlch:.rioloqcl il,\8)r1))/'lrrlui)snVrlrolsolAllso,rLllr'ros.lo.ror.rotur'puno)^oi.()oD"'ur)purrri^ru L rA\5^LrOO.r'lOcl ,.OOU lJ 1r.aOIPLrnoS sJ8.roO9 SUry to rarr-r L r,{\s.rJlLllsnqrior.ro)Ll| :-rllrlLroDl.ro-1).rlsnr 'g rrpurlloHou^oN Lrl.. nrl,!_/ (iltSI)96,1 1o3on1Ic1.ro11 cl.rr15'1pug "-Fnlnaunpod snqtuealld'8

';r'u r1rol1ooc.rsyrcou uorlclnckrcl trunl.srp ,41t1;'rt1 srql sor:rodssrLlt.lo Iulrcluur u.l.ltsa,r.qt L[]Lllsuolulls .ro.auol slrq sitllttLto.t.lll-t./.lo uorliolloatcuirll )rrno1'1Pclrlosr cq,L

'streLtnlsl.rrlLro 3ur,ruL1 ttr 'lsrd lrulr.nlolollrlad ezrsle^\ol-l ur sjcl-lrptnq cql ur \ttJDlDltp.l LIrA\pcsllluoi ,{11ucnhc.r1 uccq srq 'rr()N \'(18()l s. Dll)g g 1, O) u.Nqlr) ds \tlqtuD!id su{1purt1ur Lr.{oLrIueeq srLI :-erJecl:- srqJ

scrrcclssrql 1o sprl:xl per ,(urqsaql c4 3ur.rro1o.r'{urqs ;lurlnrJlllstiptJtu pttl,. p:r.r Sunruour .)././/r../ :Lrtr-l .qt urojg ,(folorl,rlg

'l.md llltlolll)N l.Lmqll)) Ul pellesUol JIe/r|s/'lllr,l Llotll)t\.ti)\'uoJ

'lsntloJilil 'l '(6/61 stiqtl.tDali.lsnl EZ0 _ lllt\od B rrqrno^ o,{d) I t=u laqlunu au.ro*)t.!.!o.ttlJ

iaqlua^oN,{1:uepun .raqruelde5etrl uee,\ueg pon atl 5'u u arc17

(Jf e.rnirC) putlqnrlls Ltttl.tld)JSDt$ILPII ro qluequL 'pu]s elrq'{\ 'euolspulls ,re^o pu€s ,{er8 uo s,norg rrorlnllJr1uorscrrnbcr uor1rcllolrsrql tcu3e6 tunol l uro.rl uorlialloc 'LroldlurrDroN lrunfsrp ,{:e,reuo qlr,r Io lse,\\pm? rrJEqJe)I uae,rleq str.l)O i1)l1qo4puD uotlnqt.ttsLe

(3m.'r.r,(t{) sr'lrr,rrlJ snrc;.qt to A\.r^3.rV',(].q:ri! I D Nrttuia Vol. 15,No. I (2002)

Noles.The species Ciamelaucium dilatatan Drumm., described by Drumrnond(1853: 403) as having "united vermilion petals and twenty stamens by a beautiful dark purple membrane"is probably a synonymof Pileqnthuspeduncularis as it appearsto havebeen collected in an areawhere only that speciesoccurs and because the description seems apptopriate. However, it is alsopossible that the name appliesto thespecies curently known as P . filifulius , rn wtlch caseit wouldbe the earliest description of that speciesand a new combinationwould be required. This needsto be investigatedfurther.

Two morphologicallydistinct subspecies are recognised. They are largely geographically separated but overlapbetween Kalbari andNerren Neffen Station. No interm€diateshave been recorded in the areaof overlap,but therehas beenlittle detailedcollecting in this area. Until further studiesof the variationin thisspecies are completed it seemsbest to treatthese geographical entities as subspecies.

K€y to the subspeciesof Pileanthuspeduncularis

1. Leavesglabrous. Flowers orange with a purpleor reddishbase andpurple anthers subsp.peduncularis 1. Leaveswith scatteredhairs. Flowersorange, sometimes with pinkishred anthers subsp.piliferus

8a. Pil€anthuspeduncularis Endl. subsp. peduncularis

Vegetativeleaves glabrotj's. Bracteoles glabrous. Pslals purple orreddish atthebase, orange above. AnthersPtxPle. (Figure 3 K-O)

Selectedspecimens exanlzed WESTERN AUSTRALIA: near Bruce Rock, Sep. 1929, W.E. Blackall s.n.(PERTH); 8 km E of Kalbani,L.A. Craven7063 (PERTH); 48 km S of NenenNeren Starion, 8 Nov. 1979,FL Demarz7739 (PERTH); Wongan Hills, 13Nov. 1985,H. Demarz11058 (PERTH); Mogumber,9 Nov. 1897,R. Helmss.n. (PERTH); East Yuna reserve, 70 km NE of Geraldton,12 Oct. 1976,B.G. Muir 382(PERIH); Tammin National Park, 13 Nov. 1970.R.D. Royce 9380 (PERTH).

Floweringperiod. August to November.

Distributionand haDital. Occurs from Nenen Nenen Station south to Mogumber,inland to EastYuna, WonganHills andBruce Rock, growing on sandusually in heath.(Figure 48)

Chromosoruenumber. n=1 1 (Rye 1979),voucher B.l. Powell74092.

