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A New Species of Columbarium (Gastropoda: Muricacea) from Off Eastern Australia

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A New Species of Columbarium (Gastropoda: Muricacea) from Off Eastern Australia A NEW SPECIES OF COLUMBARIUM (GASTROPODA: MURICACEA)MURICACEA) FROM OFF EASTERN AUSTRALIA M. G. Harasewych College of Marine Studies University of Delaware Newark, Delaware 19711 The genus Columbarium Martens 1881 (as(as ovate, with terminal nucleus; periostracum andand defined by Darragh, 1969) is representedrepresented inin thethe soft parts unknown. Recent fauna by eight species, which inhabitinhabit Type material - Holotype - United States Na-Na­ outer continental shelf and upper continental tional Museum, Washington, D.C. (USNM(USNM slope communities off South Africa, easterneastern 806997) length 106 mm; Paratype 1 - (USNM(USNM Australia, New Zealand and Japan. Trawling 806998) length 70 mm; ParatypeParatype 2 - Delaware operations off the southern coast of Queensland Museum of Natural History, Greenville, Dela-Dela­ have brought to light a remarkable new speciesspecies ware (DMNH 153524) length 99 mm; Paratype 3 of Columbarium, which more closely resemblesresembles - The Australian Museum, Sydney, AustraliaAustralia several Tertiary species thanthan any inin thethe RecentRecent length 73 mm. fauna.fauna~ This new species is described herein.herein. Type locality - East of Lady MusgraveMusgrave Island, Queensland, Australia, inin 140 fathomsfathoms Columbarium harrisae new species (256 meters). , Figs. 1-4 Range - Specimens have also been takentaken offoff Description - Shell large (to 110 mm), heavy,heavy, Lady Elliot Island, Queensland, Australia, inin fusiform; spire angle 37°-42°; protoconch ofof 22 150 fathoms (275 meters) (Trevor, 1982). whorls, large, bulbous, glassy, with deviated,deviated, Remarks - The large size, high spire, heavyheavy disjunct apex; transition toto teleoconchteleoconch indis-indis­ shell, stout siphonal canal and characteristic tinct, marked by gradual acquisition of aa peri-peri­ protoconch readily distinguish thisthis new speciesspecies pheral keel and axial growth lines;lines; teleoconchteleoconch from all other Recent members of thethe genus.genus. with up to 8 sharply shouldered whorls; suturesuture Columbarium harrisae most closely resemblesresembles adpressed; siphonal canal long, straight, heavy;heavy; C. vulneratum (Finlay and Marwick, 1937) fromfrom spiral scupture of 2-4 cords on body whorlwhorl andand the Paleocene of New Zealand, fromfrom whichwhich itit 20-26 finer threads on siphonal canal; axialaxial differs by its larger size and more elongate shell.shell. growth lines produce 21-2421 ~24 short, open spinesspines Young specimens of C. harrisae lacklack thethe raisedraised per whorl along the shoulder, with correspond-correspond­ peristomalperistomal plate, and bear a strong resemblanceresemblance ing scales on each of thethe spiral cordscords andand to C. rugatum (Aldrich, 1886), fromfrom thethe Lower threads; shell color white with brown spotsspots be-be­ Eocene of Alabama, but may easily be discerned tween spines and scales; aperture ovate; outerouter on the basis of protoconchs. lip smooth; columella smooth, with a raisedraised peri-peri­ The high spire, thick shell and prominent stomal plate that forms a notch belowbelow thethe spiral sculpture are primitive characters withinwithin suture, and extends along the innerinner edge ofof thethe the Columbariinae, being shared by such groups siphonal canal; operculum corneous, sharplysharply as Histricosceptrum, Peristarium and Coluzea. Vol. 97(1) January 31, 1983 THE NAUTILUS 29 FIGS. 1-4. Columbarium harrisae new species.species. 1,1, Holotype,Holo type, USNMUSNM 806997,806997, dredgeddredged easteast ofof LadyLady MusgraveMusgrave Island,Island, Queensland,QweeWaW, AwsZraZm,Australia, min 256 mefersmeters (J.AX).(1.0X). 2,2, Para^/peParatype J,1, U&ZVM#0####,USNM 806998, samesame ZocaZi^/locality aWand (ZepZ&depth (7.0%).(1. OX). 3,3, fro^oco^c^Protoconch of paratype 1 (10.OX).(1 o. OX). 4, Operculum of paratypeparatype 11 (5.0X).(5.0.LY.j. Such features of the Pacific species ofof Colum-Colum­ vided the type material. Thanks are due RichardRichard barium as enlarged protoconch, reducedreduced spiralspiral M. Kurz, Wauwatosa, Wisconsin, who providedprovided ornament, lower spire, and long,long, thinthin siphonalsiphonal additional material and information. canal are modifications thatthat have arisen sincesince .the closing of the Tethys Sea. ColumbariumColumbarium/ harrisae appears to be an offshoot fromfrom aa LITERATURE CITED primitive stock and not closely relatedrelated toto thethe Darragh, T. A. 1969. A Revision ofof thethe FamilyFamily Columbari-Columbari­ idae (Mollusca: Gastropoda). Proc. Roy.Roy. Soc.Soc. Viet. 83(1):83(1): other Australian membersme'mbers of thethe genus. 63-119. This new taxon honors Valerie Harris ofof Trevor, N. 1982. New Species of Columbariidae? KeppelKeppel Caloundra, Queensland, who generously pro-pro- Bm/Bay Tidingsr&dm#s 21(1):1.21(1):1. .
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