Anceps. Diacamma Rugosum Var

BARRY BOLTON’S ANT CATALOGUE, 2020

DIACAMMA aequale. Diacamma aequale Zettel, et al. 2016: 163, figs. 46-49 (w.) PHILIPPINES (Negros I.). Type-material: holotype worker, 3 paratype workers. Type-locality: holotype Philippines: Negros Oriental, Sibulan, Lake Balinsasayao-Lake Danae (Twin Lakes), 28-30.x. (C.V. Pangantihon); paratypes with same data. Type-depositories: NMPM (holotype); HSZC (paratypes). Distribution: Philippines (Negros). anceps. Diacamma rugosum var. anceps Matsumura & Uchida, 1926: 51. Type-material: syntype workers (number not stated). Type-localities: China: Hong Kong (Ris), Indonesia: Sumatra, Enggano I., Bua Bua, v.-vi.1891 (Modigliani), and Myanmar (“Birmania”): Carin (L. Fea). Type-depository: MSNG. [First available use of Diacamma rugosum subsp. geometricum var. anceps Emery, 1897b: 155 (w.) CHINA (Hong Kong), INDONESIA (Enggano I.), MYANMAR; unavailable (infrasubspecific) name.] As unavailable (infrasubspecific) name: Emery, 1900d: 666; Forel, 1900d: 320; Emery, 1911d: 66; Forel, 1912a: 49; Forel, 1912d: 100; Forel, 1912n: 52; Wheeler, W.M. 1921c: 529; Wheeler, W.M. 1921e: 110; Wheeler, W.M. 1923b: 1; Wheeler, W.M. 1924b: 242; Wheeler, W.M. 1928c: 5; Wheeler, W.M. 1929g: 58; Wheeler, W.M. 1930h: 58; Karavaiev, 1935a: 63; Teranishi, 1940: 56; Yasumatsu, 1940a: 67; Onoyama, 1980: 196. Junior synonym of rugosum: Lin & Wu, 2003: 67. Subspecies of rugosum: Chapman & Capco, 1951: 55; Bolton, 1995b: 169; Terayama, 2009: 103; Zhou & Ran, 2010: 104; Guénard & Dunn, 2012: 58. Distribution: China, Indonesia (Enggano, Java, Sebesi), Japan, Myanmar, Taiwan. andamane. Diacamma rugosum var. andamana Chapman & Capco, 1951: 55. Type-material: syntype worker(s), syntype male(s) (numbers not stated). Type-locality: India: S Andaman Is, Kyd I. (no collector’s name). Type-depository: MHNG. [First available use of Diacamma rugosum r. vagans var. andamana Forel, 1922: 90 (w.m.) INDIA (Andaman Is); unavailable (infrasubspecific) name (Bolton, 1995b: 169).] Distribution: India (Andaman Is). arcuatum. Diacamma rugosum subsp. arcuata Karavaiev, 1925b: 118 (w.) INDONESIA (Sumatra). Type-material: holotype worker. Type-locality: Indonesia: Sumatra, Manguinang, 25.xi.1913 (O. John). Type-depository: SIZK. As unavailable (infrasubspecific) name: Karavaiev, 1935a: 60. Subspecies of rugosum: Chapman & Capco, 1951: 56; Bolton, 1995b: 169. Distribution: Indonesia (Sumatra). assamense. Diacamma tritschleri r. assamensis Forel, in Emery, 1897b: 151 (footnote) (w.) INDIA (Assam). Type-material: syntype workers (number not stated). Type-locality: India: Assam, Nissor (Smythies). Type-depository: MHNG. Status as species: Forel, 1900d: 319; Bingham, 1903: 79; Emery, 1911d: 65; Forel, 1913e: 660; Chapman & Capco, 1951: 53; Bolton, 1995b: 169; Bharti, Guénard, et al. 2016: 49. Distribution: India, Myanmar. aureovestitum. Diacamma aureovestitum Zettel, et al. 2016: 160, figs. 42-45 (w.) PHILIPPINES (Mindanao I.). Type-material: holotype worker, 13 paratype workers. Type-locality: holotype Philippines: Mindanao, Lanao del Sur Prov., Malabang, iv.-v.1945 (J.J. Marquis); paratypes with same data. Type-depositories: CASC (holotype); CASC, HSZC, NHMW (paratypes). Distribution: Philippines (Mindanao). australe. Formica australis Fabricius, 1775: 393 (w.) AUSTRALIA (no state data, “Habitat in nova Hollandia”). Type-material: holotype worker. Type-locality: Australia (“nova Hollandia”): Mus. Banks. (no further data). Type-depository: BMNH. Forel, 1915b: 26 (m.); Wheeler, G.C. & Wheeler, J. 1952c: 609 (l.). Combination in Ponera: Smith, F. 1858b: 93; Roger, 1860: 303; combination in Diacamma: Mayr, 1862: 718. Status as species: Fabricius, 1782: 492; Fabricius, 1787: 309; Gmelin, 1790: 2801; Christ, 1791: 506; Olivier, 1792: 497; Fabricius, 1793: 359; Latreille, 1802c: 128; Fabricius, 1804: 410; Roger, 1860: 303; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Mayr, 1876: 87; Mayr, 1879: 660 (redescription); Emery, 1887b: 436 (in key, footnote); Dalla Torre, 1893: 28; Emery, 1897b: 152; Forel, 1910b: 18; Emery, 1911d: 65; Viehmeyer, 1914c: 515; Forel, 1915b: 26; Baltazar, 1966: 241; Taylor & Brown, 1985: 29; Taylor, 1987a: 24; Bolton, 1995b: 169; Shattuck & Barnett, 2006: 14 (redescription). Distribution: Australia. baguiense. Diacamma baguiensis Wheeler, W.M. & Chapman, 1925: 66, pl. 2, figs. 11, 12 (w.) PHILIPPINES (Luzon I.). Type-material: 2 syntype workers. Type-locality: Philippines: Luzon, Mountain Prov., Baguio, 1370 m., x.1924 (J.W. Chapman). Type-depositories: MCZC, MSNG. Status as species: Chapman & Capco, 1951: 53; Baltazar, 1966: 241; Bolton, 1995b: 169; Laciny, Pal & Zettel, 2015: 109 (redescription). Distribution: Philippines (Luzon). balinense. Diacamma rugosum subsp. balinensis Karavaiev, 1925b: 116 (w.) INDONESIA (Bali I.). Type-material: syntype workers (number not stated, “several”). Type-locality: Indonesia: Bali, Boeleleng (Singaradja), Nr. 2681 (W. Karawaiew). Type-depository: SIZK. Subspecies of rugosum: Chapman & Capco, 1951: 57; Bolton, 1995b: 169. Distribution: Indonesia (Bali). birmanum. Diacamma vagans var. birmanum Emery, 1887b: 441 (w.) MYANMAR. Type-material: syntype workers (number not stated). Type-locality: none given in original description. [Note: type-locality is Myanmar (“Birmania”): Minhla (Comotto), after Emery, 1897b: 157.] Type-depository: MSNG. [Misspelled as burmanum by Bingham, 1903: 82.] As unavailable (infrasubspecific) name: Emery, 1897b: 157; Forel, 1900d: 320; Forel, 1911a: 23; Emery, 1911d: 67; Forel, 1912n: 52; Forel, 1913k: 7; Viehmeyer, 1916a: 113; Wheeler, W.M. 1919e: 54; Wheeler, W.M. 1927b: 42; Chapman & Capco, 1951: 59 (second entry). Subspecies of vagans: Dalla Torre, 1893: 30; Bingham, 1903: 82. Subspecies of rugosum: Chapman & Capco, 1951: 55 (first entry); Bolton, 1995b: 169. Distribution: Indonesia (Sumatra), Malaysia (Peninsula, Sarawak), Myanmar, Singapore. bispinosum. Ponera bispinosa Le Guillou, 1842: 317 (w.) INDONESIA (Ternate I.). Type-material: holotype(?) worker. [Note: no indication of number of specimens is given.] Type-locality: Indonesia: Ternate (Moluques) (no collector’s name). Type-depository: MNHN. Combination in Diacamma: Mayr, 1862: 718. Status as species: Roger, 1860: 305; Roger, 1861b: 168; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Smith, F. 1871a: 323; Emery, 1887b: 436 (in key, footnote), 441; Dalla Torre, 1893: 28; Emery, 1897b: 152; Emery, 1911d: 65; Chapman & Capco, 1951: 53; Wilson, 1958d: 366; Bolton, 1995b: 169. Senior synonym of buruensis: Wilson, 1958d: 367; Bolton, 1995b: 169. Senior synonym of subsulcata: Wilson, 1958d: 367; Bolton, 1995b: 169. Distribution: Indonesia (Buru, Halmahera, Ternate). brevispinum. Diacamma rugosum var. brevispina Chapman & Capco, 1951: 55. Type-material: syntype workers (number not stated). Type-locality: Myanmar (“Birmanie”): Moulmein (Hodgson). Type-depository: MHNG. [First available use of Diacamma rugosum r. geometricum var. brevisipnum Forel, 1900d: 320 (w.) MYANMAR; unavailable (infrasubspecific) name (Bolton, 1995b: 169).] [Misspelled as brevisipnum by Forel, 1900d: 320.] As unavailable (infrasubspecific) name: Emery, 1911d: 66. Distribution: Myanmar. brevistriatum. Diacamma brevistriatum Laciny, Pal & Zettel, 2015: 112, figs. 35-39 (w.) WEST MALAYSIA. Type-material: holotype worker, 58 paratype workers. Type-locality: holotype Malaysia: Perak, Cameron Highland, 40 km. SE Ipoh, Banjaran Titi Wangsa, Ringlet, 900 m., 25.iii.