A Biostratigraphical Framework for Geological Correlation of the Middle Devonian Strata in the Moray-Ness Basin Project Area
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The Phylum Vertebrata: a Case for Zoological Recognition Naoki Irie1,2* , Noriyuki Satoh3 and Shigeru Kuratani4
Irie et al. Zoological Letters (2018) 4:32 https://doi.org/10.1186/s40851-018-0114-y REVIEW Open Access The phylum Vertebrata: a case for zoological recognition Naoki Irie1,2* , Noriyuki Satoh3 and Shigeru Kuratani4 Abstract The group Vertebrata is currently placed as a subphylum in the phylum Chordata, together with two other subphyla, Cephalochordata (lancelets) and Urochordata (ascidians). The past three decades, have seen extraordinary advances in zoological taxonomy and the time is now ripe for reassessing whether the subphylum position is truly appropriate for vertebrates, particularly in light of recent advances in molecular phylogeny, comparative genomics, and evolutionary developmental biology. Four lines of current research are discussed here. First, molecular phylogeny has demonstrated that Deuterostomia comprises Ambulacraria (Echinodermata and Hemichordata) and Chordata (Cephalochordata, Urochordata, and Vertebrata), each clade being recognized as a mutually comparable phylum. Second, comparative genomic studies show that vertebrates alone have experienced two rounds of whole-genome duplication, which makes the composition of their gene family unique. Third, comparative gene-expression profiling of vertebrate embryos favors an hourglass pattern of development, the most conserved stage of which is recognized as a phylotypic period characterized by the establishment of a body plan definitively associated with a phylum. This mid-embryonic conservation is supported robustly in vertebrates, but only weakly in chordates. Fourth, certain complex patterns of body plan formation (especially of the head, pharynx, and somites) are recognized throughout the vertebrates, but not in any other animal groups. For these reasons, we suggest that it is more appropriate to recognize vertebrates as an independent phylum, not as a subphylum of the phylum Chordata. -
Université Du Québec
UNIVERSITÉ DU QUÉBEC PRÉCISIONS SUR L'ANATOMIE DE L'OSTÉOLÉPIFORME EUSTHENOPTERON FOORDI DU DÉVONIEN SUPÉRIEUR DE MIGUASHA, QUÉBEC MÉMOIRE PRÉSENTÉ À L'UNIVERSITÉ DU QUÉBEC À RIMOUSKI Comme exigence partielle du programme de Maîtrise en Gestion de la Faune et de ses Habitats PAR JOËL LEBLANC Août 2005 UNIVERSITÉ DU QUÉBEC À RIMOUSKI Service de la bibliothèque Avertissement La diffusion de ce mémoire ou de cette thèse se fait dans le respect des droits de son auteur, qui a signé le formulaire « Autorisation de reproduire et de diffuser un rapport, un mémoire ou une thèse ». En signant ce formulaire, l’auteur concède à l’Université du Québec à Rimouski une licence non exclusive d’utilisation et de publication de la totalité ou d’une partie importante de son travail de recherche pour des fins pédagogiques et non commerciales. Plus précisément, l’auteur autorise l’Université du Québec à Rimouski à reproduire, diffuser, prêter, distribuer ou vendre des copies de son travail de recherche à des fins non commerciales sur quelque support que ce soit, y compris l’Internet. Cette licence et cette autorisation n’entraînent pas une renonciation de la part de l’auteur à ses droits moraux ni à ses droits de propriété intellectuelle. Sauf entente contraire, l’auteur conserve la liberté de diffuser et de commercialiser ou non ce travail dont il possède un exemplaire. 11 TABLE DES MATIÈRES TABLE DES MATIÈRES .... ..... ............................. .. ...... .. .... .. .... ........... ... ............................. .ii LISTE DES TABLEAUX .. ............ -
Tetrapods, Amphibians, and Life on Land
