Harvard University Program in Neuroscience History of Enrollment in The Program in Neuroscience July 2018 Updated each July Nicholas Spitzer, M.D./Ph.D. B.A., Harvard College Entered 1966 * Defended May 14, 1969 Advisor: David Poer A Physiological and Histological Invesgaon of the Intercellular Transfer of Small Molecules _____________ Professor of Neurobiology University of California at San Diego Eric Frank, Ph.D. B.A., Reed College Entered 1967 * Defended January 17, 1972 Advisor: Edwin J. Furshpan The Control of Facilitaon at the Neuromuscular Juncon of the Lobster _______________ Professor Emeritus of Physiology Tus University School of Medicine Albert Hudspeth, M.D./Ph.D. B.A., Harvard College Entered 1967 * Defended April 30, 1973 Advisor: David Poer Intercellular Juncons in Epithelia _______________ Professor of Neuroscience The Rockefeller University David Van Essen, Ph.D. B.S., California Instute of Technology Entered 1967 * Defended October 22, 1971 Advisor: John Nicholls Effects of an Electronic Pump on Signaling by Leech Sensory Neurons ______________ Professor of Anatomy and Neurobiology Washington University David Van Essen, Eric Frank, and Albert Hudspeth At the 50th Anniversary celebraon for the creaon of the Harvard Department of Neurobiology October 7, 2016 Richard Mains, Ph.D. Sc.B., M.S., Brown University Entered 1968 * Defended April 24, 1973 Advisor: David Poer Tissue Culture of Dissociated Primary Rat Sympathec Neurons: Studies of Growth, Neurotransmier Metabolism, and Maturaon _______________ Professor of Neuroscience University of Conneccut Health Center Peter MacLeish, Ph.D. B.E.Sc., University of Western Ontario Entered 1969 * Defended December 29, 1976 Advisor: David Poer Synapse Formaon in Cultures of Dissociated Rat Sympathec Neurons Grown on Dissociated Rat Heart Cells _______________ Professor and Director of the Neuroscience Instute Morehouse School of Medicine Peter Sargent, Ph.D. B.A., Amherst College Entered 1969* Defended March 21, 1975 Advisor: Zachary Hall Transmiers in the Leech Central Nervous System: Analysis of Sensory and Motor Cells _______________ Professor Emeritus of Cell and Tissue Biology University of California at San Francisco School of Denstry Bruce Wallace, Ph.D. B.A., Amherst College Entered 1969 * Defended September 27, 1974 Advisor: Edward Kravitz The Biosynthesis of Octopamine in the Nervous System of the Lobster, Homarus americanus _______________ Professor Emeritus University of Colorado School of Medicine Joshua Sanes, Ph.D. B.A., Yale University Entered 1970 * Defended January 15, 1976 Advisor: John Hildebrand Origin and Differenaon of Sensory Neurons in an Insect Antenna _______________ Professor of Molecular and Cellular Biology Harvard University Colin Nurse, Ph.D. B.Sc., University of Western Ontario Entered 1971 * Defended May 11, 1977 Advisor: David Poer The formaon of cholinergic synapses between dissociated rat sympathec neurons and skeletal myotubes in cell culture _______________ Professor Emeritus of Biology McMaster University Carla Shatz, Ph.D. B.A., Radcliffe College M.Phil., University College, London Entered 1971 * Defended March 25, 1976 Advisor: David Hubel A Study of Abnormal Connecons in the Visual System of the Siamese Cat, a Genec Mutant _______________ Professor of Neurobiology Stanford University King‐Wai Yau, M.D./Ph.D. A.B., Princeton University Entered 1971 * Defended October 31, 1975 Advisor: John Nicholls Recepve Fields, Geometry and Conducon Block of Sensory Cells in the Leech Central Nervous System _______________ Professor of Neuroscience Johns Hopkins University School of Medicine Linda Lai Yue Chun, Ph.D. B.S., Illinois Instute of Technology Entered 1972 * Defended May 9, 1977 Advisor: Paul Paerson The Role of Nerve Growth Factor in the Development of Rat Sympathec Neurons in vitro _______________ Assistant in Molecular Biology Seed Lab, Massachuses General Hospital Charles Gilbert, M.D./Ph.D. B.A., Amherst College Entered 1971 * Defended July 22, 1976 Advisor: David Hubel Laminar Organizaon of Cat Visual Cortex _______________ Professor of Neurosciences and Behavior Rockefeller University Duane Edgington, Ph.D. B.S., California Instute of Technology Entered 1973 * Defended July 28, 1978 Advisor: Anne Stuart Properes and Distribuon of Ionic Channels in a Photoreceptor Neuron _______________ Soware Engineer Monterey Bay Aquarium Research Instute Patrick Hogan, Ph.D. A.B., Harvard College Entered 1973 * Defended September 29, 1983 Advisor: David Poer Expression of Markers for Pain Sensory Neurons in Cell Culture _______________ Professor, Division of Signaling and Gene Expression La Jolla Instute for Allergy and Immunology Crispin Weinberg, Ph.D. S.B., S.M., University of Chicago Entered 1973 * Defended August 21, 1980 Advisor: Zachary Hall Molecular Specializaons at Mature and Developing Neuromuscular Juncons in Adult Rats _______________ President and Founder Biomedical Modeling, Inc. Kathleen Buckley, Ph.D. A.B., Radcliffe College Entered 1974 * Defended October 30, 1980 Advisor: Story Landis Morphological and Funconal Studies of Synapses and Varicosies in Cultures of Dissociated Rat Sympathec Neurons _______________ Associate Provost for Science Harvard University David Ferster, Ph.D. Sc.B., Brown University Entered 1973 * Defended October 23, 1979 Advisor: David Hubel Corcal Mechanisms of Depth Vision in the Cat _______________ Professor Emeritus Northwestern University Steve Hauger, Ph.D. B.S., Stanford University Entered 1974 * Defended January 14, 1981 Advisor: Edward Furshpan The Spaal Distribuon of Sensivity to Acetylcholine of Rat Sympathec Neurons in Cell Culture _______________ Stained Glass Arsan Margaret Livingstone, Ph.D. B.S., Massachuses Instute of Technology Entered 1974 * Defended February 20, 1980 Advisor: Edward Kravitz Monoamines in the Lobster: Biochemistry, Anatomy, and Possible Funconal Role _______________ Professor of Neurobiology Harvard Medical School Patricia Walicke, M.D./Ph.D. B.S., Massachuses Instute of Technology Entered 1975 * Defended October 11, 1979 Advisor: Paul Paerson Role of Calcium and Cyclic AMP in the Mechanism of the Development Effects of Neuronal Acvity _______________ Senior Health Sciences Administrator Naonal Instute of Neurological Disorders and Stroke Sue Schneps, A.M. B.S., Massachuses Instute of Technology Entered 1976 * Le June 1, 1980 _______________ Withdrew with Master’s degree Jewelry Designer and Owner, Sea and Star Designs Leslie Timpe, Ph.D. A.B., Princeton University Entered 1976 * Defended September 8, 1982 Advisor: Ann Stuart The Roles of Acetylcholine and Gamma‐aminobutyrin Acid in the Visual System of the Barnacle _______________ Lecturer San Francisco State University Paul Desan, Ph.D. A.B., Princeton University Entered 1977 * Defended June 27, 1984 Advisor: Simon LeVay The Organizaon of the Cerebral Cortex of the Pond Turtle, Pseudemys scripta elegans _______________ Assistant Professor of Psychiatry Yale University School of Medicine Thomas Schwarz, Ph.D. A.B., Harvard College Entered 1977 * Defended June 27, 1983 Advisor: Edward Kravitz Proctolin in the Lobster and Protein Phosphorylaon in a Lobster Muscle _______________ Professor of Neurobiology Harvard Medical School Allison Doupe, M.D./Ph.D. DCS, B.Sc., McGill University Entered 1978 * Defended April 26, 1984 Advisor: Paul Paerson Role of Glucocorcoids and Growth Factors in the Development of Neural Crest Derivaves _______________ Deceased October 24, 2014 (Formerly Professor at University of California, San Francisco) Jonathan Horton, M.D./Ph.D. A.B., Stanford University Entered 1978 * Defended February 16, 1983 Advisor: Simon LeVay Cytochrome Oxidase Patches: A New Cytoarchitectonic Feature _______________ Professor of Ophthalmology University of California at San Francisco Kathleen King Siwicki, Ph.D. Sc.B., Brown University M.Phil., Cambridge University Entered 1782 * Defended May 7, 1985 Advisor: Edward Kravitz The Neuropepde, Proctolin, in the Lobster Nervous System: Cellular Distribuon of Proctolin‐Like Angens and Biochemical Studies of Idenfied Neurons _______________ Professor of Neurobiology Swarthmore College Karla Kent, Ph.D. B.S., University of California at Santa Barbara M.A., Boston University Entered 1979 * Defended October 11, 1985 Advisor: John Hildebrand Metamorphosis of the antennal center and the influence of sensory innervaon on the formaon of glomeruli in the hawk moth Manduca sexta _______________ Professor, Integrave Biosciences Oregon Health Sciences University School of Denstry Anne Schneiderman, Ph.D. B.S., Stanford University Entered 1979 * Defended July 16, 1984 Advisor: John Hildebrand Postembryonic Development of a Sexually Dimorphic Sensory Pathway in the Central Nervous System of the Sphinx Moth, Manduca Sexta _______________ Patent Aorney Harris Beach LLC Leslie Stevens, Ph.D. A.B., University of California at Berkeley Entered 1979 * Defended March 27, 1987 Advisor: Story Landis Postembryonic Development of a Sexually Dimorphic Sensory Pathway in the Central Nervous System of the Sphinx Moth, Manduca Sexta _______________ Research Assistant Professor University of Texas at Ausn (Stein Lab) Daniel Tso, Ph.D. A.B., Harvard College Entered 1980 * Defended May 7, 1987 Advisor: Torsten Wiesel Connecvity and Funconal Organizaon in the Mammalian Visual Cortex _______________ Associate Professor Upstate Medical University of the State University of New York James Huener, Ph.D. B.S., B.A., Indiana University of Pennsylvania Entered 1981 * Defended January 15, 1987 Advisor: Robert