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Paracamelus from the Late Pliocene of the Black Sea Region

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Paracamelus from the Late Pliocene of the Black Sea Region Advances in Vertebrate Paleontology “Hen to Panta” ISSN 973-0-02910-5 :17-24 Bucharest, 2003 Paracamelus from the Late Pliocene of the Black Sea region Vadim V. Titov 347900, Taganrog, R.-Luxembourg str., 38, 20, Russia; email: [email protected], [email protected] Abstract - The small camel Paracamelus alutensis is a typical representative of East European faunas of the Late Pliocene and the Early Pleistocene. This species is considered an endemic form of the Black Sea territory. Morphological features of Paracamelus and the accompanying fauna's structure testify about the inhabitance of this animal in steppe and forest-steppe landscapes. Particularities of the Late Pliocene and the Early Pleistocene faunas from the south of Eastern Europe distinguishing the Black Sea - Kazakhstanian province in Mediterranean zoogeographical subarea. Keywords: Paracamelus, Black Sea region, Late Pliocene ÐÅÇÞÌÉ - Ìåëêèé âåðáëþä Paracamelus alutensis ÿâëÿåòñÿ òèïè÷íûì ïðåäñòàâèòåëåì âîñòî÷íîåâðîïåéñêèõ ôàóí êîíöà ïëèîöåíà - íà÷àëà ïëåéñòîöåíà. Ýòîò âèä ðàññìàòðèâàåòñÿ êàê ôîðìà, ýíäåìè÷íàÿ äëÿ òåððèòîðèè Ïðè÷åðíîìîðüÿ. Ìîðôîëîãè÷åñêèå ïðèçíàêè Paracamelus è ñîñòàâ ñîïóòñòâóþùèõ ôàóí ñâèäåòåëüñòâóþò îá îáèòàíèè ýòîãî æèâîòíîãî â óñëîâèÿõ ñòåïíîãî è ëåñîñòåïíîãî ëàíäøàôòîâ. Îñîáåííîñòè ïîçäíåïëèîöåíîâûõ è ðàííåïëåéñòîöåíîâûõ ôàóí þãà Âîñòî÷íîé Åâðîïû ïîçâîëÿþò ãîâîðèòü î âûäåëåíèè ïðè÷åðíîìîðñêî- êàçàõñòàíñêîé ïðîâèíöèè â ïðåäåëàõ Ñðåäèçåìíîìîðñêîé çîîãåîãðàôè÷åñêîé ïîäîáëàñòè. Paracamelus alutensis (ªtefãnescu) was depicted by the includes: Nyctereutes megamastoides (Pomel), Homotherium lower jaw from Romania. It was found in deposits in the range crenatidens (Fabrini), Anancus arvernensis alexeevae dating from the Upper Pliocene to the Lower Pleistocene (Bajguscheva), Archidiskodon meridionalis gromovi (Garutt et (STEFANESCU 1895, 1910, HAVESSON 1954). Since then isolated Alexeeva), Hipparion cf. moriturum Kretzoi, Equus remains of the Upper Pliocene small camels were described from livenzovensis Bajgusheva, Elasmotherium cf. caucasicum the territory of Romania (SIMIONESCU 1932, STEFANESCU 1895, Borissiak, Cervus (Rusa) cf. philisi Shaub, Eucladoceros cf. 1910, GHENEA & RADULESCU 1963, RADULESCU & SAMSON, dicranios Nesti, Libralces cf. gallicus Azzaroli and other forms. The small mammal association from the Liventsovka sand 1990, 1996, 2001, RADULESCU & BURLACU 1993), Ukraine pit (the parastratotype of Khapry faunistic complex) is (ALEXEEVA 1974, TOPACHEVSKY 1956, PIDOPLICHKO & characterized by the appearance of remains of Mimomys TOPACHEVSKY 1962, TOPACHEVSKY & NESIN, 1989), south of praepliocaenicus Rebeder, Borsodia praehungaricus Russian Plain (HAVESSON 1954, BAJGUSHEVA 1971, 1984, cotlovinensis (Topachevsky & Scorik) and Mimomys ex gr. reidi ALEXEEVA 1977a, b), Turkey (KOSTOPOULOS & SEN 1999) Hinton (TESAKOV 1993, 1995, BAJGUSHEVA et al. 2001). This (Fig.1). The most mass material was found in some localities of complex allows the correlation of the deposits with the middle the Upper Pliocene alluvial Khapry layers. These strata are Villafranchian and the lower part of MN 17 zone. Findings of exposed along the north coast of the Taganrog gulf of the Sea of "Psekups type" rodents in the upper layers of this pit testify that Azov and the left bank of the lower current of the Don River. the age of Khapry complex is older than the terminal Pliocene (VASILEV 1969, The section of newest deposits…, 1976). More (BAJGUSHEVA et al. 2001). than 200 bones of Paracamelus alutensis are known from here, In the Paracamelus alutensis remains collection, except for which is about 14% of the whole quantity of large vertebrate numerous teeth, jaws and limb bones, there is unique finding of remains of the Khapry faunal complex. This association dates to an almost whole cranium and vertebral column. There are the Late Pliocene and corresponds to the level of the typical remains of large Paracamelus cf. gigas Schlosser in collection middle Villafranchian Saint-Vallier faunal unit in the Western from Khapry deposits, too (KHOHLOVKINA 1940, GROMOV Europe (VANGENGEIM & PEVZNER 1991). These orictocenosis 1948, HAVESSON 1954, BAJGUSHEVA 1971, ALEXEEVA 1977a). 18 VADIM V. T ITOV PARACAMELUS FROM THE LATE PLIOCENE OF THE BLACK SEA REGION 19 camel remains, small mammals are known - Ochotonoides The association of small mammal (Desmana nogaica kujalnicensis Topachevsky et Scoric, Citellus nogaici Topachevsky et Paschkov, Ochotona sp., Spermophilus nogaici Topachevsky, Alactaga ucrainica Gromov & Shevireva, Topachevsky, Sicista vinogradovi Topachevsky, Cricetus Microspalax odessanus Topachevsky, Plioscirtopoda cf. nannus Schaub, Villanyia fejervaryi Kormos, Lagurodon stepanovi I. Gromov & Schevtshenko, Apodemus sp., arankae Kretzoi, Prolagurus pannonicus Kormos, Eolagurus Parapodemus sp., Euxinomys sp., Phagapodemus sp., argyropuloi adventus Rekovets, Mimomys intermedius Newton, Allocricetus cf. bursae Shaub, Promimomys stehlini, Borsodia Allophaiomys pliocaenicus) testifies to its age in the range of the petenyii Meh., B. fejervaryi, Mimomys ex gr. pliocaenicus- Nogaisk stage, at the end of the Early Pleistocene polonicus, M. hintoni Fejfar and M. reidi Hinton (Kryzhanovka- (TOPACHEVSKY 1956, PIDOPLICHKO & TOPACHEVSKY 1962, 3 by A.S. Tesakov 1993). This association belongs to Psekups ALEXEEVA 1977b, REKOVETS 1994). faunistic complex and is dating as terminal stage of the Villanyan Paracamelus cf. alutensis came in the stuff of the Late (late Villafranchian). Pliocene complex from Turkey. The association of large From the Pliocene layers near Kotlovina village (Odessa mammals, including Eucyon cf. odessanus, Pliocrocuta perrieri, region, Ukraine) the first phalanx of Paracamelus cf. alutensis Homotherium sp., Equus stenonis and Gazellospira cf. was found together with Villafranchian Anancus cf. arvernensis, torticornis allows the fauna attributed to the Sarikol Tepe Archidiskodon meridionalis gromovi, Equus ex gr. stenonis, (Ankara) to MN 17 zone (KOSTOPOULOS & SEN 1999). Stephanorhinus etruscus, Eucladoceros sp., Cervidae gen. indet., The part of a small camel's humerus in the Taman fauna Gazella sp., Vulpes sp. and Lycyaena cf. lunensis Del Kamp collection from the locality Tsymbal (Taman Peninsula) can (TOPACHEVSKY & NESIN 1989, LOGVINENKO 1998). Bones of attest that P. alutensis exists in this faunistic complex together large mammals in those localities were came chiefly from a big with one of the latest representative of Anancus (VERESTCHAGIN clastic bed. It contains the Kotlovina 2 association of small 1957, DUBROVO 1963, ALEXEEVA 1974). Rodent association mammals. Citellus cf. nogaici Topachevski, Allactaga ucrainica from this site, due of the evolutional level of Clethrionomys I. Gromov & Schevchenko, Nannospalax odessanus hintonianus (Kretzoi), is dated by A. S. Tesakov (TESAKOV (Topachevski), Borsodia petenyii, B. feiervaryi, Mimomys 1996) as early Biharian (Early Pleistocene). pliocaenicus F. Major, M. hintoni Feifar, Dolomys nehringi So the statigraphy range of Alutensis camel comprises the Kretzoi, D. milleri Nehring, Pliomys jalpugensis Nesin and Late Pliocene - the Early Pleistocene (contemporaneous with P.ucrainicus topachevskii Nesin prevail in these layers Khapry, Psekups and the beginning of Taman faunistic complexes). That correspondes to the whole MN 17 and the (TOPACHEVSKY & NESIN 1989). This association correlates with upper liventsovian microteriofauna, which is dating as the late beginning of MNQ 18 (MQ 1). However, it is possible that the first appearence of P. alutensis on the territory of the Black Sea Villanyian (TESAKOV 1995). From the middle bed of the Cherevichnoe locality, on the left region be dated to the end of the early Villafranchian (Middle Pliocene). The likely joint their findings together with Fig. 1 - Localities of Paracamelus alutensis ªtefãnescu. 1 - Khapry; 2 - Liventsovka; 3 - Morskaya; 4 - Psekups (ALEXEEVA 1977 b); 5 - Tsymbal (HAVESSON bank of Hadjibey estuary (Odessa region, Ukraine) a few findings 1954); 6 - Kairy (TOPACHEVSKIY 1956, PIDOPLICHKO & TOPACHEVSKIY 1962); 7 - Kryzhanovka (KHOMENKO 1915, HAVESSON 1954); 8 - Zhevakhova gora of P. alutensis are also known (LOGVINENKO 2000). Cherevichnoe Zygolophodon borsoni in Romania. The Alutensis camel (ALEXEEVA 1974, 1977a); 9 - Dolinskoe (ALEXEEVA 1977); 10 - Tuluceºti (GHENEA & RADULESCU 1963); 11 - Slatina, Milcovu-2 (ªTEFANESCU 1910, - 2 is characterized by rodent association of “Khapry" type with becomes the most numerous in orictocenosis of the end of GHENEA 1970, RADULESCO & SAMSON 1990); 12 - Frãteºti (RADULESCU & BURLACU 1993), 13 - Sarikol Tepe (KOSTOPOULOS & SEN 1999). the prevailing of Villanyia petenyii, V. fejervaryi, Pliomys Pliocene. ucrainicus, Mimomys cf. pliocaenicus, M.hintoni and M. reidi Paracamelus alutensis differs from the representatives of genus Camelus - C. knoblochi, Camelus bactrianus ferus by its (TOPACHEVSKY et al. 1987, REKOVETS 1994). smallest size. It resembles small individuals of C. dromedarius The range of P. alutensis distribution is considered within situated above deposits of normal polarity assigned to the Olduvai From Kujalnik in the early Pleistocene deposits, the lower jaw of small camel was described. I. P. Khomenko attributed it to L. by measurements. The cranium of little Paracamelus differs the Late Pliocene - Early Pleistocene bounds. The majority of event (RADULESCO & SAMSON 1996, 2001). Rãdulescu and from the Camelus ones by several features: Alutensis camel localities is dating within these limits. The Samson consider
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