Chapter 36 SYNAGELES 1 z 5

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Salticidae (Araneae) genera of the world - an atlas (unfinished manuscript) by Jerzy Prószy ński Professor Emeritus, Museum and Institute of Zoology, Polish Academy of Sciences, ul. Wilcza 63, 00-679 Warsaw, POLAND e-mail: [email protected] Chapter 37 HISPONINES informal group of genera Version June 3rd, 2020.

symbol of the supragroup HISPONINES Composition and searching on this page (including fossil genera): +Gorgopsidis : +Gorgopsina Hermotimus Massagris - see EUODENINES Tomobella Tomocyrba Tomomingi Exemplary representatives of the group

Source - see below HISPONINES informal group of genera

Mutual diagnostic characters of genera included . Differs from other groups by minute size of eyes posterior lateral (IInd row), sitting very closely to eyes anterior lateral, on the same black pigmented protuberance (Fig. 49J1). On the other hand, genitalic characters are insufficiently known in this group and no final conclusion could be drawn yet. Embolus bent, coiled or straight and minute, sitting atop membranous distal haematodocha. Epigyne varies extensively. Remark. Peculiar arrangement of eyes anterior lateral and posterior lateral (IInd row) on the same protuberance, followed in some genera by shallow constriction across eye field, may be misleading. The same concerns striking reduction of size of eyes posterior lateral (IInd row) in some genera classified to various groups, for instance eyes in Lystrocteissa are paired with HYLLINES[?] like palp structure (Figs 14L, 17G), likewise Tomobella fotsy Szüts, Scharff 2009 (Fig. 49I) resembles some Neon . That somatic character is apparently prone to develop parallel in various lineages of Salticidae, it is interesting that in some fossil genera (like Prolinus ) eyes posterior lateral (IInd row) are not reduced, or moderately reduced (as in amber shown on magnificent photographs by Hill - see Title page). +Gorgopsidis Type species (recognizable species). See more species at +Gorgopsidis

Gorgopsidis bechlyi [fossil] SOURCE. Wunderlich, 2004. 3B: 1790-1791, f 48-51a, photo 421-422. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy. +Gorgopsina Type species ( recognizable species). See more species at +Gorgopsina

J. Prószy ński - 2020 Salticidae (Araneae) genera of the world - an atlas Chapter 36 SYNAGELES 2 z 5

, A - Gorgopsina amabilis , B - G. constricta , C - G. expandens, D - G. flexuosa, E - G. speciosa , F - G. frenata , G-H - various fossil Gorgopsina . SOURCE. Wunderlich, 2004: Beiträge zur Araneologie 3: 1793, f 54-55, photo 423. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy. Gen. Hermotimus Simon, 1903 Type species Hermotimus coriaceus (1 recognizable species). See more species at .z Hermotimus -Q +M

A-D - Hermotimus coriaceus , E - Hermotimus sp. SOURCE. A-C - Proszynski 1987: 43, D - Szuts 2005[2007]. Opusc. zool., 36: 87-88, f 14-15, E - Maddison 2015. Journal of Arachnology. 43: 231–292, f. 39. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy . Gen. Hispo Simon, 1886 Type species Hispo cingulata (11 recognizable species). See more species at .z Hispo -Q +M

A - Hispo cingulata , B - frenata , C-E - Hispo striolata, G-J - . Hispo sp. frpm Madagascar. SOURCE. A-E - Wanless 1981a.. Bull. brit. Mus. nat. Hist. (Zool.)41 (4): 184-184, ff. 1A-B, 3A-F; 188, f.6A-G. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy . :©Photo T. Szuts. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy . Gen. Jerzego Maddison, Piascik, 2014 Type species Jerzego corticicola (4 recognizable species). See more species at .z Jerzego -Q +M

J. Prószy ński - 2020 Salticidae (Araneae) genera of the world - an atlas Chapter 36 SYNAGELES 3 z 5

A-D - Jerzego corticicola , E - Jerzego alboguttatus IMMATURE. Sarawak, F - same (as Hispo alboguttata), G - Yepoella sp. [??] from Ecuador [??], H-O - Jerzego sunillimaye . SOURCE.SOURCE. A-D - Maddison, Piascik, 2014. Male. Sarawak. Zootaxa: 571-573, f 1-13, 14-17 © Magnolia Press, F - Wanless 1981a.. Bull. brit. Mus. nat. Hist. (Zool.)41 (4): 196-197, ff. 13A-B,, H-O - Sanap, R. V., Caleb, J. T. D. & Joglekar, A. (2019). A new species of the hisponine jumping from India (Araneae: Salticidae), with some observations on its life history. Arthropoda Selecta 28 (1): 113-124. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

ATTENTION: Color pattern of immature specimen of Jerzego alboguttatus seems insufficient to identify species and to confirm conspecificity of Maddison and Wanless specimens, it resembles also coloration of Yepoella sp. (Fig. F, above) from which it differs in eyefield not only flat but even slightly depressed. Congeneric status Jerzego sunillimaye seems uncertain. . Gen. Lystrocteisa Simon, 1884 Type species Lystrocteisa myrmex [placement uncertain] (1 recognizable species) See more species at Lystrocteisa -Q +M

A-G - Lystrocteisa myrmex ( A - male by Simon, B - male by Zabka, C-G - by © Patoleta, Gardzinska 2013), H - misidentification as "cf. Lystrocteissa" (and placed in tribe "Viciriini", whithin higher taxon "Astioida"). SOURCE: L - Simon E. 1901a. Hist. Nat. Araign.: 480 (eyes) + Zabka 1988b. Ann. zool. 41 (14): 460-461, ff. 105-109. L1 - Patoleta, Gardzinska 2013. Zootaxa 3646 (5): 588-592, f 1–26. © 2013 Magnolia Press, L2 - Maddison et al, 2008 52, f. 1. All ©copyrights are retained by the original authors and copyright holders, used here by their courtesy.

COMMENT: Isolated distribution (as far as known - New Caledonia Is), together with primitive characters (eyes arrangement, transversal depression of eyefield), complicated internal structure of epigyne combined with simple palp - all these suggest that Lystrocteisa may possibly be relict. COMMENT - 2. Palp of the male and internal structure of epigyne of female look incompatible[?]. Comparison of habitus of both, shape of cephalothorax and leg I are inconclusive. Answer require study of fresh specimens, preferably live. In male location of eyes II on the same protuberance with anterior lateral, long distance to eyes posterior lateral (III row) and transverse depression between eyes II and III row,

J. Prószy ński - 2020 Salticidae (Araneae) genera of the world - an atlas Chapter 36 SYNAGELES 4 z 5 resemble strongly Hispo , Tomocyrba and related genera. Simple palps resembles ASTIAINES and SOBASINES. However there is big diversity of genitalic structures among HISPONINES. How to interpret that? Gen. Tomobella Szüts & Scharff, 2009 Type species s Tomocyrba decollata (6 recognizable species). See more species at .z Tomobella -Q +M

A-D - Tomobella fotsy, E-G - Tomobella andasibe (C - cephalothotax of Tomomingi - note size and location of eyes II). SOURCE : Szüts, Scharff 2009. Contr. Nat. Hist. 12: 1370, f 18a-d,18E-H. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy . Gen. Tomocyrba Simon, 1900 Type species s Tomocyrba decollata (6 recognizable species). See more species at .z Tomocyrba -Q +M

A-B - Tomocyrba decollata , C-E - Tomocyrba ubicki , F - Tomocyrba barbata , G -I - Tomocyrba thaleri , I-J - Tomocyrba berniae . SOURCE. T7-T8 - Szüts, Scharff 2009. Contrib. Nat. Hist. 12: 1345, f 8; 1350-1352, f 1B, 3F, 5C, 10E–H; 1354-1356, f 1C, 2D, 4F, 11A–E; 1356, f. 1D, 4E, 11F. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy. COMMENT: there are three [?] different types of internal structure of epigyne - are they congeneric? Gen. Tomomingi Szüts & Scharff, 2009 Type species Tomomingi wastani (9 recognizable species). See more species at Tomomingi -Q +M

J. Prószy ński - 2020 Salticidae (Araneae) genera of the world - an atlas Chapter 36 SYNAGELES 5 z 5

A-B - Tomomingi wastani, C - Tomomingi kikuyu , D-G - Tomomingi nywele . . SOURCE. A-B - Szüts, Scharff 2009.Contrib. Nat. Hist. 12: 1360-1362, f 1E–F, 2E–F, 4D, 5D–E,131364, f 1A–D, C - Proszynski, Zabka 1983b. Acta zool. cracov., 26: 572, f. 22. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

J. Prószy ński - 2020 Salticidae (Araneae) genera of the world - an atlas