Department of Geological Sciences | Indiana University Dinosaurs and their relatives (c) 2015, P. David Polly Geology G114 Strolling through life Tetrapods, amphibians, and life on land Tetrapods - the clade of four- limbed terrestrial vertebrates Living tetrapod groups: * amphibians * mammals (including humans) * lizards and snakes * crocodilians * birds Eurypos , early Permian temnospondyl (painting by Douglas Henderson, 1990) Department of Geological Sciences | Indiana University Dinosaurs and their relatives (c) 2015, P. David Polly Geology G114 Lobe-finned fish (Sarcopterygia) Living coelacanth Fossil sarcopterygians Late Cretaceous (ca. 65 mya) Carboniferous (ca. 300 mya) Department of Geological Sciences | Indiana University Dinosaurs and their relatives (c) 2015, P. David Polly Geology G114 Comparison of pectoral fins Actinopterygian Sarcopterygian (ray finned) (lobe finned) Scapulocoracoid Humerus Ulna Radius Department of Geological Sciences | Indiana University Dinosaurs and their relatives (c) 2015, P. David Polly Geology G114 Coelacanth pectoral fins Department of Geological Sciences | Indiana University Dinosaurs and their relatives (c) 2015, P. David Polly Geology G114 Ancestral characteristics of living tetrapods • Pelvic and pectoral girdles • Forelimb with humerus, radius, and ulna bones • Hindlimb with femur, tibia, and fibula bones • five digits on the feet • sprawling posture • undulating locomotion • skull with no fenestra Department of Geological Sciences | Indiana University Dinosaurs and their relatives (c) 2015, P. David Polly Geology G114 Tetrapoda: vertebrates more closely related to living Phylogeny of Bony Fish amphibians and amniotes than to their nearest living relatives Fossil taxa coelocanths and Fish-like amphibian-like lung fish Tetrapods Tetrapods Actinopterygia Coelocanths Dipnoans (lungfish) Osteolepis Eusthenopteron Pandericthyes Acanthostega Icthyostega tetrapods Derived Tetrapoda Sarcopterygia Osteichthyes After Coates and Ruta, 2007. -
Caithness County Council
Caithness County Council RECORDS’ IDENTITY STATEMENT Reference number: CC Alternative reference number: Title: Caithness County Council Dates of creation: 1720-1975 Level of description: Fonds Extent: 10 bays of shelving Format: Mainly paper RECORDS’ CONTEXT Name of creators: Caithness County Council Administrative history: 1889-1930 County Councils were established under the Local Government (Scotland) Act 1889. They assumed the powers of the Commissioners of Supply, and of Parochial Boards, excluding those in Burghs, under the Public Health Acts. The County Councils also assumed the powers of the County Road Trusts, and as a consequence were obliged to appoint County Road Boards. Powers of the former Police Committees of the Commissioners were transferred to Standing Joint Committees, composed of County Councillors, Commissioners and the Sheriff of the county. They acted as the police committee of the counties - the executive bodies for the administration of police. The Act thus entrusted to the new County Councils most existing local government functions outwith the burghs except the poor law, education, mental health and licensing. Each county was divided into districts administered by a District Committee of County Councillors. Funded directly by the County Councils, the District Committees were responsible for roads, housing, water supply and public health. Nucleus: The Nuclear and Caithness Archive 1 Provision was also made for the creation of Special Districts to be responsible for the provision of services including water supply, drainage, lighting and scavenging. 1930-1975 The Local Government Act (Scotland) 1929 abolished the District Committees and Parish Councils and transferred their powers and duties to the County Councils and District Councils (see CC/6). -
Of 30 Field Evidence for the Lateral Emplacement of Igneous Dykes
