Wednesday 2 September, 2.30Pm BST. Over Zoom (Details Sent to Registered Attendees) Getting Inside the Heads of Early Vertebrates

Total Page:16

File Type:pdf, Size:1020Kb

Wednesday 2 September, 2.30Pm BST. Over Zoom (Details Sent to Registered Attendees) Getting Inside the Heads of Early Vertebrates Wednesday 2 September, 2.30pm BST. Over Zoom (Details sent to registered attendees) Getting inside the heads of early vertebrates Animals with backbones (vertebrates) have an evolutionary history of nearly half a billion years, with fossils instrumental in understanding how the group became so hugely successful. Jawed bony fishes account for 99% of living vertebrate species, and over half of these are ray-finned fishes: staples of the aquarium and fishmonger, encompassing everything from goldfish to seahorses to cod. However, the double barriers of geological time and fossil preservation has led to a poor understanding of the early history and evolution of ray-fins. Many of the major innovations that drove ray-fins to be so diverse are tied up in the braincase, a bony box that sits within the head and houses the brain and sensory organs. Traditionally, these internal structures would be accessed by gradually grinding the fossil away into dust, recording the morphology through a series of drawings. By using x-ray tomography (CT scanning), it is possible to ‘virtually’ cut through the specimens without damaging the fossil. CT scanning works in exactly the same way as getting an x-ray or CAT scan at a hospital: different materials in the fossil absorb differing amounts of x-rays. This can be used to build up a picture of the fossil's internal anatomy, opening a window into the evolution of the skull and brain. Comparing these structures between key living and extinct ray-fins allows for major events to be put into context, shedding new light on innovations and evolutionary relationships. These findings also provide insight into the explosive diversification of ray-finned fishes some 350 million years ago. Sam Giles - University of Birmingham ‘The GeolSoc public lecture series represent a way to communicate my research to a broader audience. An online presentation will hopefully make it even broader still!’ Sam is a Royal Society research fellow at the University of Birmingham, prior to this she completed her MSci in Geology at the University of Bristol in 2011 and a D.Phil in Palaeobiology at the University of Oxford in 2015. She then spent two years as a Junior Research Fellow at Christ Church, Oxford. Sam's main research interests are in the relationships and morphology of bony vertebrates. She uses non-destructive x-ray imaging (CT scanning) to unlock the external and internal anatomy of living and fossil vertebrates. She focuses on the early history of bony fishes, which account for 99% of all living vertebrates. Her work provides insight into the origins and evolutionary success of different vertebrate groups. Reading list: - Friedman, M., Pierce, S. E., Coates, M. I., Giles S. 2019. Feeding structures in the ray-finned fish Eurynotus crenatus (Actinopterygii: Eurynotiformes): implications for trophic diversification among Carboniferous actinopterygians. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, https://doi.org/10.1017/S1755691018000816 - Giles, S., Xu, G.-H., Near, T. J., Friedman, M. 2017. Early members of ‘living fossil’ lineage imply later origin of modern ray-finned fishes. Nature, https://doi.org/10.1038/nature23654 - Lu, J., Giles, S., Friedman, M., Den Blaauwen, J. L., Zhu, M. 2016. The oldest actinopterygian highlights the cryptic early history of the hyperdiverse ray-finned fishes. Current Biology, http://dx.doi.org/10.1016/j.cub.2016.04.045. - Giles, S., Darras, L., Clément, G., Blieck, A., Friedman, M. 2015. An exceptionally preserved Late Devonian actinopterygian provides a new model for primitive cranial anatomy in ray-finned fishes. Proceedings of the Royal Society B, https://doi.org/10.1098/rspb.2015.1485 Popular papers from the GSL Lyell Collection: -Re-evaluation of the Permian macrofossils from the Parnaíba Basin: biostratigraphic, palaeoenvironmental and palaeogeographical implications https://sp.lyellcollection.org/content/472/1/223 -A lower Carboniferous (Visean) tetrapod trackway represents the earliest record of an edopoid amphibian from the UK https://jgs.lyellcollection.org/content/177/2/276 Finally thank you for joining in with this Public Lecture. We have more upcoming virtual events that we think maybe of interest to you: -Geo Poetry – 1 October. Register here -GSL Public Lecture - Lost in Translation: Why Talking About Geoscience is so difficult with Hazel Gibson – 12 October. Register here -Mineral Resource Estimations: Recent Advances and Current Best Practice – 19 October. Register here -GSL Public Lecture: Engaging geoscience for diplomacy – understanding Paektu volcano in North Korea with James Hammond – 9 November. Register here -GSL Public Lecture: ‘Reading the ground’ to reduce hazards and risks in engineering projects with Andrew Hart – 3 December. Register here .
