Anggrainin0511.Pdf (8.943Mb)

Total Page:16

File Type:pdf, Size:1020Kb

Anggrainin0511.Pdf (8.943Mb) CARBONATE PRESERVATION OF DINOSAUR EGGS IN THE UPPER CRETACEOUS ANACLETO FORMATION AT AUCA MAHUEVO, NEUQUÈN BASIN, ARGENTINA by Niswatin Wahida Anggraini A thesis submitted in partial of the requirements for the degree of Master of Science in Earth Sciences MONTANA STATE UNIVERSITY Bozeman, Montana January 2011 ©COPYRIGHT by Niswatin Wahida Anggraini 2011 All Rights Reserved ii APPROVAL of a thesis submitted by Niswatin Wahida Anggraini This thesis has been read by each member of the thesis committee and has been found to be satisfactory regarding content, English usage, format, citation, bibliographic style, and consistency and is ready for submission to the Division of Graduate Education. Dr. James G. Schmitt Approved for the Department of Earth Sciences Dr. Stephen G. Custer Approved for the Division of Graduate Education Dr. Carl A. Fox iii STATEMENT OF PERMISSION TO USE In presenting this thesis in partial fulfillment of the requirements for a master’s degree at Montana State University, I agree that the Library shall make it available to borrowers under rules of the Library. If I have indicated my intention to copyright this thesis by including a copyright notice page, copying is allowable only for scholarly purposes, consistent with “fair use” as prescribed in the U.S. Copyright Law. Requests for permission for extended quotation from or reproduction of this thesis in whole or in parts may be granted only by the copyright holder. Niswatin Wahida Anggraini January 2011 iv ACKNOWLEDGEMENTS I would like to thank to my advisor Jim Schmitt, and my committee members Dave Mogk and Frankie Jackson, for all of their guidance. To Jim, thank you so much for your encouragement and assistance to solve any difficulties in my work. To Dave Mogk, thanks for training me to do the treatment of thin sections and sample preparation of XRD. To Frankie, thanks for sharing articles and the detailed corrections of this thesis. I would like to thank for ExxonMobil Exploration Indonesia Inc. and ExxonMobil Houston for their grant under ExxonMobil scholarship program. Enam, thanks for your support and control about the progress of my study. Thanks to Imaging and Chemical Analysis Laboratory (ICAL) at Montana State University for providing the analytical instruments that I used in my thesis research. For my husband, Ely Setiawan and my daughter, Nafisa Setiawan for their never ending support and motivation. v TABLE OF CONTENTS 1. INTRODUCTION ..................................................................................................... 1 2. GEOLOGIC SETTING ............................................................................................. 4 Lithostratigraphy of the Upper Cretaceous Anacleto Formation............................... 5 3. METHODS ............................................................................................................... 8 4. OBSERVATION ....................................................................................................... 11 Eggshells.................................................................................................................... 11 Hand Sample......................................................................................................... 11 Thin Section.......................................................................................................... 11 Membrane .................................................................................................................. 13 Embryonic Skin ......................................................................................................... 14 Spherulites ................................................................................................................. 15 Ooids.......................................................................................................................... 16 Pellets and Peloids ..................................................................................................... 17 Microcodium.............................................................................................................. 18 Filaments.................................................................................................................... 20 Micrite and Sparite..................................................................................................... 21 Carbonate Component ............................................................................................... 21 Siliciclastic Components............................................................................................ 22 Quartz.................................................................................................................... 22 Gypsum................................................................................................................. 23 Feldspar................................................................................................................. 24 Non-Carbonate Authigenic Mineral .......................................................................... 25 5. DISCUSSION............................................................................................................ 28 Evidence of Microbial Activity ................................................................................. 28 Spherulites ........................................................................................................... 28 Microcodium........................................................................................................ 30 Ooids.................................................................................................................... 31 Precipitation of Calcium Carbonate........................................................................... 32 Microbial Mineral Precipitation................................................................................. 34 6. CONCLUSION.......................................................................................................... 42 REFERENCES CITED.................................................................................................. 44 vi LIST OF FIGURES Figure Page 2.1: Regional map and stratigraphic sections ..................................................... 7 4.1: Photo of a carbonate egg in hand sample .................................................... 11 4.2: Photomicrograph of eggshells and FEM images......................................... 12 4.3: CL photomicrograph of carbonate eggs ...................................................... 13 4.4: Photomicrograph of membrane ................................................................... 14 4.5: Photomicrograph of embryonic skin ........................................................... 15 4.6: Photomicrograph of spherulites................................................................... 16 4.7: Photomicrograph of ooids............................................................................ 17 4.8: Photomicrograph of pellets and peloids ...................................................... 18 4.9: Two different morphologies of Microcodium ............................................. 19 4.10: Photomicrograph of Microcodium............................................................. 19 4.11: FEM images of filaments........................................................................... 20 4.12: Photomicrograph of micrite and sparite .................................................... 21 4.13: XRD diagram of bulk carbonate egg samples ........................................... 22 4.14: Photomicrograph of detrital quartz............................................................ 23 4.15: EDX patterns and FEM image of gypsum................................................. 24 4.16: EDX patterns and FEM image of feldspar ................................................ 25 4.17: EDX patterns and FEM image of analcime............................................... 26 5.1: FEM image of abiotic origin of calcite crystals........................................... 30 5.2: Autotrophic pathways of carbonate bacterial .............................................. 37 vii ABSTRACT Preservation of dinosaur eggs and footprints by precipitation of calcium carbonate in the Upper Cretaceous Anacleto Formation at Auca Mahuevo, Argentina represents a relatively unusual occcurence in the fossil record. Under normal condition, eggs are readily destroyed in sediments shortly after burial by physical, chemical, and biological processes. This study attempts to determine a preservational model for carbonate eggs by characterizing their mineralogical composition and microstructures using a variety of analytical instruments including petrographic microscope, cathodoluminesce (CL) microscope, X-ray powder diffraction (XRD) and field-emission scanning electron microscope (FEM) to characterize the composition and fabric of the fossilized eggs. Several textutal features have been observed in the carbonate eggs, including membrane, embryonic skin, spherulites, ooids, peloids, Microcodium, calcified filaments, and micrite. Microbial actvity is likely responsible for the formation of these microfabric features, facilitating calcium carbonate precipitation leading to exceptional preservation of eggs. Although microbial influence in the carbonate egg preservation has not been clearly elucidated, laboratory experiments by other workers provide an argument for the role of microbes in the precipitation of calcium carbonate.
Recommended publications
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • First Ornithopod Remains from the Bajo De La Carpa Formation (Santonian, Upper Cretaceous), Northern Patagonia, Argentina
    Cretaceous Research 83 (2018) 182e193 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes First ornithopod remains from the Bajo de la Carpa Formation (Santonian, Upper Cretaceous), northern Patagonia, Argentina * Penelope Cruzado-Caballero a, , Leonardo S. Filippi b, Ariel H. Mendez a, Alberto C. Garrido c, d, Ignacio Díaz-Martínez a a Instituto de Investigacion en Paleobiología y Geología (CONICET-UNRN), Av. Roca 1242, General Roca, Río Negro, Argentina b Museo Municipal Argentino Urquiza, Jujuy y Chaco s/n, Rincon de los Sauces, Neuquen, Argentina c Museo Provincial de Ciencias Naturales “Prof. Dr. Juan Olsacher”, Direccion Provincial de Minería, Etcheluz y Ejercito Argentino, Zapala, Neuquen, Argentina d Departamento Geología y Petroleo, Facultad de Ingeniería, Universidad Nacional del Comahue, Buenos Aires 1400, Neuquen, Argentina article info abstract Article history: In the last decades, the Argentinian ornithopod record has been increased with new and diverse bone Received 29 March 2017 remains found along all the Upper Cretaceous. Most of them are very incomplete and represent taxa of Received in revised form different size. As result, the studies about the palaeobiodiversity of the Ornithopoda clade in South 24 July 2017 America are complex. In this paper, new postcranial remains of an indeterminate medium-sized Accepted in revised form 30 July 2017 ornithopod from the Santonian Bajo de la Carpa Formation (Rincon de los Sauces, Neuquen province) Available online 12 August 2017 are presented. They present diagnostic features of the Ornithopoda clade, and several characters that relate them with other Argentinian ornithopods, especially with the medium-sized members of the Keywords: Ornithischia Elasmaria clade sensu Calvo et al.
    [Show full text]
  • Gastroliths in an Ornithopod Dinosaur
    Brief report Acta Palaeontologica Polonica 53 (2): 351–355, 2008 Gastroliths in an ornithopod dinosaur IGNACIO A. CERDA Gastroliths (stomach stones) are known from many extant Institutional abbreviations.—MCSPv, Vertebrate paleontology and extinct vertebrates, including dinosaurs. Reported here collection of the Museo de Cinco Saltos, Río Negro Province, is the first unambiguous record of gastroliths in an ornitho− Argentina; MUCPv, Vertebrate paleontology collection of the pod dinosaur. Clusters of small stones found in the abdomi− Museo de la Universidad Nacional del Comahue, Neuquén nal region of three articulated skeletons of Gasparinisaura Province, Argentina. cincosaltensis were identified as gastroliths on the basis of taphonomic and sedimentologic evidence. The large number Material and geologic setting of stones found in each individual, their size, and the fact that Gasparinisaura cincosaltensis was herbivorous, all sug− Three specimens of Gasparinisaura cincosaltensis, MUCPv 213, gest that they were ingested as a result of lithophagy rather MCSPv 111, and MCSPv 112, were collected near the city of than accidental swallowing. Cinco Saltos (Río Negro Province, Patagonia, Argentina) (Fig. 1), in mudstones and sandstones of the early Campanian Anacleto Introduction Formation, in the uppermost portion of the Neuquén Group (Ramos 1981; Dingus et al. 2000). MUCPv 213 (Fig. 2A) consists Gastroliths or geo−gastroliths sensu Wings (2007) are known in of a partial skeleton that includes cranial and postcranial elements many taxa of extant and fossil vertebrates (Whittle and Everhart (see Salgado et al. 1997 for a detailed anatomical description). A 2000). Gastroliths have been occasionally reported in non−avian portion of the preserved elements (both incomplete humeri articu− dinosaurs (Wings 2004) but only few cases can withstand rigorous lated with both radii and ulnae, several posterior dorsal ribs from testing.
    [Show full text]
  • Zootaxa, Barrosasaurus Casamiquelai Gen. Et Sp. Nov., a New Titanosaur
    Zootaxa 2222: 1–16 (2009) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2009 · Magnolia Press ISSN 1175-5334 (online edition) Barrosasaurus casamiquelai gen. et sp. nov., a new titanosaur (Dinosauria, Sauropoda) from the Anacleto Formation (Late Cretaceous: early Campanian) of Sierra Barrosa (Neuquén, Argentina) LEONARDO SALGADO1 & RODOLFO A. CORIA2 1Conicet-Inibioma, Museo de Geología y Paleontología de la Universidad Nacional del Comahue, Buenos Aires 1400, 8300 Neuquén, Argentina. E-mail: [email protected] 2Conicet, