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Horned Lizards (Phrynosoma) of Sonora, Mexico: Distribution And
RESEARCH ARTICLE Horned Lizards (Phrynosoma) of Sonora, Mexico: Distribution and Ecology Cecilia Aguilar-Morales, Universidad de Sonora, Departamento de Investigaciones Científicas y Tecnológicas, Blvd. Luis Encinas y Rosales SN, Hermosillo, SON; [email protected] Thomas R. Van Devender, GreaterGood, Inc., 6262 N. Swan Road, Suite 150, Tucson, AZ; [email protected] Mexico is recognized globally as a mega-diversity of the Sierra San Javier, the southernmost Sky Island country. The state of Sonora has very diverse fauna, (Van Devender et al. 2013). The Sierra Madre Oc- flora, and vegetation. The diversity of horned lizards in cidental reaches its northern limit in eastern Sonora, the genus Phrynosoma (Phrynosomatidae) in the state with Madrean species present in the oak woodland and of Sonora is a reflection of the landscape and biotic di- pine-oak forests in the higher elevations of the Sky Is- versity. In this paper, we summarize the distribution lands. West of the Madrean Archipelago, desertscrub and ecology of eight species of Phrynosoma in Sonora. vegetation is present in the Sonoran Desert lowlands of Mexico is western and central Sonora. Methods recognized Phrynosoma records globally as a Study area Eight species of Phrynosoma are reported from So- mega-diversity The great biodiversity of Sonora is the result of nora (Enderson et al. 2010; Rorabaugh and Lemos country. The complex biogeography and ecology. The elevation in 2016). Distribution records from various sources and state of Sonora Sonora ranges from sea level at the Gulf of California many photo vouchers are publicly available in the to over 2600 m in the Sierras Los Ajos and Huachinera Madrean Discovery Expeditions (MDE) database has very diverse (Mario Cirett-G., pers. -
Blood-Squirting Variability in Horned Lizards (Phrynosoma)
Blood-Squirting Variability in Horned Lizards (Phrynosoma) Wade C. Sherbrooke; George A. Middendorf, III Copeia, Vol. 2001, No. 4. (Dec. 20, 2001), pp. 1114-1122. Stable URL: http://links.jstor.org/sici?sici=0045-8511%2820011220%293%3A2001%3A4%3C1114%3ABVIHL%28%3E2.0.CO%3B2-5 Copeia is currently published by American Society of Ichthyologists and Herpetologists. Your use of the JSTOR archive indicates your acceptance of JSTOR's Terms and Conditions of Use, available at http://www.jstor.org/about/terms.html. JSTOR's Terms and Conditions of Use provides, in part, that unless you have obtained prior permission, you may not download an entire issue of a journal or multiple copies of articles, and you may use content in the JSTOR archive only for your personal, non-commercial use. Please contact the publisher regarding any further use of this work. Publisher contact information may be obtained at http://www.jstor.org/journals/asih.html. Each copy of any part of a JSTOR transmission must contain the same copyright notice that appears on the screen or printed page of such transmission. JSTOR is an independent not-for-profit organization dedicated to and preserving a digital archive of scholarly journals. For more information regarding JSTOR, please contact [email protected]. http://www.jstor.org Sat Apr 7 17:29:06 2007 Copan. 2001(4),pp. 1111-1122 Blood-Squirting Variability in Horned Lizards (Phrynosoma) WADEC. SHERBROOKEAND GEORGEA. MIDDENDORF111 Variability within the genus Phrymmm in the occurrence of ocular-sinus blood- squirting, reportedly a defense used in canid encounters, is reviewed from the lit- erature. -
AMPHIBIANS and REPTILES of ORGAN PIPE CACTUS NATIONAL MONUMENT Compiled by the Interpretive Staff with Technical Assistance from Dr
AMPHIBIANS AND REPTILES OF ORGAN PIPE CACTUS NATIONAL MONUMENT Compiled by the Interpretive Staff With Technical Assistance From Dr. J . C. McCoy, Carnegie Museum, Pittsburg, Pa. and Dr. Robert C. Stebbins, University of California, Berkeley Comm:m Names Scientific Names Amphibians Class Amphibia Frogs ~ Toads Order Sa.;J.ienta Toads Family Bufonidae Colorado River toad Bufo alvaris (s) Great plains toad nuro co~tus (s) Red- spotted toad Biil'O puncatus (s) Sonora green toad Buf'O retiformis Spadefoot Toads Family Pelobatidae Couch's spadefoot Scaphiopus couchi (8) Reptiles Class Reptilia Turtles Order Testudinata Mud Turtles and Their Allies Family Chelydridae Sonora mud turtle Kinosternon sonorien8e (s) Land Tortoises and Their Allies Family Testudinjdae - -.;.;.;.;;;.;;;;;..--- Desert tortoise Gopherus agassizi Lizards and Snakes Order Squamata Lizards Suborder Sauria Geckos Family Gekkonidae Desert banded gecko Coleoqyx ! . variegatus (s) Iguanids Family Iguanidae Arizona zebra- tailed lizard Callisaurus draconoides ventralis (s) Western collard lizard Crotaphytus collaris bailer) ( s) Long-nosed leopard lizard Crotaphytus ! . wislizeni (s Desert iguana ~saurus d. dorsalis (s) Southern desert horned lizard osoma p!atyrhinos calidiarum Regal horned lizard Phrynosoma solara ( s) Arizona chuckwalla Sauromalus obesus tumidus (s) Desert spi~ lizard SceloEorus m. magister (s) Colorado River tree lizard Urosaurus ornatus symmetricus (s) Desert side-blotched lizard ~ stansburiana stejnegeri (s) Teids Family Teidae Red-backed whiptail Cnemidophorus burti xanthonot us ~) Southern whiptail Cnemidophorus tigris gracilis (6) Venomous Lizards Family Heloderrr~tidae Reticula.te Gila m:mster Heloderma ! .. suspectum (6) Snakes Suborder Serpentes Worm Snakes Family Leptotyphlopidae Southwestern blind snake Leptottphlops h. humilis (s) Boas Family Boidae Desert rosy boa Lichanura trivirgata gracia (s) Colubrids Family Colubridae Arizona glossy snake Arizona elegans noctivaga c. -
Preliminary Assessment of Changes in a Lizard Assemblage at an Ecotone in Southeastern Arizona
Preliminary Assessment of Changes in a Lizard Assemblage at an Ecotone in Southeastern Arizona Lawrence L. C. Jones USDA Forest Service, Coronado National Forest, Tucson, Arizona Abstract—The Madrean Archipelago and its associated valleys have the highest diversity of lizards in the United States. This is due to a convergence of ecoregions in an area that provides excellent environmental conditions for life history needs of terrestrial ectotherms. The study area, near Safford, Arizona, is known to have about 20 species of sympatric lizards, although only about one-half are common. The lizard community is represented by species of the Sonoran and Chihuahuan deserts, semi-desert grasslands, and lower Madrean and boreal woodlands. It has recently been suggested that lizard species are expected to decline globally due to climate change and other reasons. A study site representing an ecotone between desert and grassland/ montane/riparian vegetation types in a foothills situation was chosen, as ecotones are marginal habitats that are expected to be sensitive to environmental change. Study objectives were to assess the baseline lizard community and detect changes in the lizard assemblage over time due to climate change and other factors. During systematic road transects, a total of 3,889 lizards representing 13 species were recorded during 60 visits in 2003 (n = 8 visits), 2010 (n = 12,) and 2011 (n = 40). Although this represents the early stages of a long-term monitoring program, preliminary observations show differences in the lizard assemblage between years consistent with climate change predictions. Introduction Mountains. In this paper I document some changes in the lizard as- semblage, although surveys to date are only the beginning of what is The Madrean Archipelago’s “sky island” mountain ranges and intended to be a long-term monitoring project. -
Year of the Lizard News No
Year of the Lizard News No. 4 July 2012 V V V V V V V V V V www.YearoftheLizard.org Lizards Across the Land: Federal Agencies’ Role From Alaska to Hawaii to Florida, hundreds of millions of acres of our public lands are held in trust by federal land management agencies. Many of these lands support rich and diverse populations of lizards. The following collection of articles provides a sample of the outstanding scholarly and practical work being conducted on our federal public lands. Biologists at these and other federal agencies are hard at work to answer many important questions regarding A Copper-striped Blue-tailed Skink (Emoia impar) the science of lizard conservation and management and to photographed in Samoa during a USGS field survey. identify and conserve priority habitats for lizards and other Photo: Chris Brown, USGS. native wildlife. “No other landscape in these United States has —Terry Riley, National Park Service, National PARC been more impacted by extinction events and species Federal Agencies Coordinator invasions in historic times than the Hawaiian Islands, with as yet unknown long-term cascading consequences USGS Reveals “Cryptic Extinction” of Pacific to the ecosystem,” said U.S. Geological Survey director Lizard Marcia McNutt. “Today, we close the book on one more animal that is unlikely to ever be re-established in this A species of lizard is now extinct from the Hawaiian fragile island home.” Islands, making it the latest native vertebrate species to “This skink was once common throughout the become extirpated from this tropical archipelago. Hawaiian Islands, and in fact the species can still be The Copper-striped Blue-tailed Skink (Emoia impar) — found on many other island groups in the tropical a sleek lizard with smooth, polished scales and a long, sky- blue tail — was last confirmed in the Na’Pali coast of Kauai continued on p. -
Urban Naturalist No
Urban Naturalist No. 2 2014 Historic and Current Composition of Lizard Communities in Urban Preserves of Central Arizona, USA Brian K. Sullivan, David Vardukyan, and Keith O. Sullivan The Urban Naturalist . ♦ A peer-reviewed and edited interdisciplinary natural history science journal with a global focus on urban areas ( ISSN 2328-8965 [online]). ♦ Featuring research articles, notes, and research summaries on terrestrial, fresh-water, and marine organisms, and their habitats. The journal's versatility also extends to pub- lishing symposium proceedings or other collections of related papers as special issues. ♦ Focusing on field ecology, biology, behavior, biogeography, taxonomy, evolution, anatomy, physiology, geology, and related fields. Manuscripts on genetics, molecular biology, anthropology, etc., are welcome, especially if they provide natural history in- sights that are of interest to field scientists. ♦ Offers authors the option of publishing large maps, data tables, audio and video clips, and even powerpoint presentations as online supplemental files. ♦ Proposals for Special Issues are welcome. ♦ Arrangements for indexing through a wide range of services, including Web of Knowledge (includes Web of Science, Current Contents Connect, Biological Ab- stracts, BIOSIS Citation Index, BIOSIS Previews, CAB Abstracts), PROQUEST, SCOPUS, BIOBASE, EMBiology, Current Awareness in Biological Sciences (CABS), EBSCOHost, VINITI (All-Russian Institute of Scientific and Technical Information), FFAB (Fish, Fisheries, and Aquatic Biodiversity Worldwide), WOW (Waters and Oceans Worldwide), and Zoological Record, are being pursued. ♦ The journal staff is pleased to discuss ideas for manuscripts and to assist during all stages of manuscript preparation. The journal has a mandatory page charge to help defray a portion of the costs of publishing the manuscript. -
Withdrawal of Proposed Rule to List the Flat-Tailed Horned Lizard As Threatened; Proposed Rule
Vol. 76 Tuesday, No. 50 March 15, 2011 Part III Department of the Interior Fish and Wildlife Service 50 CFR Part 17 Endangered and Threatened Wildlife and Plants; Withdrawal of Proposed Rule To List the Flat-Tailed Horned Lizard as Threatened; Proposed Rule VerDate Mar<15>2010 17:35 Mar 14, 2011 Jkt 223001 PO 00000 Frm 00001 Fmt 4717 Sfmt 4717 E:\FR\FM\15MRP3.SGM 15MRP3 jlentini on DSKJ8SOYB1PROD with PROPOSALS3 14210 Federal Register / Vol. 76, No. 50 / Tuesday, March 15, 2011 / Proposed Rules DEPARTMENT OF THE INTERIOR lizard found in the Sonoran Desert of hybrids between flat-tailed and desert the southwestern United States and horned lizards have been observed in Fish and Wildlife Service northwestern Mexico. All of the species the vicinity of Ocotillo, California of lizards in the genus Phrynosoma—the (Stebbins 2003, p. 302). Additionally, 50 CFR Part 17 horned lizards—have dorso-ventrally the regal horned lizard (P. solare) also [Docket No. FWS–R8–ES–2010–0008; MO flattened, ‘‘pancake-like’’ bodies; spiny occurs in northwestern Sonora, Mexico 92210–0–0008] scales; head spines or ‘‘horns’’; cryptic (Rorabaugh 2008, p. 39); we are not coloration; and certain similar aware of hybridization with this species. RIN 1018–AX07 behavioral traits (Sherbrooke 2003, pp. 4–17; Stebbins 2003, p. 299; Leache´ and Life History Endangered and Threatened Wildlife McGuire 2006, p. 629). and Plants; Withdrawal of Proposed Flat-tailed horned lizards are Among horned lizard species, the flat- oviparous (egg-laying), are early Rule To List the Flat-Tailed Horned tailed horned lizard has particularly Lizard as Threatened maturing, and may produce multiple long and sharp horns (Funk 1981, p. -
The Effect of Prescribed Burning and Grazing on Th Threatened Texas Horned Lizard (Phrynosoma Cornutum) in the Western Rio Grande Plains
THE EFFECT OF PRESCRIBED BURNING AND GRAZING ON THE THREATENED TEXAS HORNED LIZARD (PHRYNOSOMA CORNUTUM) IN THE WESTERN RIO GRANDE PLAINS By ANNA L. BURROW Bachelor of Science University ofTennessee, Knoxville Knoxville, Tennessee 1998 Submitted to the Faculty ofthe Graduate College ofthe Oklahoma State University in partial fulfillment of the requirements for the Degree of MASTER OF SCIENCE December, 2000 h THE EFFECT OF PRESCRIBED BURNING AND GRAZING ON THE THREATENED TEXAS HORNED LIZARD (PHRYNOSOMA CORNUTUM) IN THE WESTERN RIO GRANDE PLAINS Thesis Approved: £~1f6: ~~---I~Gr~' ____ [the II L ACK OWLEDGME T I would like to thank numerous individuals for their ffort, support and motivation towards the completion ofthis project. I wish to express my appreciation to my major advisor, Dr. Eric C. Hellgren for his advice, guidance, con tructive suggestions, and quick editing. My appreciation extends to my other committee members, Dr. Chuck C. Peterson and Dr. David M. Engle, for their guidance as well. I am grateful to Dr. Richard Kazmaier for his help in every aspect of this project. His involvem nt in this project ranged from teaching me to become a seamstre s to building "the pole. 'I al a wish to recognize Donald C. Ruthven, III for his continued support and help throughout this project. Texas Parks and Wildlife Department and David Synatszke also deserve special recognition for providing me with the financial support to undertake thi project and for providing excellent accommodations throughout my field sea ons. I would also like to thank Sigma Xi Grants-in-Aid of research for partially funding thi research. -
Ecol 483/583 – Herpetology Lab 6: Reptile Diversity 1: Lizards Spring 2010
Ecol 483/583 – Herpetology Lab 6: Reptile Diversity 1: Lizards Spring 2010 P.J. Bergmann & S. Foldi (Modified from Bonine & Foldi 2008; Bonine, Smith & Stitt 2003; Edwards 2002; Prival 2000) Lab objectives The objectives of today’s lab are to: 1. Gain an appreciation for the extant diversity of lizards. 2. Learn to identify species of lizards that live in Arizona. 3. Learn what an amphisbaenian is and what one looks like. Today’s lab is the first reptile diversity. Lizards are the dominant component of herpetofauna in Arizona, so there are many species and higher taxa that you will need to learn during this lab. We start with the lizards because this is what you will see most of in the field. Tips for learning the material Continue what you have been doing in previous weeks. Examine all of the specimens on display, taking notes, drawings and photos of what you see. Attempt to identify the local species to species and the others to their higher taxon. Quiz each other to see which taxa are easy for you and which ones give you troubles, and then revisit the difficult ones. Use a key for more difficult species. This will enhance the skill with which you use a key. There will also be a keying exercise, where several unknowns will be displayed and you will be expected to identify them. Although today we are covering only one group, lizards, do not underestimate the importance of the lab. There are many local species as well as incredible global diversity. Furthermore, lizards will be the most abundant of the herpetofauna we will see in the field as the semester progresses. -
Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding
SCIENTIFIC AND STANDARD ENGLISH NAMES OF AMPHIBIANS AND REPTILES OF NORTH AMERICA NORTH OF MEXICO, WITH COMMENTS REGARDING CONFIDENCE IN OUR UNDERSTANDING SEVENTH EDITION COMMITTEE ON STANDARD ENGLISH AND SCIENTIFIC NAMES BRIAN I. CROTHER (Committee Chair) STANDARD ENGLISH AND SCIENTIFIC NAMES COMMITTEE Jeff Boundy, Frank T. Burbrink, Jonathan A. Campbell, Brian I. Crother, Kevin de Queiroz, Darrel R. Frost, David M. Green, Richard Highton, John B. Iverson, Fred Kraus, Roy W. McDiarmid, Joseph R. Mendelson III, Peter A. Meylan, R. Alexander Pyron, Tod W. Reeder, Michael E. Seidel, Stephen G. Tilley, David B. Wake Official Names List of American Society of Ichthyologists and Herpetologists Canadian Association of Herpetology Canadian Amphibian and Reptile Conservation Network Partners in Amphibian and Reptile Conservation Society for the Study of Amphibians and Reptiles The Herpetologists’ League 2012 SOCIETY FOR THE STUDY OF AMPHIBIANS AND REPTILES HERPETOLOGICAL CIRCULAR NO. 39 Published August 2012 © 2012 Society for the Study of Amphibians and Reptiles John J. Moriarty, Editor 3261 Victoria Street Shoreview, MN 55126 USA [email protected] Single copies of this circular are available from the Publications Secretary, Breck Bartholomew, P.O. Box 58517, Salt Lake City, Utah 84158–0517, USA. Telephone and fax: (801) 562-2660. E-mail: [email protected]. A list of other Society publications, including Facsimile Reprints in Herpetology, Herpetologi- cal Conservation, Contributions to Herpetology, and the Catalogue of American Amphibians and Reptiles, will be sent on request or can be found at the end of this circular. Membership in the Society for the Study of Amphibians and Reptiles includes voting privledges and subscription to the Society’s technical Journal of Herpe- tology and news-journal Herpetological Review, both are published four times per year. -
Quitobaquito Pupfish
INDEX OF SPECIES SEEN AT SCOTTSDALE COMMUNITY COLLEGE Text written by staff. Photos by Roy Barnes, Emma Olsen and Dr. John Weser. FISH SPECIES Gila Topminnow Quitobaquito Pupfish AMPHIBIAN SPECIES Couch’s Spadefoot Toad Great Plains Toad Lowland Leopard Frog Sonoran Desert Toad Woodhouse’s Toad REPTILE SPECIES SNAKES: Checkered Garter Snake Common Kingsnake (California Kingsnake) Gopher Snake Mojave Rattlesnake Red Racer (Coachwhip) Western Diamondback Rattlesnake LIZARDS: Desert Spiny Lizard Mediterranean Gecko Regal Horned Lizard Tiger Whiptail Lizard TURTLES and TORTOISES: Desert Tortoise GILA TOPMINNOW (POECILIOPSIS OCCIDENTALIS) Species Name: Poeciliopsis occidentalis Range: Currently, populations are found in several localities in the Gila River system of Mexico and Arizona. Habitat: Occupy headwater springs and vegetated margins and backwater areas of streams and rivers. They can withstand waters from near freezing to 100 degrees Fahrenheit. Diet: Omnivorous: broad spectrum diet including crustaceans and vegetable material, but will also feed on aquatic insect larvae, especially mosquitoes, when abundant. Breeding: The breeding season is April through November, but they can reproduce year-round in thermally stable springs. During breeding, some males become dark and exhibit aggressive breeding behavior. The typical brood consists of over 1000 young. The typical life span is one year. Status: In 1967, the Gila Topminnow was listed by the U.S. Fish and Wildlife Service as endangered. At one time, the Gila Topminnow was the most common fish found in the Gila River basin. Its numbers have been greatly reduced due to the introduction of other fish species, especially the mosquito fish. Photo: Taken at Scottsdale Community College on April 5, 2007. -
Phrynosoma Blainvillii) in the Southern San Joaquin Valley of California
ECOLOGY AND DEMOGRAPHY OF THE BLAINVILLE'S HORNED LIZARD (PHRYNOSOMA BLAINVILLII) IN THE SOUTHERN SAN JOAQUIN VALLEY OF CALIFORNIA SUSAN MARIE HULT A Thesis submitted to the Department of Biology California State University, Bakersfield In Partial Fulfillment of the Degree of Masters of Science JUNE 2014 iii COPYRIGHT BY SUSAN MARIE HULT 2014 ii ECOLOGY AND DEMOGRAPHY OF THE BLAINVILLE'S HORNED LIZARD (PHRYNOSOMA BLAINVILLII) IN THE SOUTHERN SAN JOAQUIN VALLEY OF CALIFORNIA Susan M. Hult This thesis has been accepted on behalf of the Department of Biology by their supervisory committee: Dr. David J. Germano Committee Chair Dr. Steve Laymon Dr. Brandon Pratt iv ACKNOWLEDGEMENTS I would sincerely like to thank my advisor, Dr. David Germano, for his patience throughout the entire processes of coursework, research, fieldwork, and writing that resulted in the completion of my degree. His professional support in the realms of writing, statistical analysis, and editing has been invaluable. I also truly appreciate Dr. Stephen Laymon, who has proven to be an incredible source of moral, professional, and field support at Atwell and in the capacity of serving on my committee. I likewise thank Dr. Brandon Pratt for his ideas, friendly support, and review of my manuscript, which added greatly to the integrity of this thesis and for serving on my committee. Without the Bureau of Land Management's (BLM) Student Career Experience Program, I most likely would not have been able to carry out this research and obtain my degree. I hope (and believe) this research serves as a valuable source of information in which they can use to make land management decisions beneficial for P.