Recommended publications
  • PROGRAM the 11Th International Congress of Vertebrate Morphology
    PROGRAM The 11th International Congress of Vertebrate Morphology 29 June – 3 July 2016 Bethesda North Marriott Hotel & Conference Center Washington, DC CONTENTS Welcome to ICVM 11 ........................ 5 Note from The Anatomical Record........... 7 Administration ............................. 9 Previous Locations of ICVM ................. 10 General Information ........................ .11 Sponsors .................................. 14 Program at-a-Glance ....................... 16 Exhibitor Listing............................ 18 Program ................................... 19 Wednesday 29th June, 2016 ................... .19 Thursday 30th June, 2016 ..................... 22 Friday 1st July, 2016 ........................... 34 Saturday 2nd July, 2016 ....................... 44 Sunday 3rd July, 2016 ......................... 52 Hotel Floor Plan ................... Back Cover Program 3 Journal of Experimental Biology (JEB)(JEB) isis atat thethe forefrontforefront ofof comparaticomparativeve physiolophysiologygy and integrative biolobiology.gy. We publish papers on the form and function of living ororganismsganisms at all levels of biological organisation and cover a didiverseverse array of elds,fields, including: • Biochemical physiology •I• Invertebratenvertebrate and vertebrate physiology • Biomechanics • Neurobiology and neuroethology • Cardiovascular physiology • Respiratory physiology • Ecological and evolutionary physiology • Sensory physiology Article types include ReseaResearchrch Articles, Methods & TeTechniques,chniques, ShoShortrt
    [Show full text]
  • The Palaeontology Newsletter
    The Palaeontology Newsletter Contents100 Editorial 2 Association Business 3 Annual Meeting 2019 3 Awards and Prizes AGM 2018 12 PalAss YouTube Ambassador sought 24 Association Meetings 25 News 30 From our correspondents A Palaeontologist Abroad 40 Behind the Scenes: Yorkshire Museum 44 She married a dinosaur 47 Spotlight on Diversity 52 Future meetings of other bodies 55 Meeting Reports 62 Obituary: Ralph E. Chapman 67 Grant Reports 72 Book Reviews 104 Palaeontology vol. 62 parts 1 & 2 108–109 Papers in Palaeontology vol. 5 part 1 110 Reminder: The deadline for copy for Issue no. 101 is 3rd June 2019. On the Web: <http://www.palass.org/> ISSN: 0954-9900 Newsletter 100 2 Editorial This 100th issue continues to put the “new” in Newsletter. Jo Hellawell writes about our new President Charles Wellman, and new Publicity Officer Susannah Lydon gives us her first news column. New award winners are announced, including the first ever PalAss Exceptional Lecturer (Stephan Lautenschlager). (Get your bids for Stephan’s services in now; check out pages 34 and 107.) There are also adverts – courtesy of Lucy McCobb – looking for the face of the Association’s new YouTube channel as well as a call for postgraduate volunteers to join the Association’s outreach efforts. But of course palaeontology would not be the same without the old. Behind the Scenes at the Museum returns with Sarah King’s piece on The Yorkshire Museum (York, UK). Norman MacLeod provides a comprehensive obituary of Ralph Chapman, and this issue’s palaeontologists abroad (Rebecca Bennion, Nicolás Campione and Paige dePolo) give their accounts of life in Belgium, Australia and the UK, respectively.