8b. Pileanthuspeduncularis subsp. piliferus Keighery,subsp. nov.

Differt a Pileanthopeduncularis slbsp. peduncularis foliis piliferis eglanduligeris,floribus auranticis,antheris roseis, bracteolis piliferis tuberculatis.

Typus:75miles [121 km] northof Camarvonon Onslowroad, Westem Australia, 24 August1963, J.S.Beard.2997 (iola; PERTH 02506882;iso: KPBG). 'ELg Ltg :LZtruotog '4,{U Io punol uoqrr4snv suoDrclldur 3|[ouox?l slr pue a?a.eu,{trAleql uI uowll?^ Fqrunu euro!-ouorqJ^(61,61)'I E (ue3llnls :lP'lsuv 'ozlun) -s8epe^ eq'lna() 8tt d., unpurpSoFueqdrrn]€ueC uorrxs-I,, O ?8uo^ tr lsod ;x{ snqtuoafd'(8061\'O'e4ul.>I 'BZL-6tL:91 undsnwuDllDtrnv udtsaA aqtIo "proJdd\eepr'e|loJ 'uolsnoH :?radoueur{H) €rle4snv ul3Neld l'l.l.c.4(Dwsaqnl) DssolS&ngIo sepads cqcolouotu ^\eu 6erql (266I ) C J '9911)01:lL uapro9 Iolaqog unoc W aqilo s/rz&y sls,{teu€olouoSol,{qd s-aBocsulw puesepu{r,{ 1986t) Dg's33ug? S V.I'uosuqo{ '9Or-g6E t9 fuDIIacslWuaprDg ^aX puo {uo1oglo lDuno[ 'puorrlllllucl 'puounmjc s,ralooli ro^lu uel SJo ,{ueloqeql ol 3ug?lor soru?I rI I [ro{ slauelsnog?^ uro{ s]ce.qxg(€98I) I '902-rsl :z0l sa1o,144tno5 ua11 [o &aDosuoeuu!1 'I 'uosuqof 's33ug aql{o s9ulpeatotd arn]JruNo.uarsorouul ruo.{ ecuepl^e:e?ec!u,{I ot{l !1lognlo^g (6161) S V 4, 5 g (uopuo'I: oJ ? e^eou) € I0A ,. stsuoltltsnv?rolC,, (198I) C'ur€rlueg

sacuaJaJau

'ua4euapuneq ot ,{pnlssnll elqEuaol aceds3u11:o,tr pue suerulcads ;o sqdz:Soloqdpepr,to;d ;o suolloellotpau€ol oq,tr ?F?qJeH s€esJe^o pue u€rl?4snv II? Jo JPls puEsroEJnc eqt luPql ol eII PIno,{\roqlnu sq1 .lsdzdsq1 w Ieluaunllsul ueaq 'e?eJeu,{I^l a,req,{uouoxzl uo suolssncslpesoq^\ pue sdpl plag Jae,lesuo u4q peluudruocc€oq,/'{ Jo 'lueqol?l dno:Esrr4 fpnls o1.roqlnu eq1 pordsur lsJrJ oq,{\ I a l^aN ot pel?clpopsl uonstnqndsrqJ

sluauraEpal,*ou:py

.SJI?q Jelncreqnl qlrrrr peJelor aJEseloelcuJq eqt pue sluq re1npue13-uouul peJeloc eJese,teel e,rpzle8e,req1 's.ret4ue par qsr-4urd :o eSue;o ;eql1e e,req feu guz16 asuq aqt 1z eldrnd :o pu ;o rnoloc Euqse.nuoc 'dsqrs .(ue lnoqlrrrr qelad eSuuro ernd Jeqlle ?u1rcrtru1 syo1ncunpad slto1ncunpad d uroq sleJJIp 'dsqns lI su .{reqEre) srroatoq s1.rc1ncunpadsm'7uoald s? u/(otnl ueeq wq selcedsqnsqqa sa7o1g

':equrelogo1 lsnEny 'poltad 34tauo1g

(gt 6ln8rd)'rleeqwtufiars a$firg ur spg fpues etlq.rJo .r\olo,{ 'perro'seunppups,rolle.(;ope:uos.rorg'rrBqIE)ot?,{lruUNwoJJSmxxo'totlqDtlpuouo!fiq!4sl6

'uJuad 'u's uosulluoJ I '9961 'uo,treureC ',(?,1^qAIH 8ny JoN luq ZZI] sallrr9Z :GJdAd) ,gggt a8rcag S V'9861xro 6Z '7961 I€ls?oJlselA rluoN uo aSpug euepg 3o N uq 8 9t:(HJuad) 98IEnuptD{) v3 3ny 37'ra,.rg 'ulooSot! '086I'^oN '}upql€>I B,4IIIrrL{ :(HJUAd)989I srrqlag E'O 0Z JoA UDI0Z :GJdAd tott 'S't '9961 'aBwU 'parlaDra proag 8nV €Z .(peuue) :VI'JV'I{IS1V NUgtrSlL\\ suawnadspapaps

'pe.rqsplurd ro e8ueJosraryJry lnoq8noJql e8ueJo slDtad sJr€qJelncJeqnl ur peJelocsaloalJDlB, sJr€qJelnpu€13-uou ur pare,rocsataal aup1a3a1

',&eq3rryI'D I9 (.eacer(lN) srytuDa d snue8aW Jo ^\ar^erV