ISSN Versión impresa 2218-6425 ISSN Versión Electrónica 1995-1043

REVIEW ARTICLE/ ARTÍCULO DE REVISIÓN CHECKLIST OF METAZOAN PARASITES OF FISHES FROM PERU

LISTA DE VERIFICACIÓN DE LOS METAZOOS PARÁSITOS DE PECES DE PERÚ

José L. Luque1*; Celso Cruces2; Jhon Chero2; Fabiano Paschoal3; Philippe V. Alves3; Ana C. Da Silva4; Lidia Sanchez5 & José Iannacone6,7

1Departamento de Parasitologia , Universidade Federal Rural do Rio de Janeiro, Caixa Postal 74.540, Seropédica, RJ, Brazil, CEP 23851-970. 2Laboratorio de Parasitología, Facultad de Ciencias Naturales y Matemática, Universidad Nacional Federico Villarreal, El Agustino, Lima, Perú. 3Programa de Pós-Graduação em Biologia Animal, Universidade Federal Rural do Rio de Janeiro, Rodovia BR 465 – Km 7, Seropédica, RJ, Brasil, CEP 23890-000. 4Programa de Pós-Graduação em Medicina Veterinária, Faculdade de Ciências Agrárias e Veterinárias, Universidade Estadual Paulista “Júlio de Mesquita Filho”, Via de Acesso Prof. Paulo Donato Castelanne s/n, Jaboticabal, SP, Brazil, CEP 14884-900. 5Departamento de Protozoología, Helmintología e Invertebrados Afines, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Perú. 6Laboratorio de Ecologia y Biodiversidad Animal, Facultad de Ciencias Naturales y Matemática, Universidad Nacional Federico Villarreal, El Agustino, Lima, Perú. 7Laboratorio de Parasitología. Facultad de Ciencias Biológicas. Universidad Ricardo Palma, Santiago de Surco, Lima, Perú. *Corresponding author: E-mail: [email protected]

Neotropical Helminthology, 2016, 10(2), jul-dic: 301-375. ABSTRACT

A checklist of all available published accounts of metazoan parasites reported from marine and freshwater fishes in Peru is provided. This list includes records of 612 different of metazoan parasites (including unidentified parasites) belonging to the following groups: Platyhelminthes (430 total), including (175), Cestoda (140), and (115); Crustacea (99); Nematoda (49); Acanthocephala (16); Myxozoa (16) and Annelida (2). These parasites were reported from 207 fish species in Peru (174 teleost, 31 elasmobranch and 2 unspecified host species). The records of parasites unidentified to the species level (182 of 612), representing 30% of all species, were also included. Among the fish parasites reported from Peru, 391 were marine, 217 were freshwater and 4 species were found in the both environments, for a total of 1015 parasite-host associations. The fish species that harbored the highest number of parasites were Merluccius gayi peruanus Ginsburg, 1954 (32), followed by Paralonchurus peruanus (Steindachner, 1875) (31), Scomber japonicus Houttuyn, 1782 (31), Sciaena deliciosa (Tschudi, 1846) (29) and Stellifer minor (Tschudi, 1846) (28), all of which are marine fish. The parasites with the highest number of records were larvae of the cestodes Adenocephalus pacificus Nybelin, 1931 (19), larvae of the nematode sp. (18), larvae of Contracaecum sp. (17) and larvae and adults of Lacistorhynchus tenuis (16), all of which were recovered from marine fish, except for Contracaecum sp. larvae, which were recovered from both marine and freshwater environments. The parasite diversity in Peruvian fishes seems to be biased by the sampling host according to researcher interests, but it is also related to the biological features of both the parasites and the hosts.

Keywords: Biodiversity – Pacific Ocean – Parasites– South American – Neotropics – Parasites –

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RESUMEN

Se proporciona una lista de verficación de todos los registros válidos publicados de los metazoos parásitos reportados en peces marinos y de agua dulce de Perú. Esta lista incluye los registros de 612 especies diferentes de metazoos parásitos (incluyendo parásitos no identificados) que pertenecen a los siguientes grupos: Platyhelminthes (430 total), incluyendo Monogenea (175), Cestoda (140), y Trematoda (115); Crustacea (99); Nematoda (49); Acanthocephala (16); Myxozoa (14) y Annelida (2). Estos parásitos fueron reportados de 207 especies de peces de Perú (174 teleósteos, 31 elasmobranquios y 2 especies de hospederos no determinados). Los registros de los parásitos no identificados a nivel de especie (182 de 612), representando el 30% de todas las especies, también fueron incluidos. Entre los parásitos de peces reportados de Perú, 391 fueron marinos, 217 de agua dulce y 4 especies se encontraron en ambos ambientes, para un total de 1015 asociaciones parásito-hospedero. Las especies de peces que albergaron el mayor número de parásitos fueron Merluccius gayi peruanus Ginsburg, 1954 (32), seguido de Paralonchurus peruanus (Steindachner, 1875) (31), Scomber japonicus Houttuyn, 1782 (31), Sciaena deliciosa (Tschudi, 1846) (29) y Stellifer minor (Tschudi, 1846) (28), todos ellos son peces marinos. Los parásitos con el mayor número de registros fueron las larvas del cestodo Adenocephalus pacificus Nybelin, 1931 (19), larvas del nematodo Anisakis sp. (18), larvas de Contracaecum sp. (17) y larvas y adultos de Lacistorhynchus tenuis (16), los cuales fueron recuperados de peces marinos, excepto las larvas de Contracaecum sp., las cuales fueron recuperados de ambos ambientes. La diversidad de parásitos en peces peruanos parece estar sesgado por el hospedero muestreado de acuerdo a los intereses del investigador, sino que también está relacionado con las características biológicas de los parásitos y de los hospederos.

Palabras clave: América del sur – Biodiversidad – Neotrópico – Oceano Pacífico – ParásitosTaxonomia

INTRODUCTION Peru is an extremely diverse country with several distinct marine and freshwater environments. The Peruvian coast borders the There is currently a consensus that parasites Pacific Ocean and comprises approximately represent a large fraction of the Earth's total 2500 km, including the Central Peru and biodiversity (Poulin, 2014), though several Humboldtian marine ecoregions (Spaldin, questions remain unanswered regarding the 2007), with a fish richness estimated at 705 magnitude of parasite diversity and its species (Froese & Pouly, 2015). distribution in the world. Luque & Poulin (2007) noted that studies on the diversity of For freshwater environments, the Peruvian fish parasites have intensified, but these Amazon, which is well known for its high organisms remain an underestimated biodiversity, comprises approximately 62% of component of the total biodiversity in many the country and includes the second-largest regions of the globe. They also suggested that portion of the Amazon rainforest after Brazil regional differences may reflect the true (IIAP, 2011). Further, Peruvian territory biological patterns, which should be includes two biodiversity hotspots with a high considered when selecting targets for local fish number of teleosts: the Tropical Andes parasitology research. (between 400 and 600 freshwater fishes, nearly

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40% of which are endemic) and Tumbes- that were not indexed by electronic databases Chocó-Magdalena (approximately 250 but were known to the authors were also species of fishes) (Anderson & Maldonado- included. Undergraduate theses and scientific Ocampo, 2011; Conservation International, meetings do not constitute formal publications 2011). and were thus not considered. Records published up to May 2016 are included. According to Froese & Pouly (2015), there are 914 species of freshwater fishes in Peru, The following parasite taxa were included: suggesting a high diversity of fish parasites as Acanthocephala, Annelida (Hirudinea), well. However, the literature on this subject is Crustacea (Malacostraca and Maxillopoda), highly dispersed, and numerous papers have Myxozoa, Nematoda and Platyhelminthes been published in regional or local journals (Trematoda, Monogenea, and Cestoda). that are not indexed. Few researchers have compiled this information, and all have The checklist follows the classification and focused on marine fish hosts (Tantaleán et al., systematic arrangement proposed by Ahyong 1982, 1985; Luque, 1991; Luque et al., 1991; et al. (2011) for Branchiura and Isopoda; Amin Tantaleán & Huiza, 1994). More recently, (2013) for Acanthocephala; Anderson et al. Luque & Poulin (2007) counted 146 and 485 (2009) and Gibbons (2010) for Nematoda; freshwater and marine parasite-host Boxshall & Halsey (2004) for Copepoda; associations from 49 and 98 freshwater and Cohen et al. (2013) for Monogenea; Davies marine fishes, respectively. (1991) for Hirudinea; Gibson et al. (2002), Jones et al. (2005) and Bray et al. (2008) for Here, we provide a checklist of metazoan Trematoda; Khalil et al. (1994) for Cestoda; parasites that are associated with both marine and Lom & Dyková (2006) for Myxozoa. and freshwater fishes from Peru, including detailed information on their locality, habitat, Parasite species are arranged by phylum, class, site of infection/infestation and life-cycle order and family and are presented in stage. The information is presented as parasite- alphabetical order, followed by information on host and host-parasite lists. We have attempted h o s t / l o c a l i t y r e c o r d s , s i t e o f to include information from all published infection/infestation, stage of development, reports. habitat (freshwater/marine) and references (between parentheses, in chronological order).

MATERIAL AND METHODS When the taxonomic status of a parasite species required clarification, the section “Notes” is included with the relevant The database was compiled from an extensive bibliographic references. bibliographical review of metazoan parasites reported from fishes in Peruvian waters using The names of host species follow Froese & two approaches. First, a search was performed Pauly (2015). The species of fishes in the host- through the Zoological Records, Biological parasite list are arranged according to the Abstracts and Helminthological Abstracts, higher-level classification of Nelson (2006). Web of Knowledge, Google Scholar, Aquatic Parasite names were updated in agreement Sciences and Fisheries Abstracts, Biological with the recent literature, but inclusion in the and Agricultural Index Plus and Scopus, using parasite or host lists does not necessarily imply a combination of several key words. Second, that authors agree with their validity. monographs and articles published in Spanish

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RESULTS PARASITE-HOST LIST PHYLUM ACANTHOCEPHALA Rudolphi, 1802 A total of 1015 parasite-host associations were Class Eoacanthocephala Van Cleave, 1936 found, including 612 named species (including Order Neoechinorhynchida Southwell & unidentified ones) of metazoan parasites MacFie, 1925 belonging to the following taxa: Monogenea (175), Cestoda (140), Trematoda (115), Family Neoechinorhynchidae (Ward, 917) C r u s t a c e a ( 9 9 ) , N e m a t o d a ( 4 9 ) , Gorytocephalus plecostomorum Nickol & Acanthocephala (16), Myxozoa (16) and Thatcher, 1971 Annelida (2). Reports of parasites that were not Pterygoplichthys multiradiatus; adult; identified to the species level constituted 182 intestine; freshwater; Loreto (Schmidt & of the 612 (30%) species included (Table 1). Hugghins, 1973a; Tantaleán et al., 2005). Neoechinorhynchus sp. Three-hundred ninety-one species were Colossoma macropomum, Prochilodus marine parasites, 217 were freshwater and 4 nigricans; adult; intestine; freshwater; species were recorded in the both habitats Loreto, Ucayali (Schmidt & Hugghins, (Table 2). A total of 207 (113 marine, 94 1973a; Tantaleán et al., 2005). freshwater) species of fishes from Peru have Pandosentis iracundus Van Cleave, 1920 been reported as hosts of metazoan parasites. Aequidens sp., Geophagus steindachneri; O f t h e m , 1 7 4 a r e t e l e o s t s , 3 1 a r e adult; intestine; freshwater; Ucayali, elasmobranchs and 2 are unspecified host Loreto (Tantaleán et al., 2005). species. The fish species with the highest number of parasites were the marine fishes Class Palaeacanthocephala Meyer, 1931 Merluccius gayi peruanus Ginsburg, 1954 Order Echinorhynchida Southwell & (32), followed by Paralonchurus peruanus MacFie, 1925 (Steindachner, 1875) (31), Scomber japonicus Houttuyn, 1782 (31), Sciaena deliciosa Family Cavisomidae Meyer, 1932 (Tschudi, 1846) (29) and Stellifer minor Paracavisoma impudica (Diesing, 1851) (Tschudi, 1846) (28). The freshwater fishes Oxydoras niger; adult; intestine; with the most parasite records were freshwater; Ucayali (Schmidt & Pseudoplatystoma fasciatum (Linnaeus, 1766) Hugghins, 1973b; Tantaleán et al., 2005). (11) and Arapaima gigas (Schinz, 1822), Potamotrygon motoro (Müller & Henle, Family Diplosentidae Tubangui & 1841), Orestias luteus Valenciennes, 1846 Masilungan, 1937 (each with 9 reports). Parasites with the most Sharpilosentis peruviensis Lisitsyna, Scholz reports were the larval stage of the & Kuchta, 2015 diphyllobothriidean Adenocephalus pacificus Duopalatinus cf. peruanus; Oxydoras Nybelin, 1931 (19), larvae of Anisakis sp. (18), niger; adult; intestine; freshwater; Loreto larvae of Contracaecum sp. (17) and adults and (Lisitsyna et al., 2015). larvae of the trypanorhynch cestode Lacistorhynchus tenuis (16), all of which are Family Rhadinorhynchidae Lühe, 1912 parasitic in marine fishes, except for Gorgorhynchus sp. Contracaecum sp., which is parasitic in Anisotremus scapularis, Hemilutjanus freshwater and marine fish. The localities with macrophthalmos; adult; intestine, the highest number of records were Lima mesentery; marine; Lima (Tantaleán & (285), Loreto (148) and La Libertad (120). Lefevre, 2004; Chero et al., 2014d).

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Rhadinorhynchus plagioscionis Thatcher, humeralis, Paralichthys adspersus, 1980 Paralonchurus peruanus, Scartichthys Plagioscion squamosissimus; adult; gigas, Sciaena deliciosa, Scomber intestine; freshwater; Loreto (Iannacone japonicus, Stellifer minor, Trachurus & Luque, 1993b). murphyi; larvae; liver, mesentery, visceral Rhadinorhynchus pristis (Rudolphi, 1802) surface; marine; Ica, La Libertad, Scomber japonicus; adult; intestine; Lambayeque, Lima, Piura, Tumbes marine; La Libertad (Cruces et al., 2014). (López & Tantaleán, 1985; Oliva et al., Rhadinorhynchus sp. 1990; Rivera & Sarmiento, 1990a; Stellifer minor, Trachurus murphyi; adult; Tantaleán & Huiza, 1994; Cabrera & intestine; marine; Lima, Lambayeque Tantaleán, 1995; Jara, 1998; Pérez et al., (Oliva et al., 1990; Luque et al., 1991; 1999; Llerena et al., 2001; Tantaleán et al., Tantaleán & Huiza, 1994; Oliva & Luque, 2005; Chero et al., 2014ac; Paschoal et al., 1998b; Tantaleán et al., 2005). 2015; Iannacone et al., 2015; Tarmeño et Tegorhynchus sp. al., 2015; Minaya et al., 2016). Cynoscion analis, Labrisomus philippii, Corynosoma sp. Paralonchurus peruanus, Sciaena Brama japonica, Cynoscion analis, deliciosa, Trachurus murphyi; adult; O d o n t e s t h e s re g i a , P a r a l a b r a x intestine; marine, Chimbote, La Libertad, humeralis, Paralichthys adspersus, Lambayeque, Lima (Luque et al., 1991; Paralonchurus peruanus, Prionotus Tantaleán & Huiza, 1994; Jara & Ponte, stephanophrys, Seriolella violácea, 1995; Oliva & Luque, 1998b; Tantaleán et Stellifer minor, Trachurus murphyi; al., 2005; Iannacone et al., 2010a). larvae; mesentery, visceral surface; marine; Lima (Armas, 1983b; Oliva et al., Order Polymorphida Petrochenko, 1956 1990; Luque et al., 1991; Luque & Oliva, 1993a; Tantaleán & Huiza, 1994; Oliva & Family Polymorphidae Meyer, 1931 Luque, 1998b; Tantaleán et al., 2005; Bolbosoma sp. Iannacone & Alvariño, 2013; Paschoal et Merluccius gayi peruanus; larvae; al., 2015). visceral surface; marine; Lima (Chero et Profilicollis altmani (Perry, 1942) al., 2014b). Anisotremus scapularis, Labrisomus Corynosoma australe Johnston, 1937 philippii; larvae; gills, intestine; marine; Labrisomus philippii, Paralonchurus Lima (Luque, 1991; Tantaleán & Huiza, peruanus, Sciaena deliciosa, Sebastes 1994; Tantaleán et al., 2005; Iannacone & capensis, Trachurus murphyi; larvae; Alvariño, 2009b, Chero et al., 2014d; abdominal cavity, gills; marine; Lima, Paschoal et al., 2015; Cruces et al., 2015). Piura (Luque & Oliva, 1993b; Oliva, P o l y m o r p h u s s p i n d l a t u s A m i m & 1999; Oliva & Luque, 1998b, 2002; Heckmann, 1991 González & Poulin, 2005; Tantaleán et al., Orestias agassizii, O. luteus, O. mulleri, 2005; González et al., 2006; Iannacone et O. olivaceus, Orestias sp., Oreochromis al., 2011b; Cruces et al., 2015). niloticus; larvae; body cavity; freshwater; Corynosoma obtuscens Lincicome, 1943 Puno (Amin, 1995; Tantaleán et al., 2005; Cilus gilberti, Coryphaena hippurus, Amin et al., 2010). C y n o s c i o n a n a l i s , G a l e i c h t h y s peruvianus, Isacia conceptionis, PHYLUM ANNELIDA Linnaeus, 1758 Labrisomus philippii, Merluccius gayi Class Hirudinea Lamarck, 1894 peruanus, Odontesthes regia, Paralabrax Order Rhynchobdellida Blanchard, 1893

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Family Piscicolidae Johnston, 1865 marine; Lima (Luque & Farfán, 1990c; Piscicolidae gen. sp. Luque et al., 1991; Tantaleán & Huiza, Cilus gilberti, Labrisomus philippii, 1994). Paralonchurus peruanus, Sciaena Bomolochus constrictus (Cressey & deliciosa; adult; gills; marine; Lima Collette, 1970) (Luque & Oliva, 1993a; Oliva & Luque, Ablennes hians, Strongylura exilis, 1998b; Iannacone et al., 2011b; Chero et Tylosurus acus acus; adult; gills; marine; al., 2014c). Piura (Cressey & Collette, 1970; Luque et Stibarobdella moorei (Oka, 1910) al., 1991; Tantaleán & Huiza, 1994). Rhinobatos planiceps; adult; skin; marine; Bomolochus ensiculus (Cressey, in Cressey Lima (Iannacone et al., 2011a) & Collette, 1970) Strongylura exilis; adult; gills; marine; PHYLUM ARTHROPODA von Siebold, Lima, Tumbes (Cressey & Collette, 1970; 1848 Luque et al., 1991; Tantaleán & Huiza, Class Maxillopoda Dahl, 1956 1994). Order Arguloida Yamaguti, 1963 Bomolochus nitidus Wilson, 1911  Mugil cephalus; adult; gills; marine; Family Leach, 1819 Lima (Iannacone & Alvariño, 2009c). Argulus peruvianus Oliva, Durán & Verano Bomolochus sp. 1980 Paralichthys adspersus, Peprilus medius; Merluccius peruanus; adult; eyes, gills, adult; gills; marine; Lima (Iannacone & opercule, skin; marine; Lima (Durán & Alvariño, 2008; Iannacone et al., 2012a). Oliva, 1980; Oliva et al., 1980; Luque et Hamaticolax galeichthyos (Luque & Bruno, al., 1991; Tantaleán & Huiza, 1994). 1990) Dolops geayi (Bouvier, 1897) Galeichthys peruvianus; adult; gills; Cichlasoma amazonarum; adult; skin; marine; Lima (Luque & Bruno, 1990b; freshwater; Loreto (Lo et al., 2011) Luque et al., 1991; Iannacone & Luque, 1993a; Tantaleán & Huiza, 1994). Order Cyclopoida Burmeister, 1834 Hamaticolax paralabracis (Luque & Bruno, 1990) Family Bomolochidae Sumpf, 1871 Paralabrax humeralis; adult; gills; Acanthocolax peruensis (Luque & Bruno, marine; Lima (Luque & Bruno, 1990b; 1990) Luque et al., 1991; Tantaleán & Huiza, M e n t i c i r r h u s o p h i c e p h a l u s , 1994). Paralonchurus peruanus, Sciaena deliciosa, Stellifer minor; adult; gills; Family Chondracanthidae Milne marine; Lima (Luque & Bruno, 1990a; Edwards, 1840 Luque et al., 1991; Luque & Oliva, 1993a; Acanthochondria sicyasis (Krøyer, 1863) Luque, 1994a). Labrisomus philippii, Scartichthys gigas, Acantholochus nudiusculus (Cressey & Sicyases sanguineus; adult; gills; marine; Cressey, 1980) Lima (Durán & Oliva, 1983; Luque et al., Scomberomorus sierra; adult; gills; 1991; Farfán et al., 1993; Tantaleán & marine; Northern coastal zone (Cressey & Huiza, 1994; Iannacone & Alvariño, 2011; Cressey, 1980; Luque et al., 1991; Iannacone et al., 2011b; Iannacone et al., Tantaleán & Huiza, 1994). 2012b). Bomolochus bellones Burmeister, 1833 Blias prionoti Krøyer, 1863 Strongylura scapularis; adult; gills; Hemanthias peruanus, Prionotus

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punctatus; adult; gills; marine; Northern Schotte, 1983 coastal zone, Piura (Ho, 1970; Luque et A n i s o t re m u s s c a p u l a r i s ; a d u l t ; al., 1991; Tantaleán & Huiza, 1994). mandibular canals; marine; Lima (Luque Chondracanthus lepophidii Ho, 1974 & Farfán, 1990c; Luque et al., 1991; Cherublemma emmelas; adult; gills; Tantaleán & Huiza, 1994). marine; Lambayeque, Northern coastal zone (Ho, 1974; Luque et al., 1991; Order Siphonostomatoida Thorell, 1859 Tantaleán & Huiza, 1994). Chondracanthus sp. Family Caligidae Burmeister, 1835 Coryphaena hippurus, Merluccius gayi Caligodes laciniatus (Krøyer, 1863) peruanus; adult; gills; marine; La Ablennes hians, Tylosurus pacificus; Libertad, Northern coastal zone (Jara & adult; mouth; marine; Tumbes (Cressey & Díaz-Limay, 1995; Vásquez-Ruiz et al., Collette, 1970; Luque et al., 1991; 2012). Tantaleán & Huiza, 1994). Protochondria longicauda Ho, 1970 Caligus bonito Wilson, 1905 Hippoglossina bollmani; adult; gills; Sarda chiliensis, Trachurus murphyi; marine; Lambayeque, Northern coastal adult; gills; marine; La Libertad, Lima, zone (Ho, 1970; Luque et al., 1991; Northern coastal zone (Durán, 1980; Tantaleán & Huiza, 1994). Durán & Oliva, 1983; Cressey & Cressey, 1980; Luque et al., 1991; Tantaleán & Family Ergasilidae von Nordmann, 1832 Huiza, 1994; Jara & Díaz-Limay, 1995). Amplexibranchius bryconis Thatcher & Caligus callaoensis Durán, 1980 Paredes, 1985 Cheilotrema fasciatum, Paralonchurus Brycon cephalus; adult; gills; freshwater; peruanus, Sciaena deliciosa; adult; gills, Loreto (Thatcher & Paredes, 1985b). internal surface of the operculum; marine; Ergasilus sp. La Libertad, Lima (Durán, 1980; Luque et Mugil cephalus; adult; gills; marine; La al., 1991; Luque & Oliva, 1993a; Luque, Libertad (Jara & Díaz-Limay, 1995. 1996; Oliva & Luque, 1998b; Tantaleán & Huiza, 1994; Jara & Díaz-Limay, 1995; Family Lernaeidae Cobbold, 1879 Iannacone et al., 2010a). Lernaea gamitanae (Thatcher & Paredes, Caligus cheilodactyli Krøyer, 1863 1985) Cheilodactylus variegatus; adult; gills, Colossoma macropomum; adult; skin; skin; marine; Lima (Luque & Farfán, freshwater; Loreto (Thatcher & Paredes, 1990c; Luque et al., 1991; Tantaleán & 1985a; Mathews et al., 2011). Huiza, 1994; Iannacone et al., 2003). Lernaea sp. Caligus coryphaenae Streenstrup & Lütken, Labrisomus philippii; adult; gills; marine; 1861 Trujillo (Jara & Díaz-Limay, 1995). Coryphaena hippurus, Katsuwonus pelamis; adult; gills; marine; Northern Family Philichthyidae Vogt, 1877 coastal zone (Cressey & Cressey, 1980; Colobomatus goodingi Cressey & Collette, Luque et al., 1991; Tantaleán & Huiza, 1970 1994; Vásquez-Ruiz et al., 2012). Tylosurus acus acus; adult; cephalic Caligus debueni Stuardo & Fagetti, 1961 channels; marine; Piura (Cressey & Merluccius gayi peruanus; adult; gills; Collette, 1970; Luque et al., 1991; marine; Lima (Durán, 1980; Durán & Tantaleán & Huiza, 1994). Oliva, 1980; Luque et al., 1991; Tantaleán Colobomatus quadrifarius Cressey & & Huiza, 1994; Chero et al., 2014b).

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Caligus omissus Cressey & Cressey, 1980 Lepeophtheirus marginatus Bere, 1936 Scomberomorus sierra; adult; gills; Galeichthys sp.; adult; skin; marine; Piura marine; Northern coastal zone (Cressey & (Shiino & Izawa, 1966; Luque et al., 1991; Cressey, 1980; Luque et al., 1991; Tantaleán & Huiza, 1994). Tantaleán & Huiza, 1994). Lepeophtheirus sp. Caligus pelamydis Krøyer, 1863 Merluccius gayi peruanus; adult; gills; Scomber japonicus; adult; unspecified site marine; Lima (Durán & Oliva, 1980; of infection; marine; Lima (Oliva et al., Tantaleán & Huiza, 1994). 2008). Caligus productus Dana, 1852 Family Dichelesthiidae Milne Edwards, Katsuwonus pelamis; adult; gills; marine; 1840 Lima, Northern coastal zone (Cressey & Anthosoma crassum (Abildgaard, 1794) Cressey, 1980; Luque et al., 1991; Isurus oxyrinchus, Prionace glauca; Tantaleán & Huiza, 1994). adult; mouth; marine; 08°47'-33°24'S, Caligus quadratus Shiino, 1954 77°02'-89°31'W (Cressey, 1968; Paralabrax humeralis, Stellifer minor, Tantaleán & Huiza, 1994). Trachinotus paitensis; adult; gills, mouth; marine; La Libertad, Lima (Luque & Family Eudactylinidae Wilson, 1992 Farfán, 1990c, 1991a; Oliva et al., 1990; Eudactylina myliobatidos Luque & Farfán, Luque et al., 1991; Luque, 1996; Oliva & 1991 Luque, 1998b; Jara & Díaz-Limay, 1995). Myliobatis chilensis; adult; gills; marine; Caligus teres Wilson, 1905 Lima (Luque & Farfán, 1991c). Merluccius gayi peruanus; adult; gills; Eudactylina peruensis Luque & Farfán, marine; Lima (Durán, 1980; Luque et al., 1991 1991; Tantaleán & Huiza, 1994). Rhinobatos planiceps; adult; gills; Caligus sp. marine; Lima (Luque & Farfán, 1991c). C o r y p h a e n a h i p p u r u s , I s a c i a Nemesis lamna lamna Risso, 1826 conceptionis, Peprilus medius, Sarda Isurus oxyrinchus; adult; gills; marine; chiliensis, Tylosurus pacificus; adult; 10°28'-33°24'S, 77°02'-89°31'W gills; marine; Ica, La Libertad, Lima, (Cressey, 1968; Tantaleán & Huiza, Piura (Chirinos de Vildoso, 1955; Cressey 1994). & Collette, 1970; Tantaleán & Huiza, 1994; Iannacone & Alvariño, 2008; Jara & Family Hatschekiidae Kabata, 1979 Díaz-Limay, 1995). Hatschekia amphiprocessa Castro-Romero Gloiopotes huttoni (Thomson 1890) & Baeza-Kuroki, 1986 Makaira sp., Xiphias gladius; adult; skin; Paralabrax humeralis; adult; gills; marine; Piura (Shiino, 1963; Luque et al., marine; Lima (Luque & Farfán, 1990c; 1991; Tantaleán & Huiza, 1994). Luque et al., 1991; Tantaleán & Huiza, Gloiopotes watsoni Kirtisinghe, 1934 1994). Makaira nigricans; adult; unspecified site Hatschekia conifera Yamaguti, 1939 of infection; marine; unspecified locality Brama japonica; adult; gills; marine; (Cressey, 1967a). Lima (Iannacone & Alvariño, 2013). Lepeophtheirus edwardsi Wilson, 1905 Paralichthys adspersus; adult; skin; Family Lernaeopodidae Milne-Edwards, marine; Lima (Luque & Farfán, 1990c; 1840 Luque et al., 1991; Tantaleán & Huiza, Brachiella sp. 1994). Merluccius gayi peruanus; adult; gills;

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marine; Lima (Durán & Oliva, 1980). & Baeza-Kuroki, 1989 Clavella applicata Castro-Romero & Baeza- Anisotremus scapularis; adult; marine; Kuroki, 1985 Lima (Iannacone & Alvariño, 2009b; Anisotremus scapularis; adult; gills; Paschoal et al., 2015). marine; Lima (Luque & Farfán 1990c; Parabrachiella chevreuxii (Van Beneden, Luque et al., 1991; Tantaleán & Huiza, 1891) 1994; Iannaconne & Alvariño, 2009b; Paralonchurus peruanus; adult; gills; Paschoal et al., 2015). marine; Lima (Luque & Farfán, 1990b; Clavella simplex Castro-Romero & Baeza- Luque et al., 1991; Luque & Oliva, 1993a; Kuroki, 1985 Tantaleán & Huiza, 1994; Luque, 1996). Isacia conceptionis; adult; caudal Parabrachiella exilis (Shiino, 1956) peduncle, fins; marine; Lima (Luque & Mugil cephalus; adult; gills; marine; Lima Farfán, 1990c; Luque et al., 1991; (Luque & Farfán, 1990c). Tantaleán & Huiza, 1994; Iannacone et Parabrachiella fasciata (Castro-Romero & al., 2015). Baeza-Kuroki, 1987) Clavella sp. Cheilotrema fasciatum; adult; gills; Stellifer minor; adult; gills; marine; Lima marine; Lima (Luque & Farfán, 1990b; (Tantaleán & Huiza, 1994). Luque et al., 1991; Tantaleán & Huiza, Clavellisa scombri (Kurz, 1877) 1994; Luque, 1996). Scomber japonicus; adult; gills; marine; Parabrachiella insidiosa (Heller, 1865) La Libertad, Lima (Cressey & Cressey, Merluccius gayi peruanus; adult; gills; 1980; Luque et al., 1991; Tantaleán & marine; Lima (Durán & Oliva, 1980, Huiza, 1994; Oliva et al., 2008; Cruces et 1983; Luque et al., 1991; Tantaleán & al., 2014). Huiza, 1994; Chero et al., 2014b). Clavellopsis sp. Parabrachiella kabatai (Luque & Farfán, Paralonchurus peruanus; adult; gills; 1991). marine; Lima (Luque & Oliva, 1993a; Isacia conceptionis; adult; fins; marine; Luque, 1996; Oliva & Luque, 1998b). Lima (Luque & Farfán, 1991b; Tantaleán Clavellotis dilatata (Krøyer, 1863) & Huiza, 1994). Cheilodactylus variegatus, Menticirrhus Parabrachiella menticirrhi (Luque & ophicephalus, Stellifer minor; adult; gills; Farfán, 1990) marine; La Libertad, Lima (Luque & Menticirrhus ophicephalus; adult; gills; Farfán, 1990b; Oliva et al., 1990; Luque et marine; Lima (Luque & Farfán, 1990b; al., 1991; Luque, 1994a, 1996; Tantaleán Luque, 1994a; 1996; Tantaleán & Huiza, & Huiza, 1994; Oliva & Luque, 1998b; 1994; Oliva & Luque, 1998b). Jara & Díaz-Limay, 1995; Iannacone et Parabrachiella oralis (Castro-Romero & al., 2003; Iannacone, 2004). Baeza-Kuroki, 1987) Clavellotis sp. Sciaena deliciosa; adult; gills; marine; Sciaena deliciosa; adult; gills, palate; Lima (Luque & Farfán, 1990b; Luque et marine; Lima (Tantaleán & Huiza, 1994). al., 1991; Tantaleán & Huiza, 1994; Naobranchia lizae (Krøyer, 1863) Luque, 1996; Oliva & Luque, 1998b). Mugil cephalus; adult; gills; marine; Lima Parabrachiella paralichthyos (Castro- (Tantaleán & Huiza, 1994). Romero & Baeza-Kuroki, 1986) Ommatokoita elongata (Grant, 1827) Paralichthys adspersus; adult; gills; Rhinobatos planiceps; adult; eyes, skin; marine; Lima (Luque & Farfán, 1990c; marine; Lima (Iannacone et al., 2011a). Luque et al., 1991; Tantaleán & Huiza, Parabrachiella anisotremi Castro-Romero 1994).

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Parabrachiella sp. Cynoscion analis, Paralonchurus Cilus gilberti; adult; gills; marine; Lima peruanus; adult; gills; marine; Lima (Chero et al., 2014c). (Luque et al., 1989, 1991; Luque & Oliva, Pseudocharopinoides myliobatides Castro- 1993a; Luque, 1996; Oliva & Luque, Romero & Baeza-Kuroki, 1987 1998b; Iannacone et al., 2001; Iannacone, Myliobatis chilensis; adult; gills; marine; 2005). Lima (Luque & Farfán, 1991c). Lernanthropus tylosuri Richiardi, 1880 Pseudocharopinus narcinae (Pillai, 1962) Ablennes hians, Strongylura exilis; adult; Myliobatis chilensis; adult; gills; marine; gills; marine; Piura (Cressey & Collette, Lima (Luque & Farfán, 1991c). 1970; Luque et al., 1991; Tantaleán & Huiza, 1994). Family Lernanthropidae Kabata, 1979 Lernanthropus sp. Lernanthropinus trachuri (Brian, 1903) Cynoscion analis, Sciaena deliciosa, Seriolella violacea, Trachurus murphyi; Trachinotus paitensis; adult; gills; marine; adult; gills; marine; La Libertad, Lima La Libertad, Lima (Tantaleán & Huiza, (Durán & Oliva, 1983; Luque et al., 1989, 1994; Jara & Díaz-Limay, 1995). 1991; Tantaleán & Huiza, 1994; Jara & Díaz-Limay, 1995; Iannacone, 2003). Family Pandaridae Milne Edwards, 1840 Lernanthropus antofagastensis Castro- Achtheinus oblongus Wilson, 1908 Romero & Baeza-Kuroki, 1985 Prionace glauca; adult; skin; marine; Anisotremus scapularis; adult; gills; Lima (Luque & Farfán, 1991c). marine; La Libertad, Lima (Luque et al., Dinemoura latifolia (Steenstrup & Lütken, 1989; Luque et al., 1991; Tantaleán & 1861) Huiza, 1994; Jara & Díaz-Limay, 1995; Isurus oxyrinchus; adult; skin; marine; Iannaconne & Alvariño, 2009b; Paschoal 12°50'-33°24'S, 77°02'-83°31'W et al., 2015). (Cressey, 1968; Tantaleán & Huiza, Lernanthropus huamani Luque & Farfán, 1994). 1990 Dinemoura producta (Muller, 1785) Cynoscion analis, Menticirrhus Isurus oxyrinchus; adult; fins; marine; ophicephalus, Sciaena deliciosa; adult; 08°47'-33°24'S, 77°08'-83°38'W gills; marine; Lima, Tumbes (Luque & (Cressey, 1968; Tantaleán & Huiza, Farfán, 1990a; Luque et al., 1991; 1994). Tantaleán & Huiza, 1994; Luque, 1996; Echthrogaleus denticulatus Smith, 1873 Oliva & Luque, 1998b; Iannacone et al., Isurus oxyrinchus; adult; skin; marine; 2010a; Minaya et al., 2016). 08°47'-33°24'S, 79°00'-83°38'W Lernanthropus pacificus Oliva & Durán, (Cressey, 1968; Tantaleán & Huiza, 1982 1994). Cilus gilberti, Mugil cephalus, Sciaena Echthrogaleus pellucidus Shiino, 1963 deliciosa; adult; gills; marine; La Alopias sp.; adult; skin; marine; 08°47'- Libertad, Lima, Northen Coastal zone 33°24'S, 77°02'-89°31'W (Cressey, (Oliva & Durán, 1982; Durán & Oliva, 1967b). 1983; Luque et al., 1991; Tantaleán & Gangliopus pyriformis Gerstaecker, 1854 Huiza, 1994; Jara & Díaz-Limay, 1995; Isurus oxyrinchus, Prionace glauca; Vásquez-Ruiz et al., 2012; Chero et al., adult; skin; marine; 32°02'S, 77°02'W 2014c). (Cressey, 1968). Lernanthropus paralonchuri Luque, Bruno Pandarus katoi Cressey, 1967 & Covarrubia, 1989 Isurus oxyrinchus; adult; skin; marine;

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08°47'-12°50'S, 81°20'-83°38'W 2010). (Cressey, 1968; Tantaleán & Huiza, 1994). Family Trebiidae Wilson, 1905 Pandarus satyrus Dana, 1849 Trebius latifurcatus Wilson, 1921 Isurus oxyrinchus; adult; skin; marine; Myliobatis chilensis; adult; gills; marine; 08°47'-33°24', 79°00'-83°33'S (Cressey, Lima (Luque & Farfán, 1991b). 1968; Tantaleán & Huiza, 1994). Pandarus smithii Rathbun, 1886 Class Malacostraca Latreille, 1802 Unspecified host (Chondrichthyes); adult; Order Isopoda Latreille, 1817 skin; marine; Tumbes (Shiino & Izawa, 1966). Family Cymothoidae Leach, 1814 Perissopus dentatus Steenstrup & Lütken, Anilocra huacho Rokicki, 1984 1861 Auxis thazard thazard, Cubiceps Galeorhinus galeus; adult; skin; marine; caeruleus; adult; skin; marine; Lima unspecified locality (Wilson, 1911). (Thatcher, 2002). Phyllothyreus cornutus (Milne Edwards, Anilocra laevis Miers, 1877 1840) Unspecified host; adult; unspecified site Isurus oxyrinchus; adult; skin; marine; of infection; marine; unspecified locality 10°28'-33°24'S, 79°00'-81°31'W (Williams & Williams, 1981). (Cressey, 1968; Tantaleán & Huiza, Asotana formosa Schiödte & Meinert, 1881 1994). Unspecified host; adult; mouth; freshwater; Amazon River Basin Family Pennellidae Burmeister, 1835 (Schiödte & Meinert, 1881). Metapeniculus anofagastensis Castro- Ceratothoa gaudichaudii (Milne Edwards, Romero & Baeza-Kuroki, 1985 1840) Anisotremus scapularis; adult; fins; Cynoscion analis, Mustelus sp., Peprilus marine; Lima (Luque et al., 1989, 1991; medius, Sarda chiliensis, Scartichthys Luque & Fárfan, 1990c; Tantaleán & gigas, Sciaenidae gen sp., Scomber Huiza, 1994; Iannacone & Alvariño, japonicus, Stromateus stellatus, 2009b; Paschoal et al., 2015). Trachurus murphyi; adult; gills, mouth; marine; Arequipa, La Libertad, Lima Family Pseudocycnidae Wilson, 1922 (Chirinos De Vildoso, 1955; Del Solar et Cybicola buccata (Wilson, 1922) al., 1970; Wrzesinski, 1982; Tantaleán & Scomberomorus sierra; adult; gills; Huiza, 1994; Iannacone et al., 2001, Oliva marine; Tumbes (Cressey & Cressey, et al., 2008; Iannacone & Alvariño, 2008; 1980; Luque et al., 1991; Tantaleán & Iannacone et al., 2010; Iannacone et al., Huiza, 1994). 2012b, Cruces et al., 2014). Pseudocycnus appendiculatus Heller, 1865 Ceratothoa sp. Thunnus albacares; adult; gills; marine; M e n t i c i r r h u s o p h i c e p h a l u s , Tumbes (Cressey & Cressey, 1980; Luque Paralonchurus peruanus, Scomber et al., 1991; Tantaleán & Huiza, 1994). japonicus; adult; gills; marine; Lima (Luque & Oliva, 1993a; Luque, 1994a; Family Sphyriidae Wilson, 1909 Oliva & Luque, 1998b; Oliva et al., 2008). Lophoura unilobulata Castro & Gonzalez, Cymothoa oestrum (Linnaeus, 1758) 2009 Unspecified host; adult; mouth; marine; Nezumia stelgidolepis; adult; near caudal unspecified locality (Del Solar et al., fin; marine; Lambayeque (Gomez et al., 1970; Tantaleán & Huiza, 1994).

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Cymothoa sp. Mugil cephalus; gills, liver, spleen, A n i s o t re m u s s c a p u l a r i s ; a d u l t ; intestine, muscles, mesentery, cartilage, unspecified site of infection; marine; gonads; marine; Ancash, Lima (Luque et Lima (Iannacone & Alvariño, 2009b; al., 1991; Tantaleán & Huiza, 1994). Paschoal et al., 2015). Myxobolus sp. Cymothoidae gen. sp. Piaractus brachypomus; skin body, fins; Stellifer minor; adult; mouth; marine; freshwater; Amazon River basin Lima (Oliva et al., 1990; Luque et al., (Vásquez, 2014). 1991; Tantaleán & Huiza, 1994; Oliva & Myxobolidae gen. sp. Luque, 1998b). Colossoma macropomum; gills, fins; freshwater; Loreto (Bences et al., 2013). Family Aegidae White, 1850 gen. sp. Rocinela sp. Labrisomus philippii, Merluccius gayi Scomber japonicus; adult; unspecified site peruanus; gills, liver, spleen, intestine, of infection; marine; Lima (Oliva et al., muscles, mesentery, cartilage, gonads; 2008). marine; La Libertad, Lima, Piura (Okada et al., 1981; Iannacone et al., 2011b). PHYLUM MYXOZOA Grassé, 1970 Thelohanellus sp. Class Myxosporea Bütschli, 1881 Carassius auratus; gills; freshwater; Lima (Gonzáles-Fernández, 2012a). Myxosporea fam. gen. sp. Crenicichla anthurus; tegument; Order Multivalvulida Shulman, 1959 freshwater; Amazon River basin (Aguinaga et al., 2015). Family Kudoidae Meglitsch, 1960 Kudoa caudata Kovaleva & Gaevskaya, Order Bivalvulida Shulman, 1959 1983 Scomber japonicus; muscles; marine; Family Ceratomyxidae Doflein, 1899 Southeastern Pacific Ocean (Kovaleva & Ceratomyxa limensis Verano, Llican & Gaevskaya, 1983; Eiras et al., 2014). Terán, 1998 Kudoa estela Verano, 1987 Merluccius gayi peruanus; gallbladder; Prionotus stephanophrys; muscles; marine; Lima, Piura (Verano et al., 1998). marine; Lima (Tantaleán & Huiza, 1994). Ceratomyxa sp. Note: This species appeared for the first Menticirrhus ophicephalus; gallbladder; time in an unpublished doctoral marine; Lima (Gonzáles & Tam, 1993). dissertation (Verano, 1987), that nomenclaturally does not constitute valid Family Myxidiidae Thélohan, 1892 publication, so the name only became Zschokkella sp. available with publication at work of Carassius auratus; vesícula biliar; Tantaleán & Huiza (1994). freshwater; Lima (Gonzáles-Fernández, Kudoa peruviana Mateo, 1972 2012a). Merluccius gayi peruanus; muscles; marine; Ancash, La Libertad, Lima, Piura Family Myxobolidae Thélohan, 1892 (Durán & Oliva, 1980; Luque et al., 1991; Henneguya sp. Jara & Escalante, 1993; Tantaleán & Paralonchurus peruanus; mesentery; Huiza, 1994; Oliva & Ballón, 2002; Eiras marine; Lima (Tantaleán & Huiza, 1994). et al., 2014; Chero et al., 2015a). Myxobolus exiguus Thélohan, 1895

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Kudoa sarmientae Verano, 1987 Coryphaena hippurus, Sarda chiliensis, Odontesthes regia; muscles; marine; Scomber japonicus, Trachurus murphyi, Lima (Tantaleán & Huiza, 1994). larvae; mesentery; marine; Lima (Pérez et Note: This species appeared for the first al., 1999; Sarmiento et al., 1999; Llerena time in an unpublished doctoral et al., 2001; Cabrera & Trillo-Altamirano, dissertation (Verano 1987), that 2004). nomenclaturally does not constitute valid (Rudolphi, 1809) publication, so the name only became Merluccius gayi peruanus, Scomber available with publication at work of japonicus, Trachurus murphyi; larvae; Tantaleán & Huiza (1994). liver, mesentery, visceral surface; marine; Kudoa sciaenae Terán, Llicán & Luque, La Libertad, Lima (Pérez et al., 1999; 1990 Sarmiento et al., 1999; Llerena et al., Cheilotrema fasciatum, Paralonchurus 2001; Oliva & Ballón, 2002; Chero et al., peruanus, Sciaena deliciosa, Stellifer 2014b). minor; muscles; marine; Lima (Terán et Anisakis sp. al., 1990; Luque et al., 1991; Oliva et al., Cilus gilberti, Coryphaena hippurus, 1992; Tantaleán & Huiza, 1994; Eiras et C y n o s c i o n a n a l i s , G a l e i c h t h y s al., 2014). peruvianus, Merluccius gayi peruanus, Kudoa sp. Mustelus dorsalis, Paralichthys Odontesthes regia; muscles; marine; adspersus, Paralonchurus peruanus, Northern coastal zone (Escalante et al., Peprilus snyderi, Prionace glauca, 1982c; Luque et al., 1991; Tantaleán & Sardinops sagax, Sciaena deliciosa, Huiza, 1994). Sciaena wieneri, Scomber japonicus, Scorpaena russula, Seriolella violacea, PHYLUM NEMATODA Rudolphi, 1808 Stellifer minor; Trachurus murphyi; Class Adenophorea Linstow, 1905 larvae; liver, mesentery, stomach, viscera, Order Enoplida Filipjev, 1929 visceral surface; marine; Ica, La Libertad, Lambayeque, Lima, Northen Coastal Family Capillariidae Railliet, 1915 zone (Tantaleán, 1972; Tantaleán & Capillaria sp. Huiza, 1994; Durán & Oliva, 1980; Brycon cephalus, Poecilia reticulata; Melendez & Vasquez, 1981; Luque et al., adult; intestine; freshwater; Ucayali 1991; Cabrera, 1994; Jara, 1998; (Tantaleán et al., 1985a; Sarmiento et al., Sarmiento et al., 199; Llerena et al., 2001; 1999). Oliva et al., 2008; Vásquez-Ruiz et al., 2012; Chero et al., 2014a; Ñacari & Class Secernentea Linstow, 1905 Sánchez, 2014). Order Ascaridida Skrjabin & Shulz, 1940 Contracaecum multipapillatum (Drasche, 1882) Family Anisakidae (Railliet & Henry, Mugil cephalus; larvae; mesentery; 1912) marine; Lima, Northen Coastal zone Acanthocheilus bicuspis (Wedl, 1855) (Iannacone & Alvariño, 2009c; Vásquez- Mustelus dorsalis, M. mento, Triakis Ruiz et al., 2012). maculata; adult; stomach; marine; La Contracaecum sp. Libertad, Lima, Piura (Tantaleán, 1991; Aequidens rivulatus; Astronotus Tantaleán & Huiza, 1994; Sarmiento et ocellatus; Brama japonica, Brycon al., 1999). atrocaudatus, Coryphaena hippurus, Anisakis physeteris (Baylis, 1923) Ethmidium maculatum, Merluccius gayi

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peruanus, Mugil cephalus, Mugil curema, freshwater; Ucayali (Tantaleán et al., Oreochromis niloticus, Orestias albus, O. 1985a; Sarmiento et al., 1999). luteus, Paralichthys adspersus, Sardinops Raphidascaris sp. sagax, Scomber japonicus, Trachurus Cilus gilberti; adult; intestine; marine; murphyi, Trichomycterus dispar; larvae; Lima (Chero et al., 2014c). kidney, mesentery, muscles, visceral surface; freshwater, marine; Ancash, Ica, Family Cosmocercidae Railliet, 1916 La Libertad, Lambayeque, Lima, Loreto, Perunema escomeli Mateo, 1970 Piura, Puno (Armas, 1979; López & Odontesthes bonariensis; adult; intestine; Tantaleán, 1985; Tantaleán et al., 1985a; freshwater; Puno (Mateo, 1970). Luque et al., 1991; Tantaleán & Huiza, Note: This was considered as genus 1994; Pérez et al., 1999; Sarmiento et al., inquirendum by Moravec (1998). 1999; Iannacone et al., 2012a; Iannacone & Alvariño, 2013; Chero et al., 2014b; Family Kathlaniidae (Lane, 1914) Cruces et al., 2014; Chiclla-Salazar & Falcaustra pumacahuai Ibañez & Córdova, Tantas-García, 2015; Serrano-Martínez et 1976 al., 2015b). Orestias luteus; adult; intestine; Goezia spinulosa (Diesing, 1839) freshwater; Puno (Ibañez & Córdova, Arapaima gigas; adult; intestine; 1976; Sarmiento et al., 1999). freshwater; Loreto (Tantaleán et al., Note: This species was considered as 1985a; Sarmiento et al., 1999). species inquirendum by Moravec (1998). Hysterothylacium aduncum (Rudolphi, Falcaustra tiahuanaquensis Ibañez & 1802) Córdova, 1976 Coryphaena hippurus, Sarda chiliensis; Orestias luteus; adult; intestine; larvae; mesentery; marine; Lima (Pérez et freshwater; Puno (Ibañez & Córdova, al., 1999; Sarmiento et al., 1999). 1976; Sarmiento et al., 1999). Hysterothylacium pelagicum Deardorff & Note: This species was considered as Overstreet, 1982 species inquirendum by Moravec (1998). Coryphaena hippurus; adult; caecum, intestine, stomach; marine; Lima Family Cucullanidae Cobbold, 1864 (Gomez-Puerta, 2014). Cucullanus (Cucullanus) grandistomis Hysterothylacium sp. (Ferraz & Thatcher, 1988) Coryphaena hippurus, Paralichthys Oxydoras niger; adult; intestine; adspersus, Peprilus snyderi, Stromateus freshwater; Loreto (Iannacone & Luque, stellatus; larvae; intestine, stomach; 1993b). marine; Lima, Northen Coastal zone Dichelyne (Cucullanellus) amaruincai (Sarmiento et al., 1999; Iannacone et al., (Freitas, Vicente & Ibañez, 1969) 2010, 2012a; Vásquez-Ruiz et al., 2012; Paralonchurus peruanus, Sciaena Ñacari & Sánchez, 2014). deliciosa; adult; intestine; marine; La Pseudoterranova decipiens (Krabbe, 1878) Libertad, Lima, Piura (Freitas et al., 1969; Scomber japonicus, Trachurus murphyi; Luque et al., 1991; Luque & Oliva, 1993a; larvae; muscles; marine; Lambayeque, Tantaleán & Huiza, 1994; Oliva & Luque, Lima (Luque et al., 1991; Tantaleán & 1998b; Sarmiento et al., 1999; Iannacone Huiza, 1994; Sarmiento et al., 1999; et al., 2010a, b; Chero et al., 2014a). Cabrera & Trillo-Altamirano, 2004). Dichelyne (Cucullanellus) elongatus Raphidascaroides sp. (Tornquist, 1931) Oxydoras niger; larvae; mesentery; Cilus gilberti; adult; intestine; marine;

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Lima (Chero et al., 2014c). Pereira, 1928 Dichelyne (Cucullanellus) sp. Triportheus angulatus; adult; intestine; Paralabrax humeralis; adult; intestine; freshwater; Ucayali (Iannacone, 2000). marine; Lima, Piura (Armas, 1983a; Procamallanus (Spirocamallanus) pereirai Luque et al., 1991; Tantaleán & Huiza, Annereaux, 1946 1994; Iannacone & Alvariño, 2009). Stellifer minor; adult; intestine; marine; Lima (Iannacone, 2004). Order Oxyurida Chabaud, 1974 Procamallanus (Spirocamallanus) rarus Travassos, Artigas & Pereira, 1928 Family Pharyngodonidae Travassos, 1919 Pimelodus maculatus; adult; intestine; Cosmoxynemoides aguirrei Travassos, 1949 freshwater; Huánuco (Tantaleán et al., Geophagus steindachneri; adult; 1983; Sarmiento et al., 1999). intestine; freshwater; Loreto (Tantaleán et Procamallanus (Spirocamallanus) sp. al., 1985a; Sarmiento et al., 1999). Aequidens rivulatus, Bryconamericus Ichthyouris ro Inglis, 1962 peruanus, Cynoscion analis, Lebiasina Geophagus steindachneri; adult; bimaculata, Polydactylus approximans, intestine; freshwater; Loreto (Tantaleán et Stellifer minor; adult; gill, intestine; al., 1985a). f r e s h w a t e r, m a r i n e ; I c a , L i m a , Lambayeque, Tumbes (Tantaleán et al., Order Spirurida Chitwood, 1933 1985a, b; Oliva et al., 1990; Luque et al., 1991; Tantaleán & Huiza, 1994; Oliva & Family Camallanidae Railliet & Henry, Luque, 1998b; Minaya et al., 2016). 1915 Camallanus (Camallanus) acaudatus Ferraz Family Acuariidae Railliet, Henry & & Thatcher, 1990 Sisoff, 1912 Osteoglossum bicirrhosum; adult; Rusguniella sp. stomach an intestine; freshwater; Amazon Peprilus snyderi; larvae; intestine; River Basin (Vásquez et al., 2007). marine; Lima (Ñacari & Sánchez, 2014). Camallanus sp. Cheilotrema fasciatum; adult; intestine; Family Philometridae Baylis & Daubney, marine; Lima (Oliva & Luque, 1998b). 1926 Procamallanus (Spirocamallanus) hilarii Alinema amazonicum (Travassos, 1960) Vaz & Pereira, 1934 juruense, Calophysus Trichomycterus piurae; adult; intestine; macropterus, Perrunichthys perruno; freshwater; Cajamarca, Lambayeque adult; mesentery, body cavity; freshwater; (Freitas & Ibañez, 1970; Rodrigues et al., Loreto (Moravec et al., 2006a, 2006b). 1991; Sarmiento et al., 1999; Moravec, Nilonema senticosum (Baylis, 1927) 1998). Arapaima gigas; adult; Intestine, swim- P r o c a m a l l a n u s ( S p i r o c a m a l l a n u s ) bladder; freshwater; Loreto (Tantaleán et chimusensis (Freitas & Ibáñez, 1968) al., 1985a; Moravec et al., 2006b; Trichomycterus díspar, T. piurae; adult; Mathews et al., 2013a; Mathews et al., intestine; freshwater; Cajamarca, 2014b; Serrano-Martínez et al., 2015). Lambayeque (Freitas & Ibañez, 1968; Philometra neptomeni Mateo, 1972 Tantaleán et al., 1985a; Sarmiento et al., Seriolella violacea; adult; gonads; 1999). marine; Lima (Mateo, 1982; Luque et al., P r o c a m a l l a n u s ( S p i r o c a m a l l a n u s ) 1991; Tantaleán & Huiza, 1994 ; Moravec inopinatus Travassos, Artigas & et al., 2011).

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Philometra sp. Family Physalopteridae Railliet, 1893 Anisotremus scapularis, Paralabrax Proleptus carvajali Fernandez & Villalba, humeralis, Paralichthys adspersus; adult; 1985 gonads; marine; Arequipa, Lima (Luque Anisotremus scapularis, Cilus gilberti, et al., 1991; Tantaleán & Huiza, 1994; Isacia conceptionis, Labrisomus Sarmiento et al., 1999; Iannacone & philippii, Rhinobatos planiceps; adult, Alvariño, 2009a; Chero et al., 2014d). larvae; general cavidad, gills, intestine, Philometridae gen sp. stomach, valve spiral; marine, Lima Cilus gilberti; adult; intestine; marine; (Iannacone et al., 2011a; Chero et al., Lima (Chero et al., 2014c). 2014c, d; Iannacone et al., 2015; Cruces et al., 2015). Family Gnathostomatidae Railliet, 1895 Proleptus obtusus Dujardin, 1845 Echinocephalus striatus Monticelli 1889 Mustelus mento, Rhinobatos planiceps, Scyllium sp.; adult; intestine; marine; Triakis maculata; adult; intestine, unspecified locality (Yamaguti, 1961). stomach; marine; La Libertad, Lima Echinocephalus sp. (Tantaleán & Rodriguez, 1987; Tantaleán Mustelus mento; adult; stomach; marine; & Huiza, 1994; Sarmiento et al., 1999). Lima (Tantaleán et al., 1983; Tantaleán & Proleptus sp. Huiza, 1994; Sarmiento et al., 1999). Galeichthys peruvianus, Labrisomus philippii, Menticirrhus ophicephalus, Family Cystidicolidae Skrjabin, 1946 Myliobatis peruvianus, Scartichthys Ascarophis sebastodis Olsen, 1952 gigas, Sciaena deliciosa, Trachinotus Sebastes capensis; adult; intestine; paitensis; larvae; intestine; mesentery; marine; Lima (González & Poulin, 2005) stomach; marine; Lima (Rivera & Ascarophis sp. Sarmiento, 1990a; Luque et al., 1991; Galeichthys peruvianus; Menticirrhus Tantaleán & Huiza, 1994; Pérez et al., ophicephalus; adult; intestine; marine; 1999; Sarmiento et al., 1999; Iannacone Lima (Luque, 1994a; Tantaleán & Huiza, et al., 2011b; Tarmeño et al., 2015). 1994; Oliva & Luque, 1998b; Sarmiento et al., 1999). Family Rhabdochonidae Travassos, Artigas & Pereira, 1928 Family Guyanemidae Petter, 1974 Johnstonmawsonia sp. Pseudodelphis sp. Scartichthys gigas; adult; intestine; Scartichthys gigas; adult; pericardial marine; Lima (Iannacone et al., 2012b). cavity; marine; Lima (Tarmeño et al., Rhabdochona sp. 2015) Paratrygon aiereba, Potamotrygon falkeri, Trichomycterus dispar; adult; Family Hedruridae Railliet, 1916 intestine; freshwater; Lambayeque, Hedruris basilichtensis Mateo, 1971 Madre Rios (Tantaleán et al., 1985a; Odontesthes bonariensis, Orestias albus, Reyda, 2008). O. luteus; adult; stomach; freshwater; Puno (Mateo, 1971, Sarmiento et al., PHYLUM PLATYHELMINTHES 1999). Gegenbaur, 1859 Hedruris orestiae Moniez, 1889 Class Cestoda Rudolphi, 1808 Orestias luteus, O. mulleri; adult; Order Bothriocephalidea Kuchta, Scholz, stomach; freshwater; Puno (Ibañez & Brabec & Bray, 2008 Córdova, 1976, Sarmiento et al., 1999).

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Family Bothriocephalidae Blanchard, hippurus, Cynoscion analis, Galeichthys 1849 peruvianus, Genypterus maculatus, Clestobothrium crassiceps (Rudolphi, 1819) Menticirrhus ophicephalus, Merluccius Merluccius gayi peruanus; adult; gayi peruanus, Paralabrax humeralis, intestine; marine; La Libertad, Lima Paralichthys adspersus, Paralonchurus (Tantaleán, 1976; Durán & Oliva, 1980; peruanus, Sarda chiliensis, Sciaena Luque et al., 1991; Tantaleán & Huiza, callaensis, Sciaena deliciosa, Scomber 1994; Jara, 1998; Chero et al., 2014b). japonicus, Scomberomorus sierra, Trachinotus paitensis, Trachurus Family Echinophallidae Schumacher, 1914 murphyi, larvae; mesentery; visceral Neobothriocephalus aspinosus Mateo & surface; marine; Ancash, La Libertad, Bullock, 1966 Lambayeque, Lima (Baer, 1969, Seriolella violacea; adult; intestine; Tantaleán, 1975b; Escalante, 1983; marine; Lima (Mateo & Bullock, 1966; Escalante & Miranda, 1986; Tantaleán & Luque et al., 1991; Tantaleán & Huiza, Huiza, 1984, 1994; Luque et al., 1991; 1994; Iannacone, 2003). Iannacone & Alvariño, 2009; Wicht et al., 2010, Chero et al., 2014a,b,c). Family Triaenophoridae Lönnberg, 1889 sp. A n c h i s t r o c e p h a l u s m i c r o c e p h a l u s Sarda chiliensis, Scomberomorus sierra, (Rudolphi, 1819) Trachurus murphyi; larvae; cavity; Mola mola; adult; intestine; marine; Lima marine; La Libertad, Lima (Baer, 1969; (Tantaleán & Huiza, 1994). Tantaleán & Huiza, 1994; Pérez et al., Fistulicola plicatus (Rudolphi, 1819) 1999; Llerena et al., 2001). Mola mola; adult; intestine; marine; Lima Diphyllobothridae gen. sp. (Tantaleán & Huiza, 1994). Merluccius gayi peruanus, Scomber Glossobothrium sp. japonicus; larvae; mesentery; marine; La Hemilutjanus macrophthalmos; adult; Libertad, Lima (Jara, 1998; Oliva et al., intestine; marine; Lima (Tantaleán et al., 2008) 1982; Luque et al., 1991; Tantaleán & Ligula intestinalis (Linnaeus, 1758) Huiza, 1994). Odontesthes bonariensis, Orestias albus, O. luteus, Orestias sp.; larvae; body Order Diphyllidea van Beneden in Carus, cavity; freshwater; Puno (Tantaleán et al., 1863 1985a).

Family Echinobothriidae Perrier, 1897 Order Proteocephalidea Mola, 1928 Echinobothrium sp. Myliobatis peruvianus; adult; spiral Family (La Rue, 1911) valvule; marine; Ica (Tantaleán & Huiza, Ageneiella sp. 1994). Ageneiosus inermis; adult; intestine; freshwater; Loreto (de Chambrier et al., Order Diphyllobothriidea Kuchta, Scholz, 2015a). Brabec & Bray, 2008 Amphoteromorphus ovalis Carfora, de Chambrier & Vaucher, 2003 Family Diphyllobothridae Lühe, 1910 Brachyplatystoma cf. filamentosum; Adenocephalus pacificus Nybelin, 1931 adult; intestine; freshwater; Loreto (de Anisotremus scapularis, Ariopsis Chambrier et al., 2015a). seemanni, Cilus gilberti, Coryphaena Amphoteromorphus peniculus Diesing, 1850

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Brachyplatystoma rousseauxii; adult; Brachyplatystoma cf. filamentosum; intestine; freshwater; Loreto (de adult; intestine; freshwater; Loreto (de Chambrier et al., 2015a, b). Chambrier et al., 2015a). Amphoteromorphus parkamoo Woodland, Endorchis sp. 1935 Pimelodus altissimus; adult; intestine; Zungaro zungaro; adult; intestine; freshwater; Loreto (de Chambrier et al., freshwater; Loreto (de Chambrier et al., 2015a). 2006, 2015a). Euzetiella tetraphylliformis de Chambrier, Amphoteromorphus piriformis Carfora, de Rego & Vaucher, 1999 Chambrier & Vaucher, 2003 Zungaro zungaro; adult; intestine; Brachyplatystoma rousseauxii; adult; freshwater; Loreto (de Chambrier et al., intestine; freshwater; Loreto (de 2006, 2015a). Chambrier et al., 2006, 2015a). Gibsoniela mandube (Woodland, 1935) Brayela karuatayi (Woodland, 1934) Ageneiosus inermis, Ageneiosus sp.; Platynematichthys notatus; adult adult; intestine; freshwater; Loreto (de (immature); intestine; freshwater; Chambrier et al., 2015a,b). Ancash, Loreto (de Chambrier et al., Harriscolex kaparari (Woodland, 1935) 2014, 2015a,b). Pseudoplatystoma fasciatum; adult; Chambriella agostinhoi (Pavanelli & Santos, intestine; freshwater; Loreto (de 1992) Chambrier et al., 2015a). Zungaro zungaro; adult; intestine; Harriscolex piramutab (Woodland, 1934) freshwater; Loreto (de Chambrier et al., Brachyplatystoma vaillantii; adult; 2006, 2015a). intestine; freshwater; Loreto (de Chambriella paranaensis (Pavanelli & Rego, Chambrier et al., 2006, 2015a). 1989) Houssayela sudobim (Woodland, 1935) Hemisorubim platyrhynchos; adult; P s e u d o p l a t y s t o m a p u n c t i f e r , intestine; freshwater; Loreto (de Sorubimichthys planiceps; adult; Chambrier et al., 2006, 2015a). intestine; freshwater; Loreto (de Chambriella sp. Chambrier & Scholz, 2005; de Chambrier B r a c h y p l a t y s t o m a v a i l l a n t i i , et al., 2006, 2015a,b). Phractocephalus hemioliopterus, Jauella glandicephalus Rego & Pavanelli, P s e u d o p l a t y s t o m a f a s c i a t u m , 1985 Sorubimichthys planiceps; adult; Zungaro zungaro; adult; intestine; intestine; freshwater; Loreto (de freshwater; Loreto (de Chambrier et al., Chambrier et al., 2006, 2015a; de 2015a). Chambrier & Scholz, 2008). Lenhataenia megacephala (Woodland, Choanoscolex abscisus (Riggenbach, 1896) 1934) Pseudoplatystoma fasciatum; adult; Sorubimichthys planiceps; adult; intestine; freshwater; Loreto (de intestine; freshwater; Loreto (de Chambrier et al., 2015a). Chambrier & Scholz 2008; de Chambrier Choanoscolex sp. et al., 2015a). Pseudoplatystoma punctifer, P. tigrinum, Manaosia bracodemoca Woodland, 1935 Sorubimichthys planiceps; adult; Sorubim lima; adult; intestine; freshwater; intestine; freshwater; Loreto (de Loreto (de Chambrier et al., 2015a). Chambrier et al., 2006, 2015a; de Mariauxiella piscatorum de Chambrier & Chambrier & Scholz, 2008). Vaucher, 1999 Endorchis piraeeba Woodland, 1934 Hemisorubim platyrhynchos; adult;

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intestine; freshwater; Loreto (de intestine; freshwater; Loreto (de Chambrier et al., 2006, 2015a). Chambrier et al., 2006, 2015a). Megathylacus jandia Woodland, 1934 Nomimoscolex sudobim Woodland, 1935 Zungaro zungaro; adult; intestine; Pseudoplatystoma fasciatum; adult; freshwater; Loreto (de Chambrier et al., intestine; freshwater; Loreto (de 2015a). Chambrier et al., 2015a). Megathylacus sp. Nomimoscolex cf. sudobim Woodland, 1935 Pseudoplatystoma fasciatum, P. Pseudoplatystoma tigrinum; adult; punctifer; adult; intestine; freshwater; intestine; freshwater; Loreto (de Loreto (de Chambrier et al., 2014, 2015a). Chambrier et al., 2006). Monticellia amazonica de Chambrier & Nomimoscolex suspectus Zehnder, de Vaucher, 1997 Chambrier, Vaucher & Mariaux, 2000 Calophysus macropterus; adult; intestine; Brachyplatystoma cf. filamentosum; freshwater; Loreto (de Chambrier et al., adult; intestine; freshwater; Loreto (de 2006, 2015a; Scholz et al., 2008). Chambrier et al., 2015a). Monticellia belavistensis Pavanelli, Nomimoscolex sp. Machado dos Santos, Takemoto & dos Pimelodus ornatus; adult; intestine; Santos 1994 freshwater; Loreto (de Chambrier et al., Pterodoras granulosus; adult; intestine; 2015a). freshwater; Loreto (de Chambrier et al., Nupelia sp. 2015a). Goeldiella eques; adult; intestine; Monticellia santafesina Arredondo & Gil de freshwater; Loreto (de Chambrier et al., Pertierra, 2010 2006, 2015a). Megalonema platycephalum; adult; Peltidocotyle lenha (Woodland, 1933) intestine; freshwater; Loreto (de Sorubimichthys planiceps, Zungaro Chambrier et. al., 2015a). zungaro; adult; intestine; freshwater; Monticellia ventrei de Chambrier & Loreto (de Chambrier et al., 2006, 2015a; Vaucher, 1999 de Chambrier & Scholz, 2008).  Pinirampus pirinampu; adult; intestine; Peltidocotyle rugosa Diesing, 1850 freshwater; Loreto (de Chambrier et al., Pseudoplatystoma fasciatum; adult; 2015a). intestine; freshwater; Loreto (de Monticelliinae gen. sp. Chambrier et al., 2015a). Phractocephalus hemioliopterus; adult; Proteocephalidea gen. sp. intestine; freshwater; Loreto (de Cichlasoma amazonarum, Paratrygon Chambrier et al., 2015a). aiereba, Potamotrygon cf. falkeri, P. Nomimoscolex admonticellia (Woodland, motoro; larvae (hyperparasite), adult; 1934) intestine; freshwater; Loreto, Madre de Pinirampus pirinampu; adult; intestine; Dios (Reyda, 2008; de Chambrier et al., freshwater; Loreto (de Chambrier et al., 2015a). 2015a). Proteocephalus gibsoni Rego & Pavanelli, Nomimoscolex lenha (Woodland, 1933) 1991 Sorubimichthys planiceps; adult; Astronotus ocellatus; adult; intestine; intestine; freshwater; Loreto (de freshwater; Loreto (de Chambrier et al., Chambrier & Scholz, 2008; de Chambrier 2006, 2015a). et al., 2015a). Proteocephalus hemioliopteri de Chambrier Nomimoscolex lopesi Rego, 1989 & Vaucher, 1997 Pseudoplatystoma fasciatum; adult; Phractocephalus hemioliopterus; adult;

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intestine; ; freshwater; Loreto (de Cichla monoculus; adult; intestine; Chambrier et al., 2015a). freshwater; Loreto (de Chambrier et al., Proteocephalus hobergi de Chambrier & 2006, 2015a,b). Vaucher, 1999 Spasskyellina lenha (Woodland, 1933) Oxydoras niger; adult; intestine; Sorubimichthys planiceps; adult; freshwater; Loreto (de Chambrier et al., intestine; freshwater; Loreto (de 2015a). Chambrier & Scholz, 2008; de Chambrier Proteocephalus kuyukuyu Woodland, 1935 et al., 2015a,b). Megalodoras uranoscopus, Pterodoras Spasskyellina spinulifera (Woodland, 1935) granulosus; Pterodoras sp.; adult; Pseudoplatystoma punctifer, P. tigrinum; intestine; freshwater; Loreto (de adult; intestine; freshwater; Loreto (de Chambrier et al., 2015a,b). Chambrier et al., 2006, 2015a). Proteocephalus macrophallus (Diesing, Spatulifer maringaensis Pavanelli & Rego, 1850) 1989 Cichla monoculus; adult; intestine; Hemisorubim platyrhynchos, Sorubim freshwater; Loreto (de Chambrier et al., lima; adult; intestine; freshwater; Loreto 2006, 2015a,b). (de Chambrier et al., 2006, 2015a). Proteocephalus microscopicus Woodland, Spatulifer rugosa (Woodland, 1935) 1935 Pseudoplatystoma punctifer; adult; Cichla monoculus; adult; intestine; intestine; freshwater; Loreto (de freshwater; Loreto (de Chambrier et al., Chambrier et al., 2006, 2015a). 2006, 2015a). Spatulifer sp. Proteocephalus sophiae de Chambrier & Pseudoplatystoma tigrinum; adult; Rego, 1994 intestine; freshwater; Loreto (de Zungaro zungaro; adult; intestine; Chambrier et al., 2006, 2015a). freshwater; Loreto (de Chambrier et al., Travassiella jandia (Woodland, 1934) 2015a). Zungaro zungaro; adult; intestine; Proteocephalus sp. freshwater; Loreto (de Chambrier et al., O r e s t i a s a l b u s , O . l u t e u s , 2006, 2015a). Phractocephalus hemioliopterus, Zygobothrium megacephalum Diesing, 1850 P i m e l o d u s b l o c h i i , P t e ro d o r a s Phractocephalus hemioliopterus; adult; granulosus; adult, larvae; intestine; intestine; freshwater; Loreto (de freshwater; Loreto (Tantaleán et al., Chambrier et al., 2006, 2015a). 1985a; de Chambrier et al., 2006, 2015a). Rudolphiella piracatinga (Woodland, 1935) Order Cathetocephalidea Schmidt & Calophysus macropterus; adult; intestine; Beveridge, 1990 freshwater; Loreto (de Chambrier et al., 2006, 2015a). Family Cathetocephalidae Dailey & Rudolphiella sp. Overstreet, 1973 Pinirampus pirinampu; adult; intestine; Cathetocephalus thatcheri Dailey & freshwater; Loreto (de Chambrier et al., Overstreet, 1973 2015a). Carcharhinus leucas, C. porosus; adult; Scholzia emarginata (Diesing, 1850) spiral valve; marine; Lima (Rivera & Phractocephalus hemioliopterus; adult; Sarmiento, 1990b; Tantaleán & Huiza, intestine; freshwater; Loreto (de 1994). Chambrier et al., 2006, 2015a). Sciadocephalus megalodiscus Diesing, 1850 Family Disculicipitidae Joyeux & Baer,

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1936 Beveridge, 2002). Disculiceps pileatus (Linton, 1891) Acanthobothrium lusarmientae Severino & Carcharhinus porosus; adult; spiral valve; Verano, 1980 marine; Lima (Tantaleán, 1991; Tantaleán Sympterygia brevicaudata; adult; spiral & Huiza, 1994). valve; marine; Lima (Severino & Verano, 1980; Tantaleán et al., 1982; Tantaleán & Order Onchoproteocephalidea Caira, Huiza, 1994; Ghoshroy & Caira, 2001; Jensen, Waeschenbach, Olson & Campbell & Beveridge, 2002; Morales et Littlewood, 2014 al., 2005). Acanthobothrium olseni Dailey & Mudry, Family Prosobothriidae Baer & Euzet, 1968 1955 Rhinobatos planiceps; adult; intestine, Prosobothrium armigerum Cohn, 1902 spiral valve; marine; Lima (Iannacone et Prionace glauca; adult; spiral valve; al., 2011a). marine; Lima (Rivera & Sarmiento, Acanthobothrium peruviense Reyda, 2008 1990b; Tantaleán & Huiza, 1994). Potamotrygon motoro; adult; spiral valve; freshwater; Madre de Dios (Reyda, 2008). Family Onchobothriidae Braun, 1900 (pro Acanthobothrium cf. peruviense Reyda, parte) 2008 Acanthobothrium batailloni Euzet, 1955 Potamotrygon cf. falkeri; adult; spiral Myliobatis chilensis; adult; spiral valve; valve; freshwater; Madre de Dios (Reyda, marine; La Libertad (Escalante, 1986; 2008). Tantaleán & Huiza, 1994). Acanthobothrium psammobati Carvajal & Acanthobothrium brevissime Linton, 1908 Goldstein, 1969 Myliobatis peruvianus; adult; spiral valve; Sympterygia brevicaudata; adult; spiral marine; Lima (Tantaleán, 1991; Tantaleán valve; marine; Lima; (Tantaleán, 1991; & Huiza, 1994; Ghoshroy & Caira, 2001; Tantaleán & Huiza, 1994). Campbell & Beveridge, 2002). Acanthobothrium cf. ramiroi Ivanov, 2005 Acanthobothrium chilensis Rego, Vicente & Potamotrygon motoro: adult; spiral valve; Herrera, 1968 freshwater; Madre de Dios (Reyda, 2008). Sarda chiliensis; adult; spiral valve; Acanthobothrium robustum Alexander, marine; Piura (Rego et al., 1968; 1953 Tantaleán & Huiza, 1994). Rhinobatos planiceps; adult; spiral valve; Acanthobothrium gonzalesmugaburui marine; Ancash, La Libertad (Escalante, Severino & Sarmiento, 1979 1986; Tantaleán & Huiza, 1994). Myliobatis peruvianus; adult; spiral valve; Acanthobothrium sp. marine; Ancash, Lima (Severino & Myliobatis chilensis; adult; spiral valve; Sarmiento, 1979; Tantaleán et al., 1982; marine; La Libertad (Tresierra et al., Tantaleán & Huiza, 1994; Ghoshroy & 1986; Tantaleán & Huiza, 1994). Caira, 2001; Campbell & Beveridge, Acanthobothroides peruensis López, 1994 2002; Morales et al., 2005). Dasyatis dipterura; adult; spiral valve; Acanthobothrium holorhini Alexander, 1953 marine; Lima (Morales et al., 2005). Myliobatis chilensis, M. peruvianus; Acanthobothroides thorsoni Brooks, 1977 adult; spiral valve; marine; Lima Dasyatis dipterura; adult; spiral valve; (Rodriguez & Tantaleán, 1980, Tantaleán marine; Ancash, Lima (López, 1994; et al., 1982; Tantaleán & Huiza, 1994; Tantaleán & Rodriguez, 1987; Tantaleán Ghoshroy & Caira, 2001; Campbell & & Huiza, 1994).

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Platybothrium auriculatum Yamaguti, 1952 Paraorygmatobothrium triacis (Yamaguti, Prionace glauca; adult; spiral valve; 1952) marine; La Libertad, Lima (Escalante, Mustelus dorsalis, M. whitneyi; adult; 1986; Tantaleán & Huiza, 1994). spiral valve; marine; Ancash, La Libertad Platybothrium sp. (Escalante & Carvajal, 1981; Tantaleán & Sphyrna zygaena; adult; spiral valve; Huiza, 1994). marine; Lima (López de McDonald & Phyllobothrium lactuca (Van Beneden, Tantaleán, 1985; Tantaleán & Huiza, 1850) 1994). Mustelus mento; adult; spiral valve; Potamotrygonocestus cf. fitzgeraldae marine; Lambayeque, Lima (Escalante & Marques, Brooks & Araujo, 2003 Carvajal, 1981; Tantaleán & Huiza, 1994). Paratrygon aiereba, Potamotrygon Phyllobothrium thridax (Van Beneden, falkeri; adult; spiral valve; freshwater; 1849) Madre de Dios (Reyda, 2008; Caira et al., Myliobatis chilensis, Myliobatis 2014) peruvianus; adult; spiral valve; marine; Potamotrygonocestus sp. Ancash, Lima (Tantaleán, 1991; Tantaleán Paratrygon aiereba, Potamotrygon cf. & Huiza, 1994). falkeri, P. motoro; adult; spiral valve; Phyllobothrium sp. freshwater; Madre de Dios (Reyda & Mustelus mento, Rhinobatos planiceps, Olson, 2003; Reyda, 2008; Caira et al., Squatina armata; adult; spiral valve; 2014). marine; La Libertad (Tresierra et al., 1986; Tantaleán & Huiza, 1994). Order Phyllobothriidea Caira, Jensen, Thysanocephalum thysanocephalum Waeschenbach, Olson & Littlewood, 2014 (Linton, 1889) Sphyrna zygaena; adult; intestine; marine; Family Phyllobothriidae Braun, 1900 Lima (López de McDonald & Tantaleán, Clistobothrium tumidum (Linton, 1922) 1985; Tantaleán & Huiza, 1994). Isurus oxyrinchus; adult; spiral valvule; marine; Lima (Tantaleán & Huiza, 1994). Order “Tetraphyllidea” Carus, 1863 Nandocestus guariticus (Marques, Brooks & Lasso, 2001) Anthobothrium laciniatum Linton, 1890 Paratrygon aiereba, Potamotrygon cf. Prionace glauca; adult; spiral valvule; falkeri; adult; spiral valve; freshwater; marine; Lima (Tantaleán & Huiza, 1994). Madre de Dios (Reyda & Olson, 2003; Anthobothrium sp. Reyda, 2008; Ruhnke, 2011; Caira et al., Myliobatis peruvianus; adult; spiral 2014). valvule; marine; Lima (Tantaleán & Orygmatobothrium musteli (Van Beneden, Huiza, 1994). 1849) Calliobothrium verticillatum (Rudolphi, Mustelus dorsalis, M. lunulatus, M. 1819) whitneyi, Prionace glauca; adult; spiral Mustelus mento; adult; spiral valve; valve; marine; Ancash, La Libertad marine; Ancash, Lambayeque (Escalante (Escalante & Carvajal, 1981; Escalante, & Carvajal, 1981; Tantaleán & Huiza, 1986; Tantaleán & Huiza, 1994). 1994). Orygmatobothrium sp. Calliobothrium sp. Mustelus mento; stomach; marine; La Mustelus mento; adult; spiral valve, Libertad (Tresierra et al., 1986; Tantaleán stomach; marine; La Libertad, Lima & Huiza, 1994). (Tresierra et al., 1986; Tantaleán & Huiza,

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1994). 2011). Caulobothrium myliobatidis Carvajal, 1977 Rhinebothrium rhinobati Dailey & Myliobatis chilensis; adult; spiral valve; Carvajal, 1976 marine; Ancash (Tantaleán, 1991; Rhinobatos planiceps; adult; spiral valve; Tantaleán & Huiza, 1994). marine; Lima (Tantaleán, 1991; Tantaleán Caulobothrium sp. & Huiza, 1994; Iannacone et al., 2011a). Rhinoptera steindachneri; adult; spiral Rhinebothrium sp. valvule; marine; Lima (Tantaleán & Myliobatis chilensis, Myliobatis Huiza, 1994). peruvianus, Paratrygon aiereba, Synodus Dinobothrium paciferum Sproston, 1948 scituliceps; adult, larvae; intestine, spiral Cetorhinus maximus; adult; spiral valve; valve; freshwater, marine; La Libertad, marine; Lima (Rivera & Sarmiento, Lima, Madre de Dios, Piura (Escalante et 1990b; Tantaleán & Huiza, 1994). al., 1987; Luque et al., 1991; Tantaleán & “Scolex pleuronectis” Müller, 1788 Huiza, 1994; Reyda & Olson, 2003; Healy Coryphaena hippurus, Ethmidium et al., 2009). maculatum, Genypterus maculatus, R h o d o b o t h r i u m m e s o d e s m a t u m Katsuwonus pelamis, Odontesthes regia, (Bahamonde & Lopez, 1962) Sarda chiliensis, Sardinops sagax, Myliobatis chilensis, M. peruvianus, Scomber japonicus, Stellifer minor, Sarda chiliensis; adult; spiral valve, Strongylura scapularis, Synodus intestine; marine; Ica, Lima (Rego et al., scituliceps, Trachurus murphyi; larve 1968; Rodriguez & Tantaleán, 1980; (metacestodes); caecum, intestine; López de McDonald & Tantaleán, 1985; marine; La Libertad, Piura (Escalante & Tresierra et al., 1986; Tantaleán & Huiza, Carjaval, 1984; Escalante et al., 1987; 1994). Luque et al., 1991; Tantaleán & Huiza, Rhodobothrium pulvinatum Linton, 1889 1994; Jara, 1998; Cruces et al., 2014). Dasyatis dipterura; adult; spiral valvule; Symcallio lintoni (Euzet, 1954) marine; Ancash (Tantaleán & Huiza, Mustelus whitneyi; adult; spiral valve; 1994). marine; La Libertad (Escalante & Rhinebothroides sp. Carvajal, 1981; Tantaleán & Huiza, 1994). Paratrygon aiereba, Potamotrygon falkeri, Potamotrygon cf. falkeri, P. Order Rhinebothriidea Healy, Caira, motoro; adult; spiral valve; freshwater; Jensen, Webster & Littlewood, 2009 Madre de Dios (Reyda & Olson, 2003; Reyda, 2008; Healy et al., 2009). Family Echeneibothriidae de Beauchamp, 1905 Order Trypanorhyncha Diesing, 1863 Echeneibothrium sp. Myliobatis chilensis; adult; spiral valvule; Trypanorhyncha gen. sp. marine; La Libertad (Tantaleán & Huiza, Paralabrax humeralis; unspecified 1994). development stage; unspecified site of infection; marine; Lima (Iannacone & Family Rhinebothriidae Euzet, 1953 Alvariño, 2009a). Rhinebothrium copianullum Reyda, 2008 Paratrygon aiereba, Potamotrygon cf. Family Lacistorhynchidae Guiart, 1937 falkeri, P. motoro, P. tatianae; adult; Dasyrhynchus pacificus Robinson, 1965 spiral intestine; freshwater; Madre de Sciaena deliciosa; larvae; mesentery; Dios (Reyda, 2008; Reyda & Marques, marine; La Libertad, Piura (Escalante &

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Carvajal, 1984; Luque et al., 1991; Mugil cephalus, Mustelus dorsalis, M. Tantaleán & Huiza, 1994; Palm, 2004). mento, M.s whitneyi, Odontesthes regia, Grillotia dollfusi Carvajal, 1971 Paralichthys adspersus, Paralonchurus Merluccius gayi peruanus; larvae; peruanus, Scartichthys gigas, Sciaena visceral surface; marine; Lima (Chero et deliciosa, Stellifer minor; adult, larvae; al., 2014b). mesentery, muscles, spiral valve, viscera; Grillotia sp. m a r i n e ; A n c a s h , L a L i b e r t a d , Paralabrax humeralis, Squatina armata; Lambayeque, Lima (Durán & Oliva, larvae; mesentery, stomach; marine; La 1980; Escalante & Carvajal, 1984; Rivera Libertad, Piura (Armas, 1983a; Tresierra & Sarmiento, 1990a; Luque et al., 1991; et al., 1986; Luque et al., 1991; Tantaleán Tantaleán & Huiza, 1994; Jara, 1998; & Huiza, 1994). Oliva & Luque, 1998b, 2002; Cruces et Pseudogrillotia peruviana Escalante & al., 2015). Carvajal, 1984 S c o m b e ro m o r u s s i e r r a ; l a r v a e ; Family Pterobothriidae Pintner, 1931 mesentery; marine; La Libertad Pterobothrium acanthotruncatum Escalante (Escalante & Carvajal, 1984; Luque et al., & Carjaval, 1984 1991; Tantaleán & Huiza, 1994). Coryphaena hippurus, Paralonchurus Callitetrarhynchus gracilis (Rudolphi, 1819) peruanus; larvae; gonads, mesentery, Cilus gilberti, Galeichthys peruvianus, swimming bladder; marine; La Libertad, H e m i l u t j a n u s m a c ro p h t h a l m o s , Lima (Tantaleán et al., 1982; Escalante & Merluccius gayi peruanus, Paralabrax Carvajal, 1984; Luque et al., 1991; humeralis, Paralonchurus peruanus, Tantaleán & Huiza, 1994). Sciaena deliciosa, Scomber japonicus, Pterobothrium sp. Trachurus murphyi; larvae; mesentery, Paralonchurus peruanus; larvae; viscera; marine; Ancash, La Libertad, mesentery; marine; Lima (Tantaleán et al., Lambayeque, Lima, Piura (Durán & 1982; Luque et al., 1991; Palm, 2004). Oliva, 1980; Escalante & Carvajal, 1984; Escalante & Miranda, 1986; Escalante et Family Eutetrarhynchidae Guiart, 1927 al., 1986; Luque et al., 1991; Tantaleán & Eutetrarhynchus sp. Huiza, 1994, Oliva & Ballón, 2002; Palm, Trachurus murphyi; larvae; mesentery; 2004; Chero et al., 2014b). marine; Lima, Piura (Oliva, 1999). Floriceps saccatus Cuvier, 1817 Prochristianella heteracantha Dailey & Centropomus nigrescens; larvae; Carvajal, 1976 mesentery; marine; Piura (Escalante & Rhinobatos planiceps; adult; intestine, Carvajal, 1984; Luque et al., 1991; spiral valve, stomach; marine; Ancash, Tantaleán & Huiza, 1994). Lima (Tantaleán & Rodriguez, 1987; Lacistorhynchus dollfusi Beveridge & Tantaleán & Huiza, 1994; Iannacone et Sakanari, 1987 al., 2011a). C h e i l o t re m a f a s c i a t u m ; l a r v a e ; mesentery; marine; Lima (Oliva & Luque, Family Otobothriidae Dollfus, 1942 1998b). Otobothrium sp. Lacistorhynchus tenuis (Van Beneden, 1858) Cynoscion analis; larvae; muscles, Anisotremus scapularis, Cheilodactylus viscera; marine; Lambayeque (Luque et variegatus, Cheilotrema fasciatum, al., 1991; Tantaleán & Huiza, 1994; Palm, Galeichthys peruvianus, Labrisomus 2004). philippii, Merluccius gayi peruanus, Poecilancistrium caryophyllum (Diesing,

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1850) al., 2015). Cilus gilberti; larvae; mesentery; marine; Tentacularia coryphaenae Bosc, 1802 La Libertad (Escalante & Carvajal, 1984; Centropomus nigrescens, Coryphaena Luque et al., 1991; Tantaleán & Huiza, h i p p u r u s , K a t s u w o n u s p e l a m i s , 1994; Palm, 2004). Merluccius gayi peruanus, Prionace glauca, Scomber japonicus, Trachurus Family Sphyriocephalidae Pintner, 1913 murphyi; adult, larvae; mesentery, spiral Hepatoxylon megacephalum (Rudolphi, valve, visceral surface; marine; La 1819) Libertad, Lambayeque, Lima, Piura Trichomycterus punctulatus; larvae; body (Durán & Oliva, 1980; Escalante & cavity; freshwater; unspecified locality Carvajal, 1984; López & Tantaleán, 1985; (Palm, 2004). Escalante, 1986; Luque et al., 1991; Hepatoxylon trichiuri (Holten, 1802) Tantaleán & Huiza, 1994; Jara, 1998; Brama japonica, Prionace glauca; adult, Oliva, 1999; Pérez et al., 1999; Llerena et larvae; mesentery, stomach, intestine; al., 2001; Palm, 2004; Oliva et al., 2008; marine; La Libertad, Lima (Escalante, Vásquez-Ruiz & Jara-Campos, 2012). 1986; Tantaleán & Huiza, 1994; Iannacone & Alvariño, 2013). Order Amphilinidea Poche, 1922 Sphyriocephalus tergestinus Pintner, 1913 Sarda chiliensis; larvae; stomach; marine; Family Schizochoeridae Poche, 1922 Lima (Chero et al., 2015b). Nesolecithus janicki (Poche, 1922) Arapaima gigas; adult; body cavity; Family Tentaculariidae Poche, 1926 freshwater; Amazon River Basin Nybelinia surmenicola Okada in Dollfus, (Thatcher, 2006). 1929 Schizochoerus liguloideus (Diesing, 1850) Trachurus murphyi; larvae; unispeficied Arapaima gigas; adult; body cavity; site of infection; marine; Piura (Oliva, freshwater; Loreto (Serrano-Martínez et 1999). al., 2015). Nybelinia sp. Brama japonica, Coryphaena hippurus, Order Gyrocotylidea Poche, 1926 Cynoscion analis, Isacia conceptionis, Isurus oxyrinchus, Merluccius gayi Family Gyrocotylidae Benham, 1901 peruanus, Mustelus mento, Myliobatis Gyrocotyle maxima MacDonagh, 1927 peruvianus, Paralonchurus peruanus, Callorhynchus callorhynchus; adult; Sarda chiliensis, Sciaena deliciosa, spiral valve; marine; Lima, La Libertad Scomber japonicus, Seriolella violacea, (Tantaleán, 1991; Tantaleán & Huiza, Stellifer minor, Trachurus murphyi; 1994). larvae; mesentery, valve, visceral surface; marine; La Libertad, Lambayeque, Lima Class Monogenea Van Beneden, 1858 (López de McDonald & Tantaleán, 1985; Order Capsalidea Lebedev, 1988 Luque et al., 1991; Tantaleán & Huiza, 1994; Jara, 1998; Pérez et al., 1999; Family Capsalidae Baird, 1853 Llerena et al., 2001; Oliva & Ballón, 2002 Benedenia sp. as N. sumernicola; Palm, 2004; Williams Isacia conceptionis; adult; gills; marine; & Williams, 2009; Vásquez-Ruiz & Jara- La Libertad (Escalante et al., 1982a; Campos, 2012; Iannacone & Alvariño, Tantaleán & Huiza, 1994; Paschoal et al., 2013; Chero et al., 2014a; Iannacone et 2015).

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Benedeniella sp. Ameloblastella sp. Myliobatis peruvianus; stomach; gills; Hassar sp., Hypophthalmus edentatus, marine; Ica (Tantaleán & Huiza, 1994). Sorubim lima; adult; gills; freshwater; Capsala gregalis (Wagner & Carter, 1967) Loreto (Mendoza-Palmero et al., 2015). Sarda chiliensis; adult; gills; marine; Amphithecium calycinum Boeger & Arequipa, Lima (Luque et al., 1991; Kritsky, 1988 Luque & Iannacone, 1991a; Tantaleán & Pygocentrus nattereri; adult; gills; Huiza, 1994). freshwater; Loreto (Iannacone & Luque, Encotyllabe antofagastensis Sepúlveda, 1993b). González & Oliva, 2014 Amphithecium camelum Boeger & Kritsky, Anisotremus scapularis; adult; gills; 1988 marine; Lima (Chero et al., 2014d). Pygocentrus nattereri; adult; gills; Encotyllabe callaoensis Tantaleán, 1974 freshwater; Loreto (Iannacone & Luque, Paralonchurus peruanus, Sciaena 1993b). deliciosa; adult; gills, mouth; marine; Anacanthorus reginae Boeger & Kritsky, Lima (Luque et al., 1991; Tantaleán & 1988 Huiza, 1994; Chero et al., 2014a). Pygocentrus nattereri; adult; gills; Encotyllabe sp. freshwater; Loreto (Iannacone & Luque, Aplodactylus punctatus, Stellifer minor; 1993b). adult; gills; marine; Ica, La Libertad, Anacanthorus spathulatus Kritsky, Lambayeque (Tantaleán & Huiza, 1994; Thatcher & Kayton 1979. Jara, 1998). Colossoma macropomum; adult; gills; Macrophyllida antarctica (Hughes, 1928) freshwater; Loreto (Soberon et al., 2014). Johnston, 1930 Anacanthorus stachophallus Kritsky, Cheilodactylus variegatus; adult; gills; Boeger & Van Every, 1992 marine; Lima (Martínez et al., 2013) Pygocentrus nattereri; adult; gills; Neobenedenia sp. freshwater; Loreto (Iannacone & Luque, Genypterus sp., Mugil cephalus, 1993b). Ophidiidae gen sp.; adult; gills; marine; Anacanthorus thatcheri Boeger & Kritsky, Ancash, Lima (Luque et al., 1991; 1988 Tantaleán & Huiza, 1994). Pygocentrus nattereri; adult; gills; freshwater; Loreto (Iannacone & Luque, Order Dactylogyridea Bychowsky, 1937 1993b). Anacanthorus sp. Family Calceostomatidae Parona & Myleus schomburgkii; adult; gills; Perugia, 1890 freshwater; Loreto (Fernández-Méndez et Calceostoma sp. al., 2015). Orthopristis chalceus; adult; gills; Ancyrocephalus sp. marine; La Libertad, Piura (Tantaleán & Strongylura scapularis; adult; gills; Huiza, 1994). marine; La Libertad, Piura (Tantaleán & Huiza, 1994). Family Dactylogyridae Bychowsky, 1933 Aphanoblastella aurorae Mendoza- Ameloblastella unapi Mendoza-Franco & Palmero, Scholz, Mendoza-Franco & Scholz, 2009 Kuchta, 2012 Calophysus macropterus; adult; gills; Goeldiella eques; adult; gills; freshwater; freshwater; Loreto (Mendoza-Franco Loreto (Mendoza-Palmero et al., 2012, & Scholz, 2009). 2015).

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Aphanoblastella sp. Dawestrema cycloancistrioides Kritsky, Goeldiella eques; adult; gills; freshwater; Thatcher & Boeger, 1985 Loreto (Mendoza-Palmero et al., 2015) Arapaima gigas; adult; gills; freshwater; Cleidodiscus sp. Loreto (Mathews et al., 2013a; Serrano- Pimelodella yuncensis; adult; gills; Martínez et al., 2015). freshwater; La Libertad (Jara & Escalante, Dawestrema cycloancistrum Price & 1984). Nowlin, 1967 Cosmetocleithrum bulbocirrus Kritsky, Arapaima gigas; adult; fins, gills, skin; Thatcher & Boeger, 1986 freshwater; Amazon River, Loreto Pterodoras granulosus; adult; gills; (Iannacone & Luque, 1991; Mathews et freshwater; Loreto (Mendoza-Palmero et al., 2007; Mathews et al., 2013a; García- al., 2012). Ruiz et al., 2013; Mathews et al., 2014a; Cosmetocleithrum confusus Kritsky, Serrano-Martínez et al., 2015a). Thatcher & Boeger, 1986 Demidospermus brevicirrus Mendoza- Oxydoras niger; adult; gills; freshwater; Palmero, Scholz, Mendoza-Franco & Loreto (Iannacone & Luque, 1991). Kuchta, 2012 Cosmetocleithrum gussevi Kritsky, Pimelodus sp.; adult; gills; freshwater; Thatcher & Boeger, 1986 Loreto (Mendoza-Palmero et al., 2012). Oxydoras niger; adult; gills; freshwater; Demidospermus centromochli Mendoza- Loreto (Iannacone & Luque, 1991). Franco & Scholz, 2009 Cosmetocleithrum sobrinus Kritsky, Centromochlus heckelii; adult; gills; Thatcher & Boeger, 1986 freshwater; Loreto (Mendoza-Franco & Oxydoras niger; adult; gills; freshwater; Scholz, 2009). Loreto (Iannacone & Luque, 1991). Demidospermus curvovaginatus Mendoza- Cosmetocleithrum sp. Palmero & Scholz, 2011 Hassar orestis; adult; gills; freshwater; Pimelodus sp., gen. sp.; Loreto (Mendoza-Palmero et al., 2015). adult; gills; freshwater; Loreto (Mendoza- Dactylogyridae gen. sp. Palmero & Scholz, 2011). Ageneiosus vittatus, Colossoma Demidospermus lebedev (Kritsky & macropomum, Hypophthalmus edentatus, Thatcher 1976) Mendoza-Palmero & Platynematichthys notatus, Platysilurus Scholz, 2011 mucosus, Pseudoplatystoma fasciatum, Pimelodus blochii, Pimelodus sp.; adult; Sorubim lima; adult; gills; freshwater; gills; freshwater; Loreto (Mendoza- Loreto (Bances et al., 2013; Alcántara- Palmero & Scholz, 2011). Bocanegra et al., 2015; Mendoza-Palmero Demidospermus macropteri Mendoza- et al., 2015; Vargas et al., 2015). Franco & Scholz, 2011 sp. Calophysus macropterus; adult; gills; Aspistogramma sp., Cichlasoma sp., freshwater; Loreto (Mendoza-Franco & Carassius auratus, Pterophyllum scalare, Scholz, 2009). Symphysodon aequifasciatus; gills; Demidospermus mortenthaleri Mendoza- tegument; freshwater; Amazon River Palmero, Scholz, Mendoza-Franco & basin, Lima (Gonzáles-Fernández, 2012a; Kuchta, 2012 Aguinaga et al., 2015). Brachyplatystoma juruense; adult; gills; Nybelin, 1924 freshwater; Loreto (Mendoza-Palmero et Cyprinus carpio carpio; adult; gills; al., 2012, 2015). freshwater; La Libertad (Jara & Escalante, Demidospermus peruvianus Mendoza- 1983a). Palmero & Scholz, 2011

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Pimelodus ornatus, P. blochii, Pimelodus freshwater; Amazon River Basin (Kritsky sp., Pimelodidae gen. sp.; adult; gills; & Boeger, 2002). freshwater; Loreto (Mendoza-Palmero & Gonocleithrum cursitans (Kritsky & Scholz, 2011). Thatcher, 1983) Demidospermus striatus Mendoza-Palmero Osteoglossum bicirrhosum; adult; gills; & Scholz, 2011 freshwater; Amazon River Basin Pimelodus blochii, Pimelodus sp.; adult; (Vásquez et al., 2007). gills; freshwater; Loreto (Mendoza- Gussevia alioides Kritsky, Thatcher & Palmero & Scholz, 2011). Boeger, 1986 Demidospermus sp. Heros severus; adult; gills; freshwater; Brachyplatystoma vaillantii; adult; gills; Loreto (Mendoza-Franco et al., 2010). freshwater; Loreto (Mendoza-Palmero et Gussevia asota Kritsky, Thatcher & Boeger, al., 2015). 1989 Euryhaliotrema chaoi Kritsky & Boeger, Astronotus ocellatus; adult; gills; 2002 freshwater; Loreto (Mendoza-Franco et Plagioscion squamosissimus; adult; gills; al., 2010; Mathews et al., 2014b). freshwater; Amazon River Basin (Kritsky Gussevia disparoides Kritsky, Thatcher & & Boeger, 2002). Boeger, 1986 Euryhaliotrema lovejoyi Kritsky & Boeger, Cichlasoma amazonarum, Heros severus; 2002 adult; gills; freshwater; Loreto (Mendoza- Plagioscion squamosissimus; adult; gills; Franco et al., 2010). freshwater; Amazon River Basin (Kritsky Gussevia longihaptor (Mizelle & Kritsky, & Boeger, 2002). 1969) Euryhaliotrema monacanthus Kritsky & Cichla monoculus; adult; gills; Boeger, 2002 freshwater; Loreto (Mendoza-Franco et Plagioscion squamosissimus; adult; gills; al., 2010). freshwater; Amazon River Basin (Kritsky Gussevia spiralocirra Kohn & Paperna, & Boeger, 2002). 1964 Euryhaliotrema paralonchuri (Luque & Pterophyllum scalare; adult; gills; Iannacone, 1989) freshwater; Loreto (Kritsky et al., 1986). Paralonchurus peruanus; adult; gills; Gussevia tucunarense Kritsky, Thatcher & marine; Lima (Luque & Iannacone, Boeger, 1986 1989a; Luque et al., 1991; Luque & Oliva, Chaetobranchus semifasciatus; adult; 1993a; Oliva & Luque, 1998b). gills; freshwater; Loreto (Mathews et al., Euryhaliotrema potamocetes Kritsky & 2013c). Boeger, 2002 Gussevia undulata Kritsky, Thatcher & Plagioscion squamosissimus; adult; gills; Boeger, 1986 freshwater; Amazon River Basin (Kritsky Cichla monoculus; adult; gills; & Boeger, 2002). freshwater; Loreto (Mendoza-Franco et Euryhaliotrema succedanus Kritsky & al., 2010; Mathews et al., 2012). Boeger, 2002 Hamatopeduncularia sp. Plagioscion squamosissimus; adult; gills; Galeichthys peruvianus; adult; gills; freshwater; Amazon River Basin (Kritsky marine; Lima (Iannacone & Luque, & Boeger, 2002). 1993a). Euryhaliotrema thatcheri Kritsky & Boeger, Haliotrema mugilinus Hargis, 1955 2002 Mugil cephalus; adult; gills; freshwater; Plagioscion squamosissimus; adult; gills; La Libertad (Jara & Escalante, 1983a).

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Jainus amazonensis Kritsky, Thatcher & (Tantaleán & Huiza 1994). Kayton, 1980 Rhinonastes pseudocapsaloideum Kritsky, Brycon cephalus; adult; gills; freshwater; Thatcher & Boeger, 1988 Loreto (Mathews et al., 2014a) Prochilodus nigricans; adult; gills; Mexicana iannaconi Chero, Cruces, Sáez & freshwater; Loreto (Mathews et al., Alvariño, 2014 2013b). Haemulon steindachneri; adult; gills; Rhinoxenus piranhus Kritsky, Thatcher & marine; Lima (Chero et al., 2014e). Boeger, 1988 Mexicana sp. Pygocentrus nattereri; adult; nasal Anisotremus scapularis; adult; gills; cavities; freshwater; Loreto (Iannacone & marine; Lima (Iannacone & Alvariño, Luque, 1993b). 2012; Chero et al., 2014d; Paschoal et al., Sciadicleithrum satanopercae Yamada, 2015). Takemoto, Bellay & Pavanelli, 2009 Mymarothecium viatorum Boeger, Piasecki Satanoperca jurupari; adult; gills; & Sobecka, 2002 freshwater; Loreto (Mendoza-Franco et Piaractus brachypomus; gills; freshwater; al., 2010). Amazon River basin (Dinis-Vásquez, Sciadicleithrum variabilum (Mizelle & 2014). Kritsky 1969) Kritsky, Thatcher & Notothecium bethae Kritsky, Boeger & Jegu Boeger, 1989 1996 Cichlasoma amazonarum; adult; gills; Pygocentrus nattereri; adult; gills; freshwater; Loreto (Mendoza-Franco et freshwater; Loreto (Gonzales et al., al., 2010). 2015). Tucunarella cichlae Mendoza-Franco, Notothecium mizellei Boeger & Kritsky, Scholz & Rozkosna, 2010 1988 Cichla monoculus; adult; gills; Myleus schomburgkii; adult; gills; freshwater; Loreto (Mendoza-Franco et freshwater; Loreto (Iannacone & Luque, al., 2010). 1993b). Unibarra paranoplatensis Suriano & Notozothecium penetratum Boeger & Incorvaia, 1995 Kritsky, 1988 Aguarunichthys torosus; adult; gills; Pygocentrus nattereri; adult; gills; freshwater; Loreto (Mendoza-Palmero et freshwater; Loreto (Iannacone & Luque, al., 2015). 1993b). Unilatus brittani Mizelle, Kritsky & Crane, Notozothecium sp. 1968 Myleus schomburgkii; adult; gills; Pterygoplichthys anisitsi; adult; gills; freshwater; Loreto (Fernández-Méndez et freshwater; Loreto (Mendoza-Palmero et al., 2015). al., 2012). Oncocleidus sp. Unilatus unilatus Mizelle & Kritsky, 1967 Lebiasina bimaculata; adult; gills; Pterygoplichthys anisitsi; adult; gills; freshwater; La Libertad (Jara & Escalante, freshwater; Loreto (Mendoza-Palmero et 1983b). al., 2012). Parancylodiscoides chaetodipteri Caballero Urocleidoides eremitus Kritsky, Thatcher & & Bravo-Hollis, 1961 Boeger, 1986 Parapsettus panamensis; adult; gills; Hoplias malabaricus; adult; gills; marine; Tumbes (Chero et al., 2015c) freshwater; Madre de Dios (Iannacone & Pseudohaliotrema sp. Luque, 1993b). Genypterus sp.; adult; gills; marine; Lima Urocleidus sp.

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Aequidens rivulatus; adult; gills; marine; Lambayeque, Lima, Tumbes freshwater; La Libertad (Jara & Escalante, (Luque & Oliva, 1993a; Tantaleán & 1983b). Huiza, 1994; Iannacone et al., 2000; Vancleaveus fungulus Kritsky, Thatcher & Iannacone et al., 2001; Minaya et al., Boeger, 1986 2016). Pseudoplatystoma fasciatum; adult; gills; jeanloui Knoff, freshwater; Loreto (Mendoza-Palmero et Cohen, Cárdenas, Cárdenas-Callirgos al., 2012). & Gomes, 2015 Vancleaveus janauacaensis Kritsky, Paranthias colonus; adult; gills; marine; Thatcher & Boeger, 1986 Lima (Knoff et al., 2015). Pterodoras granulosus; adult; gills; Rhamnocercoides menticirrhi Luque & freshwater; Loreto (Mendoza-Palmero et Iannacone, 1991 al., 2012, 2015). Menticirrhus ophicephalus; adult; gills; Vancleaveus platyrhynchi Kritsky, Thatcher marine; Lima (Luque & Iannacone, & Boeger, 1986 1991a; Luque, 1994a; Oliva & Luque, Hemisorubim platyrhynchos; adult; gills; 1998b). freshwater; Loreto (Mendoza-Palmero et Rhamnocercus oliveri Luque & Iannacone, al., 2012). 1991 Stellifer minor; adult; gills; marine; Lima Family Diplectanidae Monticelli, 1903 (Luque & Iannacone, 1991b; Iannacone, Diplectanidae gen. sp. 1 2004). Cheilotrema fasciatum; adult; gills; Rhamnocercus sp. marine; Lima (Oliva & Luque, 1998b). Stellifer minor; adult; gills; marine; La Diplectanidae gen. sp. 2 Libertad, Lambayeque, Lima (Oliva et al., Cheilotrema fasciatum; adult; gills; 1990; Luque et al., 1991; Jara, 1998; Oliva marine; Lima (Oliva & Luque, 1998b). & Luque, 1998b). Diplectanidae gen. sp. 3 R h a m n o c e r c u s s t e l l i f e r i L u q u e & Paralonchurus peruanus; adult; gills; Iannacone, 1991 marine; Lima (Oliva & Luque, 1998b). Stellifer minor; adult; gills; marine; Lima Diplectanum decorum Kritsky & Thatcher, (Luque & Iannacone, 1991b; Iannacone, 1984 2004). Plagioscion squamosissimus; adult; gills; freshwater; Loreto (Iannacone & Luque, Family Tetraonchidae Monticelli, 1903 1993b). Tetraonchus monenteron Diesing, 1858 Diplectanum pescadae Kritsky & Thatcher, Arapaima gigas; adult; gills; freshwater; 1984 Loreto (Tantaleán et al., 1985a). Plagioscion squamosissimus; adult; gills; Tylosuricola amatoi Iannacone & Luque, freshwater; Loreto (Iannacone & Luque, 1990 1993b). Strongylura scapularis; adult; gills; Diplectanum piscinarius Kritsky & marine; Lima (Iannacone & Luque, 1990). Thatcher, 1984 Plagioscion squamosissimus; adult; gills; Order Diclybothriidea Bychowsky, 1957 freshwater; Loreto (Iannacone & Luque, 1993b). Family Hexabothriidae Price, 1942 Diplectanum sp. Callorhynchocotyle marplatensis Suriano & Cynoscion analis, Paralonchurus Incorvaia, 1982 peruanus, Stellifer minor; adult; gills; Callorhynchus callorhynchus; adult; gills;

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marine; Lima (Luque et al., 1991; Luque freshwater; La Libertad (An et al., 1991). & Iannacone, 1991b; Tantaleán & Huiza, Gyrodactylus sp. 1994). Carassius auratus, Lebiasina bimaculata, Erpocotyle sp. Oreochromis spp., Trichomycterus Mustelus dorsalis; adult; gills; marine; La punctulatus; gills; tegument; freshwater; Libertad, Lima, Piura (Tantaleán & Huiza, La Libertad, Lima (Jara & Escalante, 1994). 1984; Jara, 1986; Gonzáles-Fernández, Rhinobatonchocotyle cyclovaginatus Doran, 2012ab). 1953 Gyrodactylus turnbulli Harris, 1986 Rhinobatos planiceps; adult; gills; Poecilia reticulata; adult; gills; marine; Lima (Tantaleán et al., 1998). freshwater; La Libertad (An et al., 1991). Rhinobatonchocotyle pacifica Oliva & Oögyrodactylus farlowellae Harris, 1983 Luque, 1995 Farlowella amazonum; adult; skin, fins; Rhinobatos planiceps; adult; gills; freshwater; Loreto (Harris, 1983). marine; Lima (Iannacone et al., 2011a). Scleroductus yuncensi Jara & Cone, 1989 Pimelodella yuncensis; adult; gills; Order Gyrodactylidea Bychowsky, 1937 freshwater; La Libertad (Jara & Cone, 1989). Family Gyrodactylidae Van Beneden & Hesse, 1863 Order Bychowsky, 1957 Accessorius peruensis Jara, An & Cone, 1991 Family Allopyragraphoridae Yamaguti, Lebiasina bimaculata; adult; gills; 1963 freshwater; La Libertad (Jara et al., 1990). Allopyragraphorus caballeroi (Zerecero, Anacanthocotyle sp. 1960) Bryconamericus peruanus; adult; gills; Caranx hippos; adult; gills; marine; freshwater; La Libertad (Jara, 1986). Ancash, La Libertad (Tantaleán et al., Gyrodactylidae gen. sp. 1988; Luque et al., 1991; Tantaleán & Colossoma macropomum; adult; gills; Huiza, 1994). freshwater; Loreto (Alcántara-Bocanegra et al., 2015). Family Anthocotylidae Price, 1936 Gyrodactylus bimaculatus An, Jara & Cone, Anthocotyle americana (MacCallum, 1916) 1991 Merluccius gayi peruanus; adult; gills; Lebiasina bimaculata; adult; gills; marine; Lima (Durán & Oliva, 1980; freshwater; La Libertad (An et al., 1991). Verano et al., 1990; Luque et al., 1991; Gyrodactylus lebiasinus An, Jara & Cone, Tantaleán & Huiza, 1994; Chero et al., 1991 2014b). Lebiasina bimaculata; adult; body Anthocotyle merluccii van Beneden & washings; freshwater; La Libertad (An et Hesse, 1863 al., 1991). Merluccius gayi peruanus; adult; gills; Gyrodactylus pimelodellus An, Jara & Cone, marine; Lima (Durán & Oliva, 1980; 1991 Iannacone & Luque, 1990; Oliva & Pimelodella yuncensis; adult; gills; Ballón, 2002). freshwater; La Libertad (An et al., 1991). Gyrodactylus slendrus An, Jara & Cone, Family Axinidae Monticelli, 1903 1991 Axine ibanezi Tantaleán, 1975 Lebiasina bimaculata; adult; gills; Exocoetus volitans; adult; gills; marine;

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Lima (Tantaleán, 1975a; Luque et al., adult; gills; marine; Ancash, Ica, La 1991; Tantaleán & Huiza, 1994). Libertad, Lima (Tantaleán et al., 1988; Loxura peruensis Oliva & Luque, 1995 Luque et al., 1991; Tantaleán & Huiza, Strongylura exilis, S.scapularis; adult; 1994). gills; marine; Lima, Northern coastal zone Choricotyle sp. (Escalante et al., 1982b; Luque et al., Cilus gilberti; adult; gills; marine; Lima 1991; Tantaleán & Huiza, 1994; Oliva & (Chero et al., 2014c). Luque, 1995). Hargicotyle chimbotensis (Tantaleán, 1974) Paralonchurus peruanus; adult; gills; Family Cemocotylidae Yamaguti, 1963 marine; Ancash (Tantaleán, 1974ab; Cemocotyle sp. Tantaleán & Huiza, 1994). Trachurus murphyi; adult; gills; marine; Hargicotyle louisianensis (Hargis, 1955) La Libertad, Piura (Tantaleán & Huiza, Cilus gilberti, Sciaena deliciosa; adult; 1994). gills; marine; Lima (Chero et al., 2014ac). Cemocotyle trachuri Dillon & Hargis, 1965 Hargicotyle magna Oliva & Luque, 1989  Trachurus murphyi; adult; gills; Cheilotrema fasciatum; adult; gills, marine; Piura (Oliva, 1999). mouth; marine; Lima (Oliva & Luque, 1989b; Luque et al., 1991; Tantaleán & Family Chauhaneidae Euzet & Trilles, Huiza, 1994; Oliva & Luque, 1998b). 1960 Hargicotyle menticirrhi Oliva & Luque, 1989 Oaxacotyle oaxacensis (Caballero & Bravo, Menticirrhus ophicephalus; adult; gills, 1964) Lebedev, 1984 mouth; marine; Lima (Oliva & Luque, Peprilus medius; P. snyderi; adult; gills; 1989b; Luque et al., 1991; Luque 1994a; marine; Lima (Iannacone & Alvariño, Tantaleán & Huiza, 1994; Oliva & Luque, 2008; Ñacari & Sánchez, 2014). 1998b). Hargicotyle paralonchuri Oliva & Luque, Family Diclidophoridae Cerfontaine, 1895 1989 Choricotyle anisotremi Oliva, 1987 Cynoscion analis, Paralonchurus Anisotremus scapularis; adult; gills; peruanus; adult; gills; marine; Lima marine; Lima (Iannacone & Alvariño, (Oliva & Luque, 1989b; Luque et al., 2009b; Paschoal et al., 2015). 1991; Luque & Oliva, 1993a; Tantaleán & Choricotyle caulolatili (Meserve, 1938) Huiza, 1994; Oliva & Luque, 1998b; Sproston, 1946 Iannacone et al., 2001). Caulolatilus sp.; adult; gills; marine; Hargicotyle peruensis (Tantaleán, 1974) Ancash (Tantaleán et al., 1985b; Luque et Paralonchurus peruanus, Sciaena al., 1991; Tantaleán & Huiza, 1994).  deliciosa; adult; gills; marine; Ancash Choricotyle isaciencis Oliva, González, Ruz (Tantaleán, 1974b; Tantaleán & Huiza, & Luque, 2009 1994). Isacia conceptionis; adult; gills; marine; Hargicotyle sciaenae Oliva & Luque, 1989 Lima (Iannacone et al., 2015). Sciaena deliciosa; adult; gills; marine; Choricotyle scapularis Oliva, González, Ruz Lima (Oliva & Luque, 1989b; Luque et al., & Luque, 2009 1991; Tantaleán & Huiza, 1994; Oliva & Anisotremus scapularis; adult; gills; Luque, 1998b; Iannacone et al., 2010a). marine; Lima (Chero et al., 2014d). Hargicotyle sp. Choricotyle sonorensis Caballero & Bravo- Cynoscion analis, Sciaena deliciosa; Hollis, 1962 adult; gills; marine; Ica (Tantaleán & Isacia conceptionis, Sardinops sagax; Huiza, 1994).

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Hemitagia galapagensis (Meserve, 1938) Family Price, 1943 Paralabrax humeralis; adult; gills; Pseudaxine sp. marine; La Libertad, Lima, Piura Scomberomorus sierra; adult; unspecified (Tantaleán et al., 1982; Luque et al., 1991; site of infection; marine; La Libertad Tantaleán & Huiza, 1994). (Escalante et al., 1982b). N e o h e t e r o b o t h r i u m c y n o s c i o n i (MacCallum, 1917) Family Heteraxinidae Unnithan, 1957 Cynoscion analis; adult; gills; marine; Ica, Allencotyla mcintoshi (Price, 1962) Lima, Tumbes (Tantaleán et al., 1982; Seriola lalandi; adult; gills; marine; Luque et al., 1991; Tantaleán & Huiza, Ancash (Tantaleán et al., 1988; Luque et 1994; Iannacone et al., 2001; Minaya et al., 1991; Tantaleán & Huiza, 1994). al., 2016). Cynoscionicola americanus Tantaleán, Paraeurysorchis sarmientoi (Tantaleán, Martinez & Escalante, 1987 1974) Cheilotrema fasciatum, Cilus gilbert, Seriolella violacea; adult; gills; marine; M e n t i c i r r h u s o p h i c e p h a l u s , Lima (Tantaleán, 1974b; Tantaleán et al., Paralonchurus peruanus, Stellifer minor; 1985b; Luque et al., 1991; Tantaleán & adult; gills; marine; Ancash, Lima Huiza, 1994; Iannacone, 2003). (Tantaleán et al., 1987; Luque et al., 1991; Pedocotyle annakohni Luque & Iannacone, Luque & Oliva, 1993a; Luque, 1994a; 1989 Tantaleán & Huiza, 1994; Oliva & Luque, Stellifer minor; adult; gills; marine; Lima 1998b; Chero et al., 2014c). (Luque & Iannacone, 1989b; Oliva et al., Cynoscionicola cynoscioni Tantaleán, 1990; Luque et al., 1991; Tantaleán & Martinez & Escalante, 1987 Huiza, 1994; Oliva & Luque, 1998b). Cynoscion analis; adult; gills; marine; Pedocotyle bravoi Luque & Iannacone, 1989 Ancash, Lima, Tumbes (Tantaleán et al., Stellifer minor; adult; gills; marine; La 1987; Luque et al., 1991; Tantaleán & Libertad, Lima, Northern coastal zone Huiza, 1994; Iannacone et al., 2001; (Luque & Iannacone, 1989b; Oliva et al., Iannacone, 2005 ; Minaya et al., 2016). 1990; Luque et al., 1991; Tantaleán & Cynoscionicola intermedia Tantaleán, Huiza, 1994; Jara, 1998; Oliva & Luque, Martinez & Escalante 1988 1998b). Xenistius peruanus; adult; gills; marine; Pedocotyle sp. Ancash (Tantaleán et al., 1988; Luque et Stellifer minor; adult; gills; marine; Ica, al., 1991; Tantaleán & Huiza, 1994; Lambayeque (Tantaleán & Huiza, 1994). Paschoal et al., 2015). Pseudoeurysorchis travassosi Caballero & Cynoscionicola sciaenae Tantaleán, 1974 Bravo-Hollis, 1962 Sciaena deliciosa; adult; gills; marine; Isacia conceptionis; adult; gills; marine; Ica, Lima (Tantaleán, 1974a; Luque et al., Ica, Lima (Tantaleán et al., 1985b; Luque 1991; Tantaleán & Huiza, 1994; Oliva & et al., 1991; Tantaleán & Huiza, 1994; Luque, 1998b; Iannacone et al., 2010a; Paschoal et al., 2015; Iannacone et al., Chero et al., 2014a). 2015). Cynoscionicola sp. Teleurysorchis gumercindae Gonzaléz & Paralonchurus peruanus; Stellifer minor; Sarmiento, 1990 adult; gills; marine; Ica, Lima (Oliva et al., Schedophilus haedrichi; adult; gills; 1989, 1990; Tantaleán & Huiza, 1994). marine; Ancash (González & Sarmiento, Zeuxapta seriolae (Meserve, 1938) 1990; Luque et al., 1991). Chloroscombrus orqueta; adult; gills; marine; La Libertad, Lima (Tantaleán et

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al., 1985b; Luque et al., 1991; Tantaleán Mamaev & Egorova, 1977 & Huiza, 1994). Caulolatilus sp.; adult; gills; marine; Lima (Tantaleán et al., 1982; Tantaleán & F a m i l y M a c ro v a l v i t re m a t i d a e Huiza, 1994). Yamaguti, 1963 Metamicrocotyla macracantha (Alexander, Macrovalvitrema sp. 1954) Koratha, 1955  Sciaena deliciosa; adult; gills; marine; Mugil cephalus; adult; gills; marine; Lima (Chero et al., 2014a). Ancash, Ica, Lima, Northen Coastal zone Pterinotrematoides mexicanum Caballero & (Tantaleán, 1974a; Tantaleán et al., 1982; Bravo-Hollis, 1955 Luque, 1994b; Luque et al., 1991; Micropogonias sp.; adult; gills; marine; Tantaleán & Huiza, 1994; Vásquez-Ruiz Lima (Tantaleán et al., 1985b; Tantaleán et al., 2012). & Huiza, 1994). Microcotyle nemadactylus Dillon & Hargis, 1965 Family Price, 1936 Cheilodactylus variegatus; adult; gills; Kuhnia indica Tripathi, 1959 marine; Lima (Martínez & Barrantes, Scomber japonicus; adult; gills; marine; 1977; Luque et al., 1991; Tantaleán & Arequipa (Ruelas & Cordova, 1995). Huiza, 1994; Oliva & Luque, 1998a; Kuhnia scombri (Kuhn, 1829) Sproston, Iannacone et al., 2003). 1945 Microcotyloides impudicus Caballero, Scomber japonicus; adult; gills; marine; Bravo-Hollis & Grocott, 1955 Arequipa, Lambayeque, Lima (Tantaleán Polydactylus approximans; adult; gills; et al., 1988; Luque et al., 1991; Tantaleán marine; Lima (Tantaleán & Huiza, 1994). & Huiza, 1994; Oliva et al., 2008; Ruelas Neobivagina chita Tantaleán, Morales & & Cordova, 1995). Escalante, 1998 Kuhnia sp. Anisotremus scapularis; adult; intestine; Trachinotus paitensis; adult; gills; marine; Lima (Tantaleán et al., 1998; marine; La Libertad (Jara, 1998). Iannacone & Alvariño, 2009; Chero et al., Kuhnia sprostonae Price, 1961 2014d; Paschoal et al., 2015). Scomber japonicus; adult; gills; marine; Ancash, Lambayeque, Lima (Tantaleán et Family Protomicrocotylidae Johnston & al., 1988; Luque et al., 1991; Tantaleán & Tiegs, 1922 Huiza, 1994; Oliva et al., 2008). Neomicrocotyle sp. Caranx hippos; adult; gills; marine; La Family Taschenberg, 1879 Libertad, Piura (Tantaleán & Huiza, Bicotylophora trachinoti (MacCallum, 1921) 1994). Price, 1936 Trachinotus paitensis; adult; gills; marine; Family Pyragraphoridae Yamaguti, 1963 Ica, Lima, Piura (Iannacone & Luque, Pyragraphorus pyragraphorus (MacCallum 1990; Luque et al., 1991; Tantaleán & & MacCallum, 1913) Sproston, 1946 Huiza, 1994). Trachinotus paitensis; adult; unspecified Intracotyle neghmei (Villalba, 1987) Oliva & site of infection; marine; Piura (Iannacone Luque, 1995 & Luque, 1990). Anisotremus scapularis; adult; gills; marine; Lima (Oliva & Luque, 1995; Family Thoracocotylidae Price, 1936 Paschoal et al., 2015). Mexicotyle mexicana (Meserve, 1938) Jaliscia caballeroi (Bravo-Hollis, 1960) Scomberomorus sierra; adult; gills;

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marine; Ancash, Ica (Escalante et al., Potamotrygon sp.; adult; gills; freshwater; 1982b; Tantaleán et al., 1988; Tantaleán & Loreto (Domingues & Marques, 2011). Huiza, 1994). Potamotrygonocotyle tsalickisi Mayes, Thoracocotyle crocea MacCallum, 1913 Brooks & Thorson, 1981 Scomberomorus sierra; adult; gills; Potamotrygon sp.; adult; gills; freshwater; marine; Ica, La Libertad (Tantaleán et al., Rio Madre de Dios (Domingues & 1985b; Luque et al., 1991; Tantaleán & Marques, 2011). Huiza, 1994). Class Trematoda Rudolphi, 1808 Order Monocotylidea Lebedev, 1988 Order Aspidogastrida Dollfus, 1958

Family Loimoidae Price, 1936 Family Aspidogastridae Poche, 1907 Loimos scoliodoni (Manter, 1938) Lobatostoma anisotremum Oliva & Mustelus dorsalis; adult; gills; marine; La Carvajal, 1984 Libertad, Piura (Tantaleán et al., 1988; Anisotremus scapularis; adult; intestine, Tantaleán & Huiza, 1994). stomach; marine; Central Coast, Lima, Loimos sp. Zona Sur (Moquegua-Tacna) (Ruelas & Stellifer minor; adult; gills; marine; La Córdova, 1995; Oliva & Luque, 1989a; Libertad (Jara, 1998). Luque et al., 1991; Tantaleán & Huiza, 1994; Chero et al., 2014d; Alves et al., 2015b; Paschoal et al., 2015). Family Monocotylidae Taschenberg, 1879 Lobatostoma kemostoma (MacCallum & Anoplocotyloides chorrillensis Luque & MacCallum, 1913) Iannacone, 1991 Trachinotus paitensis; adult; intestine; Rhinobatos planiceps; adult; gills; marine; Lima (Luque & Oliva, 1993b; marine; Lima (Luque & Iannacone, Alves et al., 2015b). 1991b). Lobatostoma pacificum Manter, 1940 Anoplocotyloides papillatus (Doran, 1953) Trachinotus paitensis, T. rhodopus; adult; Young, 1967 intestine; marine; Central Coast, La Rhinobatos planiceps; adult; gills; Libertad, Lima (Oliva & Luque, 1989a; marine; Lima (Luque & Iannacone, Luque et al., 1991; Tantaleán & Huiza, 1991b). 1994, Jara, 1998; Alves et al., 2015b). Heterocotyle sp. Lobatostoma sp. Rhinobatos planiceps; adult; gills; Peprilus medius, Trachinotus paitensis; marine; La Libertad (Escalante et al., adult; intestine; marine; La Libertad, 1982b; Tantaleán & Huiza, 1994). Lima (Iannacone & Alvariño, 2008). P o t a m o t r y g o n o c o t y l e c h i s h o l m a e Lobatostoma veranoi Oliva & Luque, 1989 Domingues & Marques, 2007 Menticirrhus ophicephalus; adult; Potamotrygon motoro adult; gills; intestine; marine; Central Coast, Lima freshwater; Madre de Dios (Domingues & (Oliva & Luque, 1989ab; Luque et al., Marques, 2011). 1991; Luque, 1994a; Tantaleán & Huiza, P o t a m o t r y g o n o c o t y l e d r o m e d a r i u s 1994; Alves et al., 2015b). Domingues & Marques, 2007 Potamotrygon motoro; adult; gills; Order Plagiorchiida La Rue, 1957 freshwater; Madre de Dios (Domingues & Marques, 2011). Family Allocreadiidae Looss, 1902 P o t a m o t r y g o n o c o t y l e r i o n e g r e n s e Auriculostoma foliaceum Curran, Tkach & Domingues, Pancera & Marques, 2007

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Overstreet, 2011 Notoporus sp. Bryconops caudomaculatus; adult; Orthopristis chalceus; adult; intestine; intestine; freshwater; Loreto (Curran et marine; Northern coastal zone (Escalante al., 2011). et al., 1984; Tantaleán & Huiza, 1994; Auriculostoma diagonale Curran, Tkach & Paschoal et al., 2015). Overstreet, 2011 Plagioporus sp. Stethaprion erythrops; adult; intestine; Cilus gilberti; adult; intestine; marine; freshwater; Loreto (Curran et al., 2011) Lima (Chero et al., 2014c). Pseudopecoelus sp. Family Opecoelidae Ozaki, 1925 Sebastes capensis; adult; intestine; Anisoporus sp. marine; Lima (González & Poulin, 2005; Paralichthys woolmani; adult; intestine; González et al., 2006). marine; La Libertad (Luque et al., 1991; Sphaerostoma sp. Tantaleán & Huiza, 1994). Aequidens rivulatus; adult; intestine; Bunoderina sp. freshwater; La Libertad (Jara, 1997; Jara Brycon atrocaudatus; adult; intestine; et al., 1998). freshwater; La Libertad (Jara et al., 1997). Villarrealina peruana Bolaños & Salas, 1982 Cainocreadium sp. Cilus gilberti, Sciaena deliciosa; adult; Paralichthys adspersus, Stellifer minor; intestine; marine; Lima (Luque et al., adult; intestine, pyloric caecum; marine; 1991; Tantaleán & Huiza, 1994; Chero et Lambayeque, Lima (Tantaleán & Huiza, al., 2014c). 1994; Iannacone et al., 2012a). Helicometra fasciata (Rudolphi, 1819) Family Azygiidae Lühe, 1909 Odhner, 1902 Otodistomum veliporum (Creplin, 1837) Cheilotrema fasciatum, Cilus gilberti, Stafford, 1934 Gymnothorax porphyreus, Labrisomus S q u a t i n a a r m a t a , S y m p t e r y g i a philippii, Lepophidium sp., Menticirrhus brevicaudata; adult; coelomic cavity, ophicephalus, Paralabrax humeralis, stomach; marine; Ancash, La Libertad Paralonchurus peruanus, Sciaena (Escalante & Jara, 1983; Tantaleán, 1991; deliciosa, Stellifer minor; adult; intestine; Tantaleán & Huiza, 1994). marine; Arequipa, Ica, La Libertad, Lima (Tantaleán et al., 1975; Luque et al., 1991; Family Bucephalidae Poche, 1907 Luque & Oliva, 1993ab; Luque, 1994a; Bucephalus sp. Tantaleán & Huiza, 1994; Jara, 1998; Labrisomus philippii; adult; larvae; Oliva & Luque, 1998b, 2002; Iannacone, stomach; muscles; marine; Lima (Oliva & 2004; Iannacone et al., 2010a; Iannacone Luque, 2002; Cruces et al., 2015). et al., 2011b; Chero et al., 2014ac) Dollfustrema sp. Helicometrina nimia Linton, 1910 Strongylura scapularis; intestine; marine; Sebastes capensis, Sicyases sanguineus; Northern coastal zone (Escalante et al., adult; intestine; marine; Lima (González 1984a; Tantaleán & Huiza, 1994). & Poulin, 2005; González et al., 2006; Prosorhynchoides carvajali Munoz & Bott, Iannacone & Alvariño, 2011). 2011 Helicometrina sp. Labrisomus philippii; adult; intestine; Acanthistius pictus, Paralabrax stomach; marine; Lima (Cruces et al., humeralis; adult; intestine; marine; 2015). Arequipa, Lima (Tantaleán & Huiza, Prosorhynchoides rioplatensis (Szidat, 1994). 1970) Lunaschi, 2003

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Labrisomus philippii; adult; intestine; marine; 1994). Lima (Iannacone et al., 2011b) Anaporrhutum sp. Prosorhynchoides sp. Scomberomorus sierra; adult; abdominal Labrisomus philippii, Odontesthes regia, cavity; marine; Piura (Tantaleán & Huiza, Paralichthys adspersus, Sicyases 1994). sanguineus; adult; intestine, muscles; Nagmia peruviana Tantaleán & Rodríguez, marine; Lambayeque, Lima, Northern 1990 coastal zone, Piura (Escalante et al., Mobula thurstoni; adult; visceral cavity; 1984b; Tantaleán & Huiza, 1994; Oliva & marine; Ancash, Lima (Escalante et al., Luque, 2002). 1983; Tantaleán & Rodriguez, 1990; Prosorhynchus sp. Tantaleán & Huiza, 1994). Alphestes afer, Katsuwonus pelamis, Phyllodistomum sp. Labrisomus philippii, Merluccius gayi P s e u d o p l a t y s t o m a f a s c i a t u m , peruanus, Scartichthys gigas; adult, Trachinotus paitensis; adult; kidney, larvae; caecum intestine, muscles; urynary blader; freshwater, marine; La marine; Ancash, La Libertad, Lima Libertad, Loreto (Tantaleán et al., 1985b; (Durán & Oliva, 1980; Escalante & Jara, Tantaleán & Huiza, 1994). 1983; Escalante et al., 1984a; Luque, 1991; Tantaleán & Huiza, 1994; Oliva & Family Fellodistomidae Nicoll, 1909 Luque, 2002). Monascus sp. Rhipidocotyle adbaculum Manter, 1940 Trachurus murphyi; adult; intestine; Sarda chiliensis; adult; stomach; marine; marine; Lambayeque, Piura, (Tantaleán & Lima (Luque et al., 1991; Tantaleán & Huiza, 1994). Huiza, 1994). Proctoeces lintoni Siddiqi & Cable, 1960 Rhipidocotyle sp. A n i s o t re m u s s c a p u l a r i s , I s a c i a Sarda chiliensis; Cilus gilbert; adult; conceptionis, Sicyases sanguineus; adult; intestine; marine; Arequipa (Tantaleán & intestine; marine; Lima (Luque & Oliva, Huiza, 1994). 1993b; Iannacone & Alvariño, 2009b; 2011; Paschoal et al., 2015; Iannacone et Family Echinostomatide Looss, 1899 al., 2015). Caballerotrema arapaimense Thatcher, Proctoeces sp. 1980 A n i s o t re m u s s c a p u l a r i s , I s a c i a Arapaima gigas; adult; intestine; conceptionis, Sicyases sanguineus; adult; freshwater; Loreto (Mathews et al., gills, intestine; marine; Ica, La Libertad, 2013a). Lima, (Escalante et al., 1984a; Tantaleán Caballerotrema brasiliense Prudhoe, 1960 et al., 1992; Tantaleán & Huiza, 1994; Arapaima gigas; adult; intestine; Chero et al., 2014d; Paschoal et al., 2015). freshwater; Loreto (Serrano-Martínez et al., 2015). Family Haploporidae Nicoll, 1914 Dicrogaster sp. Family Gorgoderidae Looss, 1899 Mugil cephalus; adult; intestine; Anaporrhutum albidum Brandes in freshwater; La Libertad (Tantaleán et al., Ofenheim, 1900 1985a; Jara, 1997). Myliobatis peruvianus; adult; intestine, Lecithobotrioides elongatus Thatcher, 1999 visceral cavity; marine; Ancash, Lima Prochilodus nigricans; adult; intestine; (Escalante et al., 1983; Tantaleán & freshwater; Loreto (Mathews et al., Rodriguez, 1987; Tantaleán & Huiza, 2013b).

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Metamegasolena sp. Derogenes varicus (Müller, 1784) Looss, Scartichthys gigas; adult; intestine; 1901 marine; Lima (Tarmeño et al., 2015). Merluccius gayi peruanus; adult; Saccocoelium sp. intestine; marine; Lima (Chero et al., Mugil cephalus; adult; intestine; 2014b). freshwater; La Libertad (Tantaleán et al., Genarchella overstreeti (Brooks, Mayes & 1985a). Thorson, 1979) Lunaschi, 1989 Saccocoeloides sp. Hoplias malabaricus, adult; intestine; Mugil cephalus; adult; intestine; freshwater; Loreto, Madre de Dios freshwater; La Libertad (Tantaleán et al., (Iannacone & Luque, 1993b). 1985a). Family Didymozoidae Monticelli, 1888 Family Haplosplanchnidae Poche, 1926 Didymozoidae gen. sp. Hymenocotta manteri Overstreet, 1969 Cilus gilberti; larvae; intestinr; marine; Mugil cephalus; adult; intestine; marine; Lima (Chero et al., 2014c). Ancash, Lima (González & Sarmiento, Didimozoidea gen. sp. 1986; Luque et al., 1991; Tantaleán & Scomber japonicus; adult; mesentery; Huiza, 1994). marine; La Libertad (Cruces et al., 2014). Brasicystis bennetti Thatcher, 1979 Family Accacoeliidae Odhner, 1911 Plagioscion squamosissimus; adult; Accacladocoelium alveolatum Robinson, intestine; freshwater; Loreto (Iannacone 1934 & Luque, 1993b). Mola mola; adult; intestine; marine; Lima Didymocystis sp. (Tantaleán & Huiza, 1994). Scomber japonicus; adult; unspecified Accacoelium sp. site of infection; marine; Lima (Oliva et Brama japonica; adult; operculum; al., 2008). marine; Lima (Iannacone & Alvariño, Didymoproblema sp. 2013). Katsuwonus pelamis; adult; gills; marine; North Zone (Tantaleán & Huiza, 1994). Family Bathycotylidae Dollfus, 1932 Didymozoon sp. Bathycotyle branchialis Darr, 1902 Brama japonica; Scomberomorus sierra; Coryphaena hippurus; adult; intestine; adult; intestine, operculum; marine; Lima, marine; Lima (López & Tantaleán, 1985; Northern coastal zone (Tantaleán & Luque et al., 1991; Tantaleán & Huiza, Huiza, 1994; Iannacone & Alvariño, 1994). 2013). Koellikeria filicollis (Rudolphi, 1819) Family Derogenidae Nicoll, 1910 Brama japonica; adult; general cavidad; Gonocercella pacifica Manter, 1940 marine; Lima (Iannacone & Alvariño, Trachinotus paitensis; adult; intestine; 2013). marine; Lima (Luque & Oliva, 1993b). Koellikeria sp. Gonocercella sp. Scomber japonicus; adult; mesentery; Anisotremus scapularis; adult; intestine; marine; La Libertad (Cruces et al., 2014). marine; Zona Sur (Moquegua-Tacna) Koellikerioides sp. (Ruelas & Córdova, 1995). Sarda chiliensis; adult; intestine; marine; Gonocerca sp. Arequipa (Tantaleán & Huiza, 1994). Trachinotus paitensis; adult; intestine; Maccallumtrema sp. marine, La Libertad (Jara, 1998). Scomber japonicus; adult; mesentery;

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marine; La Libertad (Cruces et al., 2014). Gymnothorax porphyreus; adult; Metanematobothrium sp. intestine; marine; Lima (Luque & Oliva, Sarda chiliensis; adult; gills; marine; 1993b). Arequipa (Tantaleán & Huiza, 1994). Lecithochirium genypteri Manter, 1954 Nematobothrium scombri (Taschenberg, Merluccius gayi peruanus; adult; 1879) intestine; marine; Lima (Chero et al., Scomber japonicus; adult; gills, ovary, 2014b). internal face of operculum; marine; La Lecithochirium macrorchis (Crowcroft, Libertad, Lima (Tantaleán & Huiza, 1994; 1946) Cruces et al., 2014). Labrisomus philippii; adult; gonads; Nematobothriinae gen. sp. intestine; stomach; marine; Lima (Cruces Scomber japonicus; adult; gonads; et al., 2015). marine; Lambayeque (Tantaleán & Huiza, Lecithochirium muraenae Manter, 1940 1994). Gymnothorax porphyreus; adult; Ovarionematobothrium saba Kamegai & stomach; marine; Lima (Tantaleán & Shimazu, 1982 Huiza, 1994). Scomber japonicus; adult; gonads; Lecithochirium sp. marine; La Libertad, Lima (Oliva et al., Centropomus nigrescens, Labrisomus 2008; Cruces et al., 2014). philippii; adult; digestive tract, intestine, Unitubulotestis pelamydis (Taschenburg, stomach; marine; Arequipa, Ancash, 1879) Lima, Moquegua, Tacna (Rivera & Sarda chiliensis; adult; gills; marine; Sarmiento, 1990a; Luque et al., 1991; Lima (Chero et al., 2015b) Tantaleán & Huiza, 1994). Unitubulotestis sp. Lecithocladium cristatum (Rudolphi, 1819) Sarda chiliensis, Scomber japonicus; Looss, 1907 adult; internal face of operculum; marine; Peprilus snyderi; Seriolella violácea, Northern coastal zone (Tantaleán & Stromateus stellatus; adult; intestine; Huiza, 1994). marine; Lima (Luque & Oliva, 1993b; Iannacone et al., 2010; Ñacari & Sánchez, Family Hemiuridae Looss, 1899 2014). Dinurus longisinus Looss, 1907 Lecithocladium sp. Coryphaena hippurus; adult; stomach; Centropomus nigrescens, Peprilus marine; Lima, Northen Coastal zone medius, Seriolella violácea, Trachinotus (López & Tantaleán, 1985; Tantaleán & paitensis; adult; intestine; marine; Huiza, 1994; Vásquez-Ruiz et al., 2012). Arequipa, Ancash, La Libertad, Lima, Dinurus sp. Moquegua, Tacna (Tantaleán & Huiza, Coryphaena hippurus, Trachinotus 1994; Jara, 1998; Iannacone & Alvariño, paitensis; adult; digestive tract, stomach; 2008). m a r i n e ; A r e q u i p a , L a L i b e r t a d , Myosaccium sp. Moquegua, Tacna (Tantaleán & Huiza, Alphestes afer; adult; intestine; marine; La 1994; Jara, 1998). Libertad (Tantaleán & Huiza, 1994). Ectenurus sp. Parahemiurus sp. Alphestes afer, Trachinotus rhodopus; Centropomus nigrescens, Trachinotus adult; intestine, stomach; marine; La rhodopus; adult; intestine, stomach; Libertad (Tantaleán & Huiza, 1994). marine; Ancash, La Libertad (Tantaleán & Lecithochirium callaoensis Luque & Oliva, Huiza, 1994). 1993

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Family Lecithasteridae Odhner, 1905 marine; Lima (Luque & Oliva, 1993b). Aponurus sp. Diphtherostomum sp. Alphestes afer; adult; intestine; marine; La Anisotremus scapularis; adult; intestine; Libertad (Tantaleán & Huiza, 1994). marine; Lambayeque, Lima, Piura Lecithasteridae gen. sp. (Tantaleán et al., 1992; Tantaleán & Sicyases sanguineus; adult; intestine; Huiza, 1994). marine; Lima (Iannacone & Alvariño, Zoogonus dextrocirrus Aldrich, 1961 2011). Labrisomus philippii, Scartichthys gigas; adult; stomach; marine; Lima (Cruces et Family Syncoeliidae Looss, 1899 al., 2015; Tarmeño et al., 2015). Copiatestes filiferus (Leuckart, 1885) Zoogonus pagrosomi Yamaguti, 1939 Hemilutjanus macrophthalmos; adult; Merluccius gayi peruanus; adult; intestine; marine; Lima (Tantaleán & intestine; marine; Lima (Rivera, 1992; Lefevre, 2004). Tantaleán & Huiza, 1994). Zoogonus rubellus (Olsson, 1868) Odhner, Family Apocreadiidae Skrjabin, 1942 1902 Myzotus sp. Labrisomus philippii, Merluccius gayi Caulolatilus affinis, Coryphaena peruanus, Scartichthys gigas; adult; hippurus; adult; intestine; marine; intestine; marine; Lima (Durán & Oliva, Northern coastal zone (Escalante et al., 1980; Luque & Oliva, 1993b; Oliva & 1984a; Tantaleán & Huiza, 1994; Luque, 2002; Oliva & Ballón, 2002; Vásquez-Ruiz et al., 2012). Iannacone & Alvariño, 2003; Iannacone et al., 2011b; Iannacone et al., 2012b). Family Lepocreadiidae Odhner, 1905 Zoogonus sp. Lepocreadium trachinoti Wang, 1987 Merluccius gayi peruanus, Scartichthys Scartichthys gigas; adult; intestine; gigas; adult; intestine; marine; La marine; Lima (Tarmeño et al., 2015). Libertad, Lima (Durán & Oliva, 1980; Lepidapedon sp. Escalante et al., 1984a; Tantaleán & Peprilus snyderi; adult; intestine; marine; Huiza, 1994). Lima (Ñacari & Sánchez, 2014). Opechona bacillaris (Molin, 1859) Dolfus, Family Monorchiidae Odhner, 1911 1927 Lasiotocus glebulentus Overstreet, 1971 Peprilus medius; adult; unspecified site of Mugil cephalus; adult; intestine; marine; infection; marine; Lima (Iannacone & Ancash, Lima (Luque et al., 1991; Luque Alvariño, 2008). & Oliva, 1993b; Tantaleán & Huiza, Opechona pharyngodactyla Manter, 1940 1994). Peprilus snyderi; adult; intestine; marine; Lasiotocus longicaecum (Manter, 1940) Lima (Ñacari & Sánchez, 2014). Yamaguti, 1954 Prodistomum orientalis (Layman, 1930) Anisotremus scapularis; adult; intestine; Bray & Gibson, 1990 marine; Lima (Luque & Oliva, 1993b; Scomber japonicus; adult; caecum; Paschoal et al., 2015). marine; La Libertad (Cruces et al., 2014). Lasiotocus sp. Anisotremus scapularis; adult; intestine; Family Zoogonidae Odhner, 1902 marine; Lima (Chero et al., 2014d). Diphtherostomum brusinae (Stossich, 1888) Monorchis latus Manter, 1942 Stossich, 1903 Anisotremus scapularis; adult; intestine; Isacia conceptionis; adult; intestine; marine; Lima (Luque & Oliva, 1993b;

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Paschoal et al., 2015). Genypterus sp.; adult; gill arteries; marine; Lima (Tantaleán & Martinez, Family Heterophyidae Leiper, 1909 1990; Tantaleán & Huiza, 1994). Ascocotyle longa Ransom, 1920 Aporocotyle wilhelmi Villalba & Fernández, Mugil cephalus; larvae; mesentery; 1986 freshwater; La Libertad, Lima (Armas, Merluccius gayi peruanus; adult; gills; 1979; Jara & Escalante, 1982; Mateo et marine; Lima (Martínez et al., 2013; al., 1985; Luque et al., 1991). Chero et al., 2014b). Centrocestus sp. Aporocotyle sp. Aequidens rivulatus, Xiphophorus Merluccius gayi peruanus; adult; gill hellerii; larvae; gills; freshwater; La arteries, heart; marine; Lima (Durán & Libertad (Pais, 1996; Jara, 1997; Jara et Oliva, 1980; Luque et al., 1991; Tantaleán al., 1998). & Huiza, 1994). Heterophyes sp. Cladocaecum tomasscholzi Orélis-Ribeiro & Mugil cephalus; larvae; muscles; Bullard, 2016 freshwater; La Libertad (Armas, 1979). Ageneiosus inermis; adult; heart; Pygidiopsis sp. freshwater; Loreto (Orélis-Ribeiro & Aequidens rivulatus, Bryconamericus Bullard, 2016). peruanus, Trichomycterus dispar; larvae; Kritskya platyrhynchi (Guidelli, Isaac & liver, mesentery; freshwater; La Libertad Pavanelli, 2002) Orélis-Ribeiro & (Jara & Esacante, 1986; Jara et al., 1990; Bullard, 2016 Jara, 1997). Hemisorubim platyrhynchos; adult; body cavity; freshwater; Loreto (Orélis-Ribeiro Family Cladorchiidae Fischoeder, 1901 & Bullard, 2016). Dadaytrema sp. Plehniella sabajperezi Orélis-Ribeiro & Piaractus brachypomus; adult; intestine; Bullard, 2015 freshwater; Loreto (Tantaleán et al., Pimelodus blochii; adult; body cavity; 1985a). freshwater; Loreto (Orélis-Ribeiro & Pseudocladorchis ferrumequinum (Diesing, Bullard, 2015). 1836) Doradidae gen. sp.; adult; intestine; Family Cyathocotylidae Mühling, 1898 freshwater; Ucayali (Tantaleán et al., Mesostephanus sp. 1974). Mugil cephalus; larvae; muscles; freshwater; La Libertad (Armas, 1979). Order Olson, Cribb, Tkach, Bray & Littlewood, 2003 Family Diplostomidae Poirier, 1886 Diplostomulum sp. Family Callodistomidae Odhner, 1910 Odontesthes bonariensis, Orestias albus, Prosthenhystera obesa (Diesing, 1850) O. luteus, Orestias sp.; larvae; brain; Travassos, 1922 freshwater; Puno (Tantaleán et al., 1974, Hoplias malabaricus; adult; Gall bladder; 1975; Hurtado et al., 1977). freshwater; Loreto (Olson et al., 2003 ; Neascus sp. Tkach & Curran, 2015). Bryconamericus peruanus; larvae; fins, skin; freshwater; Lambayeque (Tantaleán Family Aporocotylidae Odhner, 1912 et al., 1985a). Aporocotyle garciai Tantaleán & Martínez, 1990 Family Lühe, 1901

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Clinostomum marginatum (Rudolphi, 1819) antofagastensis, Gonocercella sp., Braun, 1899 Gorgorhynchus sp., Intracotyle neghmei, Tr i c h o m y c t e r u s p i u r a e ; l a r v a e ; Lacistorhynchus tenuis, Lasiotocus mesentery; freshwater; Ica, La Libertad l o n g i c a e c u m , L a s i o t o c u s s p . , (Tantaleán et al., 1974). L er nanthropus antofa g a s ten s is , Clinostomum sp. L o b a t o s t o m a a n i s o t r e m u m , Aequidens rivulatus; larvae; operculum; Metapeniculus anofagastensis, Mexicana freshwater; La Libertad (Jara, 1997; Jara sp., Monorchis latus, Neobivagina chita, et al., 1998). Parabrachiella anisotremi, Philometra sp., Proctoeces sp., Proctoeces lintoni, HOST-PARASITE LIST Profilicollis altmani, Proleptus carvajali. (ray-finned fishes) Aplodactylus punctatus Valenciennes, 1832 Ablennes hians (Valenciennes, 1846) (Aplodactylidae): Encotyllabe sp. (Belonidae): Bomolochus constrictus, A r a p a i m a g i g a s ( S c h i n z , 1 8 2 2 ) Caligodes laciniatus, Lernanthropus (Arapaimatidae): Caballerotrema tylosuri. arapaimense, C. brasiliense, Dawestrema Acanthistius pictus (Tschudi, 1846) cycloancistrioides, D. cycloancistrum, (Serranidae): Helicometrina sp. Goezia spinulosa, Nesolecithus janicki, Aequidens rivulatus (Günther, 1860) Nilonema senticosum, Schizochoerus ( C i c h l i d a e ) : U r o c l e i d u s s p . , liguloideus, Tetraonchus monenteron. . Procamallanus (Spirocamallanus) sp., Ariopsis seemanni (Günther, 1864) (Ariidae): Centrocestus sp., Clinostomum sp., Adenocephalus pacificus. Contracaecum sp., Pygidiopsis sp., Aspistogramma sp. (Cichlidae): Dactylogyrus Sphaerostoma sp. sp. Aequidens sp. (Cichlidae): Pandosentis Astronotus ocellatus (Agassiz, 1831) iracundus. (Cichlidae): Contracaecum sp., Gussevia Ageneiosus inermis (Linnaeus, 1766) asota, Proteocephalus gibsoni. (Auchenipteridae): Ageneiella sp., Auxis thazard thazard (Lacepède, 1800) Cladocaecum tomasscholzi, Gibsoniela (Scombridae): Anilocra huacho. mandube. Brachyplatystoma cf. filamentosum Ageneiosus sp. (Auchenipteridae): Gibsoniela (Lichtenstein, 1819) (Pimelodidae): mandube. Amphoteromorphus ovalis, Endorchis Ageneiosus vittatus Steindachner, 1908 piraeeba, Nomimoscolex suspectus. (Auchenipteridae): Dactylogyridae gen. Brachyplatystoma juruense (Boulenger, 1898) sp. (Pimelodidae): Alinema amazonicum, Aguarunichthys torosus Stewart, 1986 Demidospermus mortenthaleri. ( P i m e l o d i d a e ) : U n i b a r r a Brachyplatystoma rousseauxii (Castelnau, paranoplatensis. 1855) (Pimelodidae): Amphoteromorphus Alphestes afer (Bloch, 1793) (Serranidae): peniculus, A. piriformis. Aponurus sp., Ectenurus sp., Myosaccium Brachyplatystoma vaillantii (Valenciennes, sp., Prosorhynchus sp. 1840) (Pimelodidae): Chambriella sp., Anisotremus scapularis (Tschudi, 1846) Demidospermus sp., Harriscolex (Haemulidae): Adenocephalus pacificus, piramutab. Choricotyle anisotremi, Ch. scapularis, Brama japonica Hilgendorf, 1878 (Bramidae): Clavella applicata, Colobomatus Accacoelium sp., Corynosoma sp., q u a d r i f a r i u s , C y m o t h o a s p . , Contracaecum sp., Didymozoon sp., Diphtherostomum sp., Encotyllabe Hatschekia conífera, Hepatoxylon

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trichiuri, Koellikeria filicollis, Nybelinia Microcotyle nemadactylus. sp. Cheilotrema fasciatum Tschudi, 1846 B r y c o n a t ro c a u d a t u s ( K n e r, 1 8 6 3 ) (Sciaenidae): Caligus callaoensis, ( C h a r a c i d a e ) : B u n o d e r i n a s p . , Camallanus sp., Cynoscionicola Contracaecum sp., americanus, Diplectanidae gen. sp. 1, Paracamallanus sp. Diplectanidae gen. sp. 2, Hargicotyle B r y c o n c e p h a l u s ( G ü n t h e r, 1 8 6 9 ) magna, Helicometra fasciata, Kudoa (Characidae): Amplexibranchius sciaenae, Lacistorhynchus dollfusi, bryconis, Capillaria sp., Jainus Lacistorhynchus tenuis, Parabrachiella amazonensis fasciata. Bryconamericus peruanus (Müller & Cherublemma emmelas (Gilbert, 1890) Tr o s c h e l , 1 8 4 5 ) ( C h a r a c i d a e ) : ( O p h i d i i d a e ) : C h o n d r a c a n t h u s Anacanthocotyle sp., Procamallanus lepophidii. (Spirocamallanus) sp., Neascus sp., Chloroscombrus orqueta Jordan & Gilbert, Pygidiopsis sp. 1883 (Carangidae): Zeuxapta seriolae Bryconops caudomaculatus (Günther, 1864) Cichla monoculus Agassiz, 1831 (Cichlidae): (Characidae): Auriculostoma foliaceum. Gussevia longihaptor, G. undulata, Calophysus macropterus (Lichtenstein, 1819) Proteocephalus macrophallus, P. (Pimelodidae): Alinema amazonicum, m i c ro s c o p i c u s , S c i a d o c e p h a l u s Ameloblastella unapi, Demidospermus megalodiscus, Tucunarella cichlae. macropteri, Monticellia amazonica, Cichlasoma amazonarum Kullander, 1983 Rudolphiella piracatinga. (Cichlidae): Dolops geayi, Gussevia Carassius auratus (Linnaeus, 1758) disparoides, Proteocephalidea gen. sp., (Cyprinidae): Dactylogyrus sp., Sciadicleithrum variabilum. Gyrodactylus sp., Thellohanellus sp., Cichlasoma sp. (Cichlidae): Dactylogyrus sp. Zschokkella sp. Cilus gilberti (Abbott, 1899) (Sciaenidae): C a r a n x h i p p o s ( L i n n a e u s , 1 7 6 6 ) Adenocephalus pacificus, Anisakis sp., (Carangidae): Allopyragraphorus Callitetrarhynchus gracilis, Choricotyle caballeroi, Neomicrocotyle sp. s p . , C o r y n o s o m a o b t u s c e n s , C a u l o l a t i l u s a f f i n i s G i l l , 1 8 6 5 Cynoscionicola americanus, Dichelyne (Malacanthidae): Myzotus sp. ( C u c u l l a n e l l u s ) e l o n g a t u s , Caulolatilus sp. (Malacanthidae): Choricotyle Didymozoidae gen sp., Hargicotyle caulolatili, Jaliscia caballeroi. louisianensis, Helicometra fasciata, Centromochlus heckelii (De Filippi, 1853) Lernanthropus pacificus, Parabrachiella (Auchenipteridae): Demidospermus sp., Philometridae gen sp., Piscicolidae centromochli. g e n . s p . , P l a g i o p o r u s s p . , Centropomus nigrescens Günther, 1864 Poecilancistrum caryophyllum, Proleptus (Centropomidae): Floriceps saccatus, c a r v a j a l i , R a p h i d a s c a r i s s p . , Lecithochirium sp., Lecithocladium sp., Rhipidocotyle sp., Villarrealina peruana. Parahemiurus sp., Tentacularia Colossoma macropomum (Cuvier, 1816) coryphaenae. (Characidae): Anacanthorus spathulatus, Chaetobranchus semifasciatus Steindachner, Dactylogyridae gen. sp., Gyrodactylidae 1875 (Cichlidae): Gussevia tucunarense. g e n . s p . , L e r n a e a g a m i t a n a e , Cheilodactylus variegatus Valenciennes, 1833 M y x o b o l i d a e g e n . s p . , (Cheilodactylidae): Caligus cheilodactyli, Neoechinorhynchus sp. Clavellotis dilatata, Lacistorhynchus Coryphaena hippurus Linnaeus, 1758 tenuis, Macrophyllida antarctica, (Coryphaenidae): Adenocephalus

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pacificus, Anisakis sp., Anisakis physeteris, Lacistorhynchus tenuis, Proleptus sp. Bathycotyle branchialis, Caligus Galeichthys sp. (Ariidae): Lepeophtheirus c o r y p h a e n a e , C a l i g u s s p . , marginatus. Chondracanthus sp., Contracaecum sp., Genypterus maculatus (Tschudi, 1846) Corynosoma obtuscens, Dinurus sp., (Ophidiidae): Adenocephalus pacificus, Dinurus longisinus, Hysterothylacium “Scolex pleuronectis”. sp., Hysterothylacium aduncum, H. Genypterus sp. (Ophidiidae): Aporocotyle pelagicum, Myzotus sp., Nybelinia sp., g a r c i a i , N e o b e n e d e n i a s p . Pterobothrium acanthotruncatum, Pseudohaliotrema sp. “Scolex pleuronectis”, Tentacularia Geophagus steindachneri Eigenmann & coryphaenae. H i l d e b r a n d , 1 9 2 2 ( C i c h l i d a e ) : Crenicichla anthurus Cope, 1872 (Cichlidae): Cosmoxynemoides aguirrei, Ichthyouris Myxosporea fam. gen. sp. ro, Pandosentis iracundus. Cubiceps caeruleus Regan, 1914 (Nomeidae): Goeldiella eques (Müller & Troschel, 1849) Anilocra huacho. (Heptapteridae): Aphanoblastella Cynoscion analis (Jenyns, 1842) (Sciaenidae): aurorae, Aphanoblastella sp., Nupelia sp. Adenocephalus pacificus, Anisakis sp., Gymnothorax porphyreus (Guichenot, 1848) Ceratothoa gaudichaudii, Corynosoma (Muraenidae): Helicometra fasciata, o b t u s c e n s , C o r y n o s o m a s p . , Lecithochirium callaoensis, L. muraenae. Cynoscionicola cynoscioni, Diplectanum Haemulon steindachneri (Jordan & Gilbert, sp., Hargicotyle sp., Hargicotyle 1 8 8 2 ) ( H a e m u l i d a e ) : M e x i c a n a paralonchuri, Lernanthropus Huamani, iannaconi. Lernanthropus sp., L. paralonchuri, Hassar orestis (Steindachner, 1875) Neoheterobothrium cynoscioni, Nybelinia (Doradidae): Cosmetocleithrum sp. sp., Otobothrium sp., Procamallanus sp., Hassar sp. (Doradidae): Ameloblastella sp. Tegorhynchus sp. Hemanthias peruanus (Steindachner, 1875) Cyprinus carpio carpio Linnaeus, 1758 (Serranidae): Blias prionoti. (Cyprinidae): Dactylogyrus vastator Hemilutjanus macrophthalmos (Tschudi, Duopalatinus cf. peruanus Eigenmann & 1846) (Serranidae): Callitetrarhynchus A l l e n , 1 9 4 2 ( P i m e l o d i d a e ) : g r a c i l i s , C o p i a t e s t e s f i l i f e r u s , Sharpilosentis peruviensis. Glossobothrium sp. Gorgorhynchus sp. Doradidae gen. sp.: Pseudocladorchis Hemisorubim platyrhynchos (Valenciennes, ferrumequinum. 1840) (Pimelodidae): Chambriella Ethmidium maculatum (Valenciennes, 1847) paranaensis, Kritsky platyrhynchi, (Clupeidae): Contracaecum sp., “Scolex Mariauxiella piscatorum, Spatulifer pleuronectis”. maringaensis, Vancleaveus platyrhynchi. Exocoetus volitans Linnaeus, 1758 Heros severus Heckel, 1840 (Cichlidae): (Exocoetidae): Axine ibanezi. Gussevia alioides, G. disparoides. Farlowella amazonum (Günther, 1864) Hippoglossina bollmani Gilbert, 1890 ( L o r i c a r i i d a e ) : O ö g y ro d a c t y l u s (Paralichthyidae): Protochondria farlowellae. longicauda. Galeichthys peruvianus Lütken, 1874 Hoplias malabaricus (Bloch, 1794) (Ariidae): Adenocephalus pacificus, (Erythrinidae): Genarchella overstreeti, A n i s a k i s s p . , A s c a r o p h i s s p . , Prosthenhystera obesa, Urocleidoides Callitetrarhynchus gracilis, Corynosoma eremitus. obtuscens, Hamaticolax galeichthyos, Hypophthalmus edentatus Spix & Agassiz, H a m a t o p e d u n c u l a r i a s p . , 1829 (Pimelodidae): Ameloblastella sp.

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Dactylogyridae gen. sp. ( J e n y n s , 1 8 4 0 ) ( S c i a e n i d a e ) : Isacia conceptionis (Cuvier, 1830) Acanthocolax peruensis, Adenocephalus (Haemulidae): Benedenia sp., Caligus pacificus. Ascarophis sp., Ceratomyxa sp., Choricotyle isaciencis, Ch. sp., Ceratothoa sp., Clavellotis dilatata, s o n o re n s i s , C l a v e l l a s i m p l e x , Cynoscionicola americanus, Hargicotyle C o r y n o s o m a o b t u s c e n s , menticirrhi, Helicometra fasciata, Diphtherostomum brusinae, Nybelinia Lernanthropus huamani, Lobatostoma sp., Parabrachiella kabatai, Proctoeces veranoi, Parabrachiella menticirrhi, sp., Proctoeces lintoni, Proleptus Proleptus sp., Rhamnocercoides carvajali, Pseudoeurysorchis travassosi. menticirrhi. Katsuwonus pelamis (Linnaeus, 1758) Merluccius gayi peruanus Ginsburg, 1954 (Scombridae): Caligus coryphaenae, (Merluccidae): Adenocephalus pacificus, Caligus productus, Didymoproblema sp., Anisakis sp., Anisakis simplex, Prosorhynchus sp., “Scolex pleuronectis”, Anthocotyle americana, A. merluccii, Tentacularia coryphaenae. Aporocotyle sp., Aporocotyle wilhelmi, Labrisomus philippii (Steindachner, 1866) Argulus peruvianus, Bolbosoma sp., (Labrisomidae): Acanthochondria Brachiella sp., Caligus debueni, C. teres, sicyasis, Bucephalus sp., Corynosoma Ceratomyxa limensis, Corynosoma australe , C. obtuscens , Helicometra obtuscens , Callitetrarhynchus gracilis, fasciata, Lacistorhynchus tenuis, Chondracanthus sp., Clestobothrium L e c i t h o c h i r i u m m a c r o r c h i s , c r a s s i c e p s , C o n t r a c a e c u m s p . , Lecithochirium sp., Lernaea sp., Diphyllobothridae gen. sp., Derogenes Myxosporea gen. sp., Piscicolidae gen. varicus, Grillotia dollfusi, Kudoa sp., Profilicollis altmani, Proleptus peruviana, Lacistorhynchus tenuis, c a r v a j a l i , P r o l e p t u s s p . , L e c i t h o c h i r i u m g e n y p t e r i , P r o s o r h y n c h o i d e s c a r v a j a l i , Lepeophtheirus sp., Myxosporea gen. sp., Prosorhynchoides sp., P. rioplatensis, Nybelinia sp., Parabrachiella insidiosa, Prosorhynchus sp., Tegorhynchus sp., Prosorhynchus sp., Tentacularia Zoogonus dextrocirrus, Z. rubellus. coryphaenae, Zoogonus sp., Zoogonus Lebiasina bimaculata Valenciennes, 1847 pagrosomi, Z. rubellus. (Lebiasinidae): Accessorius peruensis, M i c r o p o g o n i a s s p . ( S c i a e n i d a e ) : Gyrodactylus bimaculatus, G. lebiasinus, Pterinotrematoides mexicanum. Gyrodactylus slendrus, Gyrodactylus sp., Mola mola (Linnaeus, 1758) (Molidae): Oncocleidus sp., Procamallanus A c c a c l a d o c o e l i u m a l v e o l a t u m , (Spirocamallanus) sp. Ancistrocephalus microcephalus, Lepophidium sp. (Ophidiidae): Helicometra Fistulicola plicatus. fasciata Mugil cephalus Linnaeus, 1758 (Mugilidae): Makaira nigricans Lacepède, 1802 Ascocotyle longa, (Istiophoridae): Gloiopotes watsoni. Bomolochus nitidus, Contracaecum sp., Makaira sp. (Istiophoridae): Gloiopotes Contracaecum multipapillatum, huttoni Dicrogaster sp., Ergasilus sp., Haliotrema Megalodoras uranoscopus (Eigenmann & mugilinus, Heterophyes sp., Hymenocotta Eigenmann, 1888) (Doradidae): manteri, Lacistorhynchus tenuis, Proteocephalus kuyukuyu Lasiotocus glebulentus, Lernanthropus Megalonema platycephalum Eigenmann, p a c i f i c u s , M e s o s t e p h a n u s s p . , 1912 (Pimelodidae): Monticellia M e t a m i c ro c o t y l a m a c r a c a n t h a , santafesina.Menticirrhus ophicephalus Myxobolus exiguus, Naobranchia lizae,

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Neobenedenia sp., Parabrachiella exilis, Diplostomulum sp., Ligula intestinalis, Saccocoelium sp., Saccocoeloides sp. Polymorphus spindlatus. M u g i l c u re m a Va l e n c i e n n e s , 1 8 3 6 Orthopristis chalceus (Günther, 1864) (Mugilidae): Contracaecum sp. (Haemulidae): Calceostoma sp., Myleus schomburgkii (Jardine, 1841) Notoporus sp. (Serrasalmidae): Anacanthorus sp., Osteoglossum bicirrhosum (Cuvier, 1829) Notozothecium bethae, Notozothecium sp. ( O s t e o g l o s s i d a e ) : C a m a l l a n u s Nezumia stelgidolepis (Gilbert, 1890) (Camallanus) acaudatus, Gonocleithrum (Macrouridae): Lophoura unilobulata. cursitans. Odontesthes bonariensis Valenciennes Oxydoras niger (Valenciennes, 1821) (Atherinopsidae): Diplostomulum sp., (Doradidae): Cosmetocleithrum confusus, Hedruris basilichtensis, Ligula C. gussevi, C. sobrinus, Cucullanus intestinalis, Perunema escomeli. ( C u c u l l a n u s ) g r a n d i s t o m i s , Odontesthes regia (Humboldt, 1821) Paracavisoma impudica, Proteocephalus ( A t h e r i n o p s i d a e ) : C o r y n o s o m a h o b e rg i , R a p h i d a c a ro i d e s s p . , obtuscens, Corynosoma sp., Kudoa Sharpilosentis peruviensis. sarmientae, Kudoa sp., Lacistorhynchus Paralabrax humeralis (Valenciennes, 1828) tenuis, Prosorhynchoides sp., “Scolex (Serranidae): Adenocephalus pacificus, pleuronectis”. Caligus quadratus, Callitetrarhynchus Ophidiidae gen. sp.: Neobenedenia sp. gracilis, Corynosoma obtuscens, Oreochromis niloticus (Linnaeus, 1758) C o r y n o s o m a s p . , D i c h e l y n e (Cichlidae): Contracaecum sp., (Cucullanellus) sp., Grillotia sp., Polymorphus spindlatus. Hamaticolax paralabracis, Hatschekia Oreochromis spp. (Cichlidae): Gyrodactylus amphiprocessa, Helicometra fasciata, sp. H e l i c o m e t r i n a s p . , H e m i t a g i a Orestias agassizii Valenciennes, 1846 g a l a p a g e n s i s , P h i l o m e t r a s p . , (Cyprinodontidae): Polymorphus Trypanorhyncha gen. sp. spindlatus. Paralichthys adspersus (Steindachner, 1867) Orestias albus Valenciennes, 1846 (Paralichthyidae): Adenocephalus (Cyprinodontidae): Contracaecum sp., pacificus, Anisakis sp., Bomolochus sp., D i p l o s t o m u l u m s p . , H e d r u r i s Cainocreadium sp., Contracaecum sp., basilichtensis, Ligula intestinalis, Corynosoma obtuscens, Corynosoma sp., Proteocephalus sp. Hysterothylacium sp., Lacistorhynchus Orestias luteus Valenciennes, 1846 tenuis, Lepeophtheirus edwardsi, (Cyprinodontidae): Contracaecum sp., P a r a b r a c h i e l l a p a r a l i c h t h y o s , D i p l o s t o m u l u m s p . , F a l c a u s t r a Philometra sp., Prosorhynchoides sp. pumacahuai, F. tiahuanaquensis, Paralichthys woolmani Jordan & Williams, Hedruris basilichtensis, H. orestiae, 1897 (Paralichthyidae): Anisoporus sp. Ligula intestinalis, Polymorphus Paralonchurus peruanus (Steindachner, 1875) spindlatus, Proteocephalus sp. (Sciaenidae): Acanthocolax peruensis, Orestias mulleri Valenciennes, 1846 Adenocephalus pacificus, Anisakis sp., (Cyprinodontidae): Hedruris orestiae, Caligus callaoensis, Callitetrarhynchus Polymorphus spindlatus. gracilis, Ceratothoa sp., Clavellopsis O re s t i a s o l i v a c e u s G a r m a n , 1 8 9 5 sp., Corynosoma australe, C. obtuscens, (Cyprinodontidae): Polymorphus Corynosoma sp. , Cynoscionicola sp., spindlatus Cynoscionicola americanus, Dichelyne O r e s t i a s s p . ( C y p r i n o d o n t i d a e ) : ( C u c u l l a n e l l u s ) a m a r u i n c a i ,

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Diplectanum sp., Diplectanidae gen. sp., Pimelodus altissimus Eigenmann & Pearson, Encotyllabe callaoensis, Euryhaliotrema 1942 (Pimelodidae): Endorchis sp. paralonchuri, Hargicotyle chimbotensis, Pimelodus blochii Valenciennes, 1840 H. paralonchuri, H. peruensis, (Pimelodidae): Demidospermus lebedev, Helicometra fasciata, Henneguya sp., D. peruvianus, D. striatus, Plehniella Kudoa sciaenae, Lacistorhynchus tenuis, sabajperezi, Proteocephalus sp. Lernanthropus paralonchuri, Nybelinia Pimelodus ornatus Kner, 1858 (Pimelodidae): sp. , Parabrachiella chevreuxii, D e m i d o s p e r m u s p e r u v i a n u s , Piscicolidae gen. sp., Pterobothrium Nomimoscolex sp.  acanthotruncatum, Pterobothrium sp., Pimelodus maculatus Lacepède, 1803 Tegorhynchus sp. ( P i m e l o d i d a e ) : P ro c a m a l l a n u s Paranthias colonus (Valenciennes, 1846) (Spirocamallanus) rarus. (Serranidae): Pseudorhabdosynochus Pimelodus sp. (Pimelodidae): Demidospermus jeanloui. brevicirrus, D. curvovaginatus, D. Parapsettus panamensis (Steindachner, 1876) lebedev, D. peruvianus, D. striatus. (Ephippidae): Parancylodiscoides Pinirampus pirinampu (Spix & Agassiz, 1829) chaetodipteri. (Pimelodidae): Monticellia ventrei, P e p r i l u s m e d i u s ( P e t e r s , 1 8 6 9 ) N o m i m o s c o l e x a d m o n t i c e l l i a , (Stromateidae): Bomolochus sp., Caligus Rudolphiella sp. s p . , C e r a t o t h o a g a u d i c h a u d i i , Plagioscion squamosissimus (Heckel, 1840) Lecithocladium sp., Lobatostoma sp., (Sciaenidae): Brasicystis bennetti, Oaxacotyle oaxacensis, Opechona Diplectanum decorum, D. pescadae, D. bacillaris. piscinarius, Euryhaliotrema chaoi, Peprilus snyderi Gilbert & Starks, 1904 E u r y h a l i o t r e m a l o v e j o y i , E . ( S t r o m a t e i d a e ) : A n i s a k i s s p . , m o n a c a n t h u s , E . p o t a m o c e t e s , Hysterothylacium sp., Lecithocladium Euryhaliotrema succedanus, E. thatcheri, cristatum, Lepidapedon sp. Oaxacotyle Rhadinorhynchus plagioscionis. oaxacensis, Opechona pharyngodactyla, Platynematichthys notatus (Jardine, 1841) Rusguniella sp. (Pimelodidae): Brayela karuatayi, Perrunichthys perruno Schultz, 1944 Dactylogyridae gen. sp. (Pimelodidae): Alinema amazonicum. Platysilurus mucosus (Vaillant, 1880) Phractocephalus hemioliopterus (Bloch & (Pimelodidae): Dactylogyridae gen. sp. Schneider, 1801) (Pimelodidae): Poecilia reticulata Peters, 1859 (Poeciliidae): Chambriella sp., Monticelliinae gen. sp., Capillaria sp., Gyrodactylus turnbulli. P ro t e o c e p h a l u s h e m i o l i o p t e r i , Polydactylus approximans (Lay & Proteocephalus sp., Scholzia emarginata, B e n n e t t , 1 8 3 9 ) ( P o l y n e m i d a e ) : Zygobothrium megacephalum. M i c r o c o t y l o i d e s i m p u d i c u s , Piaractus brachypomus (Cuvier, 1818) Procamallanus (Spirocamallanus) sp. ( C h a r a c i d a e ) : D a d a y t re m a s p . , Prionotus stephanophrys Lockington, 1881 Mymarothecium viatorum; Myxobolus sp. (Triglidae): Corynosoma sp., Kudoa Pimelodella yuncensis Steindachner, 1902 estela. (Heptapteridae): Cleidodiscus sp., Prionotus punctatus (Bloch, 1793) (Triglidae): Demidospermus curvovaginatus, D. Blias prionoti. peruvianus, Gyrodactylus pimelodellus, Prochilodus nigricans Spix & Agassiz, 1829 Scleroductus yuncensi. (Prochilodontidae): Lecithobotrioides Pimelodidae gen. sp.: Demidospermus elongatus, Neoechinorhynchus sp., curvovaginatus, D. peruvianus. Rhinonastes pseudocapsaloideum.

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Pseudoplatystoma fasciatum (Linnaeus, 1766) Koellikerioides sp., Metanematobothrium (Pimelodidae): Chambriella sp. sp., Nybelinia sp., Rhipidocotyle Choanoscolex abscisus, Dactylogyridae a d b a c u l u m , R h i p i d o c o t y l e s p . , gen. sp., Harriscolex kaparari, Rhodobothrium mesodesmatum, “Scolex Megathylacus sp. Nomimoscolex lopesi, p l e u ro n e c t i s ” , S p h y r i o c e p h a l u s N. sudobim, Peltidocotyle rugosa, tergestinus, Unitubulotestis sp., Phyllodistomum sp., Vancleaveus Unitubulotestis pelamydis. fungulus. Sardinops sagax (Jenyns, 1842) (Clupeidae): Pseudoplatystoma punctifer (Castelnau, 1855) Anisakis sp., Choricotyle sonorensis, (Pimelodidae): Choanoscolex sp., Contracaecum sp., Scolex pleuronectis.  Houssayella sudobim, Megathylacus sp., Satanoperca jurupari (Heckel, 1840) Spasskyellina spinulifera, Spatulifer ( C i c h l i d a e ) : S c i a d i c l e i t h r u m rugosa. satanopercae. Pseudoplatystoma tigrinum (Valenciennes, Scartichthys gigas (Steindachner, 1876) 1840) (Pimelodidae): Choanoscolex sp., (Blennidae): Acanthochondria sicyasis, Nomimoscolex cf. sudobim, Spasskyellina Ceratothoa gaudichaudii, Corynosoma spinulifera, Spatulifer sp.  obtuscens, Johnstonmawsonia sp., Pterodoras granulosus (Valenciennes, 1821) Lacistorhynchus tenuis, Lepocreadium ( D o r a d i d a e ) : C o s m e t o c l e i t h r u m trachinoti, Metamegasolena sp., bulbocirrus, Monticellia belavistensis, Proleptus sp., Prosorhynchus sp., P r o t e o c e p h a l u s k u y u k u y u , P s e u d o d e l p h i s s p . , Z o o g o n u s Proteocephalus sp., Vancleaveus dextrocirrus, Z. rubellus, Zoogonus sp. janauacaensis. Schedophilus haedrichi Chirichigno F., 1973 Pterodoras sp. (Doradidae): Proteocephalus (Centrolophidae): Teleurysorchis  kuyukuyu. gumercindae Pterophyllum scalare (Schultze, 1823) S c i a e n a d e l i c i o s a ( Ts c h u d i , 1 8 4 6 ) (Cichlidae): Dactylogyrus sp., Gussevia (Sciaenidae): Acanthocolax peruensis, spiralocirra Adenocephalus pacificus, Anisakis sp., Pterygoplichthys anisitsi Eigenmann & Caligus callaoensis, Callitetrarhynchus Kennedy, 1903 (Loricariidae): Unilatus gracilis, Clavellotis sp., Corynosoma brittani, U. unilatus.  austral, C. obtuscens, Cynoscionicola Pterygoplichthys multiradiatus (Hancock, sciaenae, Dasyrhynchus pacificus, 1828) (Loricariidae): Gorytocephalus Dichelyne (Cucullanellus) amaruincai, plecostomorum.  Encotyllabe callaoensis, Hargicotyle P y g o c e n t r u s n a t t e re r i K n e r, 1 8 5 8 louisianensis, H. peruensis, H. sciaenae, (Characidae): Amphithecium calycinum, Hargicotyle sp., Helicometra fasciata, A. camelum, Anacanthorus reginae, A. Kudoa sciaenae, Lacistorhynchus tenuis, stachophallus, A. thatcheri, Notothecium Lernanthropus huamani, L. pacificus, mizellei, N. penetratum, Rhinoxenus Lernanthropus sp., Macrovalvitrema sp., piranhus. Nybelinia sp., Parabrachiella oralis, Sarda chiliensis (Cuvier, 1832) (Scombridae): Piscicolidae gen. sp., Tegorhynchus sp., A d e n o c e p h a l u s p a c i f i c u s , Proleptus sp., Villarrealina peruana. Acanthobothrium chilensis, Anisakis Sciaena callaensis Hildebrand, 1946 physeteris, Caligus bonito, Caligus sp., (Sciaenidae): Adenocephalus pacificus. C a p s a l a g re g a l i s , C e r a t o t h o a Sciaena wieneri Sauvage, 1883 (Sciaenidae): gaudichaudii, Diphyllobothrium sp., Anisakis sp. H y s t e r o t h y l a c i u m a d u n c u m , 

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Sciaenidae gen. sp.: Ceratothoa gaudichaudi. sarmientoi, Philometra neptomeni. Scomber japonicus Houttuyn, 1782 Sicyases sanguineus Müller & Troschel, 1843 (Scombridae): Adenocephalus pacificus, (Gobiesocidae): Acanthochondria Anisakis s i c y a s i s , H e l i c o m e t r i n a n i m i a , sp., Anisakis physeteris, A. simplex, Lecithasteridae gen. sp., Proctoeces C a l i g u s p e l a m y d i s , C e r a t o t h o a lintoni, Proctoeces sp., Prosorhynchoides g a u d i c h a u d i i , C e r a t o t h o a s p . , sp. Callitetrarhynchus gracilis, Clavellisa Sorubim lima (Bloch & Schneider, 1801) s c o m b r i , C o n t r a c a e c u m s p . , (Pimelodidae): Ameloblastella sp., Corynosoma obtuscens, Didymocystis sp., Dactylogyridae gen. sp., Manaosia D i d y m o z o i d a e g e n . s p . , bracodemoca, Spatulifer maringaensis. Diphyllobothridae gen. sp., Koellikeria Sorubimichthys planiceps (Spix & Agassiz, sp., Kuhnia indica, K.scombri, Kuhnia 1829) (Pimelodidae): Chambriella sp., s p r o s t o n a e , K u d o a c a u d a t a , Choanoscolex sp., Houssayella sudobim, Maccallumtrema sp., Nematobothrium L e n h a t a e n i a m e g a c e p h a l a , scombri, Nematobothriinae gen. sp., Nomimoscolex lenha, Peltidocotyle Nybelinia sp., Ovarionematobothrium lenha, Spasskyellina lenha. s a b a , P ro d i s t o m u m o r i e n t a l i s , Stellifer minor (Tschudi, 1846) (Sciaenidae): P s e u d o t e r r a n o v a d e c i p i e n s , Acanthocolax peruensis, Anisakis sp., Rhadinorhynchus pristis, Rocinela sp., Cainocreadium sp., Caligus quadratus, “Scolex pleuronectis”, Tentacularia Clavella sp., Clavellotis dilatata, coryphaenae, Unitubulotestis sp. Corynosoma obtuscens, Corynosoma sp., Scomberomorus sierra Jordan & Starks, 1895 Cymothoidae gen. sp., Cynoscionicola ( S c o m b r i d a e ) : A c a n t h o l o c h u s americanus, Cynoscionicola sp., nudiusculus, Adenocephalus pacificus, Diplectanum sp., Encotyllabe sp., Anaporrhutum sp., Didymozoon sp., Helicometra fasciata, Kudoa sciaenae, Caligus omissus, Cybicola buccata, Lacistorhynchus tenuis, Loimos sp., Diphyllobothrium sp., Mexicotyle Nybelinia sp., Pedocotyle annakohni, P. m e x i c a n a , P s e u d a x i n e s p . , bravoi, Pedocotyle sp., Procamallanus P s e u d o g r i l l o t i a p e r u v i a n a , ( S p i r o c a m a l l a n u s ) p e r e i r a i , Thoracocotyle crocea. Procamallanus (Spirocamallanus) sp., Scorpaena russula Jordan & Bollman, 1890 Rhadinorhynchus sp., Rhamnocercus (Scorpaenidae): Anisakis sp. oliveri, Rhamnocercus sp., R. stelliferi, S e b a s t e s c a p e n s i s ( G m e l i n , 1 7 8 9 ) “Scolex pleuronectis”. (Sebastidae): Ascarophis sebastodis, S t e t h a p r i o n e r y t h ro p s C o p e , 1 8 7 0 Corynosoma australe, Helicometrina (Characidae): Auriculostoma diagonale n i m i a , N e o b e n e d e n i a m e l l e n i , S t ro m a t e u s s t e l l a t u s C u v i e r, 1 8 2 9 Pseudopecoelus sp. ( S t r o m a t e i d a e ) : C e r a t o t h o a Seriola lalandi Valenciennes, 1833 gaudichaudii; Hysterothylacium sp. (Carangidae): Allencotyla mcintoshi. Lecithocladium cristatum Seriolella violacea Guichenot, 1848 Strongylura exilis (Girard, 1854) (Belonidae): ( C e n t r o l o p h i d a e ) : A n i s a k i s s p . , Bomolochus constrictus, B. ensiculus, Corynosoma sp., Lecithocladium Lernanthropus tylosuri, Loxura c r i s t a t u m , L e c i t h o c l a d i u m s p . , peruensis. L e r n a n t h r o p i n u s t r a c h u r i , Strongylura scapularis (Jordan & Gilbert, N e o b o t h r i o c e p h a l u s a s p i n o s u s , 1882) (Belonidae): Ancyrocephalus sp. N y b e l i n i a s p . , P a r a e u r y s o rc h i s Bomolochus bellones, Dollfustrema sp.,

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Loxura peruensis, Tylosuricola amatoi, ( S p i r o c a m a l l a n u s ) h i l a r i i , P . “Scolex pleuronectis”. (Spirocamallanus) chimusensis. Symphysodon aequifasciatus Pellegrin, 1904 Trichomycterus punctulatus Valenciennes, (Cichlidae): Dactylogyrus sp. 1846 (Trichomycteridae): Hepatoxylon Thunnus albacares (Bonnaterre, 1788) megacephalum, Gyrodactylus sp. ( S c o m b r i d a e ) : P s e u d o c y c n u s Triportheus angulatus (Spix & Agassiz, 1829) appendiculatus ( C h a r a c i d a e ) : P r o c a m a l l a n u s Trachinotus paitensis Cuvier, 1832 (Spirocamallanus) inopinatus. (Carangidae): Adenocephalus pacificus, Tylosurus acus acus (Lacepède, 1803) Bicotylophora trachinoti, Caligus (Belonidae): Bomolochus constrictus, quadratus, Dinurus sp., Gonocercella Colobomatus goodingi.   pacifica, Gonocerca sp., Kuhnia sp., Tylosurus pacificus (Steindachner, 1876) Lecithocladium sp., Lernanthropus sp., (Belonidae): Caligodes laciniatus, Lobatostoma kemostoma, L. pacificum, Caligus sp. Lobatostoma sp., Phyllodistomum sp., Xenistius peruanus Hildebrand, 1946 P ro l e p t u s s p . , P y r a g r a p h o r u s ( H a e m u l i d a e ) : C y n o s c i o n i c o l a pyragraphorus. intermedia. Tr a c h i n o t u s r h o d o p u s G i l l , 1 8 6 3 Xiphias gladius Linnaeus, 1758 (Xiphiidae): ( C a r a n g i d a e ) : E c t e n u r u s s p . , Gloiopotes huttoni. Lobatostoma pacificum, Parahemiurus Xiphophorus hellerii Heckel, 1848 sp.  (Poeciliidae): Centrocestus sp. Tr a c h u r u s m u r p h y i N i c h o l s , 1 9 2 0 Zungaro zungaro (Humboldt, 1821) (Carangidae): Adenocephalus pacificus, (Pimelodidae): Amphoteromorphus Anisakis physeteris, A. simplex, Anisakis parkamoo, Chambriella agostinhoi, sp., Caligus bonito, Callitetrarhynchus Euzetiella tetraphylliformis, Jauella gracilis, Cemocotyle trachuri, glandicephalus, Megathylacus jandia, C e m o c o t y l e s p . , C e r a t o t h o a Peltidocotyle lenha, Proteocephalus gaudichaudii, Contracaecum sp., sophiae, Travassiella jandia. Corynosoma australe, C. obtuscens, Unspecified host (unspecified family): Corynosoma sp., Diphyllobothrium sp., Anilocra laevis, Asotana formosa, Eutetrarhynchus sp., Lernanthropinus Cymothoa oestrum. trachuri, Monascus sp., Nybelinia s u r m e n i c o l a , N y b e l i n i a s p . , Chondrichthyes (cartilaginous fishes) Procamallanus (Spirocamallanus) Alopias sp. (Alopiidae): Echthrogaleus chimusensis, Pseudoterranova decipiens, pellucidus. Pygidiopsis sp., Rhabdochona sp., Callorhinchus callorhynchus (Linnaeus, R h a d i n o r h y n c h u s s p . , “ S c o l e x 1 7 5 8 ) ( C a l l o r h i n c h i d a e ) : pleuronectis”, Tegorhynchus sp., Callorhynchocotyle marplatensis, Tentacularia coryphaenae. Gyrocotyle maxima. Trichomycterus dispar (Tschudi, 1846) Carcharhinus leucas (Müller & Henle, 1839) (Trichomycteridae): Contracaecum sp., (Carcharhinidae): Cathetocephalus Procamallanus (Spirocamallanus) thatcheri. c h i m u s e n s i s , P y g i d i o p s i s s p . , Carcharhinus porosus (Ranzani, 1839) Rhabdochona sp. (Carcharhinidae): Cathetocephalus Trichomycterus piurae (Eigenmann, 1922) thatcheri, Disculiceps pileatus. (Trichomycteridae): Clinostomum m a r g i n a t u m , P r o c a m a l l a n u s

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Cetorhinus maximus (Gunnerus, 1765) Pseudocharopinoides myliobatides, (Cetorhinidae): Dinobothrium paciferum. P s e u d o c h a r o p i n u s n a r c i n a e , Dasyatis dipterura (Jordan & Gilbert, 1880) Rhinebothrium sp., Rhodobothrium (Dasyatidae): Acanthobothroides mesodesmatum, Trebius latifurcatus. peruensis, A. thorsoni, Rhodobothrium Myliobatis peruvianus Garman, 1913 pulvinatum. (Myliobatidae): Acanthobothrium Galeorhinus galeus (Linnaeus, 1758) b r e v i s s i m e , A . h o l o r h i n i , A . (Triakidae): Perissopus dentatus. gonzalesmugaburui , Anaporrhutum Isurus oxyrinchus Rafinesque, 1810 a l b i d u m , A n t h o b o t h r i u m s p . , (Lamnidae): Anthosoma crassum, Benedeniella sp., Echinobothrium sp., Dinemoura, latifolia, D. producta, Eutetrarhynchus sp., Nybelinia sp., Echthrogaleus denticulatus, Gangliopus Phyllobothrium thridax, Proleptus sp., pyriformis, Nemesis lamna, Nybelinia sp., Rhinebothrium sp., Rhodobothrium P a n d a r u s k a t o i , P . s a t y r u s , mesodesmatum. Clistobothrium tumidum, Phyllothyreus Paratrygon aiereba (Müller & Henle, 1841) cornutus. (Potamotrygonidae): Nandocestus M o b u l a t h u r s t o n i ( L l o y d , 1 9 0 8 ) guariticus, Potamotrygon cf. fitzgeralde, (Myliobatidae): Nagmia peruviana. P o t a m o t r y g o n o c e s t u s s p . , Mustelus dorsalis Gill, 1864 (Triakidae): Proteocephalidea gen. sp., Rhabdochona Acanthocheilus bicuspis, Anisakis sp., sp., Rhinebothrium copianullum, Calliobothrium sp., Calliobothrium Rhinebothrium sp., Rhinebothroides sp. verticillatum, Paraorygmatobothrium Potamotrygon falkeri (Castex & Maciel, 1963) triacis, Erpocotyle sp., Lacistorhynchus ( P o t a m o t r y g o n i d a e ) : t e n u i s , L o i m o s s c o l i o d o n i , Potamotrygonocestus cf. fitzgeraldae, Orygmatobothrium musteli. Rhabdochona sp., Rhinebothroides sp. Mustelus lunulatus Jordan & Gilbert, 1882 Potamotrygon cf. falkeri (Castex & Maciel, (Triakidae): Orygmatobothrium musteli 1 9 6 3 ) ( P o t a m o t r y g o n i d a e ) : Mustelus mento Cope, 1877 (Triakidae): Acanthobothrium cf. peruviense, Acanthocheilus bicuspis, Calliobothrium N a n d o c e s t u s g u a r i t i c u s , sp., Calliobothrium verticillatum, P o t a m o t r y g o n o c e s t u s s p . , Echinocephalus sp., Lacistorhynchus Proteocephalidea gen. sp., Rhinebothrium tenuis, Nybelinia sp., Orygmatobothrium copianullum, Rhinebothroides sp. mustelid, Orygmatobothrium sp., Potamotrygon motoro (Müller & Henle, 1841) Phyllobothrium lactuca, Phyllobothrium (Potamotrygonidae): Acanthobothrium sp., Proleptus obtusus.  peruviense, Acanthobothrium cf. ramiroi, Mustelus sp. (Triakidae): Ceratothoa P o t a m o t r y g o n o c e s t u s s p . , gaudichaudii Potamotrygonocotyle chisholmae, P. Mustelus whitneyi Chirichigno F., 1973 dromedaries, Proteocephalidea gen. sp., (Triakidae): Lacistorhynchus tenuis, R h i n e b o t h r i u m c o p i a n u l l u m , Symcallio lintoni, Orygmatobothrium Rhinebothroides sp. musteli, Paraorygmatobothrium triacis. Potamotrygon tatianae Silva & Carvalho, Myliobatis chilensis Philippi, 1892 2011 (Potamotrygonidae): Rhinebothrium (Myliobatidae): Acanthobothrium batailloni, copianullum. A. holorhini, Acanthobothrium sp., Potamotrygon sp. (Potamotrygonidae): C a u l o b o t h r i u m m y l i o b a t i d i s , Potamotrygonocotyle rionegrense, P. Echeneibothrium sp., Eudactylina tsalickisi. myliobatidos, Phyllobothrium thridax, Prionace glauca (Linnaeus, 1758)

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(Carcharhinidae): Achtheinus oblongus, Scyllium sp. (Scyliorhinidae): Echinocephalus Anisakis sp., Anthobothrium laciniatum, striatus. Anthosoma crassum, Gangliopus Sphyrna zygaena (Linnaeus, 1758) pyriformis, Hepatoxylon trichiuri, (Sphyrnidae): Platybothrium sp., Paraorygmatobothrium angustum, Thysanocephalum thysanocephalum. P l a t y b o t h r i u m a u r i c u l a t u m , S q u a t i n a a r m a t a ( P h i l i p p i , 1 8 8 7 ) Prosobothrium armigerum, Tentacularia (Squatinidae): Grillotia sp., Otodistomum coryphaenae. veliporum, Phyllobothrium sp. Rhinobatos planiceps Garman, 1880 Sympterygia brevicaudata (Cope, 1877) (Rhinobatidae): Acanthobothrium olseni, ( R a j i d a e ) : A c a n t h o b o t h r i u m A . ro b u s t u m , A n o p l o c o t y l o i d e s l u s a r m i e n t a e , A . p s a m m o b a t i , chorrillensis, A. papillatus, Eudactylina Otodistomum veliporum. peruensis, Heterocotyle sp., Ommatokoita Synodus scituliceps Jordan & Gilbert, 1882 e l o n g a t a , P h y l l o b o t h r i u m s p . , (Synodontidae): Rhinebothrium sp., Prochristianella heteracantha, Proleptus “Scolex pleuronectis”. carvajali, P. obtusus, Rhinebothrium Triakis maculata Kner & Steindachner, 1867 rhinobati, Rhinobatonchocotyle (Triakidae): Acanthocheilus bicuspis, cyclovaginatus, R. pacifica, Stibarobdella Proleptus obtusus. moorei. Unspecified host (unspecified family): Rhinoptera steindachneri Evermann & Pandarus smithii. J e n k i n s , 1 8 9 1 ( M y l i o b a t i d a e ) : Caulobothrium sp.

Table 1. Nominal and undetermined species according taxonomic group recorded of fishes from Peru.

Taxonomic group Nominal sp. Undetermined sp. Total Monogenea 135 40 175 Cestoda 106 34 140 Digenea 53 57 110 Copepoda 76 12 88 Nematoda 31 18 49 Acanthocephala 10 6 16 Myxozoa 7 9 16 Isopoda 5 4 9 Aspidogastrea 4 1 5 Annelida 1 1 2 Branchiura 2 0 2

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Table 2. Environment distribution according taxonomic group recorded of fishes from Peru.

Taxonomic group Marine Freshwater Both Total environments Monogenea 90 85 0 175 Cestoda 68 71 1 140 Digenea 82 27 1 110 Copepoda 86 2 0 88 Nematoda 28 19 2 49 Acanthocephala 10 6 0 16 Myxozoa 11 5 0 16 Isopoda 8 1 0 9 Aspidogastrea 5 0 0 5 Annelida 2 0 0 2 Branchiura 1 1 0 2

DISCUSSION drainages. Recently, taxonomical studies have been performed on the parasites of fishes from t h e P e r u v i a n A m a z o n u s i n g b o t h Though many parasites have been reported morphological and molecular approaches, from fish in Peru, these reports likely represent including on cestodes (de Chambrier et al., only a small fraction of the true richness of 2015a,b), monogeneans (Mendoza-Palmero et parasite species from Peruvian fishes. There al., 2015) and acanthocephalans (Lisitsyna et are 1599 known Peruvian fish species (Froese al., 2015). However, our knowledge of the & Pauly, 2015), of which 189, i.e., less than parasite fauna of these fishes is still far from 1/8, have parasitological records. This is a complete, and most of the diversity remains negligible increase compared with the untouched. proportion of ≈ 1/10 calculated by Luque & Poulin (2007) almost 10 years ago. The current Although Peru is a megadiverse country, it was scenario for fish parasite research in Peru can not included as a fish parasite hotspot by Luque be extended to other parts of the world and to & Poulin (2007) because of the low number of other groups of vertebrates as parasite hosts studies compared to other Latin American (Madhavi, 2011; Hernández-Orts et al., 2015; countries such as Brazil, Mexico and Puerto Tavakol et al., 2015); thus, any estimates of Rico. parasite diversity are questionable (Poulin, 2014). A steadily decreasing number of The marine fishes M. gayi peruanus, S. taxonomic experts (and their uneven japonicus, P. peruanus, S. deliciosa, S. minor distribution) makes the situation even worse and T. murphyi presented the highest richness (Brooks & Hoberg, 2000). of parasites. Because Peru is one of the foremost Latin fishing nations, based mainly Despite the greater richness of freshwater on its large coast that is rich in fish species that fishes in Peru compared to marine species, few are used as basic food supplies in several parasitological surveys have focused on the regions of the country (Ibarra et al., 2000), it is fishes inhabiting Peru's large hydrological not surprising that the majority of

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parasitological studies focus on important which exhibit narrow host specificity, a highly fishery species. Moreover, there is financial specialized attachment apparatus and a direct support for parasitological studies on species life-cycle associated with numerous fish hosts that have economic importance, such as those (Whittington 1998; Cribb et al., 2002; Poulin, performed by Iannacone et al. (2009a-c), who 2002); or (2) directed studies on this parasite studied the parasitic fauna of A. scapularis, P. group performed by a single or few research humeralis and M. cephalus. teams, which is pronounced at a local scale (Poulin & Morand, 2004). The much lower The fauna of Chilean fishes is very similar to number of monogeneans reported from Chile Peru in that its largest number of parasites were (43 species) compared to Peru (175 species) reported from Sebastes oculatus Valenciennes, may reflect a lower number of studies in this 1833 (= Sebastes capensis), M. gayi gayi, T. country, which is aggravated by the lack of murphyi and C. gilberti (Muñoz & Olmos, taxonomic experts (Muñoz & Olmos, 2007). 2007, 2008). This similarity is not surprising, given the countries' obvious spatial proximity The second most species-richest group was the and their sharing of the Humboldt Current cestodes. Twelve of the 20 known orders of System, which is known for its richness of tapeworms (assuming that Proteocephalidea is nutrients and biodiversity, including parasites not included in the Onchoproteocephalidea; and fishes. Therefore, the differences in the see Alves et al., 2015a) have been reported number of hosts studied in Peru (206 species) from Peruvian fishes, including the still and Chile (approximately 100 species) seems artificial “Tetraphyllidea”, which was to arise from a reduced number of studies previously a catch-all group and is now split undertaken in the latter country and not by a into several new orders (Caira et al., 2014). To smaller number of potential hosts; 1167 fish date, the proteocephalideans presented the species are estimated to inhabit the highest richness of cestode species (46 spp.) corresponding area in the Humboldt Current and mainly parasitize (Siluriformes) System (Muñoz & Olmos, 2007; 2008; from the Peruvian Amazon (de Chambrier et Miloslavich et al., 2011). al., 2006, 2015a,b).

In the present study, the greatest richness of Concerning the trematodes, the diversity of parasites was registered among monogeneans species found in Peru, Brazil and Chile (115, (175) and cestodes (140), in contrast to the 247 and 86 species, respectively) (present observations in freshwater fishes in Mexico by study; Kohn et al., 2007; Muñoz & Olmos, Salgado-Maldonado (2013), where nematodes 2008) is highest in Brazil but may be and trematodes are the dominant group of considered to be rather low. The Digenea, parasites. In Chile, the most common fish which is the biggest trematode group to date, is parasites are crustaceans and trematodes also the largest taxon of endoparasites (Olson (Muñoz & Olmos, 2007; 2008). These et al., 2003), mostly due to the extraordinary differences, especially between Peru and diversification in its sites of infection, life Chile, which are similar in their composition of cycles and modes of feeding (Cribb et al., marine fishes, can be caused by the different 2002). Thus, the large number of fishes known foci of studies carried out by some research from Peruvian waters should serve either as teams (Poulin & Morand, 2004). definitive or intermediate hosts for these organisms. Limited information is available on The fact that monogeneans constitute the metacercariae in freshwater fishes. largest number of species could be a result of two factors: (1) the biology of these organisms, The nematodes were represented by 49

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species, almost one third of the total compiled Desenvolvimento Tecnologico (CNPq), by Luque et al. (2011), who listed 140 named Brazil; Fabiano Paschoal by a student and 75 unidentified species in fishes from f e l l o w s h i p f r o m C o o r d e n a ç ã o d e Brazil, but this number is similar to that Aperfeiçoamento de Pessoal do Ensino reported from Chile (i.e., 22 named and 17 Superior (CAPES), Brazil; Philippe V. Alves unidentified species) (Muñoz & Olmos, 2008). by a student fellowship from CNPq, and Ana Poulin (2014) stated that variation in the C. da Silva by a student fellowship from parasitic fauna among closely related Fundação de Amparo à Pesquisa do Estado de geographic areas is mainly driven by the São Paulo (FAPESP), Brazil. distribution of potential hosts. Therefore, the similarity between the nematode species in Peru and Chile might be explained by their similar fish fauna. In contrast, the Brazilian BIBLIOGRAPHIC REFERENCES coastline is the largest in South America; thus, a higher number of nematode species is Aguinaga, JF, Marcusso, PF, Claudiano, GS, expected from the numerous potential hosts. Marotta, BL, Sebastião, FA, Fernandes, JBK, Moraes, FR & Moraes, JRE. 2015. Other groups of parasites, such as Parasitic infections in ornamental Acanthocephala, Hirudinea and Myxozoa, cichlid fish in the Peruvian Amazon. were represented by only a few species, which Brazilian Journal of Veterinary corresponds to the situation in Chile (Muñoz & Parasitology, vol. 24, pp. 82-86. Olmo, 2007; 2008). Alcántara-Bocanegra, F, Verdi-Olivares, L, Murrieta-Morey, G, Rodríguez-Chu, L, The present account confirms that Peru has one Chu-Koo, F & Águila-Pizarro, M. 2015. of the biggest fish parasite fauna in South Parásitos de alevinos de gamitana America but also indicates that much more (Colossoma macropomum) y paco effort is necessary to grasp its true diversity (Piaractus brachypomus) cultivados en and might serve as a guide for further studies el C.I. Quistococha, Loreto, Perú. on this subject. Analyses based on Ciencia amazonica, vol. 5, pp. 42-49. morphological and molecular data (integrative Alves, PV, de Chambrier, A, Scholz, T & taxonomy) are pending to enrich our Luque, JL. 2015a. A new genus and knowledge of these parasites either to search species of proteocephalidean tapeworm for cryptic species and their geographical (Cestoda), first parasite found in the distributions or to link larval stages to their driftwood Tocantinsia piresi respective adult forms. (Siluriformes: Auchenipteridae) from Brazil. Folia Parasitologica, vol. 62, pp. 006. Alves, PV, Vieira, FM, Santos, CP, Scholz, T & ACKNOWLEDGEMENTS Luque, JL. 2015b. A Checklist of the Aspidogastrea (Platyhelminthes: Trematoda) of the World. Zootaxa, vol. We thank Tomás Scholz (Institute of 3918, pp. 339-396. Parasitology, Academy of Sciences, Czech Amin, OM, Heckmann, RA, Mesa, R & Mesa, Republic) for reviewing and commenting an E. 1995. Description and host early draft of the manuscript. José L. Luque relationships of cystacanths of was supported by a research fellowship from P o l y m o r p h u s s p i n d l a t u s C o n s e l h o N a c i o n a l d e P e s q u i s a e (Acanthocephala: Polymorphidae) from

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