    [Show full text]
  • University of Birmingham a Giant Dapediid from the Late Triassic Of
    University of Birmingham A giant dapediid from the Late Triassic of Switzerland and insights into neopterygian phylogeny Giles, Sam; Latimer, Ashley DOI: 10.1098/rsos.180497 License: Creative Commons: Attribution (CC BY) Document Version Publisher's PDF, also known as Version of record Citation for published version (Harvard): Giles, S & Latimer, A 2018, 'A giant dapediid from the Late Triassic of Switzerland and insights into neopterygian phylogeny', Royal Society Open Science, vol. 5, 180497. https://doi.org/10.1098/rsos.180497 Link to publication on Research at Birmingham portal General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document. When citing, please reference the published version. Take down policy While the University of Birmingham exercises care and attention in making items available there are rare occasions when an item has been uploaded in error or has been deemed to be commercially or otherwise sensitive.
    [Show full text]
  • Book of Abstracts
    THE 10th INDO-PACIFIC FISH CONFERENCE Book of Abstracts TAHITI - 2-6 October 2017 1 Table of contents A1/ Evolution and biology of `primitive' and fossil fishes 33 Adaptive radiation of Pelagia (Teleostei: Acanthomorpha) indicated by 3D mor- phometry, Hermione Beckett [et al.]......................... 34 Awesome variation in genomic organization in polyploid Acipenseriformes, Anna Barmintseva [et al.]................................... 35 Dead fish CSI: Reconstructing the enigmatic Late Cretaceous billfish analogue Protosphyraena (Teleosteomorpha: Pachycormidae), Anthony Maltese...... 36 Early members of a `living fossil' lineage and a later origin for modern ray-finned fishes, Sam Giles [et al.]................................ 37 Insight on sturgeon phylogeny and biogeography from mitogenomes and NGS- based nuclear loci sequencing., Nikolai Mugue [et al.]................ 38 Megaplanktivory, Past and Present: A comparison of Jurassic large suspension- feeders of the IndoPacific with contemporary analogues., Jeff Liston....... 39 New Data on the Endoskeletal Morphology and Evolution of Early Jawed Fishes, Mar- tin Brazeau [et al.]................................... 40 New marine fish faunas from the middle Eocene (Lutetian) of Pakistan: implica- tions for the origin of the Indo-Pacific fauna, Matt Friedman [et al.]....... 41 Polyploid evolution and functional genome diploidization in sturgeons, Victor Vasil'ev [et al.]..................................... 42 Saber-toothed fossil anchovies (Teleostei: Engrauloidea) from the early-middle Eocene of
    [Show full text]
  • Morphology and Evolutionary Significance of Phosphatic Otoliths
    Schnetz et al. BMC Evolutionary Biology (2019) 19:238 https://doi.org/10.1186/s12862-019-1568-z RESEARCH ARTICLE Open Access Morphology and evolutionary significance of phosphatic otoliths within the inner ears of cartilaginous fishes (Chondrichthyes) Lisa Schnetz1*, Cathrin Pfaff2, Eugen Libowitzky3, Zerina Johanson4, Rica Stepanek2 and Jürgen Kriwet2* Abstract Background: Chondrichthyans represent a monophyletic group of crown group gnathostomes and are central to our understanding of vertebrate evolution. Like all vertebrates, cartilaginous fishes evolved concretions of material within their inner ears to aid with equilibrium and balance detection. Up to now, these materials have been identified as calcium carbonate-bearing otoconia, which are small bio-crystals consisting of an inorganic mineral and a protein, or otoconial masses (aggregations of otoconia bound by an organic matrix), being significantly different in morphology compared to the singular, polycrystalline otolith structures of bony fishes, which are solidified bio-crystals forming stony masses. Reinvestigation of the morphological and chemical properties of these chondrichthyan otoconia revises our understanding of otolith composition and has implications on the evolution of these characters in both the gnathostome crown group, and cartilaginous fishes in particular. Results: Dissections of Amblyraja radiata, Potamotrygon leopoldi, and Scyliorhinus canicula revealed three pairs of singular polycrystalline otolith structures with a well-defined morphology within their inner ears, as observed in bony fishes. IR spectroscopy identified the material to be composed of carbonate/collagen-bearing apatite in all taxa. These findings contradict previous hypotheses suggesting these otoconial structures were composed of calcium carbonate in chondrichthyans. A phylogenetic mapping using 37 chondrichthyan taxa further showed that the acquisition of phosphatic otolith structures might be widespread within cartilaginous fishes.
    [Show full text]
  • 1 Osteichthyan-Like Cranial Conditions in an Early Devonian Stem Gnathostome Sam Giles , Matt Friedman , Martin D. Brazeau *
    Osteichthyan-like cranial conditions in an Early Devonian stem gnathostome Sam Giles1, Matt Friedman1, Martin D. Brazeau2,3* 1Department of Earth Sciences, University of Oxford, South Parks Road, Oxford, OX1 3AN, UK. 2Naturalis Biodiversity Center, P.O. Box 9517, 2300 RA Leiden, The Netherlands. 3Department of Life Sciences, Imperial College London, Silwood Park Campus, Buckhurst Road, Ascot, SL5 7PY, UK. *Corresponding author. E-mail: [email protected] [First paragraph: ca. 293 words] The phylogeny of Silurian and Devonian (443-358 million years ago [Ma]) fishes remains the foremost problem in the study of the origin of modern gnathostomes (jawed vertebrates). A central question concerns the morphology of the last common ancestor of living jawed vertebrates, with competing hypotheses advancing either a chondrichthyan-1-3 or osteichthyan-like4,5 model. Here we present Janusiscus schultzei gen. et sp. nov., an Early Devonian (ca. 415 Ma) gnathostome from Siberia previously interpreted as a ray-finned fish6, which provides important new information about cranial anatomy near the last common ancestor of 1 chondrichthyans and osteichthyans. The skull roof of Janusiscus resembles that of early osteichthyans, with large plates bearing vermiform ridges and partially enclosed sensory canals. High-resolution computed tomography reveals a braincase bearing characters typically associated with either chondrichthyans (large hypophyseal opening accommodating the internal carotid arteries) or osteichthyans (facial nerve exiting through jugular canal, endolymphatic ducts exiting posterior to the skull roof) and lacking a ventral cranial fissure, the presence of which is considered a derived feature of crown gnathostomes7,8. A conjunction of well- developed cranial processes in Janusiscus helps unify the comparative anatomy of early jawed vertebrate neurocrania, suggesting primary homologies in ‘placoderms’, osteichthyans and chondrichthyans.
    [Show full text]
  • Neurocranial Anatomy of an Enigmatic Early Devonian Fish Sheds Light on Early Osteichthyan Evolution
    RESEARCH ARTICLE Neurocranial anatomy of an enigmatic Early Devonian fish sheds light on early osteichthyan evolution Alice M Clement1,2,3†*, Benedict King1,4†, Sam Giles5†, Brian Choo1, Per E Ahlberg2, Gavin C Young6,7, John A Long1,3 1College of Science and Engineering, Flinders University, Adelaide, Australia; 2Department of Organismal Biology, Evolutionary Biology Centre, Uppsala University, Uppsala, Sweden; 3Department of Sciences, Museum Victoria, Melbourne, Australia; 4Naturalis Biodiversity Center, Leiden, Netherlands; 5Department of Earth Sciences, University of Oxford, Oxford, United Kingdom; 6Department of Applied Mathematics, Research School of Physics & Engineering, Australian National University, Canberra, Australia; 7Australian Museum Research Institute, Sydney, Australia Abstract The skull of ‘Ligulalepis’ from the Early Devonian of Australia (AM-F101607) has significantly expanded our knowledge of early osteichthyan anatomy, but its phylogenetic position has remained uncertain. We herein describe a second skull of ‘Ligulalepis’ and present micro-CT data on both specimens to reveal novel anatomical features, including cranial endocasts. Several features previously considered to link ‘Ligulalepis’ with actinopterygians are now considered generalized osteichthyan characters or of uncertain polarity. The presence of a lateral cranial canal is shown to be variable in its development between specimens. Other notable new features include the presence of a pineal foramen, the some detail of skull roof sutures, the shape of the nasal *For correspondence: capsules, a placoderm-like hypophysial vein, and a chondrichthyan-like labyrinth system. New [email protected] phylogenetic analyses place ‘Ligulalepis’ as a stem osteichthyan, specifically as the sister taxon to † These authors contributed ‘psarolepids’ plus crown osteichthyans. The precise position of ‘psarolepids’ differs between equally to this work parsimony and Bayesian analyses.