-3.iv.2002 (P. Cechovsky); paratypes: 28 workers with same data, 18 workers with same data but 25.iv.-5.v.2001(P. Cechovsky), 1 worker with same data but 25 km. NE Ipoh, Banjaran Titi Wangsa Mts, Korbu Mt., 1200 m., 27.i.-2.ii.1999 (P. Cechovsky), 1 worker Selangor, N Kuala Lumpur, Ulu Gombak, 16.ii.1993, #1 (H. Zettel), 5 workers Kelantan, 30 km. NW Gua Musang, Ulu Lalat, Kampong Sungai Om, 800-1000 m., 21.vi.-14.vii.2010 (P. Cechovsky), 1 worler as last but 22.v.-14.vi.2012, 2 workers Pahang, 50 km. NE Kuala Rompin, Endau Rompin N.P., Gunung Kerung, Kampong Tebu Hitam, 400 m., 9-30.iv.2008 (P. Cechovsky), 2 workers Pahang, 30 km. SE Ipoh, Banjaran Titi Wangsa, Tanah Rata, 1500 m., 14-15.iii.2002 (P. Cechovsky). Type-depositories: NHMW (holotype); HSCZ, NHMW (paratypes). Status as species: Zettel, et al. 2016: 166. Distribution: Malaysia (Peninsula). burmanum, misspelling, see under birmanum. buruensis. Diacamma rugosum subsp. buruensis Karavaiev, 1925b: 117 (w.) INDONESIA (Buru I.). Type-material: syntype workers (number not stated, “several”). Type-locality: Indonesia: Tifoe, Boeroe (= Buru), Nr. 2734 (W. Karawaiew). Type-depository: SIZK. Subspecies of rugosum: Karavaiev, 1935a: 70 (redescription); Chapman & Capco, 1951: 57. Junior synonym of bispinosum: Wilson, 1958d: 367; Bolton, 1995b: 169. caeruleum. Diacamma caeruleum Laciny, Pal & Zettel, 2015: 128, figs. 58-63 (w.) PHILIPPINES (Mindoro I.). Type-material: holotype worker, 17 paratype workers. Type-locality: holotype Philippines: Mindoro Oriental, 28 km. S Calapan, environment of Balete, 100-700 m., 27-29.xi.1992, #18 (H. Zettel); paratypes with same data. Type-depositories: NMPM (holotype); HSCZ, NHMW (paratypes). Distribution: Philippines (Mindoro). carbonarium. Diacamma carbonarium Laciny, Pal & Zettel, 2015: 131, figs. 57, 64-67 (w.) PHILIPPINES (Luzon I.). Type-material: holotype worker, 30 paratype workers. Type-locality: holotype Philippines: Luzon, Quezon Prov., Quezon Nat. Park, Old Zigzag Road, 27- 28.i.2002, #300 (H. Zettel); paratypes: 3 workers with same data, 6 workers with same data but 12- 13.ii.1996, #79a, 5 workers with same data but 24-30.iii.1998, #165, 1 worker with same data but 16.iii.1999, #202 (S.S. Vichozo & H. Zettel), 1 worker with same data but 9.iv.2000, #253 (H. Zettel), 5 workers with same data but 14.iii.1999, #27 (F. Seyfert), 1 worker Laguna, San Pablo, Mt Banahaw 15.xi.1992, #6 (H. Zettel), 1 worker Nueva Viscaya, Santa Fe, Imugan, 1200 m., 10.xi.2002, #202 (H. Zettel), 1 worker Laguna, Cavinti, Tibatib 26.vi.2010, #202 (C.V. Pangantihon), 1 worker Camarines Sur, 20 km. E Naga, 5 km. E Caolina, Mt Isarog, nr Malabsay Falls 4.iii.1999, #192 (H. Zettel), 2 workers Camarines Norte, SW Daet, San Vicente, Fabrica, Mananap, 6.ii.2001, #264 (H. Zettel & S.S. Vichozo), 3 workers Camarines Norte, Cabo, Tulay na Lupa, Mt Labo-Mt Bayabas area 17-18.iii.2004, #202 (H. Zettel & C.V. Pangantihon). Type-depositories: NMPM (holotype); HSZC, NHMW (paratypes). Distribution: Philippines (Luzon). celebense. Diacamma celebense Emery, 1887b: 441 (w.) INDONESIA (Sulawesi). Type-material: holotype(?) worker. [Note: no indication of number of specimens is given.] Type-locality: Indonesia: Selebes (= Sulawesi), Kandari (O. Beccari). Type-depository: MSNG. Status as species: Dalla Torre, 1893: 28. Subspecies of rugosum: Emery, 1897b: 156; Emery, 1901g: 566; Forel, 1907e: 17; Emery, 1911d: 66; Chapman & Capco, 1951: 57; Bolton, 1995b: 169. Distribution: Indonesia (Sulawesi). ceylonense. Diacamma rugosum subsp. ceylonense Emery, 1897b: 159 (w.) SRI LANKA. Type-material: syntype workers (number not stated). Type-locality: Sri Lanka (“Ceilan”): (no further data) (H. Nietner). Type-depositories: MNHU, MSNG. [Misspelled as ceylonese by Teranishi, 1940: 66.] Subspecies of rugosum: Forel, 1900d: 318; Emery, 1911d: 66; Chapman & Capco, 1951: 57; Tiwari, 1999: 25. Status as species: Bingham, 1903: 79; Teranishi, 1940: 66; Bolton, 1995b: 169; Bharti, Guénard, et al. 2016: 49; Zettel, et al. 2016: 134 (redescription). Distribution: India, Sri Lanka. coincidens. Diacamma rugosum subsp. geometrica var. coincidens Baroni Urbani, 1971b: 360. [Unavailable (infrasubspecific) replacement name for concentrica Karavaiev, 1935a: 65; unavailable (infrasubspecific) name (Bolton, 1995b: 169).] colosseense. Diacamma australe var. colosseensis Forel, 1915b: 26 (w.) AUSTRALIA (Queensland). Type-material: lectotype worker (by designation of Shattuck & Barnett, 2006: 16), 2 paralectotype workers. Type-locality: lectotype Australia: Queensland, Gin-Gin (E. Mjöberg); paralectotypes with same data. Type-depository: MHNG. Junior synonym of australe: Taylor & Brown, 1985: 29; Taylor, 1987a: 24; Bolton, 1995b: 169. Status as species: Shattuck & Barnett, 2006: 15 (redescription). Distribution: Australia. compressum. Diacamma compressum Mayr, 1879: 660 (w.) PAKISTAN (Sindh). Type-material: lectotype worker (by designation of Laciny, Pal & Zettel, 2015: 90), 1 paralectotype worker. Type-locality: lectotype Pakistan (“Ostindien”): Sindh, Plason, 1874 (no collector’s name); paralectotype with same data. Type-depository: NHMW. Status as species: Chapman & Capco, 1951: 54. Junior synonym of scalpratum: Dalla Torre, 1893: 29; Emery, 1911d: 65; Bolton, 1995b: 169; Laciny, Pal & Zettel, 2015: 90. concentricum. Diacamma palawanicum var. concentricum Wheeler, W.M. & Chapman, 1925: 64, pl. 2, figs. 7, 8 (w.) PHILIPPINES (Palawan I.). Type-material: 5 syntype workers. Type-localities Philippines: Binaluan (G. Boettcher), and Malampaya (W. Schultze). Type-depository: MCZC. Subspecies of palawanicum: Chapman & Capco, 1951: 54; Baltazar, 1966: 241; Bolton, 1995b: 169. Status as species: Laciny, Pal & Zettel, 2015: 99 (redescription). Senior synonym of sericeiventre: Laciny, Pal & Zettel, 2015: 99. Distribution: Philippines (Palawan). concentrica. Diacamma rugosum subsp. geometricum var. concentrica Karavaiev, 1935a: 65 (w.) VIETNAM; unavailable (infrasubspecific) name. [Note: material included: Vietnam (“Zentralannam”): Bana, nr Tourane, 500-800 m., Nr. 5763 (K. Davydov); 1 worker, deposited in SIZK (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 57. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 169. [Diacamma rugosum subsp. geometrica var. coincidens Baroni Urbani, 1971b: 360. Unavailable (infrasubspecific) replacement name for concentrica Karavaiev; unavailable (infrasubspecific) name.] convexa. Diacamma rugosum st. sculpturata var. convexa Santschi, 1920g: 179 (w.) VIETNAM; unavailable (infrasubspecific) name. [Note: material included: Vietnam (“Cochinchine”): Dalat (Bouvard); 2 workers, deposited in NHMB (no type status).] As unavailable (infrasubspecific) name: Santschi, 1920h: 158; Santschi, 1924c: 96; Santschi, 1928h: 124; Chapman & Capco, 1951: 59. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 169. cupreum. Ponera cuprea Smith, F. 1860b: 104 (w.) NEW GUINEA (Indonesia). Type-material: holotype worker. Type-locality: Indonesia: N. Guinea (=Irian Jaya), Dory (= Manokwari), “Dor 19” (A.R. Wallace). Type-depository: OXUM. Combination in Diacamma: Mayr, 1886c: 361. As unavailable (infrasubspecific) name: Emery, 1897b: 160; Emery, 1897d: 550; Emery, 1911b: 531; Emery, 1911d: 66; Viehmeyer, 1912: 17. Subspecies of geometricum: Emery, 1887b: 436 (in key, footnote), 438; Emery, in Dalla Torre, 1893: 28 (footnote). Subspecies of rugosum: Chapman & Capco, 1951: 55. Status as species: Smith, F. 1863: 17; Mayr, 1863: 448; Smith, F. 1865: 70; Smith, F. 1871a: 323; Mayr, 1886c: 361; Dalla Torre, 1893: 28; Donisthorpe, 1932c: 462; Wilson, 1958d: 367; Bolton, 1995b: 169. Distribution: Indonesia (Bacan, Irian Jaya, Seram), Papua New Guinea. curtula. Diacamma rugosum r. geometricum var. curtula Forel, 1912d: 100 (w.) INDONESIA (Java); unavailable (infrasubspecific) name. [Note: material included: Indonesia: Java, Nongkodjadjar, i. (E. Jacobson); unknown number of workers, deposited in MHNG (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 57. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 169. cyaneiventre. Diacamma cyaneiventre André, 1887: 293 (w.) INDIA (Tamil Nadu). Type-material: syntype workers (number not stated). Type-locality: India: Coimbatore (no collector’s name). [Note: locality Coimbatore from Emery, 1897b: 153.] Type-depository: MNHN. [Misspelled as cyaniventre by Emery, 1897b: 153; misspelled as cyaneventre by Teranishi, 1940: 65.] Subspecies of rugosum: Forel, 1900d: 318. Status as species: Dalla Torre, 1893: 28; Emery, 1897b: 153; Bingham, 1903: 78; Emery, 1911d: 65; Teranishi, 1940: 65; Chapman & Capco, 1951: 54; Bolton, 1995b: 169; Tiwari, 1999: 26; Bharti, Guénard, et al. 2016: 49. Distribution: India. debilior. Diacamma rugosum subsp. hortensis var. debilior Karavaiev, 1927e: 5 (w.) INDONESIA (Java: Panaitan (= Prinsen) I.); unavailable (infrasubspecific) name. [Note: material included: Indonesia: Prinsen I., Sunda Strait, 5.i.1913, Nr. 3017 (W. Karawajew); “several” workers, deposited in SIZK (no type status).] As unavailable (infrasubspecific) name: Karavaiev, 1935a: 64; Chapman & Capco, 1951: 58. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 169. divergens. Diacamma rugosum var. divergens Chapman & Capco, 1951: 55. Type-material: syntype workers (number not stated). Type-localities: New Guinea (“Nuova Guinea”) (no state data but probably Indonesia: Irian Jaya), and Indonesia: Halmahera (no collector’s name). Type-depository: MSNG. [First available use of Diacamma rugosum subsp. sculpturatum var. divergens Emery, 1897b: 160 (w.) NEW GUINEA (no state data), INDONESIA (Halmahera I.); unavailable (infrasubspecific) name.] As unavailable (infrasubspecific) name: Emery, 1897d: 550; Emery, 1911d: 67; Viehmeyer, 1912: 17. Junior synonym of rugosum by Wilson, 1958d: 368; Bolton, 1995b: 169; Zhou, 2001b: 45. doveri. Diacamma vagans var. doveri Mukerjee, 1934: 3 (w.) INDIA (Karnataka). Type-material: holotype worker. Type-locality: India: Karnataka, Fraserpet (= Kushalanagar), COORG (= North Coorg forest division), 7.viii.1930, ca 3700 ft, F.R.I. Sandal Insect Survey (no collector’s name). Type-depository: unknown (possibly in NZSI). Subspecies of rugosum: Bolton, 1995b: 169; Bharti, Guénard, et al. 2016: 50. Distribution: India. excellens. Diacamma excellens Zettel, et al. 2016: 155, figs. 34-37 (w.) PHILIPPINES (Biliran I.). Type-material: holotype worker, 17 paratype workers. Type-locality: holotype Philippines: Biliran, Almeria, Ilihan, 13.xi. (C.V. Pangantihon); paratypes with same data. Type-depositories: NMPM (holotype); HSZC, NHMW, TNHM (paratypes). Distribution: Philippines (Biliran). frontalis. Diacamma vagans var. frontalis Stitz, 1911a: 355 (w.) NEW GUINEA (no state data). Type-material: holotype worker. Type-locality: New Guinea: (no further data) (Werner). Type-depository: MNHU. As unavailable (infrasubspecific) name: Viehmeyer, 1912: 17. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 169. geminatum. Diacamma rugosum subsp. geminatum Emery, 1897b: 161, figs. 6, 12 (w.) INDONESIA (Halmahera I.). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Halmahera (O. Beccari). Type-depository: MSNG. Subspecies of rugosum: Emery, 1911d: 66; Chapman & Capco, 1951: 57. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 169; Zhou, 2001b: 45. generali. Diacamma generali Laciny, Pal & Zettel, 2015: 125, figs. 52-57 (w.) PHILIPPINES (Luzon I.). Type-material: holotype worker, 98 paratype workers. Type-locality: holotype Philippines: Luzon, Laguna Prov., Los Baños, Mt Makiling, Mud Springs, ca 650 m., 13.ii.1999, #2 (S. Schödl); paratypes: 4 workers with same data, 6 workers with same data but 23- 24.i.1999, #167 (H. Zettel), 3 workers with same data but 1.xii.2000, #259 (H. Zettel), 1 worker Mt Makiling, 13.xi. 1992, #2 (H. Schillhammer), 12 workers Mt Makiling, 13-18.xi.1992, #1 (H. Zettel), 2 workers UPLB University Campus, 12-14.ii.1999, #1 (S. Schödl), 7 workers Mt Makiling, Flat Rocks, 14.ii.1999, #4 (S. Schödl); 63 other workers in very extensive list, all from northern and central Luzon. Type-depositories: NMPM (holotype); FMNH, HSZC, NHMW, UPLB (paratypes). Status as species: Zettel, et al. 2016: 166. Distribution: Philippines (Luzon). geometricum. Ponera geometrica Smith, F. 1857a: 67 (w.) SINGAPORE. Type-material: holotype worker. Type-locality: Singapore: “SING 3” (A.R. Wallace). Type-depository: OXUM. Wheeler, W.M. 1919e: 54 (m.); Emery, 1899c: 5 (l.). Combination in Diacamma: Mayr, 1862: 718. Subspecies of rugosum: Emery, 1897b: 154; Forel, 1900d: 320; Emery, 1901g: 566; Forel, 1907e: 17; Emery, 1911d: 66; Forel, 1912d: 100; Wheeler, W.M. 1919e: 54; Wheeler, W.M. & Chapman, 1925: 63; Wheeler, W.M. 1927d: 1; Wheeler, W.M. 1928c: 5; Wheeler, W.M. 1930h: 58; Karaviev, 1935a: 62 (redescription); Yasumatsu, 1940a: 67; Chapman & Capco, 1951: 57; Baltazar, 1966: 242. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 169; Zhou, 2001b: 45. Status as species: Smith, F. 1858b: 86; Smith, F. 1860a: 72; Roger, 1860: 301; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Smith, F. 1871a: 321; Mayr, 1886c: 358; Emery, 1887b: 435 (in key, footnote), 436; Dalla Torre, 1893: 28; Emery, 1893e: 189; Donisthorpe, 1932c: 448; Radchenko, 1993a: 80; Laciny, Pal & Zettel, 2015: 102 (redescription); Zettel, et al. 2016: 167. Senior synonym of tritschleri: Laciny, Pal & Zettel, 2015: 102. Distribution: China, Indonesia (Ambon, Java, Prinsen, Sulawesi, Sumatra), Malaysia (Peninsula, Sabah), Philippines (Luzon, Mindanao, Negros, Panay), Singapore, Sri Lanka, Vietnam. [Note: Laciny, Pal & Zettel, 2015: 106, restrict distribution to West Malaysia, Singapore and Sumatra.] gibbosum. Diacamma rugosum subsp. gibbosum Karavaiev, 1935a: 68, fig. 4 (w.) VIETNAM. Type-material: holotype worker. Type-locality: Vietnam (“Südannam”): Dalat, Lang Biang, 1500 m., Nr. 5760 (K. Davydov). Type-depository: SIZK. Subspecies of rugosum: Chapman & Capco, 1951: 58; Bolton, 1995b: 169. Distribution: Vietnam. gusenleitneri. Diacamma gusenleitneri Zettel, et al. 2016: 132, figs. 1-4 (w.) BORNEO (Indonesia: Kalimantan). Type-material: holotype worker. Type-locality: Indonesia: “Borneo” (= Kalimantan Timur, Berau Regency, Segah), Gunung Sari, 17.viii.1956, Wegner Expd. (H.H.F. Hamann). Type-depository: OLLA. Distribution: Indonesia (Kalimantan). holosericum. Ponera holoserica Roger, 1860: 302 (w.) INDONESIA (Java). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Java (no further data). Type-depository: MNHU. [Unjustified emendation of spelling to holosericea: Mayr, 1862: 718; unjustified emendation of spelling to holosericeum: Mayr, 1863: 407, and others.] Combination in Diacamma: Mayr, 1862: 718. Status as species: Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Mayr, 1867a: 87 (redescription); Smith, F. 1871a: 323; Mayr, 1872: 149; Mayr, 1879: 660; Emery, 1887b: 435 (in key, footnote); Dalla Torre, 1893: 29; Emery, 1897b: 163 (in key); Emery, 1900d: 666; Emery, 1911d: 65; Wheeler, W.M. 1919e: 52; Crawley, 1924: 384; Stitz, 1925: 111; Karavaiev, 1925b: 115 (redescription); Chapman & Capco, 1951: 54; Bolton, 1995b: 169; Pfeiffer, et al. 2011: 55. Distribution: Malaysia (Sarawak), Indonesia (Java, Mentawei, Sumatra). holzschuhi. Diacamma holzschuhi Laciny, Pal & Zettel, 2015: 115, figs. 40-43 (w.) LAOS. Type-material: holotype worker, 23 paratype workers. Type-locality: holotype Laos: Hua Phan Prov., Ban Saleui, Phou Pan Mts, 1300-1900 m., 20°12’N, 104°01’E, 28.iv.2014 (C. Holzschuh); paratypes with same data. Type-depositories: NHMW (holotype); HSZC, NHMW (paratypes). Status as species: Jaitrong, Guénard, et al. 2016: 42. Distribution: Laos. homonyma. Diacamma rugosum subsp. sculpturata var. homonyma Baroni Urbani, 1971b: 360. [Unavailable (infrasubspecific) replacement name for papuana Karavaiev, 1935a: 67; unavailable (infrasubspecific) name (Bolton, 1995b: 170).] horizontalis. Diacamma rugosum subsp. geometricum var. horizontalis Karavaiev, 1935a: 63 (w.) VIETNAM; unavailable (infrasubspecific) name. [Note: material included: Vietnam (“Zentralannam”): Lien-Chieu, nr Tourane, base of Col de Nuage, Nr. 5636 (K. Davydov); 1 worker, deposited in SIZK (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 58. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 170. hortense. Diacamma rugosum subsp. hortensis Karavaiev, 1925b: 119. Type-material: syntype workers (number not stated). Type-locality: Indonesia: Java, Buitenzorg (= Bogor) (H. v. Buttel-Reepen). Type-depository: MHNG. [First available use of Diacamma rugosum r. vagans var. hortensis Forel, 1913k: 7 (w.) INDONESIA (Java); unavailable (infrasubspecific) name.] Subspecies of rugosum: Karavaiev, 1925b: 119 (redescription); Chapman & Capco, 1951: 58; Bolton, 1995b: 170. Distribution: Indonesia (Java). indicum. Diacamma rugosum var. indica Santschi, 1920g: 179. Type-material: syntype workers (number not stated). Type-localities: India: South Andaman Is, Lokolot Goiaba, Shaal Bay (no collector’s name), and South Andaman Is, Kyd I. (no collector’s name). Type-depository: MHNG. [First available use of Diacamma rugosum r. vagans var. indicum Forel, 1903d: 400 (w.) INDIA (Andaman Is); unavailable (infrasubspecific) name.] As unavailable (infrasubspecific) name: Forel, 1908a: 1; Emery, 1911d: 67; Forel, 1911i: 216; Donisthorpe, 1915d: 335; Donisthorpe, 1927b: 386; Chapman & Capco, 1951: 59. Subspecies of rugosum: Chapman & Capco, 1951: 55; Zhou, 2001b: 45 (error). Junior synonym of vagans: Mukerjee & Ribeiro, 1925: 205; Menozzi, 1939a: 328; Bolton, 1995b: 170. Status as species: Doums, 1999: 1958; Bharti, Guénard, et al. 2016: 50. Distribution: India (Andaman Is), Vietnam. intricatum. Ponera intricata Smith, F. 1857a: 67 (w.) BORNEO (East Malaysia: Sarawak). Type-material: 2 syntype workers. Type-locality: Malaysia (“Borneo”): Sarawak, “SAR” (A.R. Wallace). Type-depositories: BMNH, OXUM. [Note: two further worker specimens, possibly members of the original type-series, are in BMNH. One bears a Donisthorpe label, “compared with type Diacamma intricata Smith,” and is also labelled, “SAR” and “Borneo 57/36.” The second (gaster missing) has a Farren White label, and also “SAR 16.”] Combination in Diacamma: Mayr, 1862: 718. Status as species: Smith, F. 1858b: 88; Roger, 1860: 305; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Smith, F. 1871a: 321; Mayr, 1872: 150; Mayr, 1886c: 358; Emery, 1887b: 435 (in key, footnote); André, 1892b: 52; Dalla Torre, 1893: 29; Emery, 1897b: 163 (in key); Forel, 1911e: 253; Emery, 1911d: 65; Wheeler, W.M. 1919e: 52; Stitz, 1925: 111; Donisthorpe, 1932c: 448; Chapman & Capco, 1951: 54; Bolton, 1995b: 170; Jaitrong & Nabhitabhata, 2005: 19; Pfeiffer, et al. 2011: 55. Distribution: Indonesia (Kalimantan, Sumatra), Malaysia (Sabah, Sarawak), Thailand. Current subspecies: nominal plus kershawi. jacobsoni. Diacamma jacobsoni Forel, 1912d: 100 (w.) INDONESIA (Java). Type-material: holotype(?) worker. Type-locality: Indonesia: Java, Nongkodjadjar, i. (E. Jacobson). Type-depository: MHNG. Status as species: Chapman & Capco, 1951: 54; Bolton, 1995b: 170. Distribution: Java. jaitrongi. Diacamma jaitrongi Zettel, et al. 2016: 141, figs. 14-17 (w.) THAILAND. Type-material: holotype worker, 14 paratype workers. Type-locality: holotype Thailand: Chiang Mai Prov., Fang Dist., Pha Hom Pok Nat. Park, 29.v.2008, WJT08-N62 (W. Jaitrong); paratypes: 6 workers with same data, 1 worker with same data but 27.v.2008, 7 workers with same data but 28.v.2008, WJT08-N41. Type-depositories: TNHM (holotype); HSZC, NHMW, TNHM (paratypes). Distribution: Thailand. japensis. Diacamma rugosum subsp. japensis Donisthorpe, 1941h: 52 (w.) NEW GUINEA (Indonesia: Yapen I.). Type-material: holotype worker. Type-locality: Indonesia: Irian Jaya (“Dutch New Guinea”), Japen I., Mt Baduri, 1000 ft, viii.1938 (L.E. Cheesman). Type-depository: BMNH. Subspecies of rugosum: Chapman & Capco, 1951: 58. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 46. javanum. Diacamma javanum Emery, 1887b: 439, pl. 1, fig. 6 (w.) INDONESIA (Java). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Java, Buitenzorg (= Bogor) (Ferrari). Type-depository: MSNG. Status as species: Dalla Torre, 1893: 29. Junior synonym of geometrica: Emery, 1897b: 154; Forel, 1900d: 320; Emery, 1911d: 66; Wheeler, W.M. 1919e: 54. Subspecies of rugosum: Forel, 1905c: 6; Chapman & Capco, 1951: 58; Bolton, 1995b: 170. Distribution: Indonesia (Java). jerdoni. Diacamma rugosum var. jerdoni Forel, 1903d: 401 (in text) (w.) INDIA (no state data). Type-material: holotype(?) worker. [Note: no indication of number of specimens is given.] Type-locality: India: (no further data). Type-depository: MHNG. Subspecies of rugosum: Forel, 1909e: 392; Emery, 1911d: 66; Forel, 1912n: 52; Forel, 1913k: 7; Donisthorpe, 1942d: 449; Donisthorpe, 1943b: 196; Chapman & Capco, 1951: 55; Bolton, 1995b: 170; Tiwari, 1999: 25; Bharti, Guénard, et al. 2016: 50. Synonym of sculptum: Emery, 1911d: 66. [Note: Emery gives jerdoni as senior synonym, but if the synonymy is correct then sculptum has priority.] kershawi. Diacamma intricatum subsp. kershawi Wheeler, W.M. 1919e: 53 (w.) BORNEO (East Malaysia: Sabah). Type-material: holotype worker. Type-locality: Malaysia: Sabah (“British North Borneo”) (E.B. Kershaw). Type-depository: MCZC. Subspecies of intricatum: Chapman & Capco, 1951: 54; Bolton, 1995b: 170; Pfeiffer, et al. 2011: 55. Distribution: Malaysia (Sabah). latispina. Diacamma rugosum subsp. latispina Karavaiev, 1925b: 119 (w.) INDONESIA (Java). Type-material: holotype worker. Type-locality: Indonesia: Java, Tjiapus on the Salak, nr Buitenzorg (= Bogor) (W. Karawaiew). Type-depository: SIZK. Subspecies of rugosum: Karavaiev, 1935a: 67 (redescription); Chapman & Capco, 1951: 56; Bolton, 1995b: 170. Distribution: Indonesia (Java). leve. Diacamma australe var. levis Crawley, 1915a: 134 (w.) AUSTRALIA (Northern Territory). Type-material: holotype(?) worker. [Note: no indication of number of specimens is given.] Type-locality: Australia: Northern Territory, nr Adelaide Plains, 1.vi.1913 (G.F. Hill). Type-depository: BMNH or OXUM. Junior synonym of australe: Taylor & Brown, 1985: 29; Taylor, 1987a: 24; Bolton, 1995b: 170. Status as species: Shattuck & Barnett, 2006: 17 (redescription). Distribution: Australia. leyteense. Diacamma leyteense Zettel, et al. 2016: 149, figs. 26-29 (w.) PHILIPPINES (Leyte I., Samar I.). Type-material: holotype worker, 16 paratype workers. Type-locality: holotype Philippines: Leyte Prov., E Ormoc, Lake Danao, 13.ii.2000 (S. Schödl); paratypes: 3 workers with same data, 2 workers with same data but (H. Zettel), 1 worker Leyte, Baybay, Mt Pangasugan, Calbiga-a River, 50-200 m., 12.ii. (H. Zettel), 1 worker Baybay, Leyte State University, 16.x. (C.V. Pangantihon), 3 workers S Layte Prov., Ibarra, Divisoria, 21.xi. (H. Zettel & C.V. Pangantihon), 6 workers W Samar Prov., Basey, Sohoton Nat. Park, 29.i.2000 (S. Schödl). Type-depositories: NHMW (holotype); HSZC, NHMW, NMPM, TNHM (paratypes). Distribution: Philippines (Leyte, Samar). lombokense. Diacamma rugosum var. lombokensis Emery, 1897b: 154 (w.) INDONESIA (Lombok I.). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Lombok, Sambalun, 1300 m. (Fruhstorfer). Type-depository: MSNG. Subspecies of rugosum: Forel, 1907e: 17; Emery, 1911d: 66; Chapman & Capco, 1951: 56; Bolton, 1995b: 170. Distribution: Indonesia (Lombok). longiceps. Diacamma geometricum var. longiceps Santschi, 1932b: 14 (w.) VIETNAM. Type-material: 7 syntype workers. Type-locality: Vietnam: Tonkin, Hanoi (Demange). Type-depository: NHMB. As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 58. Subspecies of rugosum: Bolton, 1995b: 170. Distribution: Vietnam. longitudinale. Diacamma longitudinale Emery, 1889b: 496 (footnote) (w.) VIETNAM. Type-material: holotype worker. Type-locality: Vietnam (“Cocincina”): (no further data). Type-depository: MSNG. Status as species: Dalla Torre, 1893: 29; Emery, 1897b: 163 (in key); Forel, 1900d: 319; Emery, 1911d: 65; Forel, 1911d: 378; Forel, 1911i: 216 (footnote); Santschi, 1920h: 158; Santschi, 1924c: 95; Chapman & Capco, 1951: 54; Bolton, 1995b: 170; Jaitrong, Guénard, et al. 2016: 42. Distribution: Laos, Vietnam. macreta. Diacamma rugosum var. macreta Chapman & Capco, 1951: 56. Type-material: 3 syntype workers. Type-locality: Papua New Guinea (“Deutsch Neuguinea”): (no further data) (O. Schlaginhaufen). [Note: Wilson, 1958d: 368, provides the location data: Torricelli Mts, 640 m.] Type-depository: MNHU. [First available use of Diacamma rugosum subsp. sculpturatum var. macreta Viehmeyer, 1912: 6 (w.) NEW GUINEA (Papua New Guinea); unavailable (infrasubspecific) name.] Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. magdalenae. Diacamma magdalenae Laciny, Pal & Zettel, 2015: 86, figs. 1-6 (w.) BORNEO (East Malaysia: Sarawak; Indonesia: Kalimantan). Type-material: holotype worker, 10 paratype workers. Type-locality: holotype Malaysia (“Borneo”): Sarawak, Gunung Mulu Nat. Park, 04°02’30’’N, 114°48’46’’E, 13.xi.2009, alluvial forest, #82 (D.M. Sorger); paratypes: 3 workers with same data, 2 workers G. Mulu Nat. Park, 04°02’30’’N, 114°52’15’’E, 23-26.x.2009, dipterocarp forest, #20 (D.M. Sorger), 1 worker G. Mulu Nat. Park, 04°02’30’’N, 114°48’46’’E, 22.x.2009, alluvial/kerangas/limestone forest, along nightwalk trail, #12 (D.M. Sorger), 1 worker as last but 22.x.2009, #14, 1 worker as last but 11.xi.2009, #74, 1 worker G. Mulu Nat. Park, along way to Camp 5, 13-15.vii.2010, #GM10-H11 (D.M. Sorger), 1 worker Borneo, W Kalimantan, Kapuas, “Ban de Kapouas”, 1896 (M.A. Chaper). Type-depositories: NHMW (holotype); DMSC, HSZC, MSNG, NHMW (paratypes). Distribution: Indonesia (Kalimantan), Malaysia (Sarawak). moica. Diacamma rugosum subsp. ceylonense var. moica Forel, 1911d: 377 (w.) VIETNAM; unavailable (infrasubspecific) name. [Note: material included: Vietnam (“Cochinchine”): Pays de Mois (Dugas); 2 workers, deposited in MHNG (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 57. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 170. [Note: Zettel, et al. 2016: 136, 141, suggest that moicum was made available by Santschi, 1932b: 14. But in Santschi’s text he merely mentions “la var. moicum”, which is interpreted here as an abbreviated writing of the full quadrinomial, not as a statement of change of taxonomic status.] Material referred to orbiculatum by Zettel, et al. 2016: 136. nitidiventre. Diacamma rugosum var. nitidiventris Chapman & Capco, 1951: 56. Type-material: syntype workers (number not stated). Type-locality: Indonesia: Timor, Cupan (Loria). Type-depository: MSNG. [First available use of Diacamma rugosum subsp. timorense var. nitidiventris Emery, 1897b: 158 (w.) INDONESIA (Timor I.); unavailable (infrasubspecific) name (Bolton, 1995b: 170).] As unavailable (infrasubspecific) name: Emery, 1897d: 551; Emery, 1911d: 67. Distribution: Indonesia (Timor). oblicum. Diacamma rugosum var. oblica Chapman & Capco, 1951: 56. Type-material: holotype workers. Type-locality: Vietnam: Tonkin, Hanoi (Fouquet). Type-depository: NHMB. [First available use of Diacamma rugosum st. sculpturata var. oblica Santschi, 1920g: 178 (w.) VIETNAM; unavailable (infrasubspecific) name (Bolton, 1995b: 170).] [Misspelled as obliqua by Santschi, 1920h: 158, Santschi, 1924c: 96, Chapman & Capco, 1951: 59, second entry of name (as unavailable (infrasubspecific) name).] As unavailable (infrasubspecific) name: Santschi, 1920h: 158; Santschi, 1924c: 96. Distribution: Vietnam. orbiculatum. Diacamma ceylonensis var. orbiculatum Santschi, 1932b: 14 (w.) LAOS. Type-material: holotype worker. Type-locality: Laos: Luang Prabang (Vitalis de Salvaza). Type-depository: NHMB. As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 57. Subspecies of ceylonense: Bolton, 1995b: 170; Jaitrong, Guénard, et al. 2016: 40. Status as species: Zettel, et al. 2016: 136 (redescription). Material of the unavailable name moica referred here by Zettel, et al. 2016: 136. Distribution: Laos, Thailand, Vietnam. ovale. Diacamma rugosum var. ovalis Chapman & Capco, 1951: 56. Type-material: holotype workers. Type-locality: Vietnam (“Südannam”): Danlac Prov., Ba-Me-Thuot, 700 m., Nr.5920 (K. Davydov). Type-depository: SIZK. [First available use of Diacamma rugosum subsp. rugosum var. ovalis Karavaiev, 1935a: 62 (w.) VIETNAM; unavailable (infrasubspecific) name.] Distribution: Vietnam. palawanicum. Diacamma palawanicum Emery, 1900d: 666 (footnote) (w.) PHILIPPINES (Palawan I.). Type-material: holotype workers. Type-locality: Philippines: Palawan, 1898 (Doherty). Type-depository: MSNG. Status as species: Emery, 1911d: 65; Wheeler, W.M. & Chapman, 1925: 64; Chapman & Capco, 1951: 54; Baltazar, 1966: 242; Bolton, 1995b: 170; Laciny, Pal & Zettel, 2015: 97 (redescription). Distribution: Philippines (Palawan). pallidum. Ponera pallida Smith, F. 1858b: 85 (m.) MYANMAR. Type-material: holotype male. Type-locality: Myanmar (“Birmah”): “57/16” (Waring). Type-depository: BMNH. Combination in Leptogenys: Donisthorpe, 1932c: 474; combination in Pachycondyla: Brown, in Bolton, 1995b: 308; combination in Diacamma: Doums, 1999: 1958. Status as species: Mayr, 1863: 449; Smith, F. 1871a: 320; Dalla Torre, 1893: 40; Emery, 1911d: 116; Donisthorpe, 1932c: 474; Bolton, 1995b: 308; Guénard & Dunn, 2012: 58. Distribution: China, Myanmar. panayense. Diacamma panayensis Wheeler, W.M. & Chapman, 1925: 65, pl. 2, figs. 9, 10 (w.) PHILIPPINES (Panay I.). Type-material: 6 syntype workers. Type-locality: Philippines: Panay, Antique Prov., Culasi (McGregor). Type-depository: MCZC. Status as species: Chapman & Capco, 1951: 54; Baltazar, 1966: 242; Bolton, 1995b: 170. Distribution: Philippines (Panay). papuanum. Diacamma vagans var. papuanum Stitz, 1911a: 354, fig. 5 (w.) NEW GUINEA (Indonesia, Papua New Guinea). Type-material: syntype workers (number not stated). Type-localities: Indonesia:Irian Jaya, Mawa River (Moszkowski), and Papua New Guinea: “Kaiser Wilhelmsland” (Lauterbach). Type-depository: MNHU. As unavailable (infrasubspecific) name: Viehmeyer, 1912: 17; Donisthorpe, 1948b: 299; Chapman & Capco, 1951: 60. Subspecies of vagans: Stitz, 1912: 501. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. papuana. Diacamma rugosum subsp. sculpturatum var. papuana Karavaiev, 1935a: 67 (w.) NEW GUINEA (Papua New Guinea). [Note: material included: New Guinea, ex coll. Emery (no further data); 1 worker, deposited in SIZK (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 59. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 170. [Diacamma rugosum subsp. sculpturata var. homonyma Baroni Urbani, 1971b: 360. Unavailable (infrasubspecific) replacement name for papuana Karavaiev; unavailable (infrasubspecific) name.] paralleliruga. Diacamma rugosum subsp. geometricum var. paralleliruga Karavaiev, 1935a: 64 (w.) VIETNAM; unavailable (infrasubspecific) name. [Note: material included: Vietnam (“Südannam”): Dalat, Lang Biang, 1500 m., 2.ii.1931, Nr. 5636 (K. Davydov); 1 worker, deposited in SIZK (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 58. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 170. pubescens. Diacamma rugosum var. pubescens Chapman & Capco, 1951: 56. Type-material: syntype workers (number not stated). Type-localities: Indonesia: Mentawei (Modigliani), and Malaysia: Pulo Pinang (= Penang I.) (Modigliani). Type-depository: MSNG. [First available use of Diacamma rugosum subsp. sculpturatum var. pubescens Emery, 1897b: 160 (w.) INDONESIA (Sumatra: Mentawei Is), MALAYSIA (Penang I.); unavailable (infrasubspecific) name (Bolton, 1995b: 170).] As unavailable (infrasubspecific) name: Emery, 1900d: 666; Forel, 1901f: 338 (in text); Emery, 1911d: 67; Karavaiev, 1930a: 212. Distribution: Indonesia (Mentawei), Malaysia (Penang). pulchellum. Diacamma rugosum st. sculpturatum var. pulchellum Santschi, 1932b: 12, fig. 1 (w.) NEW GUINEA (Indonesia); unavailable (infrasubspecific) name. [Note: material included: New Guinea: Lomira, 20.iii.1929 (no collector’s name); 1 worker, deposited in NHMB (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 59. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 170. Material referred to rugosum by Wilson, 1958d: 368. purpureum. Ponera purpurea Smith, F. 1863: 18 (w.) INDONESIA (Halmahera I.). Type-material: holotype worker. Type-locality: Indonesia: Gilolo (= Halmahera), “Gil” (A.R. Wallace). Type-depository: OXUM. Combination in Diacamma: Emery, 1887b: 439. Subspecies of cupreum: Emery, 1887b: 439; Dalla Torre, 1893: 28. Status as species: Smith, F. 1871a: 323; Emery, 1897b: 152; Emery, 1911d: 66; Donisthorpe, 1932c: 470; Chapman & Capco, 1951: 54; Wilson, 1958d: 367; Bolton, 1995b: 170. Senior synonym of waigeuensis: Bolton, 1995b: 170. Distribution: Indonesia (Halmahera, Waigeu). quezonicum. Diacamma viridipurpureum subsp. quezonicum Laciny, Pal & Zettel, 2015: 122, figs. 48-51, 56, 62, 63 (w.) PHILIPPINES (Luzon I.). Type-material: holotype worker, 41 paratype workers. Type-locality: holotype Philippines: Luzon, Quezon Prov., Atimonan, Quezon Nat. Park, Old Zigzag Road, 27-28.i.2002, #300 (H. Zettel); paratypes: 12 workers with same data, 1 worker with same data but 12- 13.ii.1996, #79a, 7 workers with same data but 24-30.iii.1998, #165, 3 workers with same data but 16.iii.1999, #202, 2 workers with same data but 6.iv.2000, #253, 14 workers with same data but 30- 31.i.2001, #262, 1 worker with same data but 14.iii.1999, #27 (H. Seyfert), 1 worker with same data but 22.xi.1992, #11 (M.A. Jäch). Type-depositories: NMPM (holotype); HSZC, NHMW, NMPM, UPBL (paratypes). Subspecies of viridipurpureum: Zettel, et al. 2016: 167. Distribution: Philippines (Luzon). reductostriatum. Diacamma reductostriatum Zettel, et al. 2016: 157, figs. 38-41 (w.) PHILIPPINES (Camiguin I.). Type-material: holotype worker, 20 paratype workers. Type-locality: holotype Philippines: Camiguin, W Mambajao, Katibawasan spring area, 350-400 m., 13- 15.iii. (H. Zettel & C.V. Pangantihon); paratypes: 10 workers with same data, 4 workers Camiguin, Mambajao, Sudlon, slopes Mt Hibok-Hibok, 400 m., 14.ii. (H. Zettel), 4 workers Camiguin, Catarman, Mainit, Tuasan Flls, 23.x. (C.V. Pangantihon), 1 worker as last but 200-50 m., 11.ii. (H. Zettel), 1 worker as last but 28.ii. (A. Luckeneder). Type-depositories: NMPM (holotype); CASC, HSZC, NHMW, TNHM (paratypes). Distribution: Philippines (Camiguin). rothneyi. Diacamma rugosum var. rothneyi Bingham, 1903: 81 (in text). Type-material: syntype workers (number not stated). Type-locality: India: Cochin (Rothney). Type-depository: MHNG. [First available use of Diacamma rugosum r. rugosum var. rothneyi Forel, 1900d: 318 (diagnosis in key) (w.) INDIA (Kerala); unavailable (infrasubspecific name).] Subspecies of rugosum: Emery, 1911d: 66; Teranishi, 1940: 65; Chapman & Capco, 1951: 56; Bolton, 1995b: 170; Bharti, Guénard, et al. 2016: 50. Distribution: India. rugivertex. Diacamma rugivertex Emery, 1902c: 28 (w.) INDONESIA (Timor I.). Type-material: holotype worker. Type-locality: Indonesia: Tomor I. (no further data). Type-depository: MSNG. Status as species: Emery, 1911d: 66; Chapman & Capco, 1951: 55; Bolton, 1995b: 170. Distribution: Indonesia (Timor). rugosum. Ponera rugosa Le Guillou, 1842: 318 (w.) BORNEO (no state data). Type-material: holotype worker. Type-locality: Borneo (no further data). Type-depository: MNHN. Ogata, 1987: 111 (m.). Combination in Diacamma: Mayr, 1862: 718. Status as species: Smith, F. 1859a: 142; Roger, 1860: 305; Roger, 1861b: 167; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Mayr, 1865: 65; Mayr, 1867a: 86 (redescription); Smith, F. 1871a: 321, 323; Mayr, 1872: 149; Dalla Torre, 1893: 29; Emery, 1897b: 153; Forel, 1900d: 319; Emery, 1901f: 113; Forel, 1909d: 221; Wheeler, W.M. 1909d: 338; Emery, 1911d: 66; Wheeler, W.M. 1919e: 53; Wheeler, W.M. 1923b: 1; Wheeler, W.M. 1924b: 242; Stitz, 1925: 111; Wheeler, W.M. & Chapman, 1925: 62; Karavaiev, 1925b: 116; Karavaiev, 1926d: 417; Wheeler, W.M. 1927b: 42; Wheeler, W.M. 1927h: 84; Karavaiev, 1935a: 60; Teranishi, 1940: 65; Chapman & Capco, 1951: 55; Wilson, 1958d: 368; Baltazar, 1966: 242; Wu, J. & Wang, 1992: 1301; Radchenko, 1993a: 80; Bolton, 1995b: 170; Wu, J. & Wang, 1995: 43; Tang, J., Li, et al. 1995: 34; Tiwari, 1999: 25; Mathew & Tiwari, 2000: 274; Zhou, 2001b: 45; Lin & Wu, 2003: 67; Ghosh, et al. 2005: 7; Jaitrong & Nabhitabhata, 2005: 19; Mohanraj, et al. 2010: 6; Pfeiffer, et al. 2011: 55; Guénard & Dunn, 2012: 58; Bharti, Guénard, et al. 2016: 50; Jaitrong, Guénard, et al. 2016: 42. Senior synonym of divergens: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of frontalis: Wilson, 1958d: 368; Bolton, 1995b: 170. Senior synonym of geminatum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of japensis: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 46. Senior synonym of macreta: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of papuanum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of saussurei: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of sculpturata: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of smithi: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 46. Senior synonym of striata: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Senior synonym of versicolor: Roger, 1861b: 167; Mayr, 1863: 407; Roger, 1863b: 16; Mayr, 1865: 65; Mayr, 1872: 149; Emery, 1897b: 153; Forel, 1900d: 319; Forel, 1900d: 319; Emery, 1911d: 66; Wheeler, W.M. 1919e: 53; Wheeler, W.M. & Chapman, 1925: 62; Donisthorpe, 1932c: 447; Chapman & Capco, 1951: 60; Bolton, 1995b: 170; Zhou, 2001b: 45. Material of the unavailable name pulchellum referred here by Wilson, 1958d: 368. Distribution: China, India (+ Andaman Is), Indonesia (Aru Is, Bacan, Buru, Halmahera, Irian Jaya, Java, Kalimantan, Seram, Sumatra), Laos, Malaysia (Sarawak), Papua New Guinea, Philippines (Luzon), Sri Lanka, Thailand, Vietnam. Current subspecies: nominal plus anceps, andamane, arcuatum, balinense, birmanum, brevispinum, celebense, doveri, gibbosum, hortense, javanum, jerdoni, latispinum, lombokense, longiceps, nitidiventre, oblicum, ovale, pubescens, rothneyi, sculptum. saussurei. Diacamma bispinosum var. saussurei Forel, 1922: 89 (w.) INDONESIA (Halmahera I.). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Halmahera (H. de Saussure). Type-depository: MHNG. Subspecies of bispinosum: Chapman & Capco, 1951: 53. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. scalpratum. Ponera scalprata Smith, F. 1858b: 84, pl. 6, figs. 21, 22 (w.) INDIA (no state data, “Northern India”). Type-material: holotype worker. Type-locality: India: “N. Ind. 54/16” (Hooker). Type-depository: BMNH. Emery, 1889b: 496 (m.); Wheeler, G.C. & Wheeler, J. 1952c: 610 (l.). Combination in Diacamma: Mayr, 1862: 718. Status as species: Roger, 1860: 305; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Smith, F. 1871a: 320; Mayr, 1886c: 358; Emery, 1887b: 435 (in key, footnote); Emery, 1889b: 496; Dalla Torre, 1893: 29; Emery, 1895k: 458; Emery, 1897b: 162 (in key); Forel, 1900d: 319; Bingham, 1903: 77; Forel, 1906b: 91; Emery, 1911d: 65; Wheeler, W.M. 1927b: 42; Mukerjee, 1930: 150; Menozzi, 1939a: 328; Chapman & Capco, 1951: 60; Collingwood, 1962: 224; Bolton, 1995b: 170; Mathew & Tiwari, 2000: 275; Laciny, Pal & Zettel, 2015: 90 (redescription); Bharti, Guénard, et al. 2016: 50; Rasheed, et al. 2019: 434. Senior synonym of compressum: Dalla Torre, 1893: 29; Emery, 1911d: 65; Bolton, 1995b: 170; Laciny, Pal & Zettel, 2015: 90. Distribution: India, Myanmar, Pakistan. schoedli. Diacamma schoedli Shattuck & Barnett, 2006: 17, figs. 11-13 (w.) AUSTRALIA (Queensland, and Horn I., Torres Strait). Type-material: holotype worker, 3 paratype workers. Type-locality: holotype Australia: Queensland, 14 km. W by N Hope Vale Mission, 15°16’S, 144°59’E, 7- 10.v.1981 (J.E. Feehan); paratypes with same data. Type-depository: ANIC. Distribution: Australia. sculptum. Ponera sculpta Jerdon, 1851: 117 (w.) INDIA (Karnataka/Kerala, “Malabar”; Tamil Nadu). Type-material: syntype workers (number not stated). Type-locality: India: Malabar, sea level to top of the Neilgherries (T.C. Jerdon). Type-depository: no type-material is known to exist. [Duplicated in Jerdon, 1854b: 101.] Combination in Diacamma: Mayr, 1863: 407; Roger, 1863b: 16; Emery, in Dalla Torre, 1893: 29. Status as species: Smith, F. 1858b: 84; Smith, F. 1871a: 320; Dalla Torre, 1893: 29; Rothney, 1903: 96; Bingham, 1903: 80; Donisthorpe, 1929a: 444. Junior synonym of rugosum: Roger, 1863b: 16; Mayr, 1863: 407; Mayr, 1865: 65; Mayr, 1872: 149. Junior synonym of vagans: Emery, 1887b: 440. Subspecies of rugosum: Emery, 1897b: 156; Forel, 1900d: 320; Emery, 1900d: 666; Emery, 1901f: 113; Wheeler, W.M. 1909d: 334, 338; Yano, 1910: 419; Wheeler, W.M. & Chapman, 1925: 63; Menozzi, 1939a: 328; Teranishi, 1940: 66; Chapman & Capco, 1951: 56; Baltazar, 1966: 243; Bolton, 1995b: 170; Tiwari, 1999: 26; Bharti, Guénard, et al. 2016: 50. Synonym of jerdoni: Emery, 1911d: 66. [Note: Emery gives jerdoni as senior synonym, but if the synonymy is correct then sculptum has priority.] Distribution: India, Myanmar, Philippines (Luzon), Sri Lanka, Taiwan, Thailand. sculpturata. Ponera sculpturata Smith, F. 1859a: 142 (w.) INDONESIA (Aru Is). Type-material: holotype worker. Type-locality: Indonesia: Aroo (= Aru Is) (A.R. Wallace). Type-depository: OXUM. Karavaiev, 1926d: 417 (m.). Combination in Diacamma: Mayr, 1862: 718. Status as species: Roger, 1860: 305; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Smith, F. 1871a: 322; Mayr, 1886c: 361; Jaitrong & Nabhitabhata, 2005: 20 (error); Pfeiffer, et al. 2011: 55 (error). Subspecies of geometricum: Dalla Torre, 1893: 28. Subspecies of rugosum: Emery, 1897b: 159; Emery, 1897d: 550; Forel, 1901f: 338; Emery, 1911b: 531; Emery, 1911d: 66; Viehmeyer, 1912: 6; Crawley, 1924: 384; Karavaiev, 1925b: 120; Karavaiev, 1926d: 417; Santschi, 1928h: 124; Donisthorpe, 1932c: 454; Santschi, 1932b: 12; Karavaiev, 1935a: 65 (redescription). Donisthorpe, 1949g: 405; Donisthorpe, 1950a: 338; Chapman & Capco, 1951: 58. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. sericeiventre. Diacamma sericeiventre Stitz, 1925: 111, fig. 1 (w.) PHILIPPINES (Palawan I.). Type-material: 3 syntype workers. Type-locality: Philippines: Palawan, Binaluan (Boettcher). Type-depository: MNHU. Status as species: Chapman & Capco, 1951: 60; Baltazar, 1966: 243; Bolton, 1995b: 170. Junior synonym of concentricum: Laciny, Pal & Zettel, 2015: 99. sikkimense. Diacamma rugosum var. sikkimensis Forel, 1903d: 401 (in text) (w.) INDIA (Sikkim). Type-material: lectotype worker (by designation of Zettel, et al. 2016: 144), 4 paralectotype workers. Type-locality: lectotype India: Sikkim, 4000 ft, x.1902 (C.T. Bingham); paralectotypes with same data. Type-depository: MHNG. Subspecies of rugosum: Emery, 1911d: 66; Santschi, 1920g: 179; Santschi, 1920h: 158; Santschi, 1924c: 96; Menozzi, 1939a: 328; Chapman & Capco, 1951: 56; Bolton, 1995b: 170. Status as species: Zettel, et al. 2016: 144 (redescription). Distribution: India, Vietnam. smithi. Diacamma rugosum subsp. smithi Donisthorpe, 1943e: 115. Replacement name for Ponera tortuolosa Smith, F. 1863: 18. [Junior primary homonym of Ponera tortuolosa Smith, F. 1858b: 99.] Subspecies of rugosum: Chapman & Capco, 1951: 59. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 46. striata. Ponera striata Smith, F. 1860b: 104 (w.) INDONESIA (Bacan I.). Type-material: holotype worker. Type-locality: Indonesia: Bachian (= Bacan) “Bac” (A.R. Wallace). Type-depository: presumed lost (not in BMNH, OXUM). [Note: a single worker in OXUM fits the description reasonably well and is labelled “striata,” but its locality of “M” (Misool I.) is incorrect, not the stated type-locality of “Bac”.] Combination in Diacamma: Mayr, 1886c: 361. Status as species: Mayr, 1863: 450; Smith, F. 1865: 70; Smith, F. 1871a: 323; Mayr, 1886c: 361. Subspecies of versicolor: Emery, 1887b: 436 (footnote); Dalla Torre, 1893: 30. Subspecies of rugosum: Emery, 1897b: 161; Emery, 1911d: 67; Donisthorpe, 1932c: 461; Chapman & Capco, 1951: 59. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. subsulcata. Diacamma bispinosum var. subsulcata Emery, 1897b: 152 (w.) INDONESIA (Halmahera I.). Type-material: 2 syntype workers. Type-locality: Indonesia: Halmahera (Doherty). Type-depository: MSNG. [Misspelled as subculcatum by Chapman & Capco, 1951: 52.] Subspecies of bispinosum: Emery, 1911d: 65; Chapman & Capco, 1951: 53. Junior synonym of bispinosum: Wilson, 1958d: 367; Bolton, 1995b: 170. symposium. Diacamma symposium Zettel, et al. 2016: 152, figs. 30-33 (w.) PHILIPPINES (Mindanao I.). Type-material: holotype worker, 9 paratype workers. Type-locality: holotype Philippines: Mindanao, Bukidnon, Malaybalay, Kaamulan, 650 m., 15-20.iii. (H. Zettel); paratypes: 8 workers with same data, 1 worker with same data but 15-16.iii. Type-depositories: NMPM (holotype); HSZC, NHMW (paratypes). Distribution: Philippines (Mindanao). timorense. Diacamma javanum r. timorense Emery, 1887b: 439 (footnote) (w.) INDONESIA (Timor I.). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Timor (Vollenhoven). Type-depositories: MSNG, NHMW. Subspecies of javanum: Dalla Torre, 1893: 29. Subspecies of rugosum: Emery, 1897b: 158; Emery, 1911d: 67; Chapman & Capco, 1951: 59; Bolton, 1995b: 170. Status as species: Zettel, et al. 2016: 147 (redescription). Distribution: Indonesia (Timor). tonkina. Diacamma rugosum st. vagans var. tonkina Santschi, 1920g: 179 (w.) VIETNAM; unavailable (infrasubspecific) name. [Note: material included: Vietnam: Tonkin, Hoobint (Vitalis de Salvaza); worker(s) (no indication of number), deposited in NHMB (no type status).] As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 60; Santschi, 1920h: 158; Santschi, 1924c: 96. Declared as unavailable (infrasubspecific) name: Bolton, 1995b: 170. tortuolosa. Ponera tortuolosa Smith, F. 1863: 18 (w.q.) INDONESIA (Buru I., Seram I.). Type-material: 2 syntype workers. Type-localities: Indonesia: Seram, “Cer” (A.R. Wallace), and Buru, “B” (A.R. Wallace). Type-depository: OXUM. [Note: the Buru specimen does not match the original description, the Seram specimen does. It is assumed that the original Buru worker was lost, and later replaced with a substitute which is not conspecific.] [Junior primary homonym of Ponera tortuolosa Smith, F. 1858b: 99.] Combination in Diacamma: Mayr, 1886c: 363. Status as species: Smith, F. 1871a: 323; Forel, 1901h: 46; Dalla Torre, 1893: 29; Donisthorpe, 1932c: 471. Subspecies of geometricum: Emery, 1887b: 436 (in key, footnote), 438; Emery, in Dalla Torre, 1893: 28 (footnote). Subspecies of rugosum: Mayr, 1867a: 86 (in text); Emery, 1897b: 160; Emery, 1897d: 550; Emery, 1911d: 67; Wheeler, W.M. 1919e: 53; Santschi, 1932b: 12. Replacement name: Diacamma rugosum subsp. smithi Donisthorpe, 1943e: 115. tritschleri. Diacamma tritschleri Forel, in Emery, 1897b: 151 (footnote) (w.) INDONESIA (Sumatra). Type-material: syntype workers (number not stated). Type-locality: Indonesia: Sumatra (Tritschler), and Sumatra (Moesch). Type-depositories: BMNH, MHNG, MSNG, NHMW. Status as species: Forel, 1901b: 7; Forel, 1901f: 338; Forel, 1913k: 7; Viehmeyer, 1916a: 113; Santschi, 1928h: 123; Chapman & Capco, 1951: 60; Bolton, 1995b: 170. Junior synonym of geometricum: Laciny, Pal & Zettel, 2015: 102. vagans. Ponera vagans Smith, F. 1860b: 103 (w.) INDONESIA (Bacan I.). Type-material: holotype worker. Type-locality: Indonesia: Bachian (= Bacan) “Bac” (A.R. Wallace). Type-depository: OXUM. [Note: a supposed syntype in NHMW does not appear to be from the same series as the holotype.] [Misspelled as vayans by Forel, 1912n: 52; misspelled as vargens by Jaitrong & Nabhitabhata, 2005: 20.] Bingham, 1903: 81 (m.); Wheeler, G.C. & Wheeler, J. 1952c: 611 (l.); Imai, et al. 1984: 5 (k.). Combination in Diacamma: Mayr, 1862: 718. As unavailable (infrasubspecific) name: Emery, 1897b: 158; Wheeler, W.M. 1909d: 339; Wheeler, W.M. & Chapman, 1925: 63. Subspecies of rugosum: Forel, 1908a: 1; Forel, 1911a: 23; Forel, 1911i: 216; Emery, 1911d: 67; Forel, 1912a: 49; Forel, 1912d: 100; Forel, 1912n: 52; Forel, 1913k: 7; Viehmeyer, 1916a: 113; Wheeler, W.M. 1919e: 54; Santschi, 1920g: 179; Santschi, 1920h: 158; Forel, 1922: 90; Santschi, 1924c: 96; Wheeler, W.M. 1927b: 42; Menozzi, 1930d: 328; Donisthorpe, 1932c: 461; Donisthorpe, 1948b: 299; Chapman & Capco, 1951: 59; Baltazar, 1966: 243. Junior synonym of rugosum: Wilson, 1958d: 368; Bolton, 1995b: 170; Zhou, 2001b: 45. Status as species: Roger, 1860: 304; Mayr, 1862: 718; Mayr, 1863: 407; Roger, 1863b: 16; Mayr, 1867a: 87 (redescription); Smith, F. 1871a: 323; Smith, F. 1873: viii; Forel, 1885b: 177; Mayr, 1886c: 361; Emery, 1887b: 436 (in key, footnote); Emery, 1889b: 497; Rothney, 1889: 361; Emery, 1893e: 190; Emery, 1893g: 259; Dalla Torre, 1893: 29; Emery, 1895k: 458; Rothney, 1903: 96; Bingham, 1903: 81; Mukerjee & Ribeiro, 1925: 205; Mukerjee, 1930: 150; Mukerjee, 1934: 3; Tiwari, 1999: 25; Jaitrong & Nabhitabhata, 2005: 20; Laciny, Pal & Zettel, 2015: 106 (redescription). Distribution: India, Indonesia (Bacan), Myanmar, Philippines (Luzon, Negros), Singapore, Sri Lanka, Taiwan, Thailand, Vietnam. [Note: Laciny, Pal & Zettel, 2015: 109, cast doubt on the accuracy of this distribution.] versicolor. Ponera versicolor Smith, F. 1857a: 65 (w.) PHILIPPINES (no island nominated), BORNEO (East Malaysia: Sarawak). Type-material: 2 syntype workers. Type-localities: Philippines: “Phil. Isla. 42/22” (no collector’s name), and Malaysia (“Borneo”): Sarawak “SAR” (A.R. Wallace). [Note: the two Philippines syntypes are in BMNH. A supposed syntype specimen from “SAR” is in OXUM, but does not match the original description and is probably not versicolor.] Type-depository: BMNH. Combination in Diacamma: Roger, 1863b: 16. Status as species: Smith, F. 1858b: 87; Roger, 1860: 300; Smith, F. 1871a: 321; Dalla Torre, 1893: 30. Subspecies of geometricum: Emery, 1887b: 436 (in key, footnote), 438; Emery, in Dalla Torre, 1893: 28 (footnote); Emery, 1893e: 189; Emery, 1893f: 242. Junior synonym of rugosum: Roger, 1861b: 167; Mayr, 1863: 407; Roger, 1863b: 16; Mayr, 1865: 65; Mayr, 1872: 149; Emery, 1897b: 153; Forel, 1900d: 319; Forel, 1900d: 319; Emery, 1911d: 66; Wheeler, W.M. 1919e: 53; Wheeler, W.M. & Chapman, 1925: 62; Donisthorpe, 1932c: 447; Chapman & Capco, 1951: 60; Bolton, 1995b: 170; Zhou, 2001b: 45. violaceum. Diacamma scalpratum var. violaceum Forel, 1900d: 317 (diagnosis in key) (w.) MYANMAR. Type-material: syntype workers (number not stated). Type-localities: Myanmar (“Birmanie”): Moulmein (Hodgson), and Myanmar: Thaungyin Valley (Bingham). Type-depository: MHNG. Subspecies of scalpratum: Emery, 1911d: 65; Chapman & Capco, 1951: 60; Bolton, 1995b: 171. Status as species: Laciny, Pal & Zettel, 2015: 93 (redescription); Zettel, et al. 2016: 167. Distribution: Myanmar, Thailand. viridipurpureum. Diacamma geometricum var. viridipurpureum Emery, 1893g: 261 (w.) PHILIPPINES (Luzon I.). Type-material: lectotype worker (by designation of Laciny, Pal & Zettel, 2015: 118). Type-locality: Philippines: Luzon, Antipolo, iii.-iv.1890 (E. Simon). Type-depository: MSNG. Wheeler, G.C. & Wheeler, J. 1952c: 610 (l.). As unavailable (infrasubspecific) name: Emery, 1897b: 155; Forel, 1909d: 221; Wheeler, W.M. 1909d: 338; Forel, 1910d: 122; Emery, 1911d: 66; Wheeler, W.M. & Chapman, 1925: 64; Wheeler, W.M. 1927d: 1; Wheeler, W.M. 1930h: 59; Chapman & Capco, 1951: 58 Baltazar, 1966: 241. Subspecies of geometricum: Santschi, 1933d: 1. Subspecies of rugosum: Bolton, 1995b: 171; Zhou & Ran, 2010: 104; Guénard & Dunn, 2012: 58. Status as species: Laciny, Pal & Zettel, 2015: 118; Bharti, Guénard, et al. 2016: 50. Distribution: India, Indonesia (Sulawesi), Philippines (Bantayan, Cebu, Luzon, Marinduque & Santa Cruz, Masbate, Mindoro, Negros, Panay, Romblon, Samar). Current subspecies: quezonicum. waigeuensis. Diacamma rugosum subsp. waigeuensis Donisthorpe, 1942e: 703 (w.) NEW GUINEA (Indonesia: Waigeu I.). Type-material: 2 syntype workers. Type-locality: Indonesia: Irian Jaya (“N. Dutch New Guinea”): Waigeu I., Camp Nok, 2500 ft, v.1938 (L.E. Cheesman). Type-depository: BMNH. Subspecies of rugosum: Donisthorpe, 1943d: 438; Chapman & Capco, 1951: 60. Junior synonym of purpureum: Bolton, 1995b: 171.