Marmot Marmota Flaviventris Family: Sciuridae
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Ground Squirrels Live in Burrows That Are Litter Size: Five to Seven
L-1909 6/13 Controlling GROUND SQUIRRELDamage round squirrels are small, burrowing rodents earthen dikes with their burrows. Their burrow- found throughout the state, with the excep- ing and gnawing behavior also can cause damage G tion of extreme East Texas. There are five in irrigated areas. different species in Texas. These are the thirteen- lined ground squirrel, Mexican ground squirrel, spotted ground squirrel, rock squirrel, and the Texas antelope ground squirrel. Most ground Biology and Reproduction squirrels prefer grassy areas such as pastures, Rock squirrels golf courses, cemeteries and parks. Rock squir- Adult weight: 1½ to 1¾ pounds. rels are nearly always found in rocky cliffs, Total length: 18 to 21 inches. boulders, and canyon walls. The rock squirrel Color: Varies from dark gray to black. and thirteen-lined ground squirrel are the two species that most commonly cause damage by Tail: 7 to 10 inches, somewhat bushy. their burrowing and gnawing. Gestation period: Approximately 30 days. Ground squirrels live in burrows that are Litter size: Five to seven. usually 2 to 3 inches in diameter and 15 to 20 Number of litters: Possibly two per year, feet long. The burrow system usually has two usually born from April to August. entrances. Dirt piles around the entry holes are seldom evident. Rock squirrels and thirteen- Life span: 4 to 5 years. lined ground squirrels may hibernate during the Thirteen-lined ground squirrels coldest periods of winter. Adult weight: 5 to 9 ounces. Damage Total length: 7 to 12 inches. Ground squirrels normally do not cause Color: Light to dark brown with 13 stripes extensive damage in urban areas. -
Phylogeny, Biogeography and Systematic Revision of Plain Long-Nosed Squirrels (Genus Dremomys, Nannosciurinae) Q ⇑ Melissa T.R
Molecular Phylogenetics and Evolution 94 (2016) 752–764 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Phylogeny, biogeography and systematic revision of plain long-nosed squirrels (genus Dremomys, Nannosciurinae) q ⇑ Melissa T.R. Hawkins a,b,c,d, , Kristofer M. Helgen b, Jesus E. Maldonado a,b, Larry L. Rockwood e, Mirian T.N. Tsuchiya a,b,d, Jennifer A. Leonard c a Smithsonian Conservation Biology Institute, Center for Conservation and Evolutionary Genetics, National Zoological Park, Washington DC 20008, USA b Division of Mammals, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, Washington DC 20013-7012, USA c Estación Biológica de Doñana (EBD-CSIC), Conservation and Evolutionary Genetics Group, Avda. Americo Vespucio s/n, Sevilla 41092, Spain d George Mason University, Department of Environmental Science and Policy, 4400 University Drive, Fairfax, VA 20030, USA e George Mason University, Department of Biology, 4400 University Drive, Fairfax, VA 20030, USA article info abstract Article history: The plain long-nosed squirrels, genus Dremomys, are high elevation species in East and Southeast Asia. Received 25 March 2015 Here we present a complete molecular phylogeny for the genus based on nuclear and mitochondrial Revised 19 October 2015 DNA sequences. Concatenated mitochondrial and nuclear gene trees were constructed to determine Accepted 20 October 2015 the tree topology, and date the tree. All speciation events within the plain-long nosed squirrels (genus Available online 31 October 2015 Dremomys) were ancient (dated to the Pliocene or Miocene), and averaged older than many speciation events in the related Sunda squirrels, genus Sundasciurus. -
Distribution and Abundance of Hoary Marmots in North Cascades National Park Complex, Washington, 2007-2008
National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science Distribution and Abundance of Hoary Marmots in North Cascades National Park Complex, Washington, 2007-2008 Natural Resource Technical Report NPS/NOCA/NRTR—2012/593 ON THE COVER Hoary Marmot (Marmota caligata) Photograph courtesy of Roger Christophersen, North Cascades National Park Complex Distribution and Abundance of Hoary Marmots in North Cascades National Park Complex, Washington, 2007-2008 Natural Resource Technical Report NPS/NOCA/NRTR—2012/593 Roger G. Christophersen National Park Service North Cascades National Park Complex 810 State Route 20 Sedro-Woolley, WA 98284 June 2012 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado publishes a range of reports that address natural resource topics of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Technical Report Series is used to disseminate results of scientific studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service mission. The series provides contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. All manuscripts in the series receive the appropriate level of peer review to ensure that the information is scientifically credible, technically accurate, appropriately written for the intended audience, and designed and published in a professional manner. -
Sound Communication in the Uinta Ground Squirrel
Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 5-1965 Sound Communication in the Uinta Ground Squirrel Donna Mae Balph Utah State University Follow this and additional works at: https://digitalcommons.usu.edu/etd Part of the Environmental Public Health Commons Recommended Citation Balph, Donna Mae, "Sound Communication in the Uinta Ground Squirrel" (1965). All Graduate Theses and Dissertations. 1552. https://digitalcommons.usu.edu/etd/1552 This Thesis is brought to you for free and open access by the Graduate Studies at DigitalCommons@USU. It has been accepted for inclusion in All Graduate Theses and Dissertations by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. SOUND COMMUNICATION IN THE UINTA GROUND SQUIRREL by Donna Mae Balph A thesis submitted in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE in Wildlife Biology Approved: Major Professor ~ &1 Gradua'te Studies UTAH STATE UNIVERSITY Logan, Utah 1965 TABLE OF CONTENTS Page INTRODUCTION GENERAL LIFE HISTORY 2 METHODS AND APPARATUS 7 RESULTS 1 0 Use of calls in interaction with conspeci fics 10 Chirp 10 Chirp by males 10 Chirp by females 1 7 Churr 22 Squeal 24 Squawk 24 Teeth clatter 31 Growl 34 Use of calls by juveniles 34 Comparison of chirp and churr 36 Use of calls in interaction with other species . 43 Reaction to airborne predators 43 Reaction to predators on ground 47 Reaction to snakes 52 DISCUSSION . 55 Unspecifi c ilEilur~-.jJl call s 55 Ease of location of calls 57 CONCLUSIONS 59 LITERATURE CITED 61 LIST OF TABLES Table Page 1. -
A Review of Bristly Ground Squirrels Xerini and a Generic Revision in the African Genus Xerus
Mammalia 2016; 80(5): 521–540 Boris Kryštufek*, Ahmad Mahmoudi, Alexey S. Tesakov, Jan Matějů and Rainer Hutterer A review of bristly ground squirrels Xerini and a generic revision in the African genus Xerus DOI 10.1515/mammalia-2015-0073 Received April 28, 2015; accepted October 13, 2015; previously Introduction published online December 12, 2015 Bristly ground squirrels from the arid regions of Central Abstract: Bristly ground squirrels Xerini are a small rodent Asia and Africa constitute a coherent monophyletic tribe tribe of six extant species. Despite a dense fossil record the Xerini sensu Moore (1959). The tribe contains six species group was never diverse. Our phylogenetic reconstruction, in three genera of which Atlantoxerus and Spermophilop based on the analysis of cytochrome b gene and including sis are monotypic. The genus Xerus in its present scope all known species of Xerini, confirms a deep divergence (Thorington and Hoffmann 2005), consists of four species between the African taxa and the Asiatic Spermophilopsis. in three subgenera: X. inauris and X. princeps (subgenus Genetic divergences among the African Xerini were of a Geosciurus), X. rutilus (subgenus Xerus), and X. eryth comparable magnitude to those among genera of Holarc- ropus (subgenus Euxerus). Recent phylogenetic recon- tic ground squirrels in the subtribe Spermophilina. Evi- struction based on molecular markers retrieved Xerus to dent disparity in criteria applied in delimitation of genera be paraphyletic with respect to Atlantoxerus (Fabre et al. in Sciuridae induced us to recognize two genera formerly 2012), therefore challenging the suitability of the generic incorporated into Xerus. The resurrected genera (Euxerus arrangement of the group. -
Golden-Mantled Ground Squirrel Or Chipmunk? by Lynne Brosch
Who Is Your Pest? Golden-Mantled Ground Squirrel or Chipmunk? by Lynne Brosch Recently as I went around the lake doing talks on pest management, I had several complaints about chipmunks. People describe a lot of digging and eating of plants by these chipmunks. As I began thinking about putting out some information on how to handle the situation I thought about the golden-mantled ground squirrel, I watched eating garden plants voraciously on the Baldwin Estate grounds just yesterday. Perhaps gardeners need to know who they are dealing with. The golden-mantled ground squirrel looks a lot like a chipmunk. It has a large white stripe bordered by black on each side. The main difference between this squirrel and a chipmunk is that its stripes don’t go all the way to the face and it is a slightly larger animal. It lives along the west coast in coniferous forests and mountainous areas. It likes to eat plants, seeds, nuts, fruit and some insects. It lives in an underground burrow usually near trees or logs. Chipmunks have very similar burrows. Most common in the Tahoe basin is the Lodgepole chipmunk. Fencing can be used to protect plants from squirrels and chipmunks, but has challenges in effectiveness because of the excellent digging and climbing skills exhibited by these garden pests. Hardware cloth may be used to exclude animals from flower beds with seeds and bulbs covered by the hardware cloth and all covered with soil. This method of prevention may prove less costly and time consuming than trapping. The most successful method for control of ground squirrels and chipmunks is the use of traps. -
Action Plan for the Conservation of the Danube
Action Plan for the Conservation of the European Ground Squirrel Spermophilus citellus in the European Union EUROPEAN COMMISSION, 2013 1. Compilers: Milan Janák (Daphne/N2K Group, Slovakia), Pavel Marhoul (Daphne/N2K Group, Czech Republic) & Jan Matějů (Czech Republic). 2. List of contributors Michal Adamec, State Nature Conservancy of the Slovak Republic, Slovakia Michal Ambros, State Nature Conservancy of the Slovak Republic, Slovakia Alexandru Iftime, Natural History Museum „Grigore Antipa”, Romania Barbara Herzig, Säugetiersammlung, Naturhistorisches Museum Vienna, Austria Ilse Hoffmann, University of Vienna, Austria Andrzej Kepel, Polish Society for Nature Conservation ”Salamandra”, Poland Yordan Koshev, Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Science, Bulgaria Denisa Lőbbová, Poznaj a chráň, Slovakia Mirna Mazija, Oikon d.o.o.Institut za primijenjenu ekologiju, Croatia Olivér Váczi, Ministry of Rural Development, Department of Nature Conservation, Hungary Jitka Větrovcová, Nature Conservation Agency of the Czech Republic, Czech Republic Dionisios Youlatos, Aristotle University of Thessaloniki, Greece 3. Lifespan of plan/Reviews 2013 - 2023 4. Recommended citation including ISBN Janák M., Marhoul P., Matějů J. 2013. Action Plan for the Conservation of the European Ground Squirrel Spermophilus citellus in the European Union. European Commission. ©2013 European Communities Reproduction is authorised provided the source is acknowledged Cover photo: Michal Ambros Acknowledgements for help and support: Ervín -
Convergent Evolution of Himalayan Marmot with Some High-Altitude Animals Through ND3 Protein
animals Article Convergent Evolution of Himalayan Marmot with Some High-Altitude Animals through ND3 Protein Ziqiang Bao, Cheng Li, Cheng Guo * and Zuofu Xiang * College of Life Science and Technology, Central South University of Forestry and Technology, Changsha 410004, China; [email protected] (Z.B.); [email protected] (C.L.) * Correspondence: [email protected] (C.G.); [email protected] (Z.X.); Tel.: +86-731-5623392 (C.G. & Z.X.); Fax: +86-731-5623498 (C.G. & Z.X.) Simple Summary: The Himalayan marmot (Marmota himalayana) lives on the Qinghai-Tibet Plateau and may display plateau-adapted traits similar to other high-altitude species according to the principle of convergent evolution. We assessed 20 species (marmot group (n = 11), plateau group (n = 8), and Himalayan marmot), and analyzed their sequence of CYTB gene, CYTB protein, and ND3 protein. We found that the ND3 protein of Himalayan marmot plays an important role in adaptation to life on the plateau and would show a history of convergent evolution with other high-altitude animals at the molecular level. Abstract: The Himalayan marmot (Marmota himalayana) mainly lives on the Qinghai-Tibet Plateau and it adopts multiple strategies to adapt to high-altitude environments. According to the principle of convergent evolution as expressed in genes and traits, the Himalayan marmot might display similar changes to other local species at the molecular level. In this study, we obtained high-quality sequences of the CYTB gene, CYTB protein, ND3 gene, and ND3 protein of representative species (n = 20) from NCBI, and divided them into the marmot group (n = 11), the plateau group (n = 8), and the Himalayan marmot (n = 1). -
Prairie Dog (Cynomys Ludovicianus) in Canada
Species at Risk Act Management Plan Series Management Plan for the Black-tailed prairie dog (Cynomys ludovicianus) in Canada Black-tailed Prairie Dog June 2009 About the Species at Risk Act Management Plan Series What is the Species at Risk Act (SARA)? SARA is the Act developed by the federal government as a key contribution to the common national effort to protect and conserve species at risk in Canada. SARA came into force in 2003, and one of its purposes is “to manage species of special concern to prevent them from becoming endangered or threatened.” What is a species of special concern? Under SARA, a species of special concern is a wildlife species that could become threatened or endangered because of a combination of biological characteristics and identified threats. Species of special concern are included in the SARA List of Wildlife Species at Risk. What is a management plan? Under SARA, a management plan is an action-oriented planning document that identifies the conservation activities and land use measures needed to ensure, at a minimum, that a species of special concern does not become threatened or endangered. For many species, the ultimate aim of the management plan will be to alleviate human threats and remove the species from the List of Wildlife Species at Risk. The plan sets goals and objectives, identifies threats, and indicates the main areas of activities to be undertaken to address those threats. Management plan development is mandated under Sections 65–72 of SARA (http://www.sararegistry.gc.ca/approach/act/default_e.cfm). A management plan has to be developed within three years after the species is added to the List of Wildlife Species at Risk. -
Body Size and Diet Mediate Evolution of Jaw Shape in Squirrels (Sciuridae)
Rare ecomorphological convergence on a complex adaptive landscape: body size and diet mediate evolution of jaw shape in squirrels (Sciuridae) *Miriam Leah Zelditch; Museum of Paleontology, University of Michigan, Ann Arbor, MI 48109; [email protected] Ji Ye; Earth and Environmental Sciences, University of Michigan, Ann Arbor, MI 48109; [email protected] Jonathan S. Mitchell; Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109; [email protected] Donald L. Swiderski, Kresge Hearing Research Institute and Museum of Zoology, University of Michigan. [email protected] *Corresponding Author Running head: Convergence on a complex adaptive landscape Key Words: Convergence, diet evolution, jaw morphology, shape evolution, macroevolutionary adaptive landscape, geometric morphometrics Data archival information: doi: 10.5061/dryad.kq1g6. This is the author manuscript accepted for publication and has undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: 10.1111/evo.13168. This article is protected by copyright. All rights reserved. Abstract Convergence is widely regarded as compelling evidence for adaptation, often being portrayed as evidence that phenotypic outcomes are predictable from ecology, overriding contingencies of history. However, repeated outcomes may be very rare unless adaptive landscapes are simple, structured by strong ecological and functional constraints. One such constraint may be a limitation on body size because performance often scales with size, allowing species to adapt to challenging functions by modifying only size. When size is constrained, species might adapt by changing shape; convergent shapes may therefore be common when size is limiting and functions are challenging. -
Franklin, Richardson, Columbian, Washington, and Townsend Ground
Leonard R. Askham Professor Emeritus FRANKLIN, Department of Horticulture and Landscape Architecture Washington State University RICHARDSON, Pullman, Washington 99164-6414 COLUMBIAN, WASHINGTON, AND TOWNSEND Fig. 1. Franklin ground squirrel, Spermophilus franklinii GROUND SQUIRRELS Damage Prevention and Repellents Fumigants Control Methods None are registered. Aluminum phosphide. Toxicants Exclusion Gas cartridge. Zinc phosphide. Limited usefulness. Trapping Chlorophacinone. Cultural Methods Box traps. Diphacinone. Flood irrigation, forage removal, crop Burrow-entrance traps. rotation, and summer fallow may Note: Not all toxicants are registered Leghold traps. reduce populations and limit for use in every state. Check regis- Shooting spread. tration labels for limitations within each state. Limited usefulness. PREVENTION AND CONTROL OF WILDLIFE DAMAGE — 1994 Cooperative Extension Division Institute of Agriculture and Natural Resources University of Nebraska - Lincoln United States Department of Agriculture Animal and Plant Health Inspection Service Animal Damage Control B-159 Great Plains Agricultural Council Wildlife Committee Identification deposition of fat necessary for hiberna- tion. Most store large quantities of The Franklin ground squirrel (Spermo- food in burrow caches. Some species, philus franklinii, Fig. 1) is a rather drab like the Franklin, eat a greater amount grayish brown. Black speckling gives a of animal matter, including ground- spotted or barred effect. Head and nesting bird eggs. Insects and other body average 10 inches (25.4 cm) with animal tissue may comprise up to one- a 5- to 6-inch (12.7- to 15.2-cm) tail. fourth of their diet. Adults weigh from 10 to 25 ounces (280 to 700 g). General Biology, The Richardson ground squirrel (S. richardson) is smaller and lighter Reproduction, and colored than the Franklin. -
Alarm Calling in Yellow-Bellied Marmots: I
Anim. Behav., 1997, 53, 143–171 Alarm calling in yellow-bellied marmots: I. The meaning of situationally variable alarm calls DANIEL T. BLUMSTEIN & KENNETH B. ARMITAGE Department of Systematics and Ecology, University of Kansas (Received 27 November 1995; initial acceptance 27 January 1996; final acceptance 13 May 1996; MS. number: 7461) Abstract. Yellow-bellied marmots, Marmota flaviventris, were reported to produce qualitatively different alarm calls in response to different predators. To test this claim rigorously, yellow-bellied marmot alarm communication was studied at two study sites in Colorado and at one site in Utah. Natural alarm calls were observed and alarm calls were artificially elicited with trained dogs, a model badger, a radiocontrolled glider and by walking towards marmots. Marmots ‘whistled’, ‘chucked’ and ‘trilled’ in response to alarming stimuli. There was no evidence that either of the three call types, or the acoustic structure of whistles, the most common alarm call, uniquely covaried with predator type. Marmots primarily varied the rate, and potentially a few frequency characteristics, as a function of the risk the caller experienced. Playback experiments were conducted to determine the effects of call type (chucks versus whistles), whistle rate and whistle volume on marmot responsiveness. Playback results suggested that variation in whistle number/rate could communicate variation in risk. No evidence was found of intraspecific variation in the mechanism used to communicate risk: marmots at all study sites produced the same vocalizations and appeared to vary call rate as a function of risk. There was significant individual variation in call structure, but acoustic parameters that were individually variable were not used to communicate variation in risk.