    [Show full text]
  • Tully Monster
    University of Birmingham The ‘tully monster’ is not a vertebrate Sallan, Lauren; Giles, Sam; Sansom, Robert; Clarke, John; Johanson, Zerina; Sansom, Ivan; Janvier, Philippe DOI: 10.1111/pala.12282 License: Other (please specify with Rights Statement) Document Version Peer reviewed version Citation for published version (Harvard): Sallan, L, Giles, S, Sansom, R, Clarke, J, Johanson, Z, Sansom, I & Janvier, P 2017, 'The ‘tully monster’ is not a vertebrate: characters, convergence and taphonomy in palaeozoic problematic animals', Palaeontology, vol. 60, no. 2, pp. 149-157. https://doi.org/10.1111/pala.12282 Link to publication on Research at Birmingham portal Publisher Rights Statement: This is the peer reviewed version of the following article: Sallan, L., Giles, S., Sansom, R. S., Clarke, J. T., Johanson, Z., Sansom, I. J. and Janvier, P. (2017), The ‘Tully Monster’ is not a vertebrate: characters, convergence and taphonomy in Palaeozoic problematic animals. Palaeontology, 60: 149–157. doi:10.1111/pala.12282, which has been published in final form at 10.1111/pala.12282. This article may be used for non-commercial purposes in accordance with Wiley Terms and Conditions for Self-Archiving. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research.
    [Show full text]
  • Neurocranial Anatomy of an Enigmatic Early Devonian Fish Sheds
    Neurocranial anatomy of an enigmatic Early Devonian fish sheds light on early osteichthyan evolution Giles, Sam; Clement, Alice ; King, Ben ; Choo, Brian; Ahlberg, Per; Young, Gavin; Long, John DOI: 10.7554/eLife.34349 License: Creative Commons: Attribution (CC BY) Document Version Publisher's PDF, also known as Version of record Citation for published version (Harvard): Giles, S, Clement, A, King, B, Choo, B, Ahlberg, P, Young, G & Long, J 2018, 'Neurocranial anatomy of an enigmatic Early Devonian fish sheds light on early osteichthyan evolution', Elife, vol. 7, e34349. https://doi.org/10.7554/eLife.34349 Link to publication on Research at Birmingham portal General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document. When citing, please reference the published version.
    [Show full text]
  • ENDOSKELETAL STRUCTURE in CHEIROLEPIS (OSTEICHTHYES, ACTINOPTERYGII), an EARLY RAY-FINNED FISH by SAM GILES1, MICHAEL I
    [Palaeontology, Vol. 58, Part 5, 2015, pp. 849–870] ENDOSKELETAL STRUCTURE IN CHEIROLEPIS (OSTEICHTHYES, ACTINOPTERYGII), AN EARLY RAY-FINNED FISH by SAM GILES1, MICHAEL I. COATES2,3, RUSSELL J. GARWOOD4,5, MARTIN D. BRAZEAU6,ROBERTATWOOD7, ZERINA JOHANSON8 and MATT FRIEDMAN1 1Department of Earth Sciences, University of Oxford, South Parks Road, Oxford, OX1 3AN, UK; e-mails: [email protected], [email protected] 2Department of Organismal Biology and Anatomy, University of Chicago, 1027 E. 57th Street, Chicago, IL 60637, USA; e-mail: [email protected] 3Committee on Evolutionary Biology, University of Chicago, 1025 E. 57th Street, Chicago, IL 60637, USA 4School of Earth, Atmospheric and Environmental Sciences, The University of Manchester, Manchester, M13 9PL, UK; e-mail: [email protected] 5The Manchester X-Ray Imaging Facility, School of Materials, The University of Manchester, Manchester, M13 9PL, UK 6Department of Life Sciences, Imperial College London, Silwood Park Campus, Buckhurst Road, Ascot, SL5 7PY, UK; e-mail: [email protected] 7The Joint Engineering and Environmental Processing Beamline, Diamond Light Source, The Harwell Science and Innovation Campus, Didcot, OX11 0DE, UK; e-mail: [email protected] 8Department of Earth Sciences, Natural History Museum, Cromwell Road, London, SW7 5BD, UK; e-mail: [email protected] Typescript received 17 March 2015; accepted in revised form 30 May 2015 Abstract: As the sister lineage of all other actinopterygians, considered uncharacteristic of early actinopterygians, includ- the Middle to Late Devonian (Eifelian–Frasnian) Cheirolepis ing an uninvested dorsal aorta and imperforate propterygium, occupies a pivotal position in vertebrate phylogeny.
    [Show full text]
  • Newsletter 97 2 Editorial
    The Palaeontology Newsletter Contents 97 Editorial 2 Association Business 3 Annual Meeting 2018 3 Awards and Prizes AGM 2017 9 Diversity study update 21 PalAss needs mentors! 22 Association Meetings 23 News 27 From our correspondents Legends of Rock 34 Bug hunt 37 Career prospects for palaeontologists 42 Paradigms in palaeobiology 47 Future meetings of other bodies 50 Meeting Reports 57 Obituary: Nigel Trewin 69 Grant Reports 71 Book Reviews 90 Careering off course! 97 Palaeontology vol. 61 parts 1 & 2 99–100 Papers in Palaeontology vol. 4 part 1 101 Reminder: The deadline for copy for Issue no. 98 is 4th June 2018. On the Web: <http://www.palass.org/> ISSN: 0954-9900 Newsletter 97 2 Editorial Many are touting 2018 as the year of women, in recognition of the global #MeToo and ‘Times Up’ movements, and in the UK it marks a major centenary in the suffragette movement. Awesome women are certainly on show in this issue. Lene Delsett’s Legends of Rock piece documents the life and career of her academic grandmother, Zofia Kielan-Jaworowska. Read Rachel Warnock’s GSA meeting write-up to find out if she was successful in her quest for a Mary Anning “button”. Emma Dunne and Roxanne Armfield report on the Early Tetrapod World meeting that honoured the outstanding career of Lapworth Medal-winner Jenny Clack. Simon Wills conducted this issue’s Careers Q and A with Cathy Whitlock, which concludes with some sage advice for all of us: “[y]ou have to get out there – give talks, introduce yourself to colleagues, and participate in scientific organizations like the Palaeontological Association”.
    [Show full text]
  • Placoderm’ from Mongolia
    bioRxiv preprint doi: https://doi.org/10.1101/2020.06.09.132027; this version posted July 13, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. 1 Endochondral bone in an Early Devonian ‘placoderm’ from Mongolia 2 3 Martin D. Brazeau1,2*, Sam Giles 2,3,4, Richard P. Dearden1,5, Anna Jerve1,6, Y.A. 4 Ariunchimeg7, E. Zorig8, Robert Sansom9, Thomas Guillerme10, Marco Castiello1 5 6 1 Department of Life Sciences, Imperial College London, Silwood Park Campus, Buckhurst 7 Rd, Ascot, SL5 7PY, UK; 8 2 Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 9 5BD, UK; 10 3 School of Geography, Earth and Environmental Sciences, University of Birmingham, 11 Birmingham, UK; 12 4 Department of Earth Sciences, University of Oxford, South Parks Road, Oxford, OX1 3AN, 13 UK; 14 5 CR2P Centre de Recherche en Paléontologie – Paris, Muséum national d’Histoire 15 naturelle, Sorbonne Universités, CNRS, CP 38, 57 Rue Cuvier, 75231, Paris, Cedex 05, 16 France 17 6 Department of Organismal Biology, Subdepartment of Evolution and Development, 18 Uppsala University, Norbyvägen 18A, 752 36 Uppsala, Sweden; 19 7 Natural History Museum, P.O. Box 46/52, Ulaanbaatar 1420, Mongolia 20 8 Institute of Paleontology, Mongolian Academy of Science, P.O. Box 46/650, S. Danzan 21 Street 3/1, Chingeltei District. Ulaanbaatar 15160, Mongolia; 22 9 School of Earth and Environmental Sciences, University of Manchester, Manchester M13 23 9PT, UK; 24 10 Department of Animal and Plant Sciences, The University of Sheffield, Sheffield S10 2TN, 25 UK; bioRxiv preprint doi: https://doi.org/10.1101/2020.06.09.132027; this version posted July 13, 2020.
    [Show full text]