1 Field evidence for the lateral emplacement of igneous dykes: Implications for 3D 2 mechanical models and the plumbing beneath fissure eruptions. 3 David Healy1*, Roberto E. Rizzo1,2, Marcus Duffy1, Natalie J. C. Farrell1, Malcolm J. Hole1 & David 4 Muirhead1 5 6 1School of Geosciences, University of Aberdeen, Aberdeen AB24 3UE United Kingdom 7 2Research Complex at Harwell, Rutherford Appleton Laboratory, University of Manchester, 8 Didcot OX11 0FA United Kingdom 9 10 *Corresponding author e-mail [email protected] 11 12 Keywords: magma, relay, bridge, segment, igneous, volcanic 13 14 Abstract 15 Seismological and geodetic data from modern volcanic systems strongly suggest that magma is 16 transported significant distance (tens of kilometres) in the subsurface away from central 17 volcanic vents. Geological evidence for lateral emplacement preserved within exposed dykes 18 includes aligned fabrics of vesicles and phenocrysts, striations on wall rocks and the anisotropy 19 of magnetic susceptibility. In this paper, we present geometrical evidence for the lateral 20 emplacement of segmented dykes restricted to a narrow depth range in the crust. Near-total 21 exposure of three dykes on wave cut platforms around Birsay (Orkney, UK) are used to map out 22 floor and roof contacts of neighbouring dyke segments in relay zones. The field evidence 23 suggests emplacement from the WSW towards the ENE. Geometrical evidence for the lateral 24 emplacement of segmented dykes is likely more robust than inferences drawn from flow- 25 related fabrics, due to the prevalence of ubiquitous ‘drainback’ events (i.e. magmatic flow 26 reversals) observed in modern systems. -
The Origins of Chordate Larvae Donald I Williamson* Marine Biology, University of Liverpool, Liverpool L69 7ZB, United Kingdom
lopmen ve ta e l B Williamson, Cell Dev Biol 2012, 1:1 D io & l l o l g DOI: 10.4172/2168-9296.1000101 e y C Cell & Developmental Biology ISSN: 2168-9296 Research Article Open Access The Origins of Chordate Larvae Donald I Williamson* Marine Biology, University of Liverpool, Liverpool L69 7ZB, United Kingdom Abstract The larval transfer hypothesis states that larvae originated as adults in other taxa and their genomes were transferred by hybridization. It contests the view that larvae and corresponding adults evolved from common ancestors. The present paper reviews the life histories of chordates, and it interprets them in terms of the larval transfer hypothesis. It is the first paper to apply the hypothesis to craniates. I claim that the larvae of tunicates were acquired from adult larvaceans, the larvae of lampreys from adult cephalochordates, the larvae of lungfishes from adult craniate tadpoles, and the larvae of ray-finned fishes from other ray-finned fishes in different families. The occurrence of larvae in some fishes and their absence in others is correlated with reproductive behavior. Adult amphibians evolved from adult fishes, but larval amphibians did not evolve from either adult or larval fishes. I submit that [1] early amphibians had no larvae and that several families of urodeles and one subfamily of anurans have retained direct development, [2] the tadpole larvae of anurans and urodeles were acquired separately from different Mesozoic adult tadpoles, and [3] the post-tadpole larvae of salamanders were acquired from adults of other urodeles. Reptiles, birds and mammals probably evolved from amphibians that never acquired larvae. -
A New Osteolepidid Fish From
Rea. West. Aust. MU8. 1985, 12(3): 361-377 ANew Osteolepidid Fish from the Upper Devonian Gogo Formation, Western Australia J.A. Long* Abstract A new osteolepidid crossopterygian, Gogonasus andrewsi gen. et sp. nov., is des cribed from a single fronto-ethmoidal shield and associated ethmosphenoid, from the Late Devonian (Frasnian) Gogo Formation, Western Australia. Gogonasus is is distinguished from other osteolepids by the shape and proportions of the fronto ethmoidal shield, absence of palatal fenestrae, well developed basipterygoid pro cesses and moderately broad parasphenoid. The family Osteolepididae is found to be paraphyletic, with Gogonasus being regarded as a plesiomorphic osteolepidid at a similar level of organisation to Thursius. Introduction Much has been published on the well-preserved Late Devonian fish fauna from the Gogo Formation, Western Australia, although to date all the papers describing fish have been on placoderms (Miles 1971; Miles and Dennis 1979; Dennis and Miles 1979-1983; Young 1984), palaeoniscoids (Gardiner 1973, 1984; Gardiner and Bartram 1977) or dipnoans (Miles 1977; Campbell and Barwick 1982a, 1982b, 1983, 1984a). This paper describes the only osteolepiform from the fauna (Gardiner and Miles 1975), a small snout with associated braincase, ANU 21885, housed in the Geology Department, Australian National University. The specimen, collected by the Australian National University on the 1967 Gogo Expedition, was prepared by Dr S.M. Andrews (Royal Scottish Museum) and later returned to the ANU. Onychodus is the only other crossopterygian in the fauna. In its proportions and palatal structure the new specimen provides some additional new points of the anatomy of osteolepiforms. Few Devonian crossopte rygians are known from Australia, and so the specimen is significant in having resemblances to typical Northern Hemisphere species. -
Orcadian Basin Devonian Extensional Tectonics Versus Carboniferous
Journal of the Geological Society Devonian extensional tectonics versus Carboniferous inversion in the northern Orcadian basin M. SERANNE Journal of the Geological Society 1992; v. 149; p. 27-37 doi:10.1144/gsjgs.149.1.0027 Email alerting click here to receive free email alerts when new articles cite this article service Permission click here to seek permission to re-use all or part of this article request Subscribe click here to subscribe to Journal of the Geological Society or the Lyell Collection Notes Downloaded by INIST - CNRS trial access valid until 31/05/2008 on 31 March 2008 © 1992 Geological Society of London Journal of the Geological Society, London, Vol. 149, 1992, pp. 21-31, 14 figs, Printed in Northern Ireland Devonian extensional tectonics versus Carboniferous inversion in the northern Orcadian basin M. SERANNE Laboratoire de Gdologie des Bassins, CNRS u.a.1371, 34095 Montpellier cedex 05, France Abstract: The Old Red Sandstone (Middle Devonian) Orcadian basin was formed as a consequence of extensional collapse of the Caledonian orogen. Onshore study of these collapse-basins in Orkney and Shetland provides directions of extension during basin development. The origin of folding of Old Red Sandstone sediments, that has generally been related to a Carboniferous inversion phase, is discussed: syndepositional deformation supports a Devonian age and consequently some of the folds are related to basin formation. Large-scale folding of Devonian strata results from extensional and left-lateral transcurrent faulting of the underlying basement. Spatial variation of extension direction and distribution of extensional and transcurrent tectonics fit with a model of regional releasing overstep within a left- lateral megashear in NW Europe during late-Caledonian extensional collapse. -
Erection of One 500 Kw Wind Turbine at 550 M of Taigh Na Muir, Dunnet By
THE HIGHLAND COUNCIL Agenda Item 6.2 NORTH PLANNING APPLICATIONS COMMITTEE – Report No PLN/048/13 21 May 2013 12/03638/FUL : R R Mackay ·& Company Limited Land 550M NW of Tigh na Muir, Dunnet Report by Area Planning Manager North SUMMARY Description: Erection of 1 no 500kW wind turbine with a height to blade tip of 79.6m, 55m to hub and a rotor diameter of 48m and ancillary works at Land 550M NW of Tigh na Muir, Dunnet. Recommendation : REFUSE Ward : 04 Landward Caithness Development category : Local Pre-determination Hearing : none Reason referred to commitee ; Local member request 1. PROPOSED DEVELOPMENT 1.1 Application is made in detail for the erection of 1 no. turbine with a height to blade tip of 79.6 metres, a height to hub of 55 metres and a rotor diameter of 48 metres. The associated infrastructure includes turbine foundations; crane hardstanding; pole mounted transformer and associated cabling and an upgrading and lengthening of an existing 540 metre access track to site from the edge of the exiting private access area to ‘Tigh na Muir’. 1.2 A formal Screening Opinion was issued on 08 September 2011, advising that an Environmental Impact Assessment was not required. 1.3 It is proposed that the turbine and construction components will utilise the A836 Castletown to John O’Groats public road and the existing farm access. 1.4 The applicant has provided a number of supporting documents including a Supporting Statement, Site and Viewpoint Map, Photomontages, an Ornithology Appraisal, Technical Appendix, and Surveys. 1.5 Variations: Otter Survey Report Received 23 October 2012. -
Devonian and Carboniferous Stratigraphical Correlation and Interpretation in the Central North Sea, Quadrants 25 – 44
CR/16/032; Final Last modified: 2016/05/29 11:43 Devonian and Carboniferous stratigraphical correlation and interpretation in the Orcadian area, Central North Sea, Quadrants 7 - 22 Energy and Marine Geoscience Programme Commissioned Report CR/16/032 CR/16/032; Final Last modified: 2016/05/29 11:43 CR/16/032; Final Last modified: 2016/05/29 11:43 BRITISH GEOLOGICAL SURVEY ENERGY AND MARINE GEOSCIENCE PROGRAMME COMMERCIAL REPORT CR/16/032 Devonian and Carboniferous stratigraphical correlation and interpretation in the Orcadian area, Central North Sea, Quadrants 7 - 22 K. Whitbread and T. Kearsey The National Grid and other Ordnance Survey data © Crown Copyright and database rights Contributor 2016. Ordnance Survey Licence No. 100021290 EUL. N. Smith Keywords Report; Stratigraphy, Carboniferous, Devonian, Central North Sea. Bibliographical reference WHITBREAD, K AND KEARSEY, T 2016. Devonian and Carboniferous stratigraphical correlation and interpretation in the Orcadian area, Central North Sea, Quadrants 7 - 22. British Geological Survey Commissioned Report, CR/16/032. 74pp. Copyright in materials derived from the British Geological Survey’s work is owned by the Natural Environment Research Council (NERC) and/or the authority that commissioned the work. You may not copy or adapt this publication without first obtaining permission. Contact the BGS Intellectual Property Rights Section, British Geological Survey, Keyworth, e-mail [email protected]. You may quote extracts of a reasonable length without prior permission, provided a full acknowledgement -
Fracture Attribute Scaling and Connectivity in the Devonian Orcadian
1 Fracture attribute scaling and connectivity in the Devonian Orcadian 2 Basin with implications for geologically equivalent sub-surface 3 fractured reservoirs 4 5 6 Anna M. Dichiarante1,2, Ken J.W. McCaffrey1,3, Robert E. Holdsworth1,3, Tore I. Bjørnarå4 and 7 Edward D. Dempsey5 8 9 1 Department of Earth Sciences, Durham University, Durham DH1 3LE, UK 10 2 NORSAR, Kjeller, Norway 11 3 Geospatial Research Ltd, 1 Hawthorn Terrace, Durham, DH1 4EL, UK 12 4 NGI - Norges GeoteKnisKe Institutt, Norway 13 5 Department of Geography, Geology and Environment, University of Hull, Hull HU6 7RX, UK 14 Correspondence to: [email protected] 15 1 16 Abstract: Fracture attribute scaling and connectivity datasets from analogue systems are widely used 17 to inform sub-surface fractured reservoir models in a range of geological settings. However, 18 significant uncertainties are associated with the determination of reliable scaling parameters in 19 surface outcrops. This has limited our ability to upscale key parameters that control fluid-flow at 20 reservoir to basin scales. In this study, we present nine 1D-transect (scanline) fault and fracture 21 attribute datasets from Middle Devonian sandstones in Caithness (Scotland) that are used as an 22 onshore analogue for nearby sub-surface reservoirs such as the Clair Field, West of Shetland. By 23 taking account of truncation and censoring effects in individual datasets, our multi-scale analysis 24 show a preference for power-law scaling of fracture length over 8 orders of magnitude (10-4 to 104) 25 and kinematic aperture over 4 orders of magnitude (10-6 to 10-2). -
Contributions in BIOLOGY and GEOLOGY
MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - ---