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A Review of Southern Iraq Herpetofauna
Vol. 3 (1): 61-71, 2019 A Review of Southern Iraq Herpetofauna Nadir A. Salman Mazaya University College, Dhi Qar, Iraq *Corresponding author: [email protected] Abstract: The present review discussed the species diversity of herpetofauna in southern Iraq due to their scientific and national interests. The review includes a historical record for the herpetofaunal studies in Iraq since the earlier investigations of the 1920s and 1950s along with the more recent taxonomic trials in the following years. It appeared that, little is known about Iraqi herpetofauna, and no comprehensive checklist has been done for these species. So far, 96 species of reptiles and amphibians have been recorded from Iraq, but only a relatively small proportion of them occur in the southern marshes. The marshes act as key habitat for globally endangered species and as a potential for as yet unexplored amphibian and reptile diversity. Despite the lack of precise localities, the tree frog Hyla savignyi, the marsh frog Pelophylax ridibunda and the green toad Bufo viridis are found in the marshes. Common reptiles in the marshes include the Caspian terrapin (Clemmys caspia), the soft-shell turtle (Trionyx euphraticus), the Euphrates softshell turtle (Rafetus euphraticus), geckos of the genus Hemidactylus, two species of skinks (Trachylepis aurata and Mabuya vittata) and a variety of snakes of the genus Coluber, the spotted sand boa (Eryx jaculus), tessellated water snake (Natrix tessellata) and Gray's desert racer (Coluber ventromaculatus). More recently, a new record for the keeled gecko, Cyrtopodion scabrum and the saw-scaled viper (Echis carinatus sochureki) was reported. The IUCN Red List includes six terrestrial and six aquatic amphibian species. -
Gliding Dragons and Flying Squirrels: Diversifying Versus Stabilizing Selection on Morphology Following the Evolution of an Innovation
vol. 195, no. 2 the american naturalist february 2020 E-Article Gliding Dragons and Flying Squirrels: Diversifying versus Stabilizing Selection on Morphology following the Evolution of an Innovation Terry J. Ord,1,* Joan Garcia-Porta,1,† Marina Querejeta,2,‡ and David C. Collar3 1. Evolution and Ecology Research Centre and the School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, New South Wales 2052, Australia; 2. Institute of Evolutionary Biology (CSIC–Universitat Pompeu Fabra), Passeig Marítim de la Barceloneta, 37–49, Barcelona 08003, Spain; 3. Department of Organismal and Environmental Biology, Christopher Newport University, Newport News, Virginia 23606 Submitted August 1, 2018; Accepted July 16, 2019; Electronically published December 17, 2019 Online enhancements: supplemental material. Dryad data: https://doi.org/10.5061/dryad.t7g227h. fi abstract: Evolutionary innovations and ecological competition are eral de nitions of what represents an innovation have been factors often cited as drivers of adaptive diversification. Yet many offered (reviewed by Rabosky 2017), this classical descrip- innovations result in stabilizing rather than diversifying selection on tion arguably remains the most useful (Galis 2001; Stroud morphology, and morphological disparity among coexisting species and Losos 2016; Rabosky 2017). Hypothesized innovations can reflect competitive exclusion (species sorting) rather than sympat- have drawn considerable attention among ecologists and ric adaptive divergence (character displacement). We studied the in- evolutionary biologists because they can expand the range novation of gliding in dragons (Agamidae) and squirrels (Sciuridae) of ecological niches occupied within communities. In do- and its effect on subsequent body size diversification. We found that gliding either had no impact (squirrels) or resulted in strong stabilizing ing so, innovations are thought to be important engines of selection on body size (dragons). -
Nymphalidae, Brassolinae) from Panama, with Remarks on Larval Food Plants for the Subfamily
Journal of the Lepidopterists' Society 5,3 (4), 1999, 142- 152 EARLY STAGES OF CALICO ILLIONEUS AND C. lDOMENEUS (NYMPHALIDAE, BRASSOLINAE) FROM PANAMA, WITH REMARKS ON LARVAL FOOD PLANTS FOR THE SUBFAMILY. CARLA M. PENZ Department of Invertebrate Zoology, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, Wisconsin 53233, USA , and Curso de P6s-Gradua9ao em Biocicncias, Pontiffcia Universidade Cat61ica do Rio Grande do SuI, Av. Ipiranga 6681, FOlto Alegre, RS 90619-900, BRAZIL ANNETTE AIELLO Smithsonian Tropical Research Institute, Apdo. 2072, Balboa, Ancon, HEPUBLIC OF PANAMA AND ROBERT B. SRYGLEY Smithsonian Tropical Research Institute, Apdo. 2072, Balboa, Ancon, REPUBLIC OF PANAMA, and Department of Zoology, University of Oxford, South Parks Road, Oxford, OX13PS, ENGLAND ABSTRACT, Here we describe the complete life cycle of Galigo illioneus oberon Butler and the mature larva and pupa of C. idomeneus (L.). The mature larva and pupa of each species are illustrated. We also provide a compilation of host records for members of the Brassolinae and briefly address the interaction between these butterflies and their larval food plants, Additional key words: Central America, host records, monocotyledonous plants, larval food plants. The nymphalid subfamily Brassolinae includes METHODS Neotropical species of large body size and crepuscular habits, both as caterpillars and adults (Harrison 1963, Between 25 May and .31 December, 1994 we Casagrande 1979, DeVries 1987, Slygley 1994). Larvae searched for ovipositing female butterflies along generally consume large quantities of plant material to Pipeline Road, Soberania National Park, Panama, mo reach maturity, a behavior that may be related as much tivated by a study on Caligo mating behavior (Srygley to the low nutrient content of their larval food plants & Penz 1999). -
Pseudotrapelus Jensvindumi (Reptilia: Agamidae) in Western Asia
RESEARCH ARTICLE Effects of climate change on niche shifts of Pseudotrapelus dhofarensis and Pseudotrapelus jensvindumi (Reptilia: Agamidae) in Western Asia Iman Rounaghi, Seyyed Saeed Hosseinian Yousefkhani* Young Researchers and Elite Club, Islamic Azad University, Shirvan branch, Shirvan, Iran * [email protected] a1111111111 a1111111111 a1111111111 Abstract a1111111111 Genus Pseudotrapelus has a wide distribution in North Africa and in the Middle East. In the a1111111111 present study, we modeled the habitat suitability of two Omani species of the genus (Pseu- dotrapelus dhofarensis and Pseudotrapelus jensvindumi) to evaluate the potential effects of climate change on their distribution. Mean diurnal range and precipitation of wettest quarter are the most highly contributed variables for P. jensvindumi and P. dhofarensis, respec- OPEN ACCESS tively. The potential distribution for P. dhofarensis in the current time covers the southern Citation: Rounaghi I, Hosseinian Yousefkhani SS coastal regions of Oman, Yemen, the Horn of Africa, and Socotra Island, but the suitable (2018) Effects of climate change on niche shifts of regions were reduced in the future prediction and limited to Yemen, Socotra Island, and Pseudotrapelus dhofarensis and Pseudotrapelus jensvindumi (Reptilia: Agamidae) in Western Asia. Oman. There have not been any records of the species outside of Oman. Analysis of habitat PLoS ONE 13(5): e0197884. https://doi.org/ suitability for P. jensvindumi indicated that the species is restricted to the Al Hajar Mountain 10.1371/journal.pone.0197884 of Oman and the southeast coastal region of Iran, but there are no records of the species Editor: Luciano Bosso, Universita degli Studi di from Iran. Because mean diurnal range will not be influenced by climate change in future, Napoli Federico II, ITALY the potential distribution of the species is not expected to be changed in 2050. -
An Overview and Checklist of the Native and Alien Herpetofauna of the United Arab Emirates
Herpetological Conservation and Biology 5(3):529–536. Herpetological Conservation and Biology Symposium at the 6th World Congress of Herpetology. AN OVERVIEW AND CHECKLIST OF THE NATIVE AND ALIEN HERPETOFAUNA OF THE UNITED ARAB EMIRATES 1 1 2 PRITPAL S. SOORAE , MYYAS AL QUARQAZ , AND ANDREW S. GARDNER 1Environment Agency-ABU DHABI, P.O. Box 45553, Abu Dhabi, United Arab Emirates, e-mail: [email protected] 2Natural Science and Public Health, College of Arts and Sciences, Zayed University, P.O. Box 4783, Abu Dhabi, United Arab Emirates Abstract.—This paper provides an updated checklist of the United Arab Emirates (UAE) native and alien herpetofauna. The UAE, while largely a desert country with a hyper-arid climate, also has a range of more mesic habitats such as islands, mountains, and wadis. As such it has a diverse native herpetofauna of at least 72 species as follows: two amphibian species (Bufonidae), five marine turtle species (Cheloniidae [four] and Dermochelyidae [one]), 42 lizard species (Agamidae [six], Gekkonidae [19], Lacertidae [10], Scincidae [six], and Varanidae [one]), a single amphisbaenian, and 22 snake species (Leptotyphlopidae [one], Boidae [one], Colubridae [seven], Hydrophiidae [nine], and Viperidae [four]). Additionally, we recorded at least eight alien species, although only the Brahminy Blind Snake (Ramphotyplops braminus) appears to have become naturalized. We also list legislation and international conventions pertinent to the herpetofauna. Key Words.— amphibians; checklist; invasive; reptiles; United Arab Emirates INTRODUCTION (Arnold 1984, 1986; Balletto et al. 1985; Gasperetti 1988; Leviton et al. 1992; Gasperetti et al. 1993; Egan The United Arab Emirates (UAE) is a federation of 2007). -
Nota Lepidopterologica
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Nota lepidopterologica Jahr/Year: 2002 Band/Volume: 25 Autor(en)/Author(s): Garcia-Barros Enrique Artikel/Article: Taxonomic patterns in the egg to body size allometry of butterflies and skippers (Papilionoidea & Hesperiidae) 161-175 ©Societas Europaea Lepidopterologica; download unter http://www.biodiversitylibrary.org/ und www.zobodat.at Nota lepid. 25 (2/3): 161-175 161 Taxonomic patterns in the egg to body size allometry of butter- flies and skippers (Papilionoidea & Hesperiidae) Enrique Garcia-Barros Departmento de Biologia (Zool.), Universidad Autönoma de Madrid, E-28049 Madrid, Spain e-mail: [email protected] Summary. Former studies have shown that there is an interspecific allometric relationship between egg size and adult body size in butterflies and skippers. This is here re-assessed at the family and subfamily levels in order to determine to what extent the overall trend is uniform through different taxonomic lineages. The results suggest that different subtaxa are characterised by different allometric slopes. Al- though statistical analysis across species means is known to be potentially misleading to assess evolu- tionary relations, it is shown that the comparison of apparent patterns (based on species means) with inferred evolutionary trends (based on independent contrasts) may help to understand the evolution of egg size in butterflies. Further, intuitive reconsideration of statistically non-significant results may prove informative. As an example, argumentation in favour of a positive association between large egg size and the use of monocotyledon plants as larval food is presented. Taxa where atypical allometric trends are found include the Riodininae and Theclini (Lycaenidae), the Graphiini (Papilionidae), and the Heliconiinae (Nymphalidae). -
Selva El O C O Te
RESERVA DE LA BIOSFERA MAB SELVA EL OCOTE PRIMER REVISIÓN PERIÓDICA MÉXICO 2017 I PRIMERA REVISIÓN PERIÓDICA DE LA RESERVA DE LA BIOSFERA MAB SELVA EL OCOTE [2017] Contenido PARTE I: RESUMEN ..................................................................................................................... 1 PARTE II: INFORME DE REVISIÓN PERIÓDICA .................................................................... 5 1. RESERVA DE LA BIOSFERA ...................................................................................... 5 2. CAMBIOS SIGNIFICATIVOS EN LA RESERVA DE LA BIOSFERA DURANTE LOS DIEZ ÚLTIMOS AÑOS ............................................................ 7 3. SERVICIOS DE LOS ECOSISTEMAS ....................................................................... 32 4. FUNCIÓN DE CONSERVACIÓN .............................................................................. 35 5. FUNCIÓN DE DESARROLLO ................................................................................... 41 6. FUNCIÓN DE LOGÍSTICA ......................................................................................... 51 7. GOBERNANZA, GESTIÓN Y COORDINACIÓN DE LA RESERVA DE LA BIOSFERA ..................................................................................................... 65 8. CRITERIOS Y PROGRESOS ALCANZADOS .......................................................... 82 9. DOCUMENTOS COMPLEMENTARIOS ................................................................... 89 10. DIRECCIONES ................................................................................................. -
Volume 2. Animals
AC20 Doc. 8.5 Annex (English only/Seulement en anglais/Únicamente en inglés) REVIEW OF SIGNIFICANT TRADE ANALYSIS OF TRADE TRENDS WITH NOTES ON THE CONSERVATION STATUS OF SELECTED SPECIES Volume 2. Animals Prepared for the CITES Animals Committee, CITES Secretariat by the United Nations Environment Programme World Conservation Monitoring Centre JANUARY 2004 AC20 Doc. 8.5 – p. 3 Prepared and produced by: UNEP World Conservation Monitoring Centre, Cambridge, UK UNEP WORLD CONSERVATION MONITORING CENTRE (UNEP-WCMC) www.unep-wcmc.org The UNEP World Conservation Monitoring Centre is the biodiversity assessment and policy implementation arm of the United Nations Environment Programme, the world’s foremost intergovernmental environmental organisation. UNEP-WCMC aims to help decision-makers recognise the value of biodiversity to people everywhere, and to apply this knowledge to all that they do. The Centre’s challenge is to transform complex data into policy-relevant information, to build tools and systems for analysis and integration, and to support the needs of nations and the international community as they engage in joint programmes of action. UNEP-WCMC provides objective, scientifically rigorous products and services that include ecosystem assessments, support for implementation of environmental agreements, regional and global biodiversity information, research on threats and impacts, and development of future scenarios for the living world. Prepared for: The CITES Secretariat, Geneva A contribution to UNEP - The United Nations Environment Programme Printed by: UNEP World Conservation Monitoring Centre 219 Huntingdon Road, Cambridge CB3 0DL, UK © Copyright: UNEP World Conservation Monitoring Centre/CITES Secretariat The contents of this report do not necessarily reflect the views or policies of UNEP or contributory organisations. -
Literature Cited in Lizards Natural History Database
Literature Cited in Lizards Natural History database Abdala, C. S., A. S. Quinteros, and R. E. Espinoza. 2008. Two new species of Liolaemus (Iguania: Liolaemidae) from the puna of northwestern Argentina. Herpetologica 64:458-471. Abdala, C. S., D. Baldo, R. A. Juárez, and R. E. Espinoza. 2016. The first parthenogenetic pleurodont Iguanian: a new all-female Liolaemus (Squamata: Liolaemidae) from western Argentina. Copeia 104:487-497. Abdala, C. S., J. C. Acosta, M. R. Cabrera, H. J. Villaviciencio, and J. Marinero. 2009. A new Andean Liolaemus of the L. montanus series (Squamata: Iguania: Liolaemidae) from western Argentina. South American Journal of Herpetology 4:91-102. Abdala, C. S., J. L. Acosta, J. C. Acosta, B. B. Alvarez, F. Arias, L. J. Avila, . S. M. Zalba. 2012. Categorización del estado de conservación de las lagartijas y anfisbenas de la República Argentina. Cuadernos de Herpetologia 26 (Suppl. 1):215-248. Abell, A. J. 1999. Male-female spacing patterns in the lizard, Sceloporus virgatus. Amphibia-Reptilia 20:185-194. Abts, M. L. 1987. Environment and variation in life history traits of the Chuckwalla, Sauromalus obesus. Ecological Monographs 57:215-232. Achaval, F., and A. Olmos. 2003. Anfibios y reptiles del Uruguay. Montevideo, Uruguay: Facultad de Ciencias. Achaval, F., and A. Olmos. 2007. Anfibio y reptiles del Uruguay, 3rd edn. Montevideo, Uruguay: Serie Fauna 1. Ackermann, T. 2006. Schreibers Glatkopfleguan Leiocephalus schreibersii. Munich, Germany: Natur und Tier. Ackley, J. W., P. J. Muelleman, R. E. Carter, R. W. Henderson, and R. Powell. 2009. A rapid assessment of herpetofaunal diversity in variously altered habitats on Dominica. -
Computational Molecular Species Delimitation and Taxonomic Revision of the Gecko Genus Ebenavia Boettger, 1878
The Science of Nature (2018) 105:49 https://doi.org/10.1007/s00114-018-1574-9 ORIGINAL PAPER Computational molecular species delimitation and taxonomic revision of the gecko genus Ebenavia Boettger, 1878 Oliver Hawlitschek1 & Mark D. Scherz1,2 & Bernhard Ruthensteiner1 & Angelica Crottini3 & Frank Glaw1 Received: 22 February 2018 /Revised: 13 June 2018 /Accepted: 3 July 2018 # Springer-Verlag GmbH Germany, part of Springer Nature 2018 Abstract Cryptic species have been detected in many groups of organisms and must be assumed to make up a significant portion of global biodiversity. We study geckos of the Ebenavia inunguis complex from Madagascar and surrounding islands and use species delimitation algorithms (GMYC, BOLD, BPP), COI barcode divergence, diagnostic codon indels in the nuclear marker PRLR, diagnostic categorical morphological characters, and significant differences in continuous morphological characters for its taxonomic revision. BPP yielded ≥ 10 operational taxonomic units, whereas GMYC (≥ 27) and BOLD (26) suggested substantial oversplitting. In consequnce, we resurrect Ebenavia boettgeri Boulenger 1885 and describe Ebenavia tuelinae sp. nov., Ebenavia safari sp. nov., and Ebenavia robusta sp. nov., increasing the number of recognised species in Ebenavia from two to six. Further lineages of Ebenavia retrieved by BPP may warrant species or subspecies status, but further taxonomic conclusions are postponed until more data become available. Finally, we present an identification key to the genus Ebenavia, provide an updated distribution map, and discuss the diagnostic values of computational species delimitation as well as morphological and molecular diagnostic characters. Keywords BOLD . Operational Taxonomic Unit . Madagascar clawless gecko . Integrative taxonomy . Taxonomic inflation . Species complex Introduction taxa (Bickford et al. -
Extreme Diversity of Tropical Parasitoid Wasps Exposed by Iterative Integration of Natural History, DNA Barcoding, Morphology, and Collections
Extreme diversity of tropical parasitoid wasps exposed by iterative integration of natural history, DNA barcoding, morphology, and collections M. Alex Smith*†, Josephine J. Rodriguez‡, James B. Whitfield‡, Andrew R. Deans§, Daniel H. Janzen†¶, Winnie Hallwachs¶, and Paul D. N. Hebert* *The Biodiversity Institute of Ontario, University of Guelph, Guelph Ontario, N1G 2W1 Canada; ‡Department of Entomology, 320 Morrill Hall, University of Illinois, 505 S. Goodwin Avenue, Urbana, IL 61801; §Department of Entomology, North Carolina State University, Campus Box 7613, 2301 Gardner Hall, Raleigh, NC 27695-7613; and ¶Department of Biology, University of Pennsylvania, Philadelphia, PA 19104-6018 Contributed by Daniel H. Janzen, May 31, 2008 (sent for review April 18, 2008) We DNA barcoded 2,597 parasitoid wasps belonging to 6 microgas- A detailed recognition of species in parasitoid communities is trine braconid genera reared from parapatric tropical dry forest, cloud necessary because of the pivotal role parasitoids play in food web forest, and rain forest in Area de Conservacio´ n Guanacaste (ACG) in structure and dynamics. While generalizations about the effects of northwestern Costa Rica and combined these data with records of parasitoids on community diversity are complex (7), a common- caterpillar hosts and morphological analyses. We asked whether place predictor of the impact of a parasitoid species on local host barcoding and morphology discover the same provisional species and dynamics is whether the parasitoid is a generalist or specialist. A whether the biological entities revealed by our analysis are congruent generalist, especially a mobile one, is viewed as stabilizing food webs with wasp host specificity. Morphological analysis revealed 171 (see ref. -
Amphiesmeno- Ptera: the Caddisflies and Lepidoptera
CY501-C13[548-606].qxd 2/16/05 12:17 AM Page 548 quark11 27B:CY501:Chapters:Chapter-13: 13Amphiesmeno-Amphiesmenoptera: The ptera:Caddisflies The and Lepidoptera With very few exceptions the life histories of the orders Tri- from Old English traveling cadice men, who pinned bits of choptera (caddisflies)Caddisflies and Lepidoptera (moths and butter- cloth to their and coats to advertise their fabrics. A few species flies) are extremely different; the former have aquatic larvae, actually have terrestrial larvae, but even these are relegated to and the latter nearly always have terrestrial, plant-feeding wet leaf litter, so many defining features of the order concern caterpillars. Nonetheless, the close relationship of these two larval adaptations for an almost wholly aquatic lifestyle (Wig- orders hasLepidoptera essentially never been disputed and is supported gins, 1977, 1996). For example, larvae are apneustic (without by strong morphological (Kristensen, 1975, 1991), molecular spiracles) and respire through a thin, permeable cuticle, (Wheeler et al., 2001; Whiting, 2002), and paleontological evi- some of which have filamentous abdominal gills that are sim- dence. Synapomorphies linking these two orders include het- ple or intricately branched (Figure 13.3). Antennae and the erogametic females; a pair of glands on sternite V (found in tentorium of larvae are reduced, though functional signifi- Trichoptera and in basal moths); dense, long setae on the cance of these features is unknown. Larvae do not have pro- wing membrane (which are modified into scales in Lepi- legs on most abdominal segments, save for a pair of anal pro- doptera); forewing with the anal veins looping up to form a legs that have sclerotized hooks for anchoring the larva in its double “Y” configuration; larva with a fused hypopharynx case.