46112 درآﺑﻬﺎي اﻳﺮاﻧﻲ ﺟﻨﻮب ﺧﺰر و رﺷﺪ ﻣﺎﻫﻲ ﻛﻠﻤﻪ Rutilus Rutilus Caspius ﺑ

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46112 درآﺑﻬﺎي اﻳﺮاﻧﻲ ﺟﻨﻮب ﺧﺰر و رﺷﺪ ﻣﺎﻫﻲ ﻛﻠﻤﻪ Rutilus Rutilus Caspius ﺑ The Study of biology (age, feeding and, reproduction of Rutilus rutilus caspicus in south coast of the Caspian Sea (Iranian waters) Item Type monograph Authors Bandani, Gh.A.; Bagherzadeh, F.; Keymaram, F.; Yelghi, S.; Parafkandeh, F.; Larijani, M.; Ghorbani, R.; Tavakoli, M.; Ghasemi, Sh.; Sayad Rahim, M.; Noroozi, H.; Sabkara, J.; Daryanabard, R.; Ghadirnezhad, H. Publisher Iranian Fisheries Science Research Institute Download date 09/10/2021 23:52:15 Link to Item http://hdl.handle.net/1834/39962 وزارت ﺟﻬﺎد ﻛﺸﺎورزي ﺳﺎزﻣﺎن ﺗﺤﻘﻴﻘﺎت، آﻣﻮزش و ﺗﺮوﻳﺞﻛﺸﺎورزي ﻣﻮﺳﺴﻪ ﺗﺤﻘﻴﻘﺎت ﻋﻠﻮم ﺷﻴﻼﺗﻲ ﻛﺸﻮر – ﻣﺮﻛﺰ ﺗﺤﻘﻴﻘﺎت ذﺧﺎﻳﺮ آﺑﺰﻳﺎن آﺑﻬﺎي داﺧﻠﻲ ﻋﻨﻮان: ﺑﺮرﺳﻲ ﺑﻴﻮﻟﻮژي (ﺗﻐﺬﻳﻪ، ﺗﺨﻢ رﻳﺰي و رﺷﺪ ﻣﺎﻫﻲ ﻛﻠﻤﻪ Rutilus rutilus caspius ) درآﺑﻬﺎي اﻳﺮاﻧﻲ ﺟﻨﻮب ﺧﺰر ﻣﺠﺮي ﻣﺴﺌﻮل : ﻏﻼﻣﻌﻠﻲ ﺑﻨﺪاﻧﻲ ﺷﻤﺎره ﺛﺒﺖ 46112 وزارت ﺟﻬﺎد ﻛﺸﺎورزي ﺳﺎزﻣﺎن ﺗﺤﻘﻴﻘﺎت، آﻣﻮزش و ﺗﺮوﻳﭻ ﻛﺸﺎورزي ﻣﻮﺳﺴﻪ ﺗﺤﻘﻴﻘﺎت ﻋﻠﻮم ﺷﻴﻼﺗﻲ ﻛﺸﻮر- ﻣﺮﻛﺰ ﺗﺤﻘﻴﻘﺎت ذﺧﺎﻳﺮ آﺑﺰﻳﺎن آﺑﻬﺎي داﺧﻠﻲ ﻋﻨﻮان ﭘﺮوژه ﻣﻠﻲ : ﺑﺮرﺳﻲ ﺑﻴﻮﻟﻮژي (ﺗﻐﺬﻳﻪ، ﺗﺨﻢ رﻳﺰي و رﺷﺪ ﻣﺎﻫﻲ ﻛﻠﻤﻪ Rutilus rutilus caspius ) درآﺑﻬﺎي اﻳﺮاﻧﻲ ﺟﻨﻮب ﺧﺰر ﺷﻤﺎره ﻣﺼﻮب ﭘﺮوژه ﻣﻠﻲ : 89069 0-77-12- ﻧﺎم و ﻧﺎم ﺧﺎﻧﻮادﮔﻲ ﻧﮕﺎرﻧﺪه/ ﻧﮕﺎرﻧﺪﮔﺎن : ﻏﻼﻣﻌﻠﻲ ﺑﻨﺪاﻧﻲ ﻧﺎم و ﻧﺎم ﺧﺎﻧﻮادﮔﻲ ﻣﺠﺮي ﻣﺴﺌﻮل ( اﺧﺘﺼﺎص ﺑﻪ ﭘﺮوژه ﻫﺎ و ﻃﺮﺣﻬﺎي ﻣﻠﻲ و ﻣﺸﺘﺮك دارد ) : ﻏﻼﻣﻌﻠﻲ ﺑﻨﺪاﻧﻲ ﻧﺎم و ﻧﺎم ﺧﺎﻧﻮادﮔﻲ ﻣﺠﺮي (اﺳﺘﺎﻧﻲ): ﻛﻴﻮان ﻋﺒﺎﺳﻲ (ﭘﮋوﻫﺸﻜﺪه آﺑﺰي ﭘﺮوري آﺑﻬﺎي داﺧﻠﻲ ) ﻧﺎم و ﻧﺎم ﺧﺎﻧﻮادﮔﻲ ﻫﻤﻜﺎر(ان) : ﻣﺤﻤﻮد ﺗﻮﻛﻠﻲ-ﺳﻌﻴﺪ ﻳﻠﻘﻲ – ﻫﻴﺎت ا... ﻧﻮروزي - ﻓﺮاﻣﺮز ﺑﺎﻗﺮ زاده – ﻓﺮﻫﺎد ﻛﻴﻤﺮام – ﻓﺮخ ﭘﺮاﻓﻜﻨﺪه - ﺷﻬﺮام ﻗﺎﺳﻤﻲ- رﺳﻮل ﻗﺮﺑﺎﻧﻲ – ﻣﺼﻄﻔﻲ ﺻﻴﺎد رﺣﻴﻢ - ﻣﺤﻤﺪ ﻻرﻳﺠﺎﻧﻲ- ﺟﻠﻴﻞ ﺳﺒﻚ آرا - رﺿﺎ درﻳﺎﻧﺒﺮد- ﺣﺴﻦ ﻗﺪﻳﺮ ﻧﮋاد ﻧﺎم و ﻧﺎم ﺧﺎﻧﻮادﮔﻲ ﻣﺸﺎور(ان) : - ﻧﺎم و ﻧﺎم ﺧﺎﻧﻮادﮔﻲ ﻧﺎﻇﺮ(ان) : ﺷﻬﺮام ﻋﺒﺪاﻟﻤﻠﻜﻲ ﻣﺤﻞ اﺟﺮا : اﺳﺘﺎن ﮔﻠﺴﺘﺎن ﺗﺎرﻳﺦ ﺷﺮوع : 89/8/1 ﻣﺪت اﺟﺮا : 3 ﺳﺎل ﻧﺎﺷﺮ : ﻣﻮﺳﺴﻪ ﺗﺤﻘﻴﻘﺎت ﻋﻠﻮم ﺷﻴﻼﺗﻲ ﻛﺸﻮر ﺗﺎرﻳﺦ اﻧﺘﺸﺎر : ﺳﺎل 1395 ﺣﻖ ﭼﺎپ ﺑﺮاي ﻣﺆﻟﻒ ﻣﺤﻔﻮظ اﺳﺖ . ﻧﻘﻞ ﻣﻄﺎﻟﺐ ، ﺗﺼﺎوﻳﺮ ، ﺟﺪاول ، ﻣﻨﺤﻨﻲ ﻫﺎ و ﻧﻤﻮدارﻫﺎ ﺑﺎ ذﻛﺮ ﻣﺄﺧﺬ ﺑﻼﻣﺎﻧﻊ اﺳﺖ . «ﺳﻮاﺑﻖ ﻃﺮح ﻳﺎ ﭘﺮوژه و ﻣﺠﺮي ﻣﺴﺌﻮل / ﻣﺠﺮي» ﭘﺮوژه ﻣﻠﻲ : ﺑﺮرﺳﻲ ﺑﻴﻮﻟﻮژي (ﺗﻐﺬﻳﻪ، ﺗﺨﻢ رﻳﺰي و رﺷﺪ ﻣﺎﻫﻲ ﻛﻠﻤﻪ Rutilus rutilus caspius )درآﺑﻬﺎي اﻳﺮاﻧﻲ ﺟﻨﻮب ﺧﺰر ﻛﺪ ﻣﺼﻮب : 0-77-12-89069 ﺷﻤﺎره ﺛﺒﺖ (ﻓﺮوﺳﺖ) : 46112 ﺗﺎرﻳﺦ : 93/8/7 ﺑﺎ ﻣﺴﺌﻮﻟﻴﺖ اﺟﺮاﻳﻲ ﺟﻨﺎب آﻗﺎي ﻏﻼﻣﻌﻠﻲ ﺑﻨﺪاﻧﻲ داراي ﻣﺪرك ﺗﺤﺼﻴﻠﻲ ﻛﺎرﺷﻨﺎﺳﻲ ارﺷﺪ در رﺷﺘﻪ ﺷﻴﻼت ﻣﻲ ﺑﺎﺷﺪ. ﭘﺮوژه ﺗﻮﺳﻂ داوران ﻣﻨﺘﺨﺐ ﺑﺨﺶ ﺑﻴﻮﻟﻮژي و ارزﻳﺎﺑﻲ ذﺧﺎﻳﺮ آﺑﺰﻳﺎن در ﺗﺎرﻳﺦ 27/3/93 ﻣﻮرد ارزﻳﺎﺑﻲ و رﺗﺒﻪ ﻋﺎﻟﻲ ﺗﺄﻳﻴﺪ ﮔﺮدﻳﺪ. در زﻣﺎن اﺟﺮاي ﭘﺮوژه، ﻣﺠﺮي در : ﺳﺘﺎد □ ﭘﮋوﻫﺸﻜﺪه □ ﻣﺮﻛﺰ █ اﻳﺴﺘﮕﺎه □ ﺑﺎ ﺳﻤﺖ رﺋﻴﺲ ﺑﺨﺶ ﺑﻴﻮﻟﻮژي و ارزﻳﺎﺑﻲ ذﺧﺎﻳﺮ آﺑﺰﻳﺎن در ﻣﺮﻛﺰ ﺗﺤﻘﻴﻘﺎت ذﺧﺎﻳﺮ آﺑﺰﻳﺎن آﺑﻬﺎي داﺧﻠﻲ ﻣﺸﻐﻮل ﺑﻮده اﺳﺖ. ﻋﻨﻮان « ﻓﻬﺮﺳﺖ ﻣﻨﺪرﺟﺎت » ﺻﻔﺤﻪ ﭼﻜﻴﺪه .............................................................................................................................................................................. 1 1-ﻣﻘﺪﻣﻪ ................................................................................................................................................ 2 1-1 - ﭘﻴﺸﻴﻨﻪ ﺗﺤﻘﻴﻖ ................................................................................................................................ 3 2-1 -ﺳﻴﺴﺘﻤﺎﺗﻴﻚ ................................................................................................................................. 5 3-1 - ﭘﺮاﻛﻨﺶ ﺟﻬﺎﻧﻲ ............................................................................................................................ 6 4-1 - ﺗﻐﻴﻴﺮات زﻣﺎﻧﻲ ﺻﻴﺪ ....................................................................................................................... 7 5-1 - زﻳﺴﺘﮕﺎه ......................................................................................................................................... 8 6-1 - ﺑﻴﻮﻟﻮژي ...................................................................................................................................... 8 7-1 - اﻛﻮﻟﻮژيِ .................................................................................................................................... 9 2- ﻣﻮاد و روﺷﻬﺎ .................................................................................................................................. 11 1-2 - روش ﺗﺤﻘﻴﻖ ................................................................................................................................. 11 3- ﻧﺘﺎﻳﺞ ................................................................................................................................................ 15 1-3 - ﺻﻴﺪ ............................................................................................................................................ 15 2-3 - زﻳﺴﺖ ﺳﻨﺠﻲ ................................................................................................................................ 16 3-3 - راﺑﻄﻪ ﻃﻮل و وزن .......................................................................................................................... 17 4-3 - ﺗﺮﻛﻴﺐ ﺳﻨﻲ .................................................................................................................................. 18 5-3 - رﺷﺪ ............................................................................................................................................. 19 6-3 - ﺿﺮﻳﺐ ﭼﺎﻗﻲ ................................................................................................................................ 20 7-3 - ﻧﺴﺒﺖ ﺟﻨﺴﻲ ................................................................................................................................. 21 8-3 - ﺑﺮرﺳﻲ وﺿﻌﻴﺖ ﻏﺪدﺟﻨﺴﻲ ............................................................................................................ 22 9-3 - ﻫﻢ آوري وراﺑﻄﻪ آن ﺑﺎ ﻃﻮل و وزن ................................................................................................ 25 10-3- ﻃﻮل اوﻟﻴﻦ ﺑﻠﻮغ ﺟﻨﺴﻲ (L.M.50) ................................................................................................ 26 11-3- ﺗﻐﺬﻳﻪ ......................................................................................................................................... 28 4- ﺑﺤﺚ ................................................................................................................................................ 32 5- ﻧﺘﻴﺠﻪ ﮔﻴﺮي ....................................................................................................................................... 37 ﻣﻨﺎﺑﻊ ...................................................................................................................................................... 38 ﭼﻜﻴﺪه اﻧﮕﻠﻴﺴﻲ ....................................................................................................................................... 41 ر ژ (، ر و ر ه در .../ ١ ﭼﻜﻴﺪه درﻳﺎي ﺧﺰر ﻳﻜﻲ از ﻣﻨﺎﺑﻊ ﻣﻬﻢ آﺑﻲ ﻛﺸﻮر اﺳـﺖ ﻛـﻪ ﺑـﻪ ﻟﺤـﺎظ دارا ﺑـﻮدن ذﺧـﺎﻳﺮ ارزﺷـﻤﻨﺪ ﻣﺎﻫﻴـﺎن ﺧﺎوﻳـﺎري و اﺳﺘﺨﻮاﻧﻲ ﻣﻮرد ﺗﻮﺟﻪ ﻗﺮار ﮔﺮﻓﺘﻪ اﺳﺖ. ﻣﺎﻫﻲ ﻛﻠﻤﻪ ((.Rutilus rutilus caspicus(L) ﻳﻜﻲ از ﮔﻮن ﻫﺎي ﻣﻬﻢ ﺗﺠـﺎر ﺗﻲ در ﺳﻮاﺣﻞ ﺟﻨﻮﺑﻲ درﻳﺎي ﺧﺰر ﺑﻪ وﻳﮋه در دواﺳﺘﺎن ﮔﻠﺴﺘﺎن وﮔﻴﻼ ن ﻣﻲ ﺑﺎﺷﺪ.ﻛﻪ در 10 ﺳﺎل اﺧﻴﺮ ﺻﻴﺪ آن ﻛـﺎﻫﺶ ﻗﺎﺑﻞ ﻣﻼﺣﻈﻪ اي ﻳﺎﻓﺘﻪ اﺳﺖ. در ﺧﺼﻮص ﻇﺮﻓﻴﺖ ﺳـﺎزي ﻣﺎﻫﻴـﺎن اﺳـﺘﺨﻮاﻧﻲ ﺑـﻪ ﻟﺤـﺎظ اﻫﻤﻴـﺖ وﻳـﮋه آن درﮔﺬﺷـﺘﻪ ﺟﺎ ﻳﮕﺎه وﻳﮋه اي دارد. ﻣﺪﻳﺮﻳﺖ وﺑﻬﺮه ﺑﺮداري اﻳﻦ ﮔﻮﻧﻪ ﺧﻮد ﻧﻴﺎز ﺑﻪ ﺑﺮرﺳـﻲ ﺟـﺎﻣﻊ ودﻗﻴـﻖ ﺑﻴﻮﻟـﻮژي آن دارد . اﻳـﻦ ﻣﻄﺎﻟﻌﻪ ﺑﻪ ﻋﻨﻮان ﺑﺨﺸﻲ از ﻣﻄﺎﻟ ﻌﺎت ﺟﺎﻣﻊ ﺗﻐﺬﻳﻪ، ﺗﺨﻢ رﻳﺰي و رﺷﺪ اﻳﻦ ﮔﻮﻧﻪ ﻣﻲ ﺑﺎﺷﺪ. اﻳﻦ ﺑﺮرﺳﻲ ﺑﺎ اﺳﺘﻔﺎده از ﻧﻤﻮﻧﻪ ﻫﺎي ﺻﻴﺪ ﺷﺪه در ﺗﻮرﻫﺎي ﭘﺮه ﺳﺎﺣﻠﻲ در ﻃﻮل ﺳﻮاﺣﻞ ﺟﻨﻮﺑﻲ درﻳﺎي ﺧﺰر (آﺑﻬﺎي اﻳﺮان) ﺻﻮرت ﮔﺮﻓﺖ. اﻧﺪازه ﻃﻮل ﻓﻮرك ﻧﻤﻮﻧﻪ ﻫﺎي ﻣﻮرد ﻧﻈﺮ از /5 12 ﺗﺎ 5/29 ﺳﺎﻧﺘﻴﻤﺘﺮ و وزن ﻛـﻞ ، ﺑـﻴﻦ 29 ﺗـﺎ 293 ﮔﺮم درا ﺳﺘﺎن ﮔﻠﺴﺘﺎن و /5 10 ﺗﺎ23 ﺳﺎﻧﺘﻴﻤﺘﺮووزن ﻛﻞ ،ﺑﻴﻦ /2 17 ﮔﺮم ﺗﺎ 210 ﮔﺮم دراﺳﺘﺎن ﮔﻴﻼن ﻣﺘﻔﺎوت ﺑﻮد. ﻣﻘﺪار b راﺑﻄﻪ ﻃﻮل - وزن از 02/3 ﺗﺎ 25/3 و75/3 ﺗﺎ28/3 ﺑﺮاي ﺟﻨﺲ ﻣﺎده و ﻧﺮ ﺑـﺮاي ﮔﻠﺴـﺘﺎن وﮔـﻴﻼن ﺑـﻪ ﺗﺮﺗﻴـﺐ ﻣﺘﻐﻴﺮ ﺑﻮد. داﻣﻨﻪ ﺳﻨﻲ ﻧﻤﻮﻧﻪ ﻫﺎ از 1 ﺗﺎ 4 ﺳﺎل درﻫﺮدو اﺳﺘﺎن ﻣﺘﻔﺎوت ﺑـﻮد. اوج ﺗﺨﻤﺮﻳـﺰي در ﻫـﺮدو اﺳـﺘﺎن ،اﺳـﻔﻨﺪ وﻓﺮوردﻳﻦ ﺑﻮد. ﻫﻤﺎوري ﻣﻄﻠﻖ از 7260 ﺗﺎ 23196 ﻋﺪد ﺗﺨﻢ ﻣﺘﻔﺎوت ﺑﻮد . ﻃﻮل ﺑﻠـﻮغ 50 درﺻـﺪ ﺟﻤﻌﻴـﺖ ﻣـﺎﻫﻲ ﻛﻠﻤﻪ در اﺳﺘﺎن ﮔﻠﺴﺘﺎن وﮔﻴﻼن ﺑ ﻪ ﺗﺮﺗﻴﺐ ﻣﻌﺎدل 139و 145 ﻣﻴﻠﻴﻤﺘﺮ ﺑﻮد . ﻣﺘﻮﺳﻂ رﺷﺪ ﺑﺮ ﺣﺴﺐ ﻣﻴﻠﻲ ﻣﺘﺮ ﺑـﺮ اﺳـﺎس ﻣﺪل وان ﺑﺮﺗﺎﻻﻧﻔﻲ ﺑﺮاي ﻃﻮل ﻣﺎﻫﻲ ﺑﺮاي اﺳﺘﺎن ﮔﻠﺴﺘﺎن وﮔﻴﻼن ﺑﻪ ﺗﺮﺗﻴﺐ زﻳﺮ ﺑﺪﺳﺖ آﻣﺪ: (L(t) = 30 /94 -1(* exp (-0/42 ( t-0/18 و (L(t) = 20 /49 -1(* exp (-0/53 ( t-1 . در اﺳﺘﺎن ﮔﻠﺴﺘﺎن وﮔﻴﻼن، ﻏﺎﻟﺒﻴﺖ ﺗﻐﺬﻳﻪ ﺑﺎ ﺷﻜﻢ ﭘﺎﻳﺎن(ﮔﺎﺳﺘﺮوﭘﻮدا) ﻛﺮﻣﻬﺎي ﭘﺮﺗﺎر و ﻧﺮﻣﺘﻨـﺎن ﺑـﻮد . ﻣﻴﮕـﻮ ، ﻣـﺎﻫﻲ ، ﻛﺮﻣﻬﺎ، ﺣﺸﺮات، زﺋﻮﭘﻼﻧﻜﺘﻮن و دوﻛﻔﻪ اﻳﻬﺎ از ﻃﻌﻤﻪ ﻫـﺎي ﻧـﺎدر ﺑﻮدﻧـﺪ و ﺧﺮﭼﻨﮕﻬـﺎي ﮔـﺮد و ﮔﻴﺎﻫـﺎن ﺑـﻪ ﻋﻨـﻮان ﻏﺬاﻳﻲ ﻋﻤﻮﻣﻲ ﺑﺮاي ﻣﺎﻫﻲ ﻛﻠﻤﻪ ﺷﻨﺎﺧﺘﻪ ﺷﺪﻧﺪ. ﻛﻠﻤﺎت ﻛﻠﻴﺪي: ﻣﺎﻫﻲ ﻛﻠﻤﻪ ، ﺗﺨﻢ رﻳﺰي، ﺗﻐﺬﻳﻪ، رﺷﺪ، درﻳﺎي ﺧﺰر ٢ / ارش ح -1 ﻣﻘﺪﻣﻪ درﻳﺎي ﺧﺰر، ﺑﺎ وﺳﻌﺖ ﺣﺪود 436000 ﻛﻴﻠﻮﻣﺘﺮ ﻣﺮﺑـﻊ، 1200 ﻛﻴﻠـﻮﻣﺘﺮ ﻃـﻮل و 220 ﺗـﺎ 550 ﻛﻴﻠـﻮﻣﺘﺮ ﭘﻬﻨـﺎ دا رد. و ﺣﺠﻢ آب آن اﻓﺰون ﺑﺮ 77000 ﻛﻴﻠﻮﻣﺘﺮ ﻣﻜﻌﺐ اﺳﺖ. ﮔﻮدي اﻳﻦ درﻳﺎ در ﺑﺨﺶ ﺷﻤﺎﻟﻲ، 10 ﺗﺎ 12 ﻣﺘﺮ و در ﺑﺨـﺶ ﻣﻴﺎﻧﻲ، ﺗﺎ 770 ﻣﺘﺮ اﺳﺖ و ﮔﻮدﺗﺮﻳﻦ ﻧﻘﻄﻪ آن در ﺑﺨﺶ ﺟﻨﻮﺑﻲ، ﺗﺎ 1000 ﻣﺘﺮ ﻧﻴﺰ ﻣـﻲ رﺳـﺪ. ﺳـﻄﺢ آب درﻳﺎﭼـﻪ در ﺣﺪود 26 ﺗﺎ 28 ﻣﺘﺮ (ﺑﺮﺣﺴﺐ ﺳﺎﻟﻬﺎي ﻣﺨﺘﻠﻒ) از ﺳﻄﺢ آب درﻳﺎﻫﺎي آزاد، ﭘﺎﻳﻴﻦ ﺗﺮاﺳﺖ (ﻗﺎﺳﻤﻮف ، 1929 ) اﻳـﻦ درﻳﺎ ازﻃﺮﻳﻖ وﻟﮕﺎ و ﻫﻤﭽﻨﻴﻦ ﻛﺎﻧﺎل وﻟﮕﺎ - دن ﻛﻪ ﻣﺠﻬﺰ ﺑﻪ ﺣﻮﺿﭽﻪ ﻫﺎي ﺗﻨﻈﻴﻢ ﺳﻄﺢ آب و ﺑﺮﻗـﺮاري ﻫـﻢ ﺗـﺮازي آب اﺳﺖ، ﺑﻪ ﻃﻮر ﻏﻴﺮ ﻣﺴﺘﻘﻴﻢ ﺑﺎ درﻳـﺎي ﺑﺎﻟﺘﻴـﻚ و درﻳـﺎي ﺳـﻴﺎه ارﺗﺒـﺎط دارد . در درﻳـﺎي ﺧـﺰر 100 ﮔﻮﻧـﻪ ﻣـﺎﻫﻲ ﺷﻨﺎﺳﺎﻳﻲ ﺷﺪه اﺳﺖ ﻛﻪ از اﻳﻦ ﻣﻴﺎن، ﺷﮓ ﻣﺎﻫﻴﺎن،ﻛﭙﻮر ﻣﺎﻫﻴﺎن و ﮔﺎو ﻣﺎﻫﻴﺎن داراي ﺑﻴﺸﺘﺮﻳﻦ ﻓﺮاواﻧﻲ اﻧـﺪ و ﺑـﻴﺶ از 70 درﺻﺪ از ﻛﻞ ﻣﺎﻫﻴﺎن ﺧﺰر راﺗﺸﻜﻴﻞ ﻣﻲ دﻫﻨﺪ. ﻓﻮن ﻣﺎﻫﻴـﺎن ﺧـﺰر ﻣﺨﺘﻠـﻒ و ﻣﺘﻌﻠـﻖ ﺑـﻪ آب ﺷـﻴﺮﻳﻦ، ﻣﻬـﺎﺟﺮﻳﻦ درﻳﺎي ﺳﻴﺎه و آزوف ﻣﻲ ﺑﺎﺷﻨﺪ. ﻃﺒﻖ ﮔﺰارش ﻛﺎزاﻧﭽﻒ ( 1981 ) از ﭼﻬﺎر ﺟﻤﻌﻴـﺖ ﻣـﺎﻫﻲ ﻛﻠﻤـﻪ درﻳـﺎي ﺧـﺰر دو دو ﺟﻤﻌﻴﺖ ﻣ ﻌﺘ ﻠﻖ ﺑـﻪ ﺑﺨـﺶ ﺟﻨـﻮﺑﻲ درﻳـﺎي ﺧﺰرﻣـﻲ ﺑﺎﺷـﺪ ﻛـﻪ از آﻧﻬـﺎ ﺗﺤـﺖ ﻋﻨـﻮان ﻛﻠﻤـﻪ ﻛـﻮرا (اﻧﺰﻟـﻲ) وﻛﻠﻤـﻪ ﺗﺮﻛﻤﻨﺴﺘﺎن(ﮔﺮﮔﺎن) ﻳﺎد ﻣﻲ ﻛﻨﻨﺪ. ﻛﻠﻤﻪ ﺗﺮﻛﻤﻦ ﺑﺎ ﻧﺎم ﻋﻠﻤـﻲ Rutilus rutilus caspicus ratio parvadin knipowisitschi در ﺟﻨﻮب ﺷﺮﻗﻲ درﻳﺎي ﺧﺰر زﻳﺴﺖ ﻧﻤﻮده وﺑﻴﺸﺘﺮﻳﻦ ﺗﺮاﻛﻢ را در ﻣﺼﺐ رود ﺧﺎﻧﻪ ا ﺗﺮك دارد . ( ﻗﻠﻲ ﻳﻒ ، 1997 ) ﻣﻄﺎﺑﻖ ﻣﻄﺎﻟﻌﺎت ﭘﺮاﻓﻜﻨﺪه وﻫﻤﻜﺎران در ﺳﺎل 1382 ﻣﺎﻫﻲ ﻛﻠﻤﻪ دو ﻣﻨﻄﻘﻪ
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