Monographs of the Western North American Naturalist

Volume 7 8th California Islands Symposium Article 12

9-25-2014 Island specialists: shared flora of the Alta and Baja California Pacific slI ands Sarah E. Ratay University of California, Los Angeles, [email protected]

Sula E. Vanderplank Botanical Research Institute of Texas, 1700 University Dr., Fort Worth, TX, [email protected]

Benjamin T. Wilder University of California, Riverside, CA, [email protected]

Follow this and additional works at: https://scholarsarchive.byu.edu/mwnan

Recommended Citation Ratay, Sarah E.; Vanderplank, Sula E.; and Wilder, Benjamin T. (2014) "Island specialists: shared flora of the Alta and Baja California Pacific slI ands," Monographs of the Western North American Naturalist: Vol. 7 , Article 12. Available at: https://scholarsarchive.byu.edu/mwnan/vol7/iss1/12

This Monograph is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Monographs of the Western North American Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Monographs of the Western North American Naturalist 7, © 2014, pp. 161–220

ISLAND SPECIALISTS: SHARED FLORA OF THE ALTA AND BAJA CALIFORNIA PACIFIC ISLANDS

Sarah E. Ratay1, Sula E. Vanderplank2, and Benjamin T. Wilder3

ABSTRACT.—The floristic connection between the mediterranean region of Baja California and the Pacific islands of Alta and Baja California provides insight into the history and origin of the California Floristic Province. We present updated species lists for all California Floristic Province islands and demonstrate the disjunct distributions of 26 taxa between the Baja California and the California Channel Islands. These 26 plant taxa are found among the 16 Pacific islands without occurring on the intervening mainland of Alta California. Separate species lists for each island group (8 California Channel Islands and 8 Baja California Islands) were compiled. These lists were compared to the mainland Cali- fornia flora to identify species that occur on the California Islands and either the Baja California Pacific Islands or the mediterranean region of the Baja California Peninsula, but not the mainland of Alta California. This first compilation of the flora of the Baja California Islands and nomenclatural updates for the Channel Islands provide a platform for future research and conservation planning.

RESUMEN.—El grado de conexión florística entre la región mediterránea de Baja California y las Islas del Pacífico de Alta y Baja California proporciona un entendimiento de la historia y el origen de la Provincia Florística de California. Presentamos listados actualizados de especies de todas las islas de la Provincia Florística de California y mostramos la distribución aislada de veintiséis taxones entre la Baja California y las Islas del Canal de California. Identificamos aquel- las especies de plantas que se pueden encontrar dentro de las dieciséis islas del Pacífico, pero no se encuentran en la península de Alta California. Con este objetivo, recopilamos listados separados de las especies que habitan en cada grupo de islas (las ocho Islas del Canal de California y las ocho islas de Baja California). Estos listados se compararon con la flora existente en la península de California con el fin de identificar aquellas especies que se encuentran en las islas de California, y tanto en las Islas del Pacífico de Baja California como en la región mediterránea de la Península de Baja California, pero no en la península de Alta California. La primera recopilación de flora de las Islas de Baja Califor- nia y actualizaciones nomenclaturales en las Islas del Canal, proporcionan una plataforma para futuras investigaciones y planes de conservación.

The islands of the California Floristic Prov - well-recognized attributes that are accentuated ince (CFP) are an important component of one on the islands of the Californias. of the world’s great biodiversity hotspots The degree of floristic similarity of 16 off- (Myers et al. 2000). The CFP has commonly shore islands, and their apparent role as refugia been referred to as an environmental island for species once present on the mainland, has due to its regionally unique mediterranean-type been a persistent biogeographical puzzle. The climate, isolation (via oceanic and orographic CFP includes 16 islands from the adjacent boundaries), and corresponding globally sig- coast of Alta and Baja California and extends nificant levels of endemism. The plant com- south to the 28th parallel, including the CFP munity has had ample time to evolve since vegetation found on Cedros and Guadalupe the emergence of wet winters and dry sum- Islands (Raven and Axelrod 1978, Moran 1996, mers at least 5–10 Mya (Axelrod 1973) and Oberbauer 2002a). The CFP has an increasing perhaps earlier (Ackerly 2009, Keeley et al. gradient of aridity from north to south, but 2011). The remarkable diversity and endemism vegetation distributions are heavily influenced of the CFP has been attributed to relative cli- by fog presence; this fog provides additional matic stability during glacial-interglacial tran- moisture and reduces radiant loadings, signifi- sitions of the Pleistocene (Lancaster and Kay cantly changing conditions for plants (Dawson 2012, Sniderman et al. 2013). The antiquity 1998, Fischer et al. 2008, Vanderplank 2013). and distinctiveness of the California flora are Physical data suggest that ocean currents have

1University of California, Los Angeles, Department of Ecology and Evolutionary Biology, 612 Charles E. Young Drive East, Box 957246, Los Angeles, CA 90095. E-mail: [email protected] 2Botanical Research Institute of Texas, 1700 University Dr., Fort Worth, TX 76107. 3University of California, Riverside, Department of Botany and Plant Sciences, 900 University Avenue, Riverside, CA 92507.

161 162 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7

Fig. 1. California Channel and Baja California Pacific Islands with Diegan Scrub as defined by Axelrod (1978). Map by B.T. Wilder, adapted from Google Earth™ imagery. been stable for millions of years (Jacobs et al. nomenclatural updates to the flora of the 8 2004), which indicates the presence of fog California Channel Islands, and (3) the plants during the development of the CFP. that occur on the California Channel Islands Within the large number of plant species and in the Diegan Scrub but not the rest of on the CFP islands and the mainland of Alta mainland California, hereafter referred to as and Baja California is the occurrence of dis- “exemplar taxa.” junct Baja California taxa on the California Channel Islands. These species are seen today METHODS only on the CFP islands and a distinct portion Development of Checklists of the California mainland termed the Diegan Scrub, the San Diego region southward into We assembled all existing species checklists coastal northwestern Baja California (Fig. 1; for the 2 island groups (Alta California and Axelrod 1978). However, the number and iden- Baja California) into 2 lists, then compared tity of this group of disjunct species has re - those lists with the flora of the California mained unknown. Floristic Province for both mainland Alta and To identify the species that define this pat- Baja California to generate the list of exemplar tern, we present (1) the first compiled check- taxa. The nomenclatural updates presented here list of the 8 CFP Baja California Islands, (2) are based on our best understanding of the 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 163 island floras at the date of publication and fol- National Park Service checklist from Junak et low the models of the sources indicated below al. (1997) for San Miguel, Santa Rosa, Santa for the 2 lists. Species now thought to be extir- Cruz, Anacapa, and Santa Barbara islands. For pated from the islands are denoted by an San Clemente Island we included updates ampersand (&). Unvouchered taxa (including from Ross et al. (1996) and for San Nicholas reports, photo vouchers, and personal commu- Island updates from Junak (2008). Additional nications) are indicated by a question mark (?). taxa and island records are included based on Nonnative species are indicated by an asterisk recent collections on Santa Catalina Island by (*). When a subspecies was not recorded for Sarah Ratay, observations and collections on all the islands and some ambiguity remains, San Clemente Island by Emily Howe (Soil the taxon is recorded at the species level only. Ecology and Restoration Group, SDSU), and Exotic species that were planted or are not herbarium specimens found in the Consor- naturalized (e.g., Nerium oleander) were deleted tium of California Herbaria (CCH 2014), which from checklists. are listed in footnotes. The nomenclatural re - vision of this checklist is consistent with tax- Baja California Islands Checklist onomy of the new Jepson Manual (Baldwin et The most recent species lists for each of al. 2012) through the use of the dynamic con- the 8 Baja California Islands (Appendix 1; cordance tool provided on the Jepson e-flora Coronados, Todos Santos, San Martín, Jeron- website (Jepson Flora Project 2013). This imo, Guadalupe, San Benito, Cedros, and checklist was developed to identify distribu- Natividad) were compiled from the following tion patterns in the flora, and though the data sources: nomenclature has been updated, we have not confirmed identifications of questionable her - Coronados—Oberbauer 2002b barium specimens, nor have we diligently pur- Todos Santos—Junak and Philbrick 1994a sued the lowest taxonomic rank beyond the San Martín—Junak and Philbrick 1994b published sources. We are also aware of addi- Guadalupe—Moran 1996, Rebman 2006, Rebman et tional updates to the flora of many of the al. 2007 San Benito—Junak and Philbrick 2002b, Rebman 2007a Channel Islands in process by Steve Junak, Cedros—Oberbauer 1987, Rebman 2007b including publication of the flora of Catalina Natividad—Junak and Philbrick 2002a Island; Junak’s updates are not included here. Exemplar Taxa The list of Jeronimo island flora, consisting of 8 taxa, was provided via personal communica- The identification of exemplar taxa was tion from Steve Junak. We verified mainland generated through a comparison of the 2 newly Baja California flora distributions with the generated island checklists (Appendixes 1, 2) unpublished data of Jon Rebman and Bart with the Jepson Manual (Baldwin et al. 2012) O’Brien (personal communication 2013). Sev- and the inventory of rare and endemic plants eral taxa were added to the checklists based of CFP Baja California (O’Brien et al. 2014). A on recent collections, herbarium specimens large number of species from CFP Baja Cali- recently encountered, and recent reports by fornia extend just slightly into southern Califor- Sula Vanderplank, Jon Rebman, and Steve nia and are considered near-endemic to Baja Junak; these taxa are indicated by footnotes. California (O’Brien et al. 2014). As such, we The taxonomy for this checklist follows the include the near-endemic species of CFP Checklist of Baja California Plants in prepara- Baja California as ‘absent’ in the mainland of tion by Jon Rebman. Alta California (with the exception of Crosso- soma californicum, which occurs on the Palos California Channel Islands Checklist Verde Peninsula, and Euphorbia misera, which The updated flora of the 8 California Chan- occurs in the South Coast region of California nel Islands (Appendix 2; San Clemente, San and beyond; Baldwin et al. 2012). These few Nicolas, Santa Catalina, Santa Barbara, Santa species are consistent with the concept of Rosa, Santa Cruz, Anacapa, and San Miguel) is Diegan Scrub as proposed by Axelrod (1978). based on the master checklist complied by Diegan Scrub is named for its occurrence in Gary Wallace (1985). We included published southern San Diego County and south into updates to individual island floras found in the Baja California. It is a coastal scrub, rich in 164 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7

TABLE 1. The 26 exemplar taxa as they occur on 8 Baja California Pacific Islands and the 8 Channel California Islands.a CRPR column provides the California Rare Plant List ranking, and the BC column provides the State of Baja California Family Exemplar taxa Habit NAT CED BEN GUA JER MAR Apiaceae Lomatium insulare Herb GUA Asteraceae Hazardia cana Shrub GUA Asteraceae Senecio lyonii Shrub BEN MAR Boraginaceae Phacelia floribunda Herb GUA Cactaceae Bergerocactus emoryi Succulent MAR Convolvulaceae Calystegia macrostegia Vine GUA MAR subsp. macrostegia Crossosomataceae Crossosoma californicum Shrub GUA Euphorbiaceae Euphorbia misera Shrub NAT CED BEN GUA MAR Fabaceae Acmispon argophyllus Shrub GUA subsp. argenteus Fabaceae Lupinus guadalupensis Herb GUA Fabaceae Trifolium palmeri Herb GUA Fagaceae Quercus tomentella Tree GUA Grossulariaceae Ribes viburnifolium Shrub CED Lamiaceae Salvia brandegeei Shrub Papaveraceae Eschscholzia ramosa Herb NAT CED BEN GUA MAR Phmyraceae Mimulus latifolius Herb GUA Plantaginaceae Gambelia speciosa Shrub GUA Poaceae Dissanthelium californicum Herb GUA Polemoniaceae Gilia nevinii Herb GUA Polemoniaceae Leptosiphon pygmaeus Herb GUA subsp. pygmaeus Rhamnaceae Ceanothus arboreus Shrub GUA Rhamnaceae Rhamnus pirifolia Shrub GUA Rosaceae Prunus ilicifolia subsp. lyonii Tree Saxifragaceae Jepsonia malvifolia Herb GUA Scrophulariaceae Scrophularia villosa Shrub GUA Solanaceae Solanum wallacei Shrub GUA NUMBER OF SPECIES 23321 05 aNatividad [NAT], Cedros [CED], San Benito [BEN], Guadalupe [GUA], Jeronimo [JER], San Martín [MAR], Todos Santos [TOS], Coronados [COR], San Clemente succulents, that includes many endemic and (1) multiple California Channel Islands and near-endemic species from CFP Baja; it also Guadalupe Island, 13 taxa includes the coastal succulent scrub and coastal (2) Guadalupe Island–San Clemente Island only, sage scrub ecoregions (González-Abraham et 4 taxa (Hazardia cana, Leptosiphon pyg- al. 2010). maeus ssp. pygmaeus, Lupinus guadalupen- sis, Phacelia floribunda) RESULTS (3) Diegan Scrub and the Channel Islands only, 2 taxa (Prunus ilicifolia ssp. lyonii and The California Pacific Islands in total contain Salvia brandegeei) 1239 unique taxa. The California Islands separately contain 976 taxa and the Baja Califor- (4) widespread, 1 taxon (Eschscholzia ramosa) nia Islands 535 taxa. Twenty-six of these taxa have occurs on 14 of 16 islands a disjunct occurrence between the Baja Cali- (5) certain Channel and Baja California Islands fornia Pacific Islands and the California Chan- and Diegan Scrub, 4 taxa (Bergerocactus nel Islands, absent from Alta California out- emoryi, Crossosoma californicum, Euphor- side the Diegan Scrub (Table 1). These exem- bia misera, Ribes viburnifolium) plar taxa represent 16 families and 26 unique (6) certain Channel and Baja California Islands, genera. No one growth form predominates; of 2 taxa (Calystegia macrostegia ssp. macro - the 26 taxa, there are 12 shrubs, 10 herba- stegia and Senecio lyonii) ceous annuals, 2 trees, 1 vine, and 1 xerophytic succulent (Table 1). Fabaceae is the largest DISCUSSION family with 3 species on the exemplar taxa list. These 26 taxa fall into 6 biogeographic sub- The complex biogeography of the Califor- patterns: nia Floristic Province can be explained in part 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 165

Islands are listed left to right by increasing latitude. The Mainland column indicates continental occurrences. The protected rankings. TOS COR Mainland CLE NIC CAT BAR ROS CRU ANA MIG # ISL CRPR BC No CLE NIC 3 1B.2 1b No CLE 2 1B.2 1b Baja only CLE CAT 4 2a No CLE 2 1B.2 1b TOS COR Baja & CA CLE CAT 5 2.2 4 No CAT ROS CRU ANA MIG 7 2a

CA only CLE CAT 3 1B.2 1b TOS COR Baja & CA CLE CAT CRU 10 2.2 No CLE NIC CAT 4 2a

No CLE 2 1B.2 1b No CLE NIC CAT BAR ANA 5 4.2 2a No CLE CAT ROS CRU ANA 6 4.2 2a Baja & CA CAT 2 1B.2 4 Baja only ROS 1 1B.2 1b TOS COR No CLE NIC CAT BAR ROS CRU 13 4.3 2a No CAT CRU 3 1A 1b No CLE CAT BAR 4 1B.2 1b No CLE CAT 3 1B.2 1a No CLE NIC CAT BAR ROS CRU ANA 8 4.3 2a No CLE 2 1B.2 1b

No CAT ROS CRU 4 2a No CLE CAT ROS CRU MIG 6 4.2 2a Baja only CLE CAT ROS CRU ANA 5 No CLE NIC CAT ROS CRU 6 4.2 2a No CLE CAT 3 1B.2 1b No CAT ROS CRU 4 1B.1 1b 33 20 6 20 4 10 11 52 21 24

[CLE], San Nicolas [NIC], Santa Catalina [CAT], Santa Barbara [BAR], Santa Rosa [ROS], Santa Cruz [CRU], Anacapa [ANA], and San Miguel [MIG]. by analyzing unusual floristic patterns seen moved north, fog-moderated climates retained on the California Pacific Islands. The 26 taxa species native to that region. shared by the Baja California and Channel A number of the exemplar taxa have distri- Islands indicate a clear connection between butions on the Channel Islands and farther these isolated and disjunct regions. What pre- south in Baja California, suggestive of the cludes these taxa from occurring on the Alta refugium hypothesis (i.e., these species were California mainland, and what does this sug- once more widespread). Epling and Lewis gest for the past extent of the CFP? Did these (1942) proposed that the diverse Diegan Scrub taxa evolve lo cally and then migrate, or does has many taxa derived from Miocene vegeta- this pattern instead reflect a distant legacy? tion that came from the north Mexican plateau. Three current hypotheses attempt to ex - Axelrod (1978) states that Diegan Scrub was plain the presence of these disjunct popula- more restricted in the Pleistocene and ex - tions on the California Pacific Islands: (1) panded in the Holocene as arid-adapted taxa these species have the ability to disperse long moved northward into the southern Channel distances yet for some reason have failed to Islands. Many CFP species are found in dis- establish on the California mainland; (2) these junct occurrences, scattered across Arizona species were his torically more widespread, and Mexico, not unlike the presence of these connected by former geologic linkages, and 26 species on the islands (Valiente-Banuet et have been unable to persist on the Alta Cali- al. 1998, Bhaskar et al. 2007). Small-scale fornia mainland, remaining in refugia on the topo graphic niches and facilitation by other islands and in Baja California; or (3) the Chan- species often support these occurrences. Axel- nel Islands originally occurred farther south rod (1978) also points out that the most impor- near San Diego (Atwater 1998) and as they tant conditions for Diegan Scrub vegetation 166 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 appear to be the absence of frost and the pres- introduced species. Presumably, the lower ence of a degree of summer moisture (in the proportion of invasive species on the Baja form of fog). The cold California current and California Islands is due to less human use dense fog banks suggest that similarities in cli- and fewer impacts to date. mate may help to explain the similarities in The exemplar taxa identified by this process the floras (Fischer et al. 2008). can now serve as the focal point of research to On the other hand, a number of the exem- better understand the region’s historical bio- plar taxa are perhaps better explained as hav- geography. Such research can incorporate ing evolved on the islands. This is especially phylogenetic relationships and their importance true of many island Lotus species (Acmispon; in community assembly (Webb et al. 2008). McGlaughlin et al. 2011, McGlaughlin personal Functional trait analyses would help deter- communication), Eschscholzia ramosa (Still and mine if these taxonomically diverse species Potter 2013), and Crossosoma californicum share inherent traits that explain their disjunct (Wallace and Helenurm 2009), among others distributions. Comparative phylogeographic (Baldwin, 2007). For this pattern, we hypothe- studies of these shared taxa would test the size local evolution and subsequent dispersal specific hypothesis discussed above, among to similar niches on the CFP Baja Islands. others, and offer a glimpse into the past. Addi- Guadalupe Island poses a particularly in - tionally, the knowledge of patterns and modes teresting study system. As the most isolated of dispersal would provide valuable natural island, located roughly 270 km offshore, history information. Further genetic work to Guada lupe is a true oceanic island never con- resolve the origin of these island specialist nected to the Baja California Peninsula. Under- species would help to elucidate questions of standing the colonization of Guadalupe may timing, and definitively separate neo- and paleo - contribute to understanding the dispersal pat- endemics in the exemplar list. terns that we see today. Of 26 exemplar taxa, Conservation activity in Baja California has 21 of them occur on Guadalupe. Floristic recently expanded, yet knowledge of plant similarities to the Channel Islands flora (at distributions and ecology remains limited. least 400 km north) may be due to the Califor- We hope that these updated checklists of the nia current moving species from the Califor- California Pacific Islands will facilitate nia islands southward. Guadalupe Island has species status and distribution updates on been above water for millions of years, and its both sides of the border, and foster well- current flora could represent the accumula- informed conservation decisions that promote tion of species from the possible sources over expanded science and management for the this time. transborder ecoregion. In addition to a better understanding of the origin and evolution of the CFP, these species ACKNOWLEDGMENTS lists allow for improved conservation planning. Twenty-one of the 26 species are listed by the This work is based on the legacy of bo - California Rare Plant Ranking (CNPS 2013) tanists who have explored and documented yet currently have no protection with the Mexi - the floristic diversity of California and the can government. Twenty-five of the 26 species Pacific Islands. In particular, we thank the fol- are listed in the rare plant ranking for Baja lowing individuals who have directly aided California (O’Brien et al. in press). These rank- this paper. Steve Junak’s expertise has been ings are utilized in conservation planning in invaluable; we are most grateful for his per- California and would be useful for land plan- sonal communications and guidance. His dedi - ning in Baja California. These combined cation and role as an information clearing- islands species lists provide important infor- house for the island floras is vital for island- mation about the level of invasive species in focused plant work. We are extremely grateful the re gions. Of the 535 taxa on the Baja Cali- to Jon Rebman of the San Diego Natural His- fornia Islands, roughly 14% (74) are intro- tory Museum (SD) for his assistance with the duced species (excluding planted species, Baja California Island checklist, as well as his which were not included in our analyses). In additional data, nomenclatural reference works, comparison, of the 976 species on the Califor- and taxonomic assistance. Karen Rich and Judy nia Channel Islands, roughly 28% (278) are Gibson at SD also provided information. Bart 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 167

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______. 2002b. Vegetation and flora of Islas Los Coronados, flora under a non–Mediterranean-type climate. PNAS Baja California, México. Pages 212–223 in H.W. 110:3423–3428. Chaney, K.L. Mitchel, and D.R. Browne, editors, STILL, S.M., AND D. POTTER. 2013. California poppy Proceedings of the Fifth California Islands Sympo- conundrums: insights into relationships within tribe sium: 29 March to 1 April 1999. Santa Barbara Eschscholtzieae (Papaveraceae). Systematic Botany Museum of Natural History, Santa Barbara, CA. 38:104–117. O’BRIEN, B., J. DELGADILLO-RODRÍQUEZ, S.A. JUNAK, T.A. VALIENTE-BANUET, A., N. FLORES-HERNÁNDEZ, M. VERDÚ, OBERBAUER, J.P. REBMAN, H. RIEMANN, AND S.E. AND P. DÁVILA. 1998. The chaparral vegetation of VANDERPLANK. In press. The rare, endangered and Mexico under nonmediterranean climate: the con- endemic plants of the California Floristic Province vergence and Madrean–Tethyan hypotheses recon- portion of Baja California, Mexico. Aliso. sidered. American Journal of Botany 85:1398–1408. RAVEN, P.H., AND D.I. AXELROD. 1978. Origin and rela- VANDERPLANK, S. 2013. Endemism in an ecotone: from tionships of the California flora. University of Cali- chaparral to desert in Baja California, Mexico. In: C. fornia Publications in Botany 72:1–134. Hobhom, editor, Vascular plant endemism. Springer- REBMAN, J.P. 2006. The flora of Guadalupe Island, Mexico. Verlag, Dresden. [Accessed 26 April 2013]. Available from: http://baja WALLACE, G.D. 1985. Vascular plants of the Channel flora.org/floras/GuadalupeEndemics1.htm Islands of southern California and Guadalupe Island, ______. 2007a. The flora of San Benitos Island, Mexico. Baja California, México. Contributions in Science [Accessed 26 April 2013]. Available from: http://baja 365, Natural History Museum of Los Angeles County. flora.org/Floras/SanBenitosIsland.htm WALLACE, L.E., AND K. HELENURM. 2009. Has herbivory ______. 2007b. The flora of Cedros Island, Mexico. [Ac- negatively impacted genetic variability in the flora cessed 26 April 2013]. Available from: http://baja of the California Channel Islands? Insights from flora.org/Floras/CedrosIsland.htm Crossosoma californicum (Crossosomataceae). Inter- REBMAN, J.P., T.A. OBERBAUER, AND J.L. LEÓN DE LA LUZ. national Journal of Plant Sciences 170:311–322. 2007. La flora de Isla Guadalupe y sus islotes adya- WEBB, C.O., D.D. ACKERLY, AND S.W. KEMBEL. 2008. Phy- centes, Baja California, México. [Accessed 26 April locom: software for the analysis of phylogenetic com - 2013]. Available from: http://www2.ine.gob.mx/pub munity structure and trait evolution. Bioinformatics licaciones/libros/477/cap5.html 24:2098–2100. ROSS, T.S., S. BOYD, AND S. JUNAK. 1996. Additions to the vascular flora of San Clemente Island, Los Angeles Received 27 April 2013 County, California, with notes on clarifications and Accepted 27 May 2014 deletions. Aliso 15(1):27–40. Early online 25 September 2014 SNIDERMAN, J.M.K., G.J. JORDAN, AND R.M. COWLING. 2013. Fossil evidence for a hyperdiverse sclerophyll

Appendix 1 on page 169. Appendix 2 on page 188. 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 169 CED CED CED CED CED CED CED? CED CED CED GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA COR COR TOS COR COR TOS MAR JER GUA BEN CED NAT (J.T. Howell) Yatsk., (J.T. & E.Wollenw. & E. Wollenw. & E. Wollenw. Windham ssp. leucophylla Windham mucronata var. Windham Yatsk., (Weath.) ssp. maxonii Windham, Eaton) Yatsk., ssp. viscosa (D.C. binata (Engelm.) Lemmon var. ssp. californica ssp. triangularis ssp. semipallida Howell) Axelrod cedrosensis (J.T. var. ssp./var. and infraname ssp./var. S. Watson S. 1 1. Plant species checklist for the 8 Baja California islands (Cedros [CED], Coronados [COR], Jeronimo [JER], Guadalupe [GUA], Natividad [NAT]; San Benito [BEN], (Kaulf.) Yatskievych et al. Yatskievych (Kaulf.) et al. Yatskievych (Kaulf.) (Kaulf.) Yatskievych et al Yatskievych (Kaulf.) et al Yatskievych (Kaulf.) Bartel Watson) (S. D.C. Eaton Cheilanthes brandegeei D.C. Cheilanthes newberryi Domin Eaton Notholaena californica D.C. Fee andromedifolia (Kaulf.) Pellaea Eaton mucronata D.C. Pellaea triangularis Pentagramma triangularis Pentagramma Watson Ephedra aspera S. Don Pinus radiata D. Polypodium californicum Kaulf. Polypodium scouleri Hook. & Grev. Polypodium Adiantum capillus-veneris L. Eaton Notholaena californica D.C. triangularis Pentagramma triangularis Pentagramma Don Pinus radiata D. (Kaulf.) C. Presl munitum (Kaulf.) Polystichum Hesperocyparis guadalupensis Juniperus californica Carrière Watson S. ex Brahea edulis H. Wendl. PPENDIX Synonym: Cupressus guadalupensis Pteridaceae Ephedraceae Pinaceae Polypodiaceae Pteridaceae Dryopteridaceae Cupressaceae Arecaceae 1 ONOCOTS YMNOSPERMS ERNS M G A Family/species F San Martín [MAR], and Santos Todos [TOS]). An ampersand (&) indicates an extirpated species; a question mark (?) nonnative species. indicates an unvouchered taxon; and an asterisk (*) indicates a 170 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 NAT CED CED CED CED CED CED CED CED CED? CED CED CED? CED CED? BEN CED NAT GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA MAR MAR MAR TOS TOS CORCOR TOS TOSCOR MAR TOSCORCOR MAR TOS TOS MAR COR MAR COR COR TOS COR COR COR TOS COR TOS MAR JER GUA BEN CED NAT ssp. leopoldii bufonius var. congestus Wahlenb var. ssp. rubens (L.) Husnot ssp./var. and infraname ssp./var. pectinacea var. 6 5 Englem. (J. Presl) Á. Löve (J. * 7 3 4 2 1. Continued H.L. Bell & Columbus Juncus bufonius L. Arundo donax L. * Avena barbata Brot. * Avena fatua L. * Bromus berteroanus Colla Bromus carinatus Hook. & Arn. Bromus diandrus Roth.* Bromus hordeaceus L. * Bromus tectorum L. * Distichlis spicata (L.) Greene Distichlis littoralis (Engelm.) Juncus bufonius L. Calochortus splendens Dougl. Rydb Watson) Piperia cooperi (S. Agrostis pallens Trin. Aristida adscensionis L. Bromus madritensis L. * * Cynodon dactylon (L.) Pers. Agave sebastiana Greene Agave shawii Englem. valida Brandegee Yucca Carex spissa L.H. Bailey Eleocharis geniculata (L.) Roemer & Schultes Schoenoplectus californicus (C.A. Mey) Soják Juncus acutus L. Elymus Presl condensatus J. Eragrostis pectinacea (Michx.) Nees bromoides L. Festuca PPENDIX Report confirmed, herbarium specimen @ SD New record; herbarium specimen @ RSA, SD New record; herbarium specimen @ RSA, SD Synonym: Monanthochloe littoralis Synonym: Leymus condensatus bromoides (L.) Gray Synonym: Vulpia Liliaceae Orchidaceae Poaceae A Agavaceae Cyperaceae Juncaceae 2 3 4 5 6 7 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 171 NAT NAT CED CED? CED? CED? CED? CED NAT CED CED CED CED CED CED CED CED CED CED CED CED GUA GUA GUA GUA GUA GUAGUA BEN CED GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA MAR MAR JER MAR 11 TOS TOS TOS TOS MAR TOS COR TOS COR COR TOS MAR COR CORCOR TOS TOSCOR MAR COR MAR TOS TOS MAR MAR COR TOS MAR COR TOS MAR JER GUA BEN CED NAT (Piper) Henrard ssp. capitatum ssp. leporinum (Link) Arcang ssp. secunda ssp./var. and infraname ssp./var. pauciflora Scribn. var. octoflora var. ssp. glaucum (Steud.) Tzvelev. var. hirtella var. V. o. and V. * 13 14 8 (Trin.& Rupr.) Barkworth Rupr.) (Trin.& (Forssk.) C. Chr. (Forssk.) 10 15 12 (Walter) Rydb. var. octoflora var. Rydb. (Walter) * (Hitchc.) Barkworth 9 1. Continued Festuca octoflora Walt. Stipa diegoensis Swallen Stipa pulchra Hitchc. aestivum L. * Triticum Zea mays L. * Dichelostemma capitatum Alph. Wood latifolia L. Typha Phyllospadix scouleri Hook. Poa thomasii Refulio Polypogon viridis (Gouan) Breistr. Lenz guadalupensis L.W. Triteleia Hoover palmeri (S.Watson) Triteleiopsis Hordeum murinum L. * Lamarckia aurea (L.) Moench * Melica frutescens Scribner Melica imperfecta Trin. Muhlenbergia microsperma (DC.) Kunth Phalaris caroliniana Walt. Piptochaetium pringlei (Beal) Parodi annua L. * Poa Festuca myuros L. Phalaris minor Retz. * Presl secunda J.S. Poa Festuca microstachys Nutt. Hordeum murinum L. * Polypogon monspeliensis* (L.) Desf. Polypogon Schismus barbatus (L.) Thell. * Stipa & Rupr. speciosum Trin. Sorghum bicolor (L.) Moench * Stipa lepida A. Hitchc. PPENDIX Vulpia octoflora Synonyms: Vulpia New record; herbarium specimen @ SD (Thorne 53921) Synonym: Stenochloa californica Nutt. Synonym: Agrostis semiverticillata Synonym: Achnatherum speciosum Synonym: Nassella lepida Vulpia microstachys (Nutt.) Murno Synonym: Vulpia myuros (L.) C.C. Gmel. Synonym: Vulpia Themidaceae Typhaceae Zosteraceae A 8 9 10 11 12 13 14 15 Family/species 172 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CED CED CEDCED NAT CED? CED CED? CED CED CED? CED CED CED? CEDCED NAT CED GUA GUA GUA BENGUA CED?GUA NAT GUA BEN GUA GUA GUA GUAGUA BEN CED NAT MAR JER GUA MAR MAR MAR? TOS TOS TOS MAR TOS CORCOR TOS COR TOS TOS MAR MAR JER GUA BEN CED NAT COR TOS COR TOS COR TOS MAR COR TOS MAR JER GUA BEN CED NAT (Kell.) Gentry veatchiana (Kell.) var. ssp./var. and infraname ssp./var. 16 17 lentii (Hook. & Arn.) Koso-Pol. × 1. Continued ICOTS D Nerium oleander L. * Asclepias subulata Decne. in A. DC. Hydrocotyle umbellata Pachycormus discolor (Benth.) Cov. Pachycormus Watson Rhus integrifolia (Nutt.) Brewer & S. Rhus integrifolia (Nutt. in T. & G.) Nutt. ex Abrams Nutt. ex & G.) Malosma laurina (Nutt. in T. S. Watson Phyllospadix torreyi S. Anemopsis californica (Nutt.) H. & A. * Br. Carpobrotus chilensis (Molina) N.E. Mesembryanthemum crystallinum L. * Mesembryanthemum nodiflorum L. * Zostera marina L. Bowlesia incana Ruiz & Pav. Daucus pusillus Michaux (Eastw.) Munz. Lomatium insulare (Eastw.) microcarpa Yabea Rhus lentii Kell. Coriandrum sativum L. * vulgare Mill. * Foeniculum & A. Gray Apiastrum angustifolium Nutt. in Torrey Ambrosia chamissonis (Less.) Greene Ambrosia chenopodiifolia (Benth.) Payne Ambrosia magdalenae (Brandegee) Payne Amauria rotundifolia Benth. Amblyopappus pusillus Hook. & Arn. Ambrosia camphorata (Greene) Payne Agoseris heterophylla (Nutt.) Greene PPENDIX New record; herbarium specimen @ RSA, SD 1988) New record; herbarium specimen @ SD (166694, Amadeo Rea 1568, 22 Feb Apocynaceae Araliaceae Anacardiaceae A Saururaceae Aizoaceae 16 17 Apiaceae Asteraceae UDICOTS ASAL Family/species E B 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 173 NAT NAT NAT CED CED CED CED CED? CED? CED CEDCED? NAT CED CED CED BEN BEN CED GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA TOS TOS TOS TOS TOS TOS TOS MAR 22 COR COR COR COR TOS COR COR COR TOS COR TOS MAR JER GUA BEN CED NAT var. grandiflorum var. var. glabriuscula var. ssp./var. and infraname ssp./var. var. juncea var. * 24 (Moran) B.L. Turner (Moran) B.L. 25 L. * 19 23 18 20 (Moran) Rose ssp. peninsularis D. greeneana D. A. Gray × (A. Gray) Greene 21 1. Continued B.G. Baldwin B.G. (Pursh) J. Forbes Eriophyllum lanatum (Pursh) J. Eriophyllum confertiflorum (DC.) A. Gray Spach Glebionis coronarium (L.) Cass. ex (Rose) B.G. Baldwin Deinandra greeneana (Rose) B.G. Deinandra palmeri (Rose) B.G.Baldwin Deinandra palmeri Coreocarpus involutus Greene Deinandra fasciculata (DC.) Greene Baldwin Deinandra frutescens (A. Gray) B.G. Chaenactis lacera Greene Baccharis sarothroides A. Gray Howell Baeriopsis guadalupensis J.T. Bahiopsis laciniata A. Gray Artemisia californica Less. Baldwin Deinandra streetsii (A. Gray) B.G. Blake) Clark & Kyhos Encelia asperifolia (S.F. Encelia californica Nutt. Hypochaeris glabra L. * Greenella ramulosa Greene Gutierrezia sarothrae (Pursh) Britt. & Rusby Hazardia berberidis (A. Gray) Greene Hazardia cana (A. Gray) Greene Hazardia orcuttii Deinandra peninsularis Brickellia microphylla (Nutt.) A. Gray Brickellia Centaurea melitensis L. * Chaenactis glabriuscula DC. Encelia stenophylla Greene Ericameria brachylepis (A. Gray) Hall Erigeron sumatrensis Retz. Bebbia juncea (Benth.) Greene & Rose Encelia palmeri Vasey PPENDIX Viguiera laciniata Synonym: Viguiera New record; herbarium specimen @ SD New record; herbarium specimen @ SD Synonym: Hemizonia greeneana New record; herbarium specimens@ SD Synonym: Conyza floribunda Kunth Synonym: Chrysanthemum coronarium s.n. 1920-specimen recently identified) New record; herbarium specimen @ SD (5248, Ralph Sumner, A 18 19 20 21 22 23 24 25 Family/species 174 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CED CED CED? CED CED CED CED? CED CED CED CED CED BEN BEN CED 31 32 GUA GUA GUAGUA BEN CEDGUA NAT GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA TOS TOS TOS MAR TOS COR TOS MAR COR TOS MAR COR TOS COR TOS COR TOS MAR COR COR COR COR COR COR TOS var. odorata var. ssp./var. and infranamessp./var. menziesii var. COR TOS MAR JER GUA BEN CED NAT 29 27 * 28 30 26 33 (Greene) Rydb. Fisch. & C.A. Mey. Fisch. (Nutt.) Hook. f. (Kellogg) H.M. Hall H.M. (Kellogg) Nutt. (Nutt.) A.Gray A. Gray (Hook. & Arn.) Morefield (A. Gray) Holub 1. Continued (Kunth) Anderberg (Kunth) G.L. Nesom G.L. Burtt Hilliard & B.L. Porophyllum gracile Benth. Porophyllum beneolens Davidson Pseudognaphalium Micropus californicus californica Benth. Perityle incana A. Gray Perityle Pluchea odorata (L.) Cass. Perityle emoryi Torrey Perityle Logfia filaginoides Rafinesquia californica Nutt. Senecio aphanactis Greene Senecio cedrosensis Greene Leptosyne maritima stramineum Pseudognaphalium (Fisch. & Mey.) A. Gray & Mey.) Layia platyglossa (Fisch. Lasthenia gracilis (DC.) Greene Leptosyne gigantea Kellogg Isocoma menziesii (Hook. & Arn.) Iva hayesiana A. Gray Lasthenia coronaria (Nutt.) Ornduff Logfia arizonica biolettii Anderberg Pseudognaphalium luteoalbum (L.) Pseudognaphalium microcephalum Nutt. Pseudognaphalium ramosissimum Nutt. Pseudognaphalium sp. nova Pseudognaphalium Malacothrix clevelandii A. Gray Matricaria occidentalis Greene Malacothrix foliosa A. Gray Malacothrix insularis Greene Malacothrix similis Davis & Raven PPENDIX Synonym: Coreopsis gigantea Synonym: Coreopsis maritima arizonica Synonym: Filago californica Synonym: Filago Synonym: Chamomilla occidentalis New record; herbarium specimen @ RSA, SD New record; herbarium specimen @ RSA, SD New record; herbarium specimen @ RSA, SD (description pending) A 26 27 28 29 30 31 32 33 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 175 CED CED CED CED CED CED? CED NAT CED CED CED? CED CED CED? CED CED NAT CED? BEN& CED BEN& CED NAT BEN BEN GUA GUA GUA BEN CED NAT GUA GUA GUAGUA BENGUA CEDGUA NAT GUA GUA GUAGUA BEN GUA GUA GUA MAR MAR MAR TOS TOS TOS TOS COR TOSCOR MAR COR TOS MAR CORCOR TOS TOSCOR MAR MAR TOS COR TOS COR TOS COR TOS MAR JER GUA BEN CED NAT var. barbigera (A. Gray) Greene var. Johnston cryptochaeta I.M. var. cedrosensis (Greene) Jtn. var. maritima Greene var. ssp./var. and infraname ssp./var. californica var. penduliflora var. Constance bipinnatifida (Torr.) var. var chrysanthemifolia Johnst. ssp. ferocula I.M. Howell hispida (A. Gray) J. var. I. M. Johnston; herbarium specimen @ SD (Moran 25430) I. M. 34 1. Continued & M.G. Simpson & M.G. J.F. Macbr. Amsinckia inepta J.F. Amsinckia menziesii (Lehm.) Nels. & Macbr. Cryptantha barbigera Cryptantha clevelandii Greene Cryptantha foliosa Reiche & R.) Macr. Cryptantha grayi (V. Cryptantha intermedia (A. Gray) Greene Cryptantha maritima (Greene) Cryptantha maritima (Greene) Cryptantha patula Greene Uropappus lindleyi (DC.) Nutt. Xylothamia diffusa (Benth.) Nesom Emmenanthe penduliflora Benth. Senecio lyonii A. Gray Senecio sylvaticus L. * Sonchus oleraceus L. * Sonchus tenerrimus L. * Stebbinsoseris heterocarpa (Nutt.) Chambers Stephanomeria diegensis Gottlieb Stephanomeria guadalupensis Brandegee californica Kell. Trixis dissita A. Gray Verbesina hastata Kell. Verbesina A. Gray lanata (Kell.) Viguiera Eucrypta chrysanthemifolia (Benth.) Greene Eucrypta chrysanthemifolia (Benth.) Greene Harpagonella palmeri A. Gray Heliotropium curassavicum L. Johnston) Hasenstab Johnstonella angelica (I.M. linearis DC Pectocarya Johnst. recurvata I.M. Pectocarya Phacelia cedrosensis Rose Phacelia cicutaria Greene Senecio palmeri A. Gray Phacelia distans Benth. Phacelia floribunda Greene Phacelia crenulata Torr. PPENDIX Synonym: Cryptantha angelica Boraginaceae A 34 Family/species 176 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 NAT CED NAT CED CED CED CED CED? CED NAT CED CEDCED NAT CED CED CED CED CED CED CED BEN CED NAT BEN CED NAT 35 GUA GUA GUAGUA BEN? GUA GUA GUA GUA GUA GUA GUA GUAGUA BENGUA CEDGUA GUA NAT GUA GUA MAR? MAR MAR TOS TOS MAR TOS MAR TOS TOS MAR TOS TOS CORCOR TOS TOS MAR COR TOS MAR COR TOS COR TOS MAR JER GUA BEN CED NAT var. heterophyllus var. var. latifolium var. insulare C.L.Hitchc. var. ssp. sativa var. californicus var. gracilis Jtn. var. ssp./var. and infraname ssp./var. var. brachycarpa (Richardson) Detling var. (Woot. & Standl.) Detl.ssp. glabra (Woot. COR ssp. halictorum (Cockerell) Detling ssp. menziesii (DC.) Detling var. integrifolia var. 36 * 38 (L.) Desv. 37 (Hook. & Arn.) Greene 1. Continued I.M. Johnst. I.M. Caulanthus heterophyllus Payson Capsella bursa-pastoris (L.) Medik. * Torrey & A. Gray Lepidium nitidum Torrey Lepidium lasiocarpum Nutt. Lepidium oblongum Small Plagiobothrys acanthocarpus (Piper) Pholistoma auritum (Lindl.) Lilja Pholistoma racemosum (Nutt.) Const. Jhtn. I.M. Plagiobothrys collinus (Philbr.) Jhtn. I.M. Plagiobothrys collinus (Philbr.) Phacelia hirtuosa A. Gray Kell. Phacelia ixodes Phacelia parryi Torr. Phacelia phyllomanica A. Gray Brassica tournefortii Gouan * Cakile maritima Scop. * Caulanthus lasiophyllus (Hook. & Arn.) Payson Brassica rapa L. * Athysanus pusillus (Hook.) Greene * Koch Brassica nigra (L.) W.D.J. (Walt.) Britt. Descurainia pinnata (Walt.) Sisymbrium irio L. * Sisymbrium orientale L. * Watson Thysanocarpus erectus S. Thysanocarpus laciniatus Nuttall Raphanus sativus L. * Greene Watson) Sibara angelorum (S. Erysimum moranii Rollins Hornungia procumbens (L.) Hayek Descurainia pinnata Descurainia pinnata (Cockerell) Detling Descurainia pinnata (Cockerell) Detling S. Watson Draba cuneifolia S. Eruca vesicaria (L.) Cav. PPENDIX New record; herbarium specimen @ RSA, SD Synonym: Guillenia lasiophylla New record; herbarium specimen @ SD (John Brown, s.n., April 4 1983) Synonym: Hutchinsia procumbens A 35 36 37 38 Brassicaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 177 NAT CED? NAT CED CED CED CED NAT CED? CEDCED NAT NAT CED? CED CED? CED?CED NAT BEN CEDBEN? CED NAT? BEN CED BEN? GUA GUA GUA GUA aff. 40 41 42 43 MAR MAR MAR MAR MAR MAR TOS MAR COR TOS MAR COR TOS MAR COR TOS MAR? COR TOS MAR COR COR TOS MAR COR TOS MAR JER GUA BEN CED NAT Rebman) G.E. Linds. grandiflorus G.E. var. schottii var. shurliana Gates var. Dawson rectispina E.Y. var. goodridgei var. ssp./var. and infraname ssp./var. (Torr.) Benson reticulata (Torr.) var. Iltis angustata (Parish) var. globosa (Coville) Iltis var. var. guadalupensis (Morin) Lammers var. Parish 45 46 39 Nutt. ssp. angustata Planch. 44 1. Continued Gibson & K.E. Horak (M.E. Jones) K. Schum.Echinocereus maritimus (M.E. chrysacanthus (Orcutt) Britt. & Rose Ferocactus fordii (Orcutt) Britt. & Rose Ferocactus & R. Lophocereus schottii (Engelm.) Br. maritimus var. Mammillaria blossfeldiana Boedeker Brandegee Mammillaria dioica M.K. Mammillaria goodridgei Scheer Mammillaria goodridgei Scheer Mammillaria hutchisoniana (Gates) Boed. Linds Mammillaria louisae G.E. Cylindropuntia sp. nova (Engelm.) Br. & R. Bergerocactus emoryi (Engelm.) Br. Cochemiea pondii Walton Knuth F.M. Weber) Cylindropuntia alcahes (F.A.C. Knuth F.M. Weber) Cylindropuntia cholla (F.A.C. Cylindropuntia prolifera (Engelm.) F.M.Knuth alcahes var. Craig Mammillaria neopalmeri R.T. Myrtillocactus cochal (Orcutt) Britt. & Rose Opuntia ficus-indica (L.) Miller Opuntia littoralis (Engelm.) Cockerell Peritoma arborea (Nutt.) Iltis Peritoma arborea (Nutt.) Iltis Peritoma (Ruiz & Pav.) Greene biflora (Ruiz & Pav.) Triodanis Celtis laevigata (S. Watson) Britton & Rose Watson) pringlei (S. Pachycereus Stenocereus gummosus (Engelm.) A.C. Githopsis diffusa A. Gray Opuntia oricola Philbrick PPENDIX New record; pers. comm. Jon Rebman, SD (includes Cylindropuntia cedrosensis New record; herbarium specimen @ RSA, SD hutchinsonia Re-identification, changed from M. New record; herbarium specimen @ RSA New record; herbarium specimen @ RSA Synonym: Celtis douglasii Synonym: Isomeris arborea New record; herbarium specimen @ SD Rebman 6230 A Cactaceae 39 40 41 42 43 44 45 46 Capparaceae Cannabaceae Campanulaceae Family/species 178 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 aff. × NAT NAT CED CED? CEDCED NAT CED NAT CED NAT CED CED CED CED CED BEN CED? NAT? GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA BEN CED NAT GUAGUA BEN CED? NAT aff. 50 52 MAR MAR MAR MAR MAR TOS TOS TOS MAR COR TOS COR TOS MAR JER GUA BEN CED NAT COR COR COR TOS MAR COR COR COR COR COR TOS MAR COR TOS A. Gray var. davidsonii (Standl.) Munzvar. COR (Greene) B.L. Rob. leucantha (Greene) B.L. var. var. denudata Rehder var. holosteoides var. cinerea (DC.) Coutinho var. ssp./var. and infraname ssp./var. talinum (Greene) Jeps. var. ssp. canescens ssp. laciniata 51 49 C.L. Hitch. & Maguire 47 Parish (Lindl.) Douglas ex Torr. & A. Gray var. vacillans & A. Gray var. Torr. (Lindl.) Douglas ex 48 Cav. ssp. major Cav. 1. Continued (Hornem. ex Cham. & Schltdl.) Heynh. (Hornem. ex (Hornem. ex Cham. & Schltdl.) Heynh. (Hornem. ex Polycarpon depressum Nutt. Polycarpon Silene antirrhina L. Silene gallica L. * Silene laciniata Cav. J.& K. Presl Spergularia marina J.& (Moq.) D. Dietr. Atriplex coulteri (Moq.) D. Dietr. Atriplex leucophylla (Moq. in DC.) D. Atriplex pacifica Nels. * Atriplex semibaccata R. Br. Abrams Atriplex serenana A. Nelson ex S. Watson Atriplex julacea S. Spergularia bocconii (Scheele) Merino * Spergularia macrotheca Lonicera subspicata Hook. & Arn. & A. Gray Achyronychia cooperi Torrey Cerastium glomeratum Thuill. * Drymaria holosteoides Benth. Herniaria hirsuta L. Camb. * Spergularia villosa (Pers.) * Stellaria media (L.) Vill. Stellaria pallida (Dumort.) Crép * Stellaria nitens Nutt. Kuntze Watson) Allenrolfea occidentalis (S. Aphanisma blitoides Nutt. Standl. Arthrocnemum subterminale (Parish) Lonicera hispidula Dougl. Spergularia macrotheca Atriplex californica Moq. Atriplex canescens (Pursh) Nutt. Atriplex barclayana (Benth.) Dietr. PPENDIX Synonym: Lonicera hispidula Synonym: Silene laciniata New record; herbarium specimen @ RSA, SD New record; herbarium specimen @ RSA, SBBG Synonym: Salicornia subterminalis New record; herbarium specimen @ RSA, potentially new species Caryophyllaceae Chenopodiaceae A Caprifoliaceae 47 48 49 50 51 52 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 179 NAT NAT CED CED CED? CED CED? aff CED BEN BEN? CED NAT BEN BEN? GUA BEN CED GUA GUA GUA GUA BEN? CED GUA GUA 55 MAR JER MAR MAR MAR MAR MAR MAR MAR MAR MAR TOS TOS 53 56 COR COR COR TOS COR? COR TOS COR TOS COR TOS MAR COR TOS COR TOS MAR JER GUA BEN CED NAT COR TOS COR COR COR TOS MAR JER GUA BEN CED NAT ssp. intermedia ssp. macrostegia ssp./var. and infraname ssp./var. ssp. salina 54 57 L. D. cultrata Rose D. × 1. Continued A. Berger Calystegia macrostegia (Greene) Brummitt Calystegia macrostegia (Greene) Brummitt (Rafinesque) J.F. Macbride Suaeda nigra (Rafinesque) J.F. Suaeda taxifolia (Standl.) Standl. Cuscuta corymbosa Ruiz & Pavon Dudleya albiflora Rose Dudleya anomala (Davidson) Moran Dudleya anthonyi Rose Dudleya anthonyi Calystegia macrostegia (Greene) Brummitt Cuscuta californica Hook. & Arn. Cuscuta salina Engelm. Dudleya acuminata Rose Atriplex suberecta Verdoorn * Salsola australis R.Br. Dichondra occidentalis House Crassula connata (Ruiz Lopez & Pavon) Atriplex watsonii A. Nelson Chenopodium album L. (S. Watson) S. Watson S. Watson) Chenopodium californicum (S. Chenopodium flabellifolium Standl. Chenopodium murale L. * Salicornia pacifica Standl. (S. Watson) Moran Watson) Dudleya attenuata (S. Johans. Dudleya brittonii D.A. Dudleya candida Britton Dudleya cedrosensis Moran Dudleya cultrata Rose Dudleya guadalupensis Moran Dudleya lanceolata (Nutt.) Britton & Rose Dudleya linearis (Greene) Britt. & Rose Dudleya pachyphytum Moran & Benedict Dudleya ingens Rose PPENDIX New record; pers. comm. Jon Rebman Synonym: Salicornia virginica New record; herbarium specimen @ SD New record; pers. comm. Jon Rebman New record; herbarium specimen @ RSA, SD Moran 20315 Convolvulaceae A Crassulaceae 53 54 55 56 57 Family/species 180 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 NAT CED NAT CED CED CED CED CED? CED NAT CED NAT CED CED CED CED CED CED BEN CED NAT GUA GUA GUA GUA GUA GUA BEN CED NAT GUA GUA MAR TOS TOS MAR COR? COR TOS COR TOS MAR COR TOS MAR COR TOS MAR JER GUA BEN CED NAT (Greene) P.H. Raven (Greene) P.H. ssp./var. and infraname ssp./var. ssp. extima Moran var. minimus var. ssp. argenteus (Dunkle) grandiflorus var. ssp. brevivexillus (Ottley) Brouillet var. insularis var. (A. Gray) Greene ssp. ornithopus 63 59 62 Ottley 60 64 66 65 61 58 (Millsp.) Millsp. Greene ssp. brevivexillus (Benth.) Greene Greene (Greene) Brouillet (Torr. & A. Gray) Ottley (Torr. (Nutt.) Greene (Greene) Greene (Benth.) Greene gambelianus E. Sheld. semiteres (Rose) Moran × 1. Continued Acmispon argophyllus (A. Gray) Brouillet Dudleya Dudleya virens (Rose) Moran Acmispon grandiflorus (Benth.) Brouillet Crossosoma californicum Nutt. Watson S. Echinopepon minimus (Kell.) Marah guadalupensis Greene Arctostaphylos sp . Acalypha californica Benth. Andrachne ciliato-glandulosa (Millsp.) Croizat & A. Gray * Euphorbia albomariginata Torr. Euphorbia bartolomaei Greene Euphorbia crenulata Engelm. Euphorbia misera Benth. Euphorbia pondii Millsp. Marah macrocarpus (Greene) Greene Xylococcus bicolor Nutt. Euphorbia polycarpa Benth. (Vogel) Brouillet Acmispon glaber (Vogel) Acmispon rigidus (Benth.) Brouillet Acmispon strigosus (Nutt.) Brouillet Acmispon flexuosus Sokoloff Acmispon maritimus (Nutt.) D.D. Acmispon nudatus (Greene) Brouillet Astragalus aff. Jones M.E. Astragalus fastidius (Kell.) Astragalus insularis Kell. PPENDIX Synonym: Chamaesyce pondii Synonym & per Jon Rebman may be recognized as: Lotus argophyllus Synonym: Lotus cedrosensis Synonym: Lotus scoparius Synonym: Lotus grandiflorus Synonym: Lotus salsuginosus Synonym: Lotus nudatus Synonym: Lotus rigidus Synonym: Lotus strigosus Fabaceae A 58 59 60 61 62 63 64 65 66 Crossosomataceae Cucurbitaceae Ericaceae Euphorbiaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 181 NAT NAT CED NAT CED? CED CED CED CED CED CED BEN? CED BEN CED NAT BEN? CED NAT 69 GUA? GUA? GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA BEN CED MAR TOS MAR TOS TOS TOS TOS 70 COR TOS COR COR TOS COR COR COR TOS MAR JER GUA BEN CED NAT var. minuta var. (M.E. Jones) Seigler & Ebinger minuta (M.E. var. ssp./var. and infraname ssp./var. magdalenae var. Jones cedrosensis M.E. var. Jones) Barneby lonchus (M.E. var. ssp. concinnus (Greene) Martin ex Isely truncatum (Greene) Martin ex var. ssp. ludoviciana (Greene) C. P. Smith pondii (Greene) C. P. var. 68 * 71 72 Standl. (A. Gray) B.L. Rob. (A. Gray) B.L. (Vasey & Rose) Greene (Vasey 67 1. Continued Britton & Rose Johnst. I.M. Astragalus magdalenae Greene Astragalus nuttallianus DC. Astragalus trichopodus A. Gray Dalea mollis Benth. Ebenopsis confinis (Stand.) Errazurizia benthamii (Brandegee) Lupinus concinnus Agardh. & A. Gray gracilentum Torr. Trifolium Desv. depauperatum Desv. Trifolium microcephalum Pursh Trifolium Watson palmeri S. Trifolium willdenovii Sprengel Trifolium Watson hassei S. Vicia ludoviciana Nutt. Vicia & Arn. farnesiana (L.) Wight Vachellia Quercus cedrosensis C.H. Müll. Quercus tomentella Engelm. S. Watson palmeri S. Frankenia (Molina) I.M. Johnst. salina (Molina) I.M. Frankenia Garrya veatchii Kell. Zeltnera Mans. venusta (A. Gray) G. Lupinus guadalupensis Greene Lupinus succulentus Dougl. Lupinus truncatus Hook. & Arn. Medicago polymorpha L. * Lupinus Watson niveus S. Lupinus sparsiflorus Benth. Melilotus indicus (L.) All. * Phaseolus filiformis Benth. (Vasey & Rose) Brand Syrmatium watsonii (Vasey PPENDIX Synonym: Pithecellobium confine Synonym: Lotus watsonii New record; pers. comm. Jon Rebman New record; herbarium specimen @ SD New record; pers. comm. Jon Rebman, photovoucher @ SD; synonym: Acacia farnesiana Synonym: Centaurium venustum A 67 68 69 70 71 72 Frankeniaceae Garryaceae Gentianaceae Family/species 182 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 NAT NAT CED CED CED CED CED CED CED CED CED? NAT? CED? CED? CED NAT BEN? BEN CED? NAT BEN CED BEN CED? NAT GUA GUA GUA GUA GUA GUA GUA BEN? GUA GUA GUA GUA BEN? CED? NAT JER TOS TOS TOS TOS TOS COR TOS MAR COR TOS MAR COR COR COR TOS? MAR COR TOS MAR JER GUA BEN CED NAT var. nesiotes var. ssp./var. and infraname ssp./var. ssp. assurgentiflora * 74 77 73 75 Kellogg 76 S. Watson S. Moran (A. Gray) Briq. S. Watson S. hortorum L.H. Bailey * × 1. Continued Torr. & A. Gray Torr. (Kellogg) M.F. Ray M.F. Malva assurgentiflora (Kellogg) Ribes tortuosum Benth. Ribes viburnifolium A. Gray Clinopodium palmeri (A. Gray) Kuntze Pogogyne tenuiflora A. Gray Pogogyne Salvia columbariae Benth. glandulosum Kell. Teucrium Eucnide cordata Kell. Mentzelia adhaerens Bentham Watson Mentzelia hirsutissima S. Mentzelia micrantha (Hook. & Arn.) Marrubium vulgare L. * Monardella thymifolia Greene Salvia cedrosensis Greene linearis Greene Petalonyx Ray Malva pacifica M.F. Abutilon californicum Benth. Eremalche exilis (A. Gray) Greene (L.) L’Her. ex Ait. * ex Erodium moschatum (L.) L’Her. Pelargonium Sphaeralcea palmeri Jeps. (L.) L’Her. ex Ait. * ex Erodium cicutarium (L.) L’Her. Erodium texanum A. Gray Sphaeralcea fulva Greene Watson Sphaeralcea sulphurea S. (Cav.) Bertol * Erodium botrys (Cav.) Thell. * Erodium brachycarpum (Godr.) Malva parviflora L. * (S. Watson) M.F. Ray M.F. Watson) Malva occidentalis (S. (Moran) M.F. Ray Malva lindsayi (Moran) M.F. PPENDIX Synonym: Satureja palmeri Synonym: Lavatera lindsayi Synonym: Lavatera occidentalis Synonym: Lavatera venosa Synonym: Lavatera assurgentifolia Grossulariaceae Lamiaceae Loasaceae Malvaceae A Geraniaceae 73 74 75 76 77 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 183 CED CED? CED? NAT CED CED CED CED? NAT GUA BENGUA GUA CED NAT GUA GUA BEN?GUA CED? NAT GUA GUA GUA GUA GUA GUA MAR MAR TOS MAR TOS MAR COR TOS MAR COR TOS MAR COR TOS MAR JER GUA BEN CED NAT (S. Watson) P.H. Raven P.H. Watson) (S. W.L. Wagner & Hoch Wagner W.L. Wagner & Hoch Wagner var. crassifolia (Choisy) Spellenb. var. (Rydberg) Miller (Chambers)ssp. mexicana (Rydberg) COR TOS MAR W.L. Watson) ssp. suffruticosa (S. ssp./var. and infraname ssp./var. ssp. parviflora ssp. cedrosensis (Greene) 84 79 80 83 (Hornem. ex Spreng.) Raim. ssp. suffruticosa Spreng.) (Hornem. ex S. Watson) P.H. Raven ssp. guadalupensis P.H. Watson) S. 78 (Greene) P.H. Raven Raven ssp. cedrosensis (Greene) P.H. P.H. (Torr.) (Nutt. ex Torr. & A. Gray) P.H. Raven & A. Gray) P.H. Torr. (Nutt. ex 81 85 (Greene) P.H. Raven (Greene) P.H. Nutt. P.H. Raven P.H. 82 1. Continued Carolin ex Hershkovitz Carolin ex & Hoch Wagner W.L. Spreng.) ex & Hoch Wagner W.L. & Hoch Wagner Carolin ex Hershk. Carolin ex W.L. Wagner & Hoch Wagner W.L. Claytonia spathulata Douglas Anagallis arvensis L. * Mirabilis laevis (Benth.) Curran Domm.-Smith & RoseXylonagra arborea (Kell.) ssp. wigginsii Castilleja attenuata (A. Gray) Chuang & Heckard Dougl. ex Hook. Claytonia parviflora Dougl. ex & Chambers Claytonia perfoliata Willd. Watson S. Abronia maritima Nutt. ex Hesperelaea palmeri A.Gray Camissoniopsis cheiranthifolia (Hornem. & A. Gray Torr. Eulobus californica Nutt. ex Domm.-Smith & RoseXylonagra arborea (Kell.) ssp. arborea (Ruiz Lopez & Pavon) DC. Calandrinia ciliata (Ruiz Lopez & Pavon) Watson) Camissoniopsis guadalupensis (S. Raven) W.L. Camissoniopsis robusta (P.H. Small ssp. guadalupensis Chylismia cardiophylla (Torr.) & A. Gray) Suksdorf Epilobium foliosum (Torr. Cistanthe guadalupensis (Dudley) Cistanthe maritima (Nutt.) Eulobus crassifolius (Greene) PPENDIX Synonym: Calandrinia maritima New record; herbarium specimen @ RSA, SD Synonym: Camissonia cheiranthifolia Synonym: Camissonia guadalupensis Synonym: Camissonia robusta Synonym: Camissonia cardiophylla Synonym: Camissonia californica Synonym: Camissonia crassifolia Myrsinaceae Orobanchaceae Nyctaginaceae Oleaceae Onagraceae A Montiaceae 78 79 80 81 82 83 84 85 Family/species 184 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CED? CED CED CED CED CED GUA GUA GUA BEN?GUA CED? NAT GUA GUA GUA GUA GUA GUA GUA GUAGUA BEN CED NAT GUA GUA BEN CED NAT GUA GUA TOS TOS TOS TOS? COR TOS MAR COR TOS COR TOS COR TOS MAR COR COR COR TOS MAR JER GUA BEN CED NAT ssp. insularis (Nutt. ex Greene) V.E. Grant Greene) V.E. ssp. abrotanifolia (Nutt. ex COR ssp./var. and infraname ssp./var. ssp. exserta ssp. subinclusa var. heterophylla var. (A. Gray) D.M. Thomps.ssp. subsessile (A. Gray) D.M. COR TOS MAR ssp. pygmaeus 87 92 (L.) Dum. Cours. 93 (A. Gray) G.L. Nesom (Douglas ex Benth.) Spach Nesom (Douglas ex (A. Gray) G.L. 90 86 (Benth.) G. Taylor (Benth.) G. 91 Curtis var. aurantiacus Curtis var. (Brand) J.T. Howell ssp. pygmaeus (Brand) J.T. 89 Douglas ex Benth. Douglas ex A. Gray, Diplacus latifolius A. Gray, (Nutt.) A. Gray (Benth.) Ball Scheele and Linaria canadensis 88 (McMinn ex Munz) Tulig australis (McMinn ex × 1. Continued Chuang & Heckard G.L. Nesom G.L. (Scheele) D.A. Sutton Nuttallanthus texanus (Scheele) D.A. Penstemon cedrosensis Kell. Penstemon Plantago ovata Forssk. (Benth.) A. & V. Grant Allophyllum gilioides (Benth.) A. & V. Grant Gilia angelensis V.E. Gilia capitata Sims Gilia nevinii A. Gray Howell Leptosiphon pygmaeus (Brand) J.T. Castilleja exserta (A. Heller) Diplacus Erythranthe cardinalis Castilleja fruticosa Moran Castilleja guadalupensis Brandegee Castilleja subinclusa Greene Eschscholzia californica Cham. Eschscholzia elegans Greene. Eschscholzia palmeri Rose Eschscholzia ramosa Greene Platystemon californicus Benth. heterophyllum A. Gray Papaver Vasey & Rose Antirrhinum watsonii Vasey Collinsia heterophylla Buist. Sutton Gambelia juncea (Benth.) D.A. Linanthus dianthiflorus (Benth.) Greene Antirrhinum nuttallianum Benth. Gambelia speciosa Nutt. Diplacus brandegeei (Pennell) PPENDIX Synonym: Stylomecon heterophylla Synonym: Mimulus aurantiacus Synonym: Mimulus latifolius Synonym: Mimulus cardinalis Synonym: Galvezia juncea Synonym: Galvezia speciosa Synonyms: Linaria texana Synonym: Linanthus pygmaeus Polemoniaceae A Phrymaceae 86 87 88 89 90 91 92 93 Papaveraceae Plantaginaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 185 NAT CED? CED? CED? CED? CED? CED CED CED CED NAT CED CED CED? CED CED CED? CED? CED? CED? GUA GUA GUA GUA GUA GUA GUA GUA BENGUA GUA CEDGUA NAT TOS COR TOS COR TOS COR TOS COR TOS COR TOS MAR COR TOS MAR JER GUA BEN CED NAT (Trel.) Jeps. (Trel.) A. Gray var. perplexans A. Gray var. ssp. microphylla var. testudinum Reveal var. fruticosa (Greene) Reveal var. var. taxifolium (Greene) Parish var. ssp./var. and infraname ssp./var. Microsteris gracilis (Hook.) Greene (H.J. Thomps.) Reveal ssp. insulare (H.J. 94 95 Goodman 1. Continued Greene & Parry Porter & L.A. Johnson Porter Rhamnus insula Kellogg Rhamnus pirifolia Greene Adenostoma fasciculatum Hook. & Arn. Wats. obtusifolium S. var. Ceanothus verrucosus Nutt. Ziziphus parryi Torr. Eriogonum fasciculatum Benth. Eriogonum grande Greene Eriogonum intricatum Benth. Eriogonum molle Greene Benth. ex Eriogonum pondii Greene Torrey Eriogonum zapatoense Moran Harfordia macroptera (Benth.) Lastarriaea coriacea (Goodman) Hoover Eriogonum wrightii Linanthus uncialis (Brandegee) Moran J.M. Linanthus veatchii (C. Parry) Portulaca oleracea L. * Portulaca Dodecatheon clevelandii Greene Delphinium cardinale Hook. Delphinium parryi A. Gray Myosurus minimus L. Macbr. Oligomeris linifolia (Vahl.) Clematis pauciflora Nutt. Ranunculus hebecarpus H. & A. Ceanothus arboreus Greene Ceanothus crassifolius Torr. Ceanothus cuneatus (Hook.) Nutt. Ceanothus perplexans Trel. F. & M. Pterostegia drymarioides F. PPENDIX Synonym: Chorizanthe coriacea SBBG; synonym: Ceanothus greggii New record; herbarium specimen @ RSA, SD, Rosaceae Polygonaceae A 94 95 Portulacaceae Primulaceae Resedaceae Ranunculaceae Rhamnaceae Family/species 186 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 NAT NAT CED NAT CED CED CED? NAT CED? CED? CED CED NAT CED? BEN CED? BEN? GUA BEN? GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA GUA MAR MAR MAR TOS MAR TOS TOS MAR JER 96 COR COR TOS COR TOS MAR COR TOS MAR COR COR TOS COR COR COR COR COR COR TOS MAR JER GUA BEN CED NAT var. crassifolia var. Hilend & J. Howell eremicum Hilend & J. var. brevipes var. ssp. angustifolium ssp./var. and infraname ssp./var. 1. Continued Lycium exsertum A. Gray Lycium fremontii A. Gray Nicotiana glauca Grah. * Solanum americanum Miller * Solanum nodiflorum Jacq. * Lycopersicon esculentum L. * Nicotiana attenuata Torr. Nicotiana clevelandii A. Gray Physalis crassifolia Benth. Solanum douglasii Dunal Solanum hindsianum Benth. Lycium californicum Nutt. Galium angulosum A. Gray Galium coronadoense Dempster Galium stellatum Kellogg Ruta chalepensis L. Datura discolor Bernh. Datura wrightii Regel Lycium andersonii A. Gray Lycium brevipes Benth. (Lindl.) M. Roem. Heteromeles arbutifolia (Lindl.) M. Galium angustifolium Nutt. Galium aparine L. Aesculus parryi A. Gray Jepsonia malvifolia (Greene) Small Small. Jepsonia parryi (Torr.) Ngaio. * f. Myoporum laetum Forst. Scrophularia villosa Pennell Simmondsia chinensis (Link) C. Schneider Aphanes occidentalis Rydberg PPENDIX New record; herbarium specimen @ SD. Rubiaceae Rutaceae Solanaceae Sapindaceae Saxifragaceae Saxifragaceae Scrophulariaceae Simmondsiaceae A 96 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 187 CED CED? CEDCED NAT CED GUA GUA GUA GUA TOS MAR TOS MAR 129 145 107 8 232 57 266 79 COR COR TOS COR TOS MAR JER GUA BEN CED NAT ssp./var. and infraname ssp./var. ssp. clokeyi var. geniculata var. 97 Green 1. Continued Hesperocnide tenella Torrey Urtica urens L. * Glandularia lilacina (Greene) Umber. Tamarix ramosissima Ledeb. * Tamarix Hinton hespera B.D. Parietaria Vasey & Rose Solanum palmeri Vasey Solanum wallacei (A. Gray) Parish (Kellogg) Greene geniculata (Kellogg) Viscainoa Fagonia laevis Standl. Fagonia Torr. ex Trel. ex Phoradendron densum Torr. PPENDIX Verbena lilacina Synonym: Verbena Urticaceae Verbenaceae Tamaricaceae A Zygophyllaceae Viscaceae UMBER OF SPECIES Family/species 97 N 188 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 ROS CRU 2 ROS CRU CRU CAT ROS CRU ANA CAT CRU ROS CRU CRU CAT CAT ROS CRU CAT CAT ROS CRU 1 3

CRU CLE NIC CAT BAR ROS CRU ANA MIG CLE NIC CAT ROS CRU ANA MIG CLE?

ssp./var. and infraname ssp./var. (D.C. Eaton) Domin(D.C. CLE 2. Plant species checklist for the 8 California Channel Islands (Anacapa [ANA], San Clemente [CLE], San Miguel [MIG], San Nicolas [NIC], Santa Barbara [BAR], Santa D.C. Eaton D.C. Windham, & E. Wollenw. Windham, & E. Wollenw. Windham, & Roy L. Taylor Underw. pubescens Underw. var. Pteridium aquilinum (L.) Kuhn CAT ROS CRU Maxon Dryopteris arguta (Kaulf.) CLE C. Presl munitum (Kaulf.) CAT Polystichum ROS CRU ANA MIG Azolla filiculoides Lam. fimbriata Sm. Woodwardia CLE? (Kaulf.) Yatsk., al. Yatsk., CLE triangularis (Kaulf.) Eaton) Yatsk.et Pentagramma ssp. viscosa (D.C. CAT ROS CRU Selaginella bigelovii Underw. CLE CAT ROS CRU ANA (Kaulf.) Yatsk., Yatsk., triangularis (Kaulf.) Pentagramma ssp. triangularis Equisetum hyemale L. ssp. affine (Engelm.) Calder ROS CRU Equisetum laevigatum A. Braun Equisetum telmateia Ehrh. Milde) Hauke ssp. braunii (J. CAT CRU californicum Kaulf. Polypodium CLE NIC CAT BAR scouleri Hook. & Grev. Polypodium ROS CRU ANA C.A. Paris Adiantum aleuticum (Rupr.) Adiantum capillus-veneris L. Adiantum jordanii Müll. Hal. CLE NIC CAT Aspidotis californica (Hook.) Copel. ROS CRU ANA MIG Eaton Cheilanthes clevelandii D.C. Eaton Cheilanthes cooperae D.C. Cheilanthes newberryi (Kaulf.) Fée andromedifolia (Kaulf.) Pellaea CLE NIC CAT Eaton) ROS mucronata (D.C. mucronata CRU ANA Pellaea var. MIG PPENDIX Pers. com. E. Howe Pers. Ratay collection S. com. E. Howe Pers. Dennstaedtiaceae Dryopteridaceae A Azollaceae Blechnaceae 1 2 3 Selaginellaceae Equisetaceae Polypodiaceae Pteridaceae ERNS F Family/species Catalina [CAT], Santa Cruz [CRU], and Santa Rosa [ROS]). An ampersand (&) indicates an extirpated species; a question mark (?) indicates an indicates a nonnative species. unvouchered taxa; and an asterisk (*) 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 189 CRU CAT ROS CRU 4 ROS CRU ROS CRU CLE NIC CAT BAR ROS CRU ANA MIG

ssp./var. and infraname ssp./var. 2. Continued Rupr. cyclosorum Rupr. (L.) Roth Athyrium filix-femina var. Cystopteris fragilis (L.) Bernh. ROS CRU CRU Bartel * Hesperocyparis macrocarpa (Hartw.) CAT ROS CRU ANA muricata Don var. Pinus muricata D. Pinus pinea L. * Don * Pinus radiata D. Carrière ex CRU Pinus torreyana Parry Haller ssp. insularis J.R. ROS MIG Agave americana L. * Chlorogalum pomeridianum (DC.) Kunth lacunosum Watson var. Allium lacunosum S. Allium praecox Brandegee CLE CAT ROS CRU MIG Amaryllis belladonna L. * Narcissus tazetta L. * ANA MIG CRU Lemna minor L. * Zantedeschia aethiopica (L.) Spreng. MIG filifera (André) de Bary Washingtonia MIG Asparagus asparagoides (L.) Druce * CRU MIG NIC Asparagus officinalis L. * CAT Asphodelus fistulosus L. * CLE Koyama Bolboschoenus maritimus (L.) Palla ssp. paludosus (A. Nelson) T. Carex barbarae Dewey CRU MIG Carex globosa Boott CAT Carex gracilior Mack. ROS CRU Carex harfordii Mack. ROS ROS CRU PPENDIX NY: T.S. Ross, 6137, 04 13 1992 T.S. NY: A Woodsiaceae Cupressaceae Pinaceae Agavaceae Agavaceae Alliaceae Amaryllidaceae Araceae Arecaceae Asparagaceae Asphodelaceae Cyperaceae 4 ONOCOTS YMNOSPERMS G M Family/species 190 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CRU ROS 5 6 7 ROS CLE NIC CAT BAR ROS CRU ANA MIG (Fernald & Weigand) & Weigand) ssp. pacificus (Fernald ROS ssp./var. and infraname ssp./var. ROS CRU 8 E. Mey.; taxonomic issues taxonomic E. Mey.; 2. Continued (Pers.) Schinz & R. Keller (Pers.) Piper & Beattie Eleocharis macrostachya Britton CLE NIC CAT Roem. & Schult. Isolepis cernua (Vahl) ROS Schoenoplectus americanus NIC CAT ROS MIG Soják Schoenoplectus californicus (C.A. Mey.) Sm. longispicatus (Britton) S.G. Palla Schoenoplectus pungens (Vahl) var. ROS CRU ROS CRU Watson Sisyrinchium bellum S. CAT ROS CRU MIG Juncus acutus L. Snogerup ssp. leopoldii (Parl.) CAT Juncus balticus Willd. Snogerup ssp. ater (Rydb.) CAT Juncus bufonius L. ROS CLE NIC MIG CAT Juncus effusus L. ROS CRU ANA MIG Juncus mexicanus Willd. CAT Juncus patens E. Mey. ROS CRU MIG CLE phaeocephalus Juncus phaeocephalus Engelm. var. ROS CRU MIG Juncus textilis Buchenau CAT Juncus xiphioides E. Mey. Buchenau Luzula Watson) subsessilis (S. CAT CRU Carex pansa L.H. Bailey Boott Carex praegracilis W. ROS Carex senta Boott CAT MIG ROS Carex subbracteata Mack. MIG CRU Carex triquetra Boott ROS CRU Carex tumulicola Mack. CAT CLE Cyperus eragrostis Lam. ROS CRU Cyperus esculentus L. CAT CLE CAT Cyperus involucratus Rottb. * Cyperus odoratus L. * CRU CAT Calochortus albus (Benth.) Benth. Watson Calochortus catalinae S. CAT ROS CRU CAT Calochortus luteus Lindl. ROS CRU Calochortus splendens Benth. CRU Lilium humboldtii Duch. CAT Thorne ssp. ocellatum (Kellogg) ROS CRU ANA PPENDIX RSA: Mark Hoefs, Steven A. Junak, Janet Takara, M. Gay, 2384, Jul 14 1995 Gay, M. RSA: Mark Hoefs, Steven A. Junak, Janet Takara, Thorne, 2458, Jul 26 1995 Hoefs, R.F. SBBG: M.L. Jul 26 1995 2482 Thorne, Janet Takara RSA: Mark Hoefs, R.F. Synonym: Luzula comosa Iridaceae Juncaceae A 5 6 7 8 Liliaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 191 ROS CRU 10 CRU CRU CAT CAT ROS CRU

NIC CAT CRU CAT NIC CRU 9 11 13 CLE NIC CAT BAR ROS CRU ANA MIG CLE? NIC CAT ROS CRU ANA MIG ssp./var. and infraname ssp./var. * CAT 15 CLE NIC CAT BAR ROS CRU ANA MIG 12 CLE NIC CAT BAR ROS CRU ANA MIG 14 L. Desv.; Per Jepson Interchange: molecular evidence needed to resolve question of nativity Per Desv.; 2. Continued Trin. Agrostis pallens Trin. CLE CAT Agrostis stolonifera* L. ROS CRU CAT Ammophila arenaria (L.) Link * NIC ANA Campb. scabriglumis C.S. Britton et al. var. Andropogon glomeratus (Walter) Aristida adscensionis L. CLE CRU CAT gentilis (Henrard) Allred var. Aristida ternipes Cav. ROS CRU Arundo donax L. * CRU NIC Avena barbata Link * CLE NIC ROS CRU CAT BAR MIG ROS CRU ANA Avena fatua L. * MIG CLE NIC CAT BAR ROS CRU ANA Avena sativa L. * MIG CLE CAT Herter Bothriochloa barbinodis (Lag.) ROS CRU * Beauv. CAT Brachypodium distachyon (L.) P. CLE? (Greene) Rydb. Piperia michaelii (Greene) Rydb. Aegilops cylindrica Host * Agrostis exarata Trin. *Bromus diandrus Roth * CLE NIC CAT BAR ROS CRU ANA MIG Bromus hordeaceus L. * (Torr.) Rydb. Rydb. fremontii (Torr.) Toxicoscordion ROS CRU ANA MIG Epipactis gigantea Hook. Rydb. Watson) Piperia cooperi (S. CLE? CRU Briza minor L. * Bromus arizonicus (Shear) Stebbins CLE NIC CAT BAR ROS CRU ANA Bromus berteroanus Colla MIG ANA Rydb. Piperia elongata Rydb. CAT Hook. & Arn. var. carinatus Bromus carinatus Hook. & Arn. var. Calamagrostis rubescens Buckley CRU Bromus maritimus (Piper) Hitchc. * Bromus catharticus Vahl CLE ROS CRU ANA MIG Bromus laevipes Shear * CAT Bromus madritensis L. * CLE NIC CAT ROS CRU BAR ROS CRU ANA MIG Bromus vulgaris (Hook.) Shear PPENDIX SBBG: S.A. Junak Apr 30 1998 SCa-568 SBBG: S.A. com. E. Howe Pers. Synonym: Bromus trinii Apr 11 1992 6122 Ross, E. Kellogg SBBG: T.S. Synonym: Bromus mollis UCSB: Dan Hancock 03/30/1961 146 Pers. com. E. Howe Pers. Poaceae Poaceae A Melanthiaceae Orchidaceae 9 10 11 12 13 14 15 Family/species 192 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CRU ROS CRU MIG 18 20 21 22 24 NIC CAT CRU MIG CAT 17 19 16 23 CAT ROS CRU CLE NIC CAT BAR ROS CRU ANA MIG CLE NIC CAT ROS CRU MIG ssp./var. and infraname ssp./var. CLE? 2. Continued Bell & Columbus Asch. & Graebn. * Elymus glaucus Buckley ssp. glaucus Elymus repens (L.) Gould * Presl Elymus condensatus J. CLE NIC CAT ROS CRU ANA MIG MIG Elymus pacificus Gould NIC Elymus triticoides Buckley ROS NIC CAT MIG arundinacea Schreb. * Festuca ROS CRU ANA MIG * Sm. CLE perennis (L.) Columbus & J.P. NIC Festuca CAT BAR ROS CRU MIG * Sm. CLE temulenta (L.) Columbus & J.P. Festuca CAT ROS CRU bromoides L. * Festuca CLE CAT ROS CRU ANA microstachys Nutt. CLE Festuca MIG CAT myuros L. * CRU ANA Festuca CLE NIC MIG CAT BAR ROS CRU ANA MIG octoflora Walter CLE Festuca NIC CAT BAR ROS CRU ANA MIG Gastridium phleoides (Nees & Meyen) C.E. Hubb. * CLE CAT ROS CRU Greuter * Hainardia cylindrica (Willd.) CLE * Chloris virgata Sw. CLE Cortaderia selloana (Schult. & Schult. f.) CLE? Crypsis alopecuroides (Piller & Mitterp.) Schrad. * * Cynodon dactylon (L.) Pers. CLE NIC CAT BAR Dactylis glomerata L. * ROS CRU ANA MIG CLE CAT Danthonia californica Bol. CRU Munro Deschampsia danthonioides (Trin.) CLE ROS * Desmazeria rigida (L.) Tutin Digitaria sanguinalis (L.) Scop. * CAT Dissanthelium californicum (Nutt.) Benth. CAT Distichlis spicata (L.) Greene CAT CLE NIC CAT Distichlis littoralis (Engelm.) H.L. ROS CRU ANA MIG CAT ROS * Beauv. Echinochloa crus-galli (L.) P. CLE CAT Ehrharta calycina* Sm. CLE CAT Ehrharta erecta Lam. * CAT (Forssk.) Opiz * Crypsis vaginiflora (Forssk.) CAT PPENDIX CAS: Steven A. Junak 9 Nov 1990 SCI-143 com. E. Howe Pers. Gay Jul 27 1995 2491 M. Thorne, Janet Takara, RSA: Mark Hoefs, R.F. com. E. Howe Pers. H. Raven 22 May 1962 17826 CAS: Peter UCD: Beecher Crampton 08 10 1968 8322 Junak Jun 13 1998 SCa-743 SSBG: S.A. Junak May 14 1985 SCl-71 SBBG: S.A. L. Hoefs Jun 4 1997 SCa-421 Junak, M. SBBG: S.A. A 16 17 18 19 20 21 22 23 24 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 193 ROS CRU ANA BAR ROS CRU ANA MIG CRU ANA ROS CRU ANA MIG 25 2 7 30 31 33 CRU ROS MIG NIC CAT NIC ROS CRU CAT BAR ROS CRU ANA MIG NIC CAT 26 28 29 32

CLE CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. 2. Continued Bothmer et al. MIG (Scribn. & J.G. Sm.) Rydb. Sm.) Rydb. Hordeum depressum (Scribn. & J.G. Hordeum intercedens Nevski CLE CRU Hordeum marinum Huds. * Thell. ssp. gussoneanum (Parl.) CLE annua L. * Poa CLE NIC CAT BAR ROS CRU ANA palustris L. * Poa MIG & Scribn. howellii Vasey Poa CAT douglasii Nees Poa Presl secunda J. Poa ssp. secunda * viridis (Gouan) Breistr. Polypogon CAT BAR ROS CRU ANA * interruptus Kunth Polypogon CLE CAT ROS CRU MIG Hordeum murinum L. * ssp. glaucum (Steud.) Tzvelev CLE NIC CAT BAR ROS CRU ANA Hordeum murinum L. * MIG CLE ssp. leporinum (Link) Arcang. NIC CAT BAR ROS CRU ANA MIG Hordeum vulgare L. * CLE NIC CAT ROS CRU Koeleria macrantha (Ledeb.) Schult. Shinners * Koeleria gerardii (Vill.) ROS CRU Lamarckia aurea (L.) Moench * CLE NIC CAT BAR CRU ROS Melica imperfecta Trin. CRU ANA MIG CLE NIC CAT BAR ROS CRU ANA Goodd. Muhlenbergia appressa C.O. MIG CLE Muhlenbergia microsperma (DC.) Kunth CLE incurva (L.) C.E. Hubb. * Parapholis CLE NIC CAT BAR ROS * CRU ANA dilatatum Poir. Paspalum CLE MIG CAT distichum L. Paspalum CAT * clandestinum Chiov. Pennisetum CRU * NIC CLE Chiov. setaceum (Forssk.) Pennisetum CAT ROS CRU ANA MIG Hordeum brachyantherum Nevski ssp. californicum (Covas & Stebbins) NIC CAT Phalaris aquatica L. * NIC CAT Phalaris canariensis L. * CRU ANA CAT * Phalaris caroliniana Walter CLE NIC CAT BAR Phalaris lemmonii Vasey CRU CLE CAT ROS Phalaris minor Retz. * CLE NIC CAT BAR ROS CRU ANA Phalaris paradoxa* L. CLE MIG NIC CAT PPENDIX SBBG: L.W. Nuttall May 12 1920 190 SBBG: L.W. Munz Apr 8 1923 6613 P.A. POM: Junak May 3 2001 SCa-1464 SBBG: S.A. Raven May 10 1962 17738 SBBG: P.H. CAS: E.R. Blakley 7 Dec. 1963 6327 Ratay collection S. Hoefs Jun 5 1997 SCa-427 Junak, M.L. SBBG: S.A. H. Raven Apr 12 1962 17332 UC: Peter Mar 20 1931 S4310 Fosberg UC: F.R. A 25 26 27 28 29 30 31 32 33 Family/species 194 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 43 CRU ROS CRU ANA BAR CRU ROS CRU ANA MIG 35 36 40 42 CAT ROS CRU ANA CAT ROS CRU NIC CAT ROS CRU MIG NIC CAT 37 41 34 39 CAT ROS CRU CLE NIC CAT CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. ? CLE 38 2. Continued Phyllospadix scouleri Hook. CLE NIC BAR CAT Watson Phyllospadix torreyi S. CRU ANA MIG CLE NIC BAR CAT ROS CRU ANA MIG Zostera marina L. CLE? Schismus barbatus (L.) Thell. * CAT * Kuntze Stenotaphrum secundatum (Walter) CRU NIC Stipa diegoensis Swallen NIC Stipa cernua Stebbins & Love CLE NIC CAT ROS CRU ANA MIG ROS CRU ANA Stipa lepida Hitchc. MIG CLE NIC BAR CAT ROS CRU ANA Stipa pulchra Hitchc. MIG CLE NIC BAR CAT ROS CRU ANA miliacea MIG Stipa miliacea (L.) Hoover * var. aestivum L. * Triticum CLE NIC crispus L. * Potamogeton pectinata (L.) Börner Stuckenia CAT CLE? Typha latifolia L. CLE Typha NIC CAT CRU Zannichellia palustris L. CAT * monspeliensis (L.) Desf. Polypogon CLE NIC BAR CAT ROS CRU ANA MIG Schismus arabicus Nees * CLE S. Watson Watson Zostera pacifica S. NIC CAT Ruppia maritima L. CLE NIC CAT ROS CRU crocea Coville var. Bloomeria crocea (Torr.) Brodiaea jolonensis Eastw. Niehaus Brodiaea kinkiensis T.F. CLE CAT ROS CRU MIG Dichelostemma capitatum (Benth.) Alph. Wood CLE NIC BAR CAT ROS CRU ANA MIG clementina Hoover Triteleia CLE hyacinthina (Lindl.) Greene Triteleia angustifolia L. Typha CRU Pers. domingensis Pers. Typha CLE PPENDIX RSA: E. Kellogg Jun 12 1992 s.n. RSA: E. Kellogg Junak Mar 22 1998 SCa-512 SBBG: S.A. Thorne Sep 14, 1966 36643 SD: Robert F. com. E. Howe Pers. com. E. Howe Pers. H. Raven Jul 11 1962 18018 SBBG: P. May 12 1932 Wolf UC: Carl B. com. E. Howe Pers. Thorne Sep 16 1966 36714 RSA: R.F. Zosteraceae Potamogetonaceae Zannichelliaceae A 34 35 36 37 38 39 40 41 42 Ruppiaceae Themidaceae Typhaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 195 ROS CRU ANA 45 CAT CRU NIC CAT NIC ANA 47 44 46 ssp./var. and infranamessp./var. CLE NIC CAT BAR ROS CRU ANA MIG 2. Continued ICOTS J.M. Coult. & Rose J.M. D (Kensit) L. Bolus NIC Delosperma litorale* (Kensit) Malephora crocea* (Jacq.) Schwantes CLE Mesembryanthemum crystallinum L. * CLE NIC BAR CAT ROS CRU ANA MIG Mesembryanthemum nodiflorum L. * CLE NIC BAR CAT ROS CRU ANA Sesuvium verrucosum Raf. MIG CLE * Kuntze tetragonioides (Pall.) Tetragonia NIC ROS CRU ANA MIG Amaranthus albus L. * CAT Watson Amaranthus blitoides S. ROS CRU ANA Amaranthus deflexus L. * ROS CRU CAT Watson Amaranthus powellii S. CRU Malosma laurina (Nutt.) Abrams CLE CRU ANA CAT Rhus integrifolia (Nutt.) Rothr. CLE CAT Watson Rhus ovata S. ROS CRU ANA CLE? MIG Anemopsis californica (Nutt.) Hook. & Arn. CLE NIC CAT CRU Sambucus nigra L. Bolli ssp. caerulea (Raf.) CLE CAT ROS CRU * Br. Carpobrotus chilensis (Molina) N.E. CLE NIC BAR CAT ROS CRU ANA MIG Br. Carpobrotus edulis* (L.) N.E. CLE Schinus molle L. * CLE NIC CAT & A. Gray) Greene diversilobum (Torr. CLE ROS CRU Toxicodendron NIC CAT ROS CRU ANA MIG Apiastrum angustifolium Nutt. CLE NIC BAR CAT ROS CRU ANA MIG Apium graveolens L. * CLE NIC Berula erecta (Huds.) Coville ROS CRU NIC Bowlesia incana Ruiz & Pav. CLE CRU MIG CAT Conium maculatum L. * ROS CRU NIC CAT CRU Daucus carota L. * NIC Daucus pusillus Michx. CLE NIC BAR CAT ROS CRU ANA vulgare Mill. * MIG Foeniculum CLE NIC CAT ROS CRU Lomatium caruifolium (Hook. & Arn.) MIG ROS CRU MIG PPENDIX RSA: M.B. Dunkle Aug 20 1940 7671 Dunkle Aug RSA: M.B. Raven Jul 12 1962 18045 RSA: P.H. 4732, Apr 29 1931 Fosberg, F.R. POM: 26 1997 SCl-936 Junak Aug SBBG: S.A. com. E. Howe Pers. Amaranthaceae A Saururaceae Adoxaceae Aizoaceae 43 44 45 46 47 Apiaceae UDICOTS ASAL Family/species B E 196 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CRU ROS CRU 50 51 CAT CAT CAT BAR 48 49 53 ROS CRU ANA ssp./var. and infranamessp./var. CLE NIC CAT BAR ROS CRU ANA MIG CLE CAT ROS CRU MIG 52 DC. 2. Continued Shan & Constance Sanicula hoffmannii (Munz) R.H. ROS CRU arvensis (Huds.) Link * Torilis nodosa (L.) Gaertn. * Torilis NIC CAT CRU ROS CRU microcarpa (Hook. & Arn.) Koso-Pol. MIG Yabea CLE Acourtia microcephala DC. CAT apargioides Agoseris apargioides (Less.) Greene var. ROS CRU Agoseris grandiflora (Nutt.) Greene Agoseris heterophylla (Nutt.) Greene ROS CRU MIG Amblyopappus pusillus Hook. & Arn. CLE NIC CAT ROS BAR ROS CRU ANA MIG Ambrosia chamissonis (Less.) Greene CLE NIC CAT Ambrosia psilostachya DC. ROS CRU ANA NIC CAT MIG ROS Anthemis cotula L. * MIG CAT Arctium minus (Hill) Bernh. * ROS CRU Argyranthemum frutescens (L.) Sch. Bip. * ROS CAT Artemisia californica Less. CLE NIC CAT BAR ROS CRU ANA Artemisia douglasiana Besser MIG CAT Artemisia dracunculus L. ROS CRU CAT Raven Artemisia nesiotica P.H. CLE NIC BAR Baccharis glutinosa Pers. Asclepias fascicularis Decne. CAT Krings hartwegii (Vail) cynanchoides (Decne.) Schltr. var. Funastrum ROS CRU CAT major L. * Vinca CAT CRU Achyrachaena mollis Schauer CLE CAT (Eastw.) Munz Lomatium insulare (Eastw.) CLE NIC Coult. & Rose Sanicula arguta J.M. CLE NIC CAT ROS CRU ANA Sanicula crassicaulis DC. MIG CLE? Baccharis pilularis DC. CLE NIC CAT BAR ROS CRU ANA Baccharis plummerae A. Gray ssp. plummerae MIG Torr. & A. Gray Baccharis salicina Torr. CLE PPENDIX Pers. com. E. Howe Pers. Munz Apr 9 1923 6652 P.A. POM: Thorne 04 28 1966 36224 UCD: Robert F. Stacey Otte 24 April 1993 6946 Takara, Ross, J. CAS: Tim Synonym: Baccharis douglasii 22, 1949 11961 Feb. DS: Ira L. Wiggins Apocynaceae Asteraceae Achillea millefolium L. CLE NIC CAT BAR ROS CRU ANA MIG A 48 49 50 51 52 53 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 197 ROS CRU ANA MIG 59 ROS CRU ANA MIG ROS MIG ROS CRU ROS CRU MIG ROS CRU ANA MIG 55 58 60 61 64 65 68 NIC CAT NIC CAT CAT CRU ANA NIC CAT CAT BAR ROS CRU ANA 67 54 57 62 63 66 CRU CLE ssp./var. and infranamessp./var. CLE NIC CAT BAR ROS Hall lanosa (DC.) H.M. var. CRU ROS ANA MIG ? 56 2. Continued Schilling & Panero & C.E. Turner Keil D.J. (Torr. & A. Gray) Gray californica (Torr. Brickellia CLE (Hook.) S.F. Blake var. nanum Blake var. Blennosperma nanum (Hook.) S.F. (Ruiz & Pav.) Pers. Pers. Baccharis salicifolia (Ruiz & Pav.) CLE Centromadia fitchii (A. Gray) Greene Chaenactis glabriuscula CRU Cichorium intybus L. * Cirsium brevistylum Cronquist CRU Cirsium occidentale (Nutt.) Jeps. californicum (A. Gray) var. CAT CRU Cirsium occidentale (Nutt.) Jeps. occidentale var. CLE Cirsium occidentale (Nutt.) Jeps. venustum (Greene) var. Cirsium ochrocentrum A. Gray * ROS * Cirsium vulgare (Savi) Ten. ROS CAT Constancea nevinii (A. Gray) B.G.Baldwin ROS CRU CLE MIG CAT BAR Corethrogyne filaginifolia (Hook. & Arn.) Nutt. CAT Baccharis sarothroides A. Gray CAT Bahiopsis laciniata (A. Gray) E.E. CLE Centaurea melitensis L. * CLE NIC CAT BAR Cirsium occidentale (Nutt.) Jeps. coulteri var. * Hook. f. Cotula australis (Spreng.) CLE Calendula officinalis L. * NIC Carduus pycnocephalus L. * CAT Cotula coronopifolia L. * NIC CAT Cynara cardunculus L. * CLE? ROS CRU ANA MIG Centaurea solstitialis L. * NIC CAT Centaurea benedicta (L.) L. * CRU Centromadia pungens (Hook. & Arn.) Greene CAT CRU PPENDIX SD: Peter H. Raven May 08, 1962 17629 SD: Peter HSC: James Henrickson Jan 1973 8113 com. E. Howe Pers. CAS: Steven A. Junak 2 Oct 1996 SCI- 703 Philbrick, SCa-1532, May 18 2001 Hoefs, R.N. Junak, M.L. SBBG: S.A. Thorne, 37492, Apr 28 1968, suspected incorrect georeferencing RSA: R.F. RSA:Mark Hoefs, 309, Jun 4 1973 Everett Jun 23, 1965 35015 Thorne, P. SD: Robert F. Hammitt May 20 1991 5325 Mistretta, M. Ross, O. SBBG: T. Ross, Laurel Arnseth Apr 7 1990 4271 UCR: Steve Boyd, T.S. Apr 29 1931 S-4709 Fosberg UCR: F.R. SBBG: E.R. Blakley Sep 23 1961 4740 25 1997 SCl-724 Junak Feb SBBG: S.A. com. E. Howe Pers. Hoefs, K. Kirkland Junak, M.L. SBBG: S.A. A 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 Family/species 198 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 BAR ROS CRU ANA MIG ROS CRU ANA MIG CRU ANA BAR ROS CRU ANA MIG 70 72 73 77 79 ROS 75 NIC CAT NIC CAT BAR NIC CAT ROS 69 71 76 78 80 ROS CRU CAT ROS CRU ANA MIG CLE CAT CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. CLE NIC CAT CRU 74 L. 2. Continued Erigeron canadensis L. CLE Nutt. var. foliosus Erigeron foliosus Nutt. var. Gawl. Erigeron glaucus Ker Watson Erigeron sanctarum S. ROS CRU ANA MIG Eriophyllum confertiflorum (DC.) A. Gray confertiflorum var. ROS CRU Eriophyllum staechadifolium Lag. Nesom Eurybia radulina (A. Gray) G.L. ROS CRU ANA MIG Euthamia occidentalis Nutt. ROS CRU Gamochaeta ustulata (Nutt.) Holub CAT ANA Glebionis coronaria (L.) Spach * ROS CRU ANA MIG Helminthotheca echioides (L.) Holub * NIC CAT Hesperevax sparsiflora (A. Gray) Greene CRU CLE Grindelia stricta DC. Lane platyphylla (Greene) M.A. var. Hazardia cana (A. Gray) Greene ROS CRU ANA CLE MIG? Hazardia detonsa (Greene) Greene Clark Hazardia squarrosa (Hook. & Arn.) Greene grindelioides (DC.) W.D. var. ROS CRU ANA CAT Hedypnois cretica (L.) Dum. Cours. * ROS CRU ANA MIG CLE Helenium puberulum DC. CAT (Brandegee) B.G. Baldwin Deinandra clementina (Brandegee) B.G. CLE NIC CAT BAR CRU ANA Deinandra fasciculata (DC.) Greene CLE CAT BAR ROS Baldwin CRU ANA B.G. Keck) ssp. increscens Deinandra increscens (D.D. Delairea odorata Lem. * Encelia californica Nutt. CLE ROS NIC CAT BAR ANA ROS CRU ANA Ericameria ericoides (Less.) Jeps. Nesom Hall) G.L. pachylepis (H.M. Hall Ericameria palmeri (A. Gray) H.M. var. CAT CRU Erigeron bonariensis L. * CLE Helianthus annuus L. * CLE Gnaphalium palustre Nutt. CLE NIC CAT Grindelia camporum Greene CRU CAT Grindelia hirsutula Hook. & Arn. ROS CRU ANA NIC PPENDIX SBBG: S.A. Junak, SCl-681, Sep 29 1996 SBBG: S.A. 5364, Jul 10 1931 Fosberg, F.R. POM: Junak Oct 28 1997 SCl-981 SBBG: S.A. Bushman Oct 21 1998 SCa-813 Junak, K. Kirkland, M. SBBG: S.A. Propst Mar 14 1973 119 RSA: Mark Hoefs, D. Synonym: Chrysanthemum coronarium Jun 29 1978 Beauchamp and H.A. Wier UC: R.M. CAS: Steven A. Junak 16 May 1996 SCI-470 McClatchie Sept. 1893 s.n. DS: A.J. Beauchamp Sep 15 1979 14 R.M. SBBG: H.L. Ferguson, Nuttall, s.n., May 22 1920 L.W. POM: Junak Mar 13 1997 SCl-784 SBBG: S.A. A 69 70 71 72 73 74 75 76 77 78 79 80 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 199 ROS CRU ANA MIG ROS CRU MIG 81 83 84 85 NIC CAT NIC CAT ROS CRU ANA MIG CAT BAR MIG CAT ROS CRU 82 86 87 88 CLE CLE NIC CAT BARCLE ROS CRU ANA MIG ssp./var. and infraname ssp./var. 2. Continued (Nutt.) G.L. Nesom vernonioides (Nutt.) G.L. Nesom Isocoma menziesii (Hook. & Arn.) G.L. var. CLE NIC CAT ROS CRU ANA MIG Jaumea carnosa (Less.) A. Gray CAT Lactuca saligna L. * ROS CRU MIG CAT Lactuca serriola L. * CLE Laennecia Nesom coulteri (A. Gray) G.L. CLE Lasthenia californica Lindl. CLE CAT BAR Lasthenia glabrata Lindl. ssp. coulteri (A. Gray) Ornduff ROS CRU ANA MIG LastheniaROS gracilis (DC.) Greene NIC CAT MIG A. Gray & C.A. Mey.) Layia platyglossa (Fisch. CLE CAT Lepidospartum squamatum (A. Gray) A. Gray ROS CRU MIG Leptosyne gigantea Kellogg CLE NIC CRU CAT BAR ROS CRU ANA Logfia arizonica (A. Gray) Holub MIG CLE CAT Logfia filaginoides (Hook. & Arn.) Morefield BAR CLE CAT BAR Logfia gallica (L.) Coss. & Germ. * ROS CRU ANA MIG CLE Madia exigua (Sm.) A. Gray CAT Madia gracilis (Sm.) Applegate CRU CAT Madia sativa Molina * CRU CLE CAT Malacothrix clevelandii A. Gray CRU & A. Gray Malacothrix coulteri Harv. ROS Malacothrix foliosa A. Gray Davis ssp. crispifolia W.S. ROS CRU Malacothrix foliosa A. Gray ssp. foliosa ANA Malacothrix foliosa A. Gray Davis ssp. philbrickii W.S. BAR Malacothrix foliosa A. Gray Davis NIC ssp. polycephala W.S. & A. Gray Malacothrix incana (Nutt.) Torr. CLE NIC Malacothrix indecora Greene ROS MIG Davis Malacothrix junakii W.S. ROS CRU MIG ANA Heterotheca grandiflora Nutt. CLE NIC CAT Hieracium argutum Nutt. ROS CRU Hypochaeris glabra L. * CLE ROS CRU CAT Hypochaeris radicata L. * ROS CRU ANA CLE NIC Nesom decumbens (Greene) G.L. Nesom Isocoma menziesii (Hook. & Arn.) G.L. var. CLE CAT menziesii Nesom Isocoma menziesii (Hook. & Arn.) G.L. var. Nesom sedoides (Greene) G.L. Nesom Isocoma menziesii (Hook. & Arn.) G.L. var. CAT PPENDIX Collections of this taxa have been made on Catalina, but there are taxonomic issues Collections of this taxa have been made on Catalina, but there are taxonomic 25 1894 Brandegee Aug UC: T.S. SD: E.R. Blakley Sep 24, 1961 4781 Apr 29 1931 S4707 Fosberg UC: F.R. Junak, K. Kirkland Jul 17 1998 SCa-777 SBBG: S.A. RSA: Steven A. Junak Oct 2 1996 SCI700 Junak Sep 28 1996 SCl-673 SBBG: S.A. Junak May 15 1985 SCl-74 SBBG: S.A. A 81 82 83 84 85 86 87 88 Family/species 200 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 ROS CRU ANA MIG 93 ROS CRU ANA ROS CRU 90 91 92 NIC CAT BAR ROS CRU ANA MIG NIC CAT CAT BAR ROS CRU ANA 89 94 95 96 CAT CLE NIC CLE CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. A. Gray CLE CAT 2. Continued K.L. Chambers K.L. Chambers Turner & T.M. Barkley & T.M. Turner Fisch. & C.A. Mey. var. californicus var. & C.A. Mey. Micropus californicus Fisch. Microseris douglasii (DC.) Sch. Bip. ssp. douglasii Microseris douglasii (DC.) Sch. Bip. ssp. tenella (A. Gray) NIC ROS CRU MIG Microseris douglasii (DC.) Sch. Bip. ssp. platycarpha (A. Gray) CLE CAT Microseris elegans A. Gray CLE NIC Raven CLE Johnst.) P.H. Munzothamnus blairii (Munz & I.M. CRU MIG lyonii A. Gray Pentachaeta CAT emoryi Torr. Perityle CLE CAT BAR ROS CRU ANA Pluchea odorata (L.) Cass. CAT Pluchea sericea (Nutt.) Coville CRU CAT biolettii Anderb. Pseudognaphalium CLE NIC CAT BAR Pseudognaphalium californicum (DC.) Anderb. Pseudognaphalium CLE W.S. Davis & P.H. Raven Davis & P.H. Malacothrix similis W.S. Malacothrix squalida Greene CRU MIG Matricaria occidentalis Greene CRU ANA CAT beneolens (Davidson) Anderb. Pseudognaphalium CLE NIC CAT ROS CRU MIG microcephalum (Nutt.) Anderb. Pseudognaphalium CLE Burtt * luteoalbum (L.) Hilliard & B.L. Pseudognaphalium CLE NIC CAT BAR ROS CRU ANA MIG ramosissimum (Nutt.) Anderb. Pseudognaphalium Anderb. stramineum (Kunth) Pseudognaphalium CRU NIC CAT ROS brevissimus Nutt. CRU ANA Psilocarphus brevissimus var. MIG (Eastw.) H.M. Hall H.M. implicata (Eastw.) & A. Gray var. Malacothrix saxatilis (Nutt.) Torr. NIC tenuifolia (Nutt.) A. Gray ROS & A. Gray * var. CLE CRU ANA Malacothrix saxatilis (Nutt.) Torr. MIG Matricaria discoidea DC. * CAT tenellus Nutt. Psilocarphus CLE CAT Rafinesquia californica Nutt. ROS CRU CLE CAT * BAR Senecio glomeratus Poir. ROS CRU ANA Senecio aphanactis Greene CRU MIG CAT Senecio flaccidus Less. douglasii (DC.) B.L. CLE var. ROS CRU CAT CRU Senecio lyonii PPENDIX RSA: Steven A. Junak, Mary C. Hochberg, H.L. Ferguson Jul 28 1981 sci13 H.L. Ferguson RSA: Steven A. Junak, Mary C. Hochberg, Thorne Apr 29, 1966 36278 SD: Robert F. Thorne Apr 29 1966 36290 SBBG: R.F. Otte Apr 24 1993 6957 S. Ross, Janet Takara, RSA: Tim flora checklist NPS Park Ross Apr 16 1992 6207 UC: Tim Beauchamp March 19, 1967 315 R. M. SDSU: Ross, Orlando Mistretta, Mike Hammitt May 18 1991 5184 RSA: Tim A 89 90 91 92 93 94 95 96 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 201 102 ROS CRU 97 ROS CRU CRU ROS CRU MIG 99 101 103 CAT ROS CRU CAT CRU ANA MIG 100 107 CLE CAT ROS CRU CLE CAT CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. CLE NIC CAT ROS CRU ANA 98 CLE NIC CAT BAR ROS CRU ANA MIG 104 NIC CAT CRU CAT BAR& ROS CRU MIG 106 (Nutt.) K.L. Chambers (DC.) Sch. Bip. 105 2. Continued Stephanomeria virgata Benth. ssp. virgata Stylocline gnaphaloides Nutt. CLE (Nees) G.L. Nesom Symphyotrichum chilense (Nees) G.L. Nesom Symphyotrichum subulatum (Michx.) G.L. CLE ROS CRU erythrospermum Besser * Taraxacum ROS CRU CAT spectabilis var. & C.A. Mey. CLE Amsinckia spectabilis Fisch. NIC BAR ROS CRU ANA MIG intermedia Cryptantha intermedia (A. Gray) Greene var. Solidago velutina DC. ssp. californica (Nutt.) Semple CAT ROS CRU Sonchus asper (L.) Hill * CLE NIC CAT Sonchus oleraceus L. * ROS CRU ANA CLE MIG NIC CAT BAR ROS CRU ANA Sonchus tenerrimus L. * MIG CLE NIC CAT BAR Stebbinsoseris heterocarpa (Nutt.) K.L. Chambers Senecio vulgaris L. * CLE NIC CAT Silybum marianum (L.) Gaertn. * ROS CRU ANA NIC MIG CAT BAR * Wigg; officinale F.H. Taraxacum NIC CAT Xanthium strumarium L. Venegasia carpesioides DC. Venegasia Xanthium spinosum L. ROS CRU Batis maritima L. CLE Berberis pinnata Lag. ssp. insularis Munz ROS CRU ANA & C.A. Mey. Amsinckia intermedia Fisch. CLE NIC CAT BAR CLE Macbr. ROS CRU ANA Amsinckia menziesii (Lehm.) A. Nelson & J.F. MIG * Kuntze Thelesperma megapotamicum (Spreng.) CAT porrifolius L. * Tragopogon CLE NIC Uropappus lindleyi (DC.) Nutt. CRU Stephanomeria cichoriacea A. Gray Stephanomeria diegensis Gottlieb ROS CRU CLE CAT Stephanomeria exigua Benth. ssp. coronaria (Greene) Gottlieb ROS CRU CAT RSA: Orlando Mistretta Apr 5 1992 206 Thorne Sep 15, 1966 36698 SD: Robert F. Everett, 36823, Apr 18, 1967 Thorne, P. SD: Robert F. Thorne Apr 6 1996 2665 Hoefs, R.F. SBBG: M.L. Synonym: Microseris linearifolia Native status on islands unknown Native status on islands unknown Ekhoff May 16 2001 s.n. UCR: J.R. PPENDIX SBBG: R.N. Philbrick Mar 18 1968 B68-20 SBBG: R.N. Synonym: Microseris heterocarpa 4460, Apr 3 1931 Fosberg, RSA: F.R. A 97 98 99 100 101 102 103 104 105 106 107 Bataceae Berberidaceae Boraginaceae Family/species 202 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 114 BAR CRU 108 109 110 115 NIC CAT ROS CRU ANA 111 112 CLE CAT CRU CAT CLE CAT BAR ROS CRU ANA MIG ROS MIG CLE? CAT BAR CRU CLE NIC CAT BAR ROS CRU ANA MIG CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. CRU 113 (J.F. Macbr.) J.T. Howell J.T. Macbr.) (J.F.

Greene var. hubbyi Greene var. 2. Continued Tidestr. Johnst. ex I.M. A. Nelson & J.F. Macbr. A. Nelson & J.F. Plagiobothrys canescens Benth. CLE CAT ROS CRU ANA Pectocarya penicillata (Hook. & Arn.) A. DC. Pectocarya Howell CAT CLE hispida J.T. Phacelia cicutaria Greene var. CAT CRU Garrison L.M. Macbr.) ROS CRU ANA Phacelia hubbyi (J.F. Phacelia distans Benth. Phacelia floribunda Greene CLE Phacelia grandiflora (Benth.) A. Gray CAT ROS CRU Cryptantha clevelandii Greene CLE CAT Greene BAR & C.A. Mey.) ROS Cryptantha leiocarpa (Fisch. CRU ANA MIG Cryptantha maritima (Greene) Greene CLE ROS NIC CAT MIG Johnst. Cryptantha wigginsii I.M. CAT Emmenanthe penduliflora Benth. penduliflora var. Eriodictyon traskiae Eastw. ssp. traskiae Eucrypta chrysanthemifolia (Benth.) Greene chrysanthemifolia var. Harpagonella palmeri A. Gray CAT oculatum (A. Heller) Heliotropium curassavicum L. var. CLE NIC CAT ROS CRU ANA MIG Nama stenocarpum A. Gray CLE Nemophila menziesii Hook. & Arn. CAT Nemophila pedunculata Benth. NIC DC. Johnst.) Thorne CLE ssp. ferocula (I.M. linearis (Ruiz & Pav.) Pectocarya ROS CRU MIG insularis Phacelia insularis Munz var. Phacelia lyonii A. Gray CLE Lehm. Phacelia ramosissima Douglas ex CAT Phacelia viscida (Lindl.) Torr. ROS CRU MIG CAT Pholistoma auritum (Lindl.) Lilja auritum var. ROS CRU ANA MIG Pholistoma racemosum (A. Gray) Constance CLE CAT BAR Johnst. Plagiobothrys acanthocarpus (Piper) I.M. CRU CAT Cryptantha micromeres (A. Gray) Greene CAT ROS Cryptantha microstachys (A. Gray) Greene CRU CAT Cryptantha muricata (Hook. & Arn.) CRU Johnst. CLE Cryptantha traskiae I.M. NIC CAT UC: T.S. Brandegee May 20 1890 UC: T.S. Douglas Mar 24, 1972 3222 SD: R. Mitchel Beauchamp, M. Simpson, Lori L. Simpson April 21 2012 3682 UC: Michael G. Jun 28 1992 s.n. RSA: E. Kellogg Junak Mar 7 1997 SCl-732 SBBG: S.A. Synonym: Phacelia cicutaria 1985 but not NPS list In Wallace Ross Apr 22 1993 6895 UCR: T.S. PPENDIX A 108 109 110 111 112 113 114 115 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 203 118 CRU MIG 117 120 123 NIC BAR ROS CRU ANA MIG CAT NIC CAT MIG 122 116 125 CLE NIC CAT BAR ROS CRU ANA MIG (Thell.) C.L. Hitchc. (A. Gray) Higgins CLE var. robinsonii var. ssp./var. and infraname ssp./var. CLE CAT BAR ROS CRU ANA MIG 119 * CLE NIC CAT ROS CRU MIG 121 (Greene) Thell.; Lepidium virginicum NIC CAT CRU 126 * ssp. menziesii (DC.) Thell. CLE CAT CRU var. pubescens var. (Hook. & Arn.) Greene 124 (Desf.) Benth. (Desf.) 2. Continued Hornungia procumbens (L.) Hayek CLE? Lepidium draba L. * CAT Lepidium lasiocarpum Nutt. ROS CRU CLE NIC CAT Lepidium latipes Hook. CLE CAT Lepidium nitidum Nutt. CRU CLE NIC CAT BAR Lepidium ROS oblongum Small CRU ANA MIG CLE NIC CAT Rattan * Lepidium Watson) strictum (S. ROS CRU ANA MIG CAT (Phil.) I.M. Johnst. Plagiobothrys collinus (Phil.) I.M. Johnst.) Higgins gracilis (I.M. var. CLE CAT Johnst. Plagiobothrys collinus (Phil.) I.M. californicus var. CRU ANA MIG Athysanus pusillus (Hook.) Greene CLE CAT Boechera hoffmannii (Munz) Al-Shehbaz CRU * Koch Brassica nigra (L.) W.D.J. CLE NIC CAT BAR ROS CRU ROS CRU ANA MIG Brassica tournefortii Gouan * NIC Brassica rapa L. * CLE CAT Cakile edentula (Bigelow) Hook. * ROS CRU MIG NIC Cakile maritima Scop. * ROS CLE NIC CAT MIG Capsella bursa-pastoris (L.) Medik. * ROS CRU ANA MIG CLE CAT Cardamine californica (Nutt.) Greene ROS CRU MIG CAT Cardamine oligosperma Nutt. ROS CRU ANA MIG Caulanthus lasiophyllus (Hook. & Arn.) Payson CRU Lepidium virginicum L. (Walter) Britton Descurainia pinnata (Walter) CLE CAT Dithyrea maritima (Davidson) Davidson ROS CRU ANA NIC CAT * Lobularia maritima (L.) Desv. CLE W.T. Aiton Nasturtium officinale W.T. Nutt. ex Torr. & A. Gray CLE NIC Torr. Draba cuneifolia Nutt. ex CAT CRU Erysimum ammophilum A. Heller Erysimum cheiri (L.) Crantz * ROS CAT Erysimum insulare Greene MIG Hirschfeldia incana (L.) Lagr.-Fossat ROS CRU ANA MIG POM: P.A. Munz Apr 10 1923 6705 P.A. POM: Propst, R. Carolin, 36761, Mar 19 1967 Thorne, R.C. Rollins, D. RSA: R.F. Brandegee, Apr 1888 UC: T.S. Synonym: Guillenia lasiophylla from Catalina? per: http://www.rareplants.cnps.org/detail/571.html No specimens in CCH, extirpated Synonym: Brassica geniculata com. E. Howe Pers. Mar 28 1997 SCa-388 Takara Hoefs, J. Junak, M.L. SBBG: S.A. Synonyms: Lepidium virginicum Jul 28 1981 SCl-17 H. Ferguson Hochberg, Junak, M.C. SBBG: S.A. Native status on islands unknown, doubtful PPENDIX A Brassicaceae 116 117 118 119 120 121 122 123 124 125 126 Family/species 204 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 BAR ROS MIG 132 CAT CRU CAT CRU NIC CAT ANA CAT CRU 129 127 130 131 CLE NIC CAT BAR ROS CRU ANA MIG CRU ssp./var. and infraname ssp./var. * CLE? (DC.) L.C. Wheeler 128 2. Continued & J.F. Macbr. & J.F. Loeflingia squarrosa Nutt. NIC & A. Gray) Mattf. Minuartia douglasii (Torr. CLE CAT depressum Nutt. ROS Polycarpon CRU CLE CAT CRU MIG Sisymbrium altissimum L. * Sisymbrium irio L. * CLE NIC CAT Sisymbrium officinale (L.) Scop. * ANA CAT Sisymbrium orientale L. * CLE CRU MIG Raphanus raphanistrum L. * CLE CAT Raphanus sativus L. * ROS CRU CLE NIC CAT Sibara filifolia (Greene) Greene ROS CRU MIG CLE Sinapis arvensis L. Thysanocarpus conchuliferus Greene Thysanocarpus curvipes Hook. CLE Thysanocarpus laciniatus Nutt. CRU CLE CAT gracile Hook. Tropidocarpum ROS CRU CLE CAT glabra L. Turritis CRU Bergerocactus emoryi (Engelm.) Britton & Rose CLE Knuth CAT Cylindropuntia prolifera (Engelm.) F.M. CLE NIC CAT BAR ROS CRU ANA Opuntia ficus-indica (L.) Mill. * CLE NIC CAT Opuntia littoralis (Engelm.) Cockerell CRU ANA CLE NIC CAT BAR ROS CRU ANA Opuntia oricola Philbrick MIG CLE NIC CAT BAR ROS CRU ANA MIG Githopsis diffusa A. Gray ssp. diffusa Greene biflora (Ruiz & Pav.) Triodanis CAT ROS CRU & A. Gray Lonicera hispidula (Lindl.) Torr. CLE CAT Lonicera interrupta Benth. ROS CRU Lonicera subspicata Hook. & Arn. CAT Symphoricarpos mollis Nutt. CRU ROS CRU CAT ROS CRU A. Nelson Cardionema ramosissimum (Weinm.) ROS CRU MIG Cerastium glomeratum Thuill. * CLE CAT Herniaria hirsuta L. * cinerea (DC.) Loret & Barrandon var. ROS CRU ANA CLE NIC CAT MIG RSA: Orlando Mistretta Apr 5 1992 142 Synonym: Brassica kaber com. E. Howe Pers. Junak May 29 1996 SCl-545 SBBG: S.A. Ross Apr 14 1992 6153 UC: Tim Junak Mar 27 1997 SCa-346 SBBG: S.A. PPENDIX A 127 128 129 130 131 132 Cactaceae Campanulaceae Caprifoliaceae Caryophyllaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 205 ROS CRU 133 CLE CAT ROS CRU ANA MIG CLE NIC CAT BAR ROS CRU ANA MIG NIC CAT CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. CLE ROS 2. Continued S.L. Welsh & Reveal Welsh S.L. Polycarpon tetraphyllum (L.) L. * Polycarpon CAT Sagina apetala Ard. NIC ROS CRU ANA MIG & A. Gray ROS Crow G.E. CRU Watson) ssp. occidentalis (S. Sagina decumbens (Elliott) Torr. MIG CAT ROS CRU MIG Silene antirrhina L. CLE CAT Silene gallica L. * ROS CRU CLE NIC MIG CAT BAR ROS Silene laciniata Cav. CRU ANA ssp. laciniata MIG Silene coniflora Otth CAT Spergula arvensis L. * CRU CAT Spergularia bocconi (Scheele) Graebn. * CLE ROS NIC CAT BAR CRU ANA Spergularia macrotheca (Cham. & Schltdl.) Heynh. ssp. macrotheca Spergularia marina (L.) Besser CLE NIC CAT BAR ROS Cambess. * CRU ANA Spergularia villosa (Pers.) CLE CAT * Stellaria media (L.) Vill. ROS CRU CLE CAT Stellaria nitens Nutt. ROS CRU ANA MIG CAT Aphanisma blitoides Moq. CLE ROS CRU NIC CAT BAR ROS CRU ANA MIG Standl. Arthrocnemum subterminale (Parish) CLE CAT Watson) Atriplex argentea Nutt. ssp. expansa (S. CLE NIC CAT ROS CRU MIG ROS CRU Atriplex californica Moq. CLE NIC CAT BAR ROS Atriplex canescens (Pursh) Nutt. * CRU ANA canescens MIG var. Dietr. Atriplex coulteri (Moq.) D. CLE Hall & Clem. H.M. CAT Watson) (S. Watson S. CLE NIC Atriplex lentiformis (Torr.) CAT ROS CRU ANA MIG CRU ANA Dietr. Atriplex leucophylla (Moq.) D. CLE NIC CAT ROS CRU ANA Atriplex pacifica A. Nelson MIG CLE NIC CAT BAR * Atriplex semibaccata R. Br. ROS CRU ANA CLE NIC CAT BAR ROS CRU ANA Atriplex serenana Abrams davidsonii (Standl.) Munz MIG var. CAT Atriplex serenana Abrams serenana var. Atriplex prostrata DC. * NIC Atriplex watsonii Abrams CLE CRU NIC CAT MIG * Kuntze Bassia hyssopifolia (Pall.) ROS CRU ANA MIG CLE NIC CAT Beta vulgaris L. * ssp. maritima (L.) Arcang. CLE CAT Chenopodium berlandieri Moq. ROS CRU ANA NIC MIG CLE Watson S. NIC Watson) Chenopodium californicum (S. ROS CAT CRU ANA BAR ROS CRU ANA MIG Chenopodium murale L. * CLE NIC CAT BAR ROS CRU ANA Dysphania ambrosioides (L.) Mosyakin & Clemants * NIC MIG CAT ROS CRU Dysphania multifida (L.) Mosyakin & Clemants * Monolepis nuttalliana (Schult.) Greene CLE Salicornia depressa Standl. BAR ROS CRU MIG NIC Salicornia pacifica Standl. CLE CAT ROS CRU ANA MIG RSA: Blanche Trask Mar 1901 s.n. RSA: Blanche Trask PPENDIX A 133 Family/species 206 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 137 135 138 CAT ROS CRU 136 CLE NIC CAT BAR ROS CRU ANA MIG ROS CRU ANA CRU CAT ROS CRU ANA MIG ssp./var. and infraname ssp./var. * CLE NIC CAT ROS CRU 134 L. 2. Continued Salsola australis R. Br. Suaeda taxifolia (Standl.) Standl. CLE NIC CAT BAR ROS CRU ANA MIG Rob. Helianthemum greenei B.L. CAT Helianthemum scoparium Nutt. ROS CRU MIG& CLE? Cuscuta campestris Yunck. CAT Cuscuta occidentalis Millsp. CAT pacifica var. Wright Cuscuta pacifica Costea & M. Dichondra occidentalis House CAT Ipomoea cairica (L.) Sweet * ROS CRU MIG CAT Cornus glabrata Benth. CAT Crassula aquatica (L.) Schönl. CAT A. Berger Crassula connata (Ruiz & Pav.) CLE NIC CAT BAR ROS CRU ANA Crassula ovata (Mill.) Druce * MIG CLE Moran Dudleya blochmaniae (Eastw.) ssp. blochmaniae Moran Dudleya blochmaniae (Eastw.) ssp. insularis (Moran) Moran Britton & Rose Dudleya caespitosa (Haw.) ROS Dudleya candelabrum Rose ANA Dudleya greenei Rose ROS CRU MIG CAT McCabe Dudleya gnoma S.W. ROS CRU MIG ROS Peritoma arborea (Nutt.) H.H. Iltis Peritoma CLE CAT ROS Calystegia macrostegia (Greene) Brummitt ssp. amplissima Brummitt CLE NIC BAR Calystegia macrostegia (Greene) Brummitt ssp. cyclostegia (House) Brummitt CAT Calystegia macrostegia (Greene) Brummitt ssp. intermedia (Abrams) Brummitt CAT Calystegia macrostegia (Greene) Brummitt ssp. macrostegia Calystegia malacophylla (Greene) Munz * ssp. pedicellata (Jeps.) Munz NIC& * Calystegia soldanella (L.) R. Br. CLE NIC CAT Convolvulus arvensis L. * ROS CRU NIC MIG CAT Perry Convolvulus simulans L.M. ROS CRU CLE CAT Cressa truxillensis Kunth CLE NIC CRU CAT Cuscuta californica Hook. & Arn. ROS CRU ANA MIG CLE ROS CRU Suaeda calceoliformis (Hook.) Moq. CAT Synonym: Salsola tragus Junak Oct 22 1998 SCa-823 SBBG: S.A. com. E. Howe Pers. Benedict Sep 30 1969 SBBG: M.R. Grant 4541 RSA: George B. PPENDIX A Cistaceae Cornaceae Crassulaceae 134 135 136 137 138 Cleomaceae Convolvulaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 207 142 CRU MIG 141 CAT 140 BAR CRU ANA NIC 144 139 143 CLE NIC CAT NIC CAT CLE BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. 2. Continued (Kunth) Small * Chamaesyce serpens (Kunth) CLE Small Chamaesyce serpyllifolia (Pers.) CAT Croton setigerus Hook. CLE CAT Euphorbia crenulata Engelm. ROS CRU CAT Euphorbia misera Benth. CLE CAT Euphorbia peplus L. * CLE CRU CAT Euphorbia spathulata Lam. ROS CRU CLE CAT Ricinus communis L. * CLE NIC CAT Watson Stillingia linearifolia S. CRU Don * Acacia cyclops G. CLE? Dudleya nesiotica (Moran) Moran Dudleya traskiae (Rose) Moran CRU BAR Dudleya virens (Rose) Moran ssp. hassei (Rose) Moran CAT Dudleya virens (Rose) Moran ssp. insularis Dudleya virens (Rose) Moran ssp. virens Crossosoma californicum Nutt. CLE NIC CAT Cucurbita foetidissima Kunth Marah fabacea (Naudin) Greene CLE CRU Marah macrocarpa (Greene) Greene CLE NIC CAT BAR ROS CRU ANA MIG Arbutus menziesii Pursh Wells Arctostaphylos catalinae P.V. CAT CRU Arctostaphylos confertiflora Eastw. Arctostaphylos insularis Parry ROS et al. Parker V.T. Wells) ssp. insulicola (P.V. Arctostaphylos crustacea Eastw. ROS CRU CRU et al. Parker V.T. ssp. subcordata (Eastw.) Arctostaphylos crustacea Eastw. ROS CRU McMinn Arctostaphylos viridissima (Eastw.) Greene Comarostaphylis diversifolia (Parry) Wallace ssp. planifolia (Jeps.) G.D. CRU CAT ROS CRU ANA ovatum Pursh Vaccinium Xylococcus bicolor Nutt. ROS CRU CAT Chamaesyce maculata (L.) Small * CLE RSA: F.H. Elmore Feb 19 1939 422 Elmore Feb RSA: F.H. Dunkle Jul 26 1939 8359 SBBG: M.B. 24, 1968 4486 Howe Feb SD: Darley F. Dunkle Jul 31 1939 8402 SBBG: M.B. RSA: Steven A. Junak Nov 9 1990 SCI144 com. E. Howe Pers. PPENDIX Fabaceae A Crossosomataceae Cucurbitaceae Ericaceae Euphorbiaceae 139 140 141 142 143 144 Family/species 208 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 147 MIG 146 153 CAT CAT CRU

CLE NIC CAT BAR ROS CRU ANA MIG CLE NIC CAT ROS CRU ANA MIG CAT CLE CAT ROS CRU ANA CAT ROS CRU NIC CAT ROS CRU ANA MIG ssp./var. and infraname ssp./var. Lag., misappl. Lag., CLE CAT ROS CRU on Cruz, but NPS Park Flora did not on Cruz, but NPS Park

150 CRU 148 maritimus var. 151 149 var. americanus var. 145 Fisch. & C.A. Mey.; Lotus subpinnatus & C.A. Mey.; Fisch. Greene Greene; Wallace listed L. humistratus Greene; Wallace Benth. Clem. & E.G. Clem. Clem. & E.G. Greene 152 2. Continued D.D. Sokoloff D.D. Acmispon strigosus (Nutt.) Brouillet CLE CAT & C.A. Mey.) Acmispon wrangelianus (Fisch. ROS CAT CRU ANA MIG ROS CRU ANA MIG (Benth.) D.D. Sokoloff Acmispon parviflorus (Benth.) D.D. * Acacia decurrens Willd. CAT Acacia dealbata Link * * Acacia melanoxylon R. Br. CAT CRU Acmispon americanus (Nutt.) Rydb. CRU Acmispon argophyllus (A. Gray) Brouillet niveus (Greene) var. Sokoloff Acmispon brachycarpus (Benth.) D.D. CRU Benth. * Albizia lophantha (Willd.) Astragalus curtipes A. Gray CRU didymocarpus Astragalus didymocarpus Hook. & Arn. var. ROS Astragalus gambelianus E. Sheld. MIG CAT Astragalus miguelensis Greene CLE CRU Astragalus nevinii A. Gray CLE ROS CRU ANA MIG Astragalus traskiae Eastw. NIC BAR Astragalus trichopodus (Nutt.) A. Gray Jones) Barneby lonchus (M.E. var. CAT Astragalus trichopodus (Nutt.) A. Gray ROS trichopodus CRU var. ANA * Caesalpinia spinosa (Molina) Kuntze CAT Coronilla valentina L. * Genista linifolia L. * CAT CAT Johnson * Genista monspessulana (L.) L.A.S. CAT Acmispon argophyllus (A. Gray) Brouillet adsurgens (Dunkle) CLE var. Acmispon argophyllus (A. Gray) Brouillet argenteus (Dunkle) var. CLE NIC CAT BAR ROS Acmispon dendroideus (Greene) Brouillet dendroideus var. Acmispon dendroideus (Greene) Brouillet veatchii (Greene) Brouillet var. Acmispon dendroideus (Greene) Brouillet traskiae (Abrams) Brouillet var. CLE grandiflorus Acmispon grandiflorus (Benth.) Brouillet var. MIG Brouillet Acmispon heermannii (Durand & Hilg.) CAT Sokoloff Acmispon maritimus (Nutt.) D.D. (Torr. & A. Gray) Brouillet Acmispon micranthus (Torr. Synonym: Lotus purshianus 30 June, 1931 239 CAS: L.R. Abrams, I.L. Wiggins DS: E.R. Blakley 3 April 1962 5081 Synonym: Lotus humistratus Synonym: Lotus salsuginosus Synonym: Lotus hamatus Synonym: Lotus micranthus Synonym: Lotus wrangelianus Ross Apr 23 1993 6934 RSA: Tim PPENDIX A 145 146 147 148 149 150 151 152 153 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 209 CAT ROS CRU 156 CAT ROS CRU 154 155 CRU CLE NIC CATCLE CAT ROS CRU ANA BAR NIC CAT ROS ROS CRU ANA MIG CRU ANA MIG ssp./var. and infraname ssp./var. in TJM2) by Michael Vincent in 2005. in TJM2) by Michael Vincent T. dichotomum (= T. 2. Continued Torr. barbigerum Torr. Trifolium ciliolatum Benth. Trifolium ROS CAT & A. Gray) McDermott CLE depauperatum Desv. amplectens (Torr. Trifolium var. MIG ROS CAT CRU depauperatum Desv. truncatum (Greene) Isely CLE Trifolium var. NIC CAT ROS fucatum Lindl. CRU ANA Trifolium MIG CLE & A. Gray CAT gracilentum Torr. Trifolium CLE NIC ROS CAT BAR CRU ANA MIG ROS CRU ANA hirtum All. Trifolium MIG CLE macraei Hook. & Arn. Trifolium CAT microcephalum Pursh Trifolium ROS CRU CLE NIC CAT microdon Hook. & Arn. Trifolium ROS NIC CRU ANA CAT MIG Watson palmeri S. Trifolium CLE NIC CAT BAR repens L. * Trifolium ANA CAT Lupinus guadalupensis Greene CLE Lupinus hirsutissimus Benth. CLE CAT Lupinus microcarpus Sims ROS CRU ANA Lupinus succulentus K. Koch ROS CRU CLE NIC CAT MIG Lupinus truncatus Nutt. ROS CLE CRU ANA MIG CAT Lupinus variicolor Steud. ROS CRU ANA Medicago polymorpha L. * CLE NIC CAT BAR Medicago sativa L. * ROS CRU ANA MIG CLE NIC MIG CAT Melilotus albus Medik. * ROS CLE CRU NIC ANA MIG CAT Melilotus indicus (L.) All. * CRU ANA CLE NIC CAT BAR montana Nutt. ROS Pickeringia montana var. CRU ANA MIG Pisum sativum L. * Robinia pseudoacacia L. * NIC CRU Spartium junceum L. * NIC CRU CAT & A. Gray albopurpureum Torr. Trifolium NIC CRU Lathyrus odoratus L. * CLE Lathyrus tingitanus L. * CAT White) Isley Lathyrus vestitus Nutt. alefeldii (T.G. var. CAT Lathyrus vestitus Nutt. vestitus var. Lotus corniculatus L. * CAT Sm. Lupinus albifrons Benth. Agardh) C.P. douglasii (J. var. CRU Lupinus arboreus Sims Lupinus bicolor Lindl. ROS CLE NIC CRU CAT Lupinus chamissonis Eschsch. MIG ROS CRU ANA MIG Agardh Lupinus concinnus J. CLE ROS MIG SD: Steven A. Junak May 19, 1997 SCl877 Benedict Jun 26 1971 SBBG: M.R. dichotomum JEPS93564 (SN-846) det. var. PPENDIX A 154 155 156 Family/species 210 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 BAR ROS CRU ANA MIG CRU ANA ROS CRU ANA ROS ROS CRU 158 159 161 162 163 164 166 CAT CAT CAT 157 160 165 CLE NIC CAT BAR ROS CRU ROS CRU ANA MIG CLE NIC ROS CRU ANA ssp./var. and infraname ssp./var. macdonaldii Greene CAT ROS CRU × 2. Continued Trifolium variegatum Nutt. major Loja. Trifolium var. willdenovii Spreng. Trifolium CRU CLE agrifolia Quercus agrifolia Née var. Quercus chrysolepis Liebm. CLE CAT Quercus douglasii Hook. & Arn. ROS CRU ANA CAT Quercus engelmannii Greene CRU CAT Newb. Quercus kelloggii ROS Quercus lobata Née CRU CAT Quercus ROS CRU Berol. * Erodium botrys (Cav.) CLE NIC CAT Nixon & C.H. Mull. Quercus pacifica Nixon CAT ROS parvula Quercus parvula Greene CRU var. CRU Quercus tomentella Engelm. CLE CAT ROS CRU ANA Johnst. salina (Molina) I.M. Frankenia CLE NIC CAT ROS CRU ANA MIG Garrya veatchii Kellogg CRU Mans. Zeltnera davyi (Jeps.) G. CLE Mans. Zeltnera venusta (A. Gray) G. NIC ROS CAT CRU Aldasoroet California macrophylla (Hook. & Arn.) J.J. CAT (Godr.) Thell. * Erodium brachycarpum (Godr.) CLE Willd. americana Willd. Vicia benghalensis L. * Vicia ROS CRU MIG CAT Watson hassei S. Vicia CLE NIC CAT & A. Gray ssp. ludoviciana ludoviciana Torr. Vicia ROS CRU ANA MIG sativa L. * Vicia nigra (L.) Erhart ssp. sativa L. * Vicia sativa ssp. ROS villosa Roth Vicia (Host) Corb. NIC CAT Erodium cicutarium (L.) Aiton * CLE NIC CAT BAR Erodium moschatum (L.) Aiton * ROS CRU ANA MIG CLE NIC CAT BAR ROS CRU ANA MIG RSA: P.H. Raven May 8 1962 17614 RSA: P.H. Mar 25 1931 s4367 Fosberg UC: F.R. Raven May 20 1962 17789 SBBG: P.H. Ross, L. Arnseth Apr 7 1990 4260 UCR: Steve Boyd, T.S. Thorne Mar 7 1966 35954 SBBG: R.F. Propst May 29, 1968 37691 Thorne, D. SD: Robert F. Wallace GH: W.A. May 16 1998 SCa-697 Junak, K. Kirkland, L. Vorobik SBBG: S.A. Hammitt May 18 1991 5191 Mistretta, M. Ross, O. SBBG: T. Hoefs, K. Kirkland Apr 5 1998 SCa-549 Junak, M.L. SBBG: S.A. PPENDIX A 157 158 159 160 161 162 163 164 165 166 Frankeniaceae Garryaceae Gentianaceae Geraniaceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 211 168 CAT CRU ANA NIC CAT ROS CRU MIG 169 167 CLE NIC CAT BAR ROSCLE ROS CRU CRU ANA ANA MIG ssp./var. and infraname ssp./var. hortorum L.H. Bailey * CLE × 2. Continued Scutellaria tuberosa Benth. ROS CRU Stachys ajugoides Benth. MIG Stachys bullata Benth. lanceolatum Benth. Trichostema ROS CRU ANA CAT Hesperolinon micranthum (A. Gray) Small CAT Mentzelia affinis Greene CLE NIC CAT ROS & A. CRU Mentzelia micrantha (Hook. & Arn.) Torr. CLE CAT CRU Ammannia robusta Heer & Regel CAT Erodium texanum A. Gray CAT Geranium carolinianum L. CAT Geranium dissectum L. * ROS CRU Geranium molle L. * ROS Pelargonium * CRU peltatum (L.) L’Hér. Pelargonium NIC malvaceum Ribes malvaceum Sm. var. Ribes viburnifolium A. Gray CAT Ribes thacherianum (Jeps.) Munz CRU * Watson Juglans californica S. CAT Juglans regia L. * CRU Clinopodium douglasii (Benth.) Kuntze CAT ROS CRU Lamium amplexicaule L. * Lepechinia fragrans (Greene) Epling CAT ROS CRU CRU Marrubium vulgare L. * CLE NIC CAT ROS CRU Mentha aquatica L. * MIG CAT Mentha spicata L. * CAT CRU Nepeta cataria L. * CAT Salvia apiana Jeps. CAT Salvia brandegeei Munz ROS Salvia columbariae Benth. CLE CAT ROS CRU Salvia leucophylla Greene Salvia mellifera Greene CRU CLE? SBBG: T. Ross May 22 1991 5435 SBBG: T. Philbrick Mar 25 1966 B66-91 SBBG: R.N. com. E. Howe Pers. PPENDIX Linaceae Loasaceae Lythraceae A Grossulariaceae Juglandaceae Lamiaceae 167 168 169 Family/species 212 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 ROS CRU BAR ROS CRU ANA MIG ROS CRU BAR CRU MIG 170 172 174 176 177 ROS ANA CAT CRU MIG NIC CAT 171 173 175 ROS CRU MIG NIC CAT CLE CAT ROS CRU CLE * NIC * ROS CRU ANA CLE MIG CAT CRU * CLE NIC CAT BAR ROS CRU ANA MIG ssp. assurgentiflora ssp. glabra (Munz & I.M. ssp./var. and infraname ssp./var. CAT 2. Continued & K.L. Chambers Malva nicaeensis All. * Malva parviflora L. * CLE NIC ROS CAT CRU BAR ROS CRU ANA Malvella leprosa (Ortega) Krapov. MIG CLE CAT Sidalcea malviflora (DC.) A. Gray ssp. malviflora ROS Watson Calandrinia breweri S. & C.A. Mey. Calandrinia ciliata Fisch. CLE NIC CAT ROS CRU BAR ROS CRU ANA MIG Cistanthe maritima (Nutt.) Hershk. CLE CAT Claytonia parviflora Hook. BAR ROS ssp. parviflora CRU ANA Anagallis arvensis L. * CLE NIC CAT BAR ROS CRU ANA Anagallis minima (L.) E.H.L. Krause MIG ROS Eucalyptus camaldulensis Dehnh. * CAT Alcea rosea L. * CAT Eremalche exilis (A. Gray) Greene CLE CRU CAT Lavatera assurgentiflora CRU Claytonia perfoliata Willd. Mill. John M. ssp. mexicana (Rydb.) CLE Claytonia perfoliata Willd. ssp. perfoliata Montia fontana L. CAT CRU carica L. * Ficus CLE Muell. * Eucalyptus cladocalyx F. Lavatera assurgentiflora Malacothamnus clementinus Johnst.) Kearney CLE Kearney catalinensis (Eastw.) & A. Gray) Greene var. Malacothamnus fasciculatus (Torr. CAT Rob.) Kearney nesioticus (B.L. CRU & A. Gray) Greene var. Malacothamnus fasciculatus (Torr. & Berthel. * CLE Malva pseudolavatera Webb Torr. & A. Gray Lythrum californicum Torr. Lythrum hyssopifolia L. * CRU CLE NIC CAT Eucalyptus globulus Labill. * CLE CAT BAR ROS CRU ANA RSA: Mark Hoefs, Steven A. Junak, Janet Takara, M. Gay Jul 14 1995 2401 M. RSA: Mark Hoefs, Steven A. Junak, Janet Takara, SD: Steven A. Junak Apr 23, 1997 SCI858 Junak, K. Kirkland Apr 13 2001 SCa-1421 SBBG: S.A. Ross, Orlando Mistretta, Mike Hammitt May 19 1991 5286 SBBG: Tim SBBG: E.R. Blakley Apr 7 1963 5537 Junak May 20 1996 SBBG: S.A. Ross Apr 23 1993 6926 UCR: T.S. 25 1968 s.n. Clarke Feb UCR: O.F. PPENDIX Montiaceae Myrsinaceae Myrtaceae Malvaceae Moraceae A 170 171 172 173 174 175 176 177 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 213 ROS CRU ANA MIG 178 181 CRU CRU 179

CLE NIC CAT BAR ROS CRU ANACLE NIC MIG BAR ROS CRU MIG CLE CAT CRU ANA CAT ssp./var. and infraname ssp./var. ssp. angustifolium 180 2. Continued W.L. Wagner & Hoch Wagner W.L. & Hoch & Hoch W.L.Wagner Wagner W.L. & Hoch Wagner & Hoch W.L. Wagner W.L. & Hoch Wagner W.L. & Hoch Wagner W.L. & Hoch Wagner W.L. & Hoch Wagner W.L. Lewis H. Lewis & M. Eucalyptus polyanthemos Schauer * CAT * Eucalyptus sideroxylon Woolls Eucalyptus tereticornis Sm. * CRU ROS CRU Abronia latifolia Eschsch. Watson Abronia maritima S. CLE NIC CAT Abronia umbellata Lam. ROS CRU ANA MIG MIG CLE NIC CAT Mirabilis laevis (Benth.) Curran ROS CRU crassifolia (Choisy) Spellenb. var. CLE MIG CAT BAR ROS CRU ANA Olea europaea L. * BAR CRU ANA Raven & C.A. Mey.)Camissonia strigulosa (Fisch. P.H. Camissoniopsis cheiranthifolia (Spreng.) ssp. cheiranthifolia ROS Camissoniopsis cheiranthifolia (Spreng.) Watson) ssp. suffruticosa (S. NIC Raven) CLE Watson) ssp. clementina (P.H. Camissoniopsis guadalupensis (S. Camissoniopsis hirtella (Greene) & Hoch Wagner Camissoniopsis ignota (Jeps.) W.L. Raven) Camissoniopsis intermedia (P.H. ROS CRU CAT CRU MIG Camissoniopsis micrantha (Spreng.) CLE CAT ROS CRU ANA MIG Raven) CLE Camissoniopsis robusta (P.H. CAT CRU MIG Lewis Clarkia davyi (Jeps.) H. Lewis & M. & A. Gray) Nelson J.F. CLE Clarkia epilobioides (Torr. ROS CAT ROS CRU ANA Lewis Clarkia prostrata H. Lewis & M. Macbr. ssp. quadrivulnera (Lindl.) Clarkia purpurea (Curtis) A. Nelson & J.F. CAT ROS CRU Clarkia unguiculata Lindl. CAT Epilobium brachycarpum C. Presl CLE? CRU (Greene) P.H. Raven ssp. canum Epilobium canum (Greene) P.H. (Greene) P.H. Raven Epilobium canum (Greene) P.H. UCR: T.S. Ross, Janet Takara, Stacey Otte Apr 24 1993 6964 Ross, Janet Takara, UCR: T.S. com. E. Howe Pers. This taxa unresolved 8, 1966 54 Beauchamp August R.M. SDSU: PPENDIX A Nyctaginaceae Oleaceae Onagraceae 178 179 180 181 Family/species 214 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 CRU NIC CAT ROS CRU NIC CAT ROS CRU MIG 182 183 184 CAT CRU MIG CRU CAT ROS CRU ANA MIG CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. 2. Continued Oxalis articulata Savigny * ssp. rubra (A. St.-Hil.) Lourteig ROS Oxalis californica (Abrams) R. Knuth CAT Oxalis corniculata L. * CRU CLE Raf. Epilobium ciliatum Raf. ssp. ciliatum & A. Gray Eulobus californicus Torr. Raven ssp. peploides CAT P.H. Ludwigia peploides (Kunth) ROS CRU * & Hoch Wagner W.L. Oenothera xenogaura & Hoch * Wagner Oenothera sinuosa W.L. CRU CAT Dietr. Oenothera elata Kunth. W. Watson) ssp. hirsutissima (S. CRU Castilleja affinis Hook. & Arn. ssp. affinis Chuang & Heckard Castilleja attenuata (A. Gray) T.I. Chuang & Heckard NIC Castilleja densiflora (Benth.) T.I. CRU ROS CRU MIG Chuang & Heckard Castilleja exserta (A. Heller) T.I. CAT ROS CRU Castilleja foliolosa Hook. & Arn. MIG CAT Castilleja grisea Dunkle CLE Castilleja hololeuca Greene Castilleja mollis Pennell ROS CRU ANA MIG Kopsiopsis strobilacea (A. Gray) Beck ROS MIG Orobanche bulbosa Beck ROS CAT ROS CRU Orobanche californica Cham. & Schltdl. ssp. grandis Heckard Orobanche fasciculata Nutt. ROS CLE NIC CAT ROS CRU ANA Orobanche parishii (Jeps.) Heckard MIG ssp. brachyloba Heckard NIC CAT ROS CRU MIG Orobanche uniflora L. CLE? Oxalis pes-caprae L. * CLE NIC CAT CRU Dendromecon harfordii Kellogg CLE CAT Ehrendorferia ochroleuca (Engelm.) Fukuhara ROS CRU Eschscholzia californica Cham. CRU CLE Eschscholzia ramosa (Greene) Greene CLE NIC CAT BAR ROS CRU Meconella denticulata Greene californicum A. Gray Papaver CRU heterophyllum (Benth.) Greene Papaver ROS CRU CLE CAT BAR somniferum L. * ROS Papaver CRU ANA MIG CAT Platystemon californicus Benth. NIC CAT BAR CRU ROS CRU ANA MIG * Romneya coulteri Harv. CAT Pers. com. E. Howe Pers. RSA: Steven A. Junak May 14 1985 SC1-56 Dunkle May 27 1928 1983 RSA: M.B. PPENDIX Oxalidaceae A Orobanchaceae 182 183 184 Papaveraceae Papaveraceae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 215 CRU CRU MIG 185 187 188 189 190 CAT ROS 186 CLE NIC CAT BAR ROS CRU ANA MIG CAT ssp./var. and infraname ssp./var. 2. Continued D.A. Sutton D.A. Mimulus aurantiacus Curtis aurantiacus var. Thomps. Mimulus aurantiacus Curtis parviflorus (Greene) D.M. var. CLE Thomps. D.M. Mimulus aurantiacus Curtis pubescens (Torr.) ROS var. CRU ANA Thomps. ROS Mimulus aurantiacus Curtis puniceus (Nutt.) D.M. CRU var. CAT Mimulus cardinalis Benth. CAT Mimulus floribundus Lindl. CLE CRU * Willd. Linaria bipartita (Vent.) CAT Nuttallanthus texanus (Scheele) CLE CAT ROS CRU ANA MIG anagallis-aquatica L. * Veronica Plantago coronopus L. * CRU NIC CAT Plantago elongata Pursh ANA Plantago erecta E. Morris MIG ROS CRU CLE MIG CAT Plantago lanceolata L. * ROS CRU ANA CLE CAT Mimulus guttatus DC. CLE CAT Mimulus latifolius A. Gray ROS CRU CAT Mimulus traskiae A.L. Grant CAT * Pittosporum undulatum Vent. CAT Greene Antirrhinum kelloggii CAT Antirrhinum multiflorum Pennell CRU Antirrhinum nuttallianum A. DC. Thomps. CLE ssp. subsessile (A. Gray) D.M. CAT CRU ROS CRU ANA Callitriche marginata Torr. MIG CLE CAT Collinsia heterophylla Graham CLE Gambelia speciosa Nutt. CLE ROS CAT BAR Keckiella cordifolia (Benth.) Straw CLE CAT Kickxia elatine (L.) Dumort. * ROS CRU ANA CAT Plantago major L. * NIC CAT Plantago maritima L. CRU Meyers & A. Liston S.C. insularis (Eastw.) Plantago ovata Forssk. CLE var. NIC CAT BAR ROS ROS CRU ANA MIG * Plantago subnuda Pilg. ROS CRU MIG Platanus racemosa Nutt. CAT* CRU OBI: John Knapp, Denise 7, April 13, 2003 Ross, L. Arnseth Apr 8 1990 4291 Boyd, T. SBBG: S. Thompson 2012. Mimulus , in Jepson Flora Project (eds.) eFlora, accessed on May 7 2014 Dave M. Propst 1974-09-12 45100 Thorne & A.D. GH: R.F. Jul 25 1995 2438 Takara Thorne, J. Hoefs, R.F. SBBG: M.L. Thorne Apr 6 1996 2671 Hoefs, R.F. SBBG: M.L. PPENDIX A Phrymaceae 185 186 187 188 189 190 Pittosporaceae Plantaginaceae Plantanaceae Family/species 216 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 193 ROS * 192 191 CAT CRU* CAT CLE CLE CAT ROS CRU ANA CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. 2. Continued (Boiss.) A. Heller * Grant Grant & V.E. & Standl. Wooton & L.A. Johnson Porter J.M. Hook. & Arn. var. fasciculatum Eriogonum fasciculatum Benth. * var. (Mill.) Willd. Armeria maritima (Mill.) Willd. ssp. californica (Boiss.) A.E. Porsild Limonium californicum CLE? ROS Hubb. * Limonium perezii (Stapf) F.T. CLE CAT Limonium sinuatum (L.) Mill. * CAT ANA& Allophyllum glutinosum (Benth.) A.D. MIG& CLE CAT Eriastrum filifolium (Nutt.) CLE CAT CRU Grant Grant & A.D. Gilia achilleifolia Benth. ssp. multicaulis (Benth.) V.E. Grant Gilia angelensis V.E. ROS CLE CAT Grant Gilia capitata Sims ssp. abrotanifolia (Greene) V.E. CRU CAT Grant Gilia clivorum (Jeps.) V.E. CRU CAT Gilia nevinii A. Gray ROS CRU ANA CLE MIG NIC CAT BAR Grant ROS CRU Grant & V.E. ANA A.D. Gilia tenuiflora Benth. ssp. hoffmannii (Eastw.) Leptosiphon bicolor Nutt. ROS CLE CAT Leptosiphon parviflorus Benth. Leptosiphon pygmaeus (Brand) ssp. pygmaeus ROS CRU MIG Linanthus dianthiflorus (Benth.) Greene CAT CRU Navarretia atractyloides (Benth.) CLE CAT ROS CRU Navarretia hamata Greene ssp. leptantha (Greene) H. Mason CLE CAT californica Nutt. Polygala Chorizanthe staticoides Benth. CRU Watson Chorizanthe wheeleri S. CAT Eriogonum arborescens Greene ROS CRU Eriogonum cinereum Benth. ROS CRU ANA NIC & A. Gray polifolium (Benth.) Torr. NIC Eriogonum fasciculatum Benth. * var. compactum Dunkle Watson Eriogonum giganteum S. var. BAR formosum K. Brandegee Watson Eriogonum giganteum S. var. CLE giganteum Watson Eriogonum giganteum S. var. grande Eriogonum grande Greene var. Pers. com. E. Howe Pers. 21 1990 SN-432, Feb Murphey, Junak, T. SBBG: S.A. Haefs Sep 12 1974 45110 Propst, M. Thorne, D. HSC: R.F. PPENDIX A Plumbaginaceae Polemoniaceae Polygalaceae 191 192 193 Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 217 CAT ROS CRU 194 CAT CRU CLE CAT CRU CRU CLE CAT ROS CRU CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. CLE & C.A. Mey. NIC Fisch. CAT BAR ROS CRU ANA MIG 2. Continued Rumex crispus L. * CLE NIC CAT Rumex fueginus Phil. ROS CRU ANA MIG CAT Rumex obtusifolius L. * NIC Rumex pulcher L. * MIG Rumex salicifolius Weinm. CLE NIC CRU CAT ROS CRU MIG oleracea L. * Portulaca CAT CRU Thomps. Dodecatheon clevelandii Greene ssp. insulare H.J. CLE NIC CAT ROS CRU ANA MIG Clematis lasiantha Nutt. Clematis ligusticifolia Nutt. CRU CAT Clematis pauciflora Nutt. ROS CRU Delphinium parryi A. Gray Warnock ssp. maritimum (Davidson) M.J. CAT CRU ROS CRU ANA MIG Delphinium parryi A. Gray ssp. parryi & A. Gray Warnock CLE ssp. kinkiense (Munz) M.J. Delphinium variegatum Torr. & A. Gray ssp. thornei Munz Delphinium variegatum Torr. CLE Ranunculus californicus Benth. Ranunculus hebecarpus Hook. & Arn. CAT ROS CRU CAT MIG Macbr. Oligomeris linifolia (Hornem.) J.F. CLE NIC CAT BAR ROS CRU ANA MIG Reseda odorata L. * CAT Ceanothus arboreus Greene CAT Munz CLE insularis (Eastw.) Ceanothus megacarpus Nutt. var. ROS CRU CAT ROS CRU ANA MIG& megacarpus Ceanothus megacarpus Nutt. var. californica (Eschsch.) A. Gray Frangula californica ssp. Rhamnus pirifolia Greene CLE CAT ROS CRU MIG& Adenostoma fasciculatum Hook. & Arn. fasciculatum var. Eriogonum grande Greene rubescens (Greene) Munz var. Eriogonum grande Greene timorum Reveal NIC var. ROS CRU ANA MIG Lastarriaea coriacea (Goodman) Hoover CAT lapathifolia (L.) Gray Persicaria ROS CRU * argyrocoleon Kunze Polygonum CLE CRU NIC CAT aviculare L. * ssp. depressum (Meisn.) Arcang. Polygonum CLE NIC CRU CAT ROS CRU ANA Pterostegia drymarioides Rumex acetosella L. * Rumex conglomeratus Murray * CLE ROS CRU SD: Steven A. Junak May 18, 1996 SCI488a PPENDIX Portulacaceae Portulacaceae Primulaceae Ranunculaceae Resedaceae Rhamnaceae Rosaceae A 194 Family/species 218 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 ROS 196 197 CAT CAT CAT CAT CAT ROS CRU CAT CRU CLE NIC CAT BAR ROS CRU ANA MIG ssp./var. and infraname ssp./var. CAT 195 Lindl. 2. Continued Dempster & Stebbins Dempster & Stebbins Hook. & Arn. var. prostratum Dunkle Adenostoma fasciculatum Hook. & Arn. var. CAT ROS CRU Aphanes occidentalis (Nutt.) Rydb. CLE CAT betuloides Cercocarpus betuloides Nutt. var. ROS CRU blancheae (C.K. Schneid.) Little Cercocarpus betuloides Nutt. var. CAT Cercocarpus traskiae Eastw. ROS CRU Drymocallis glandulosa (Lindl.) Rydb. CAT Raven CLE P.H. ssp. lyonii (Eastw.) Dietr. Prunus ilicifolia (Hook. & Arn.) D. CAT ROS CRU ANA Rosa californica Cham. & Schltdl. CAT ROS CRU Rubus ursinus Cham. & Schltdl. CAT ROS CRU MIG Howell) Galium angustifolium subsp. ssp. foliosum (Hilend & J.T. ROS CRU ANA Galium angustifolium A. Gray ssp. angustifolium Galium aparine L. CLE NIC CAT BAR ROS CRU ANA Galium buxifolium Greene MIG Galium californicum Hook. & Arn. ssp. flaccidum (Greene) Dempster & Stebbins CRU MIG Galium californicum Hook. & Arn. ssp. miguelense (Greene) CRU ROS MIG Galium catalinense A. Gray ssp. acrispum Dempster CLE Galium catalinense A. Gray ssp. catalinense Galium nuttallii A. Gray ssp. insulare Ferris CAT ROS CRU Galium parisiense L. * CAT (Lindl.) M. Roem. Heteromeles arbutifolia (Lindl.) M. CLE CAT Holodiscus discolor (Pursh) Maxim. ROS CRU ANA MIG CAT Raven Lyonothamnus floribundus A. Gray CLE ssp. aspleniifolius (Greene) P.H. CRU Lyonothamnus floribundus A. Gray ssp. floribundus ROS CRU anserina L. Potentilla ssp. pacifica (Howell) Rousi sanguisorba L. * Poterium CRU MIG CAT Galium porrigens Dempster porrigens var. Ruta chalepensis L. * CAT ssp. fremontii Watson fremontii S. Populus trichocarpa Hook. Populus Salix exigua Nutt. CAT NIC ROS CRU Salix gooddingii C.R. Ball CRU CLE Salix laevigata Bebb CAT CRU Potentilla glandulosa Synonym: Potentilla Ross, 6923, Apr 22 1993 UC: Tim Hoefs, K. Kirkland Jul 17 1998 SCa-769 Junak, M.L. SBBG: S.A. PPENDIX A Rubiaceae 195 196 197 Rutaceae Salicaeae Family/species 2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 219 ANA 209 BAR& CRU ANA& MIG& BAR CRU ANA 204 205 207 NIC CAT NIC CAT CRU NIC CAT ROS CRU ANA MIG * CAT ROS CRU ROS CRU NIC CRU& NIC ROS 198 202 203 206 200 201 208 CRU CLE 199 Greene 2. Continued Cham. & Schltdl. Benth. var. lasiandra Salix lasiandra Benth. var. Salix lasiolepis Benth. CLE? parviflora Juss. Petunia Solanum americanum Mill. * CLE ROS * Forst. Myoporum laetum G. CLE Scrophularia californica ROS Scrophularia villosa Pennell CLE CAT thapsus L. * Verbascum CRU Datura wrightii Regel CLE? Solanum douglasii Dunal CLE NIC CAT BAR ROS CRU ANA MIG Acer macrophyllum Pursh CRU Heuchera maxima Greene Jepsonia malvifolia (Greene) Small ROS CRU ANA CLE NIC CAT ROS CRU Lithophragma affine A. Gray & A. Gray CAT Lithophragma cymbalaria Torr. Lithophragma maximum Bacig. ROS CRU CLE Micranthes californica (Greene) Small Nicotiana clevelandii A. Gray CAT Nicotiana glauca Graham * CLE? Lycium brevipes Benth. CLE NIC CAT& Lycium californicum Nutt. CLE NIC CAT BAR CRU ANA Lycium fremontii A. Gray MIG Lycium verrucosum Eastw. NIC& ROS Lycopersicum esculentum Mill. * CLE? Nicotiana quadrivalvis Pursh CAT Pers. com. E. Howe Pers. Synonym : Saxifraga californica Junak Mar 13 1997 SCl-770 SBBG: S.A. Junak Jul 31 1981 SCl-53 SBBG: S.A. com. E. Howe Pers. com. E. Howe Pers. Thorne, 47381, Sep 4 1975 RSA: R.F. Mar 1901 s.n. RSA: Blanche Trask com. E. Howe Pers. Everett, 34883, Jun 22, 1965 Thorne, P. SD: Robert F. Beauchamp Sep 15 1979 R.M. SBBG: H.L. Ferguson, Junak Sep 23 1998 SR-896 SBBG: S.A. PPENDIX A 198 199 200 201 202 203 204 205 206 207 208 209 Scrophulariaceae Solanaceae Sapindaceae Saxifragaceae Family/species and infraname Family/species CLE ssp./var. NIC CAT BAR ROS CRU ANA MIG 220 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7 214 ANA 211 216 CAT BAR& CAT CRU CAT ROS 210 212 213 215 CLE NIC CAT 436 279 633 150 519 662 271 296 2. Continued * Solanum elaeagnifolium Cav. CAT Solanum wallacei (A. Gray) Parish ROS CRU CAT ROS CRU aphylla (L.) H. Karst. * Tamarix * chinensis Lour. Tamarix ROS CLE? parviflora DC. * Tamarix CAT & Rodr. bracteata Lag. Verbena CLE Ulmus xhollandica Mill. * CRU Hesperocnide tenella Torr. CLE CAT BAR ROS CRU hespera Hinton Parietaria CLE NIC CAT BAR ROS CRU ANA Soleirolia soleirolii (Req.) Dandy * MIG NIC Urtica dioica L. ssp. holosericea (Nutt.) Thorne CAT ROS CRU Urtica urens L. * ANA CAT ROS CRU MIG Centranthus ruber (L.) DC. * CAT Plectritis ciliosa (Greene) Jeps. CRU CRU Phyla nodiflora (L.) Greene * CLE Verbena lasiostachys Link lasiostachys Verbena var. lasiostachys Link scabrida Moldenke Verbena var. CAT ROS CRU MIG & A. Gray pedunculata Torr. Viola CLE CAT ROS CRU girdiana Munson Vitis CAT terrestris L. * Tribulus CAT Tamarix ramosissima Ledeb. * Tamarix CLE NIC ROS CRU MIG Samolus parviflorus Raf. CRU majus L. * Tropaeolum CLE Pers. com. E. Howe Pers. Thorne, 45759, Mar 22 1975 RSA: R.F. Ross, 5434, May 22 1991 UCR: T.S. Mar 12 1981 234 SBBG: H.L. Ferguson SC-63, Hochberg, May 9 1979 Philbrick, M.C. Junak, R.N. SBBG: S.A. Jun 28 1992 s.n. RSA: E. Kellogg Bushman Oct 21 1998 SCa-815 Junak, K. Kirkland, M. SBBG: S.A. PPENDIX A Tamaricaceae 210 211 212 213 214 215 216 Ulmaceae Urticaceae Valerianaceae Verbenaceae Violaceae Violaceae Vitaceae Zygophyllaceae Theophrastaceae Tropaeolaceae UMBER OF SPECIES Family/species ssp./var. and infraname ssp./var. Family/species CLE NIC CAT BAR ROS CRU ANA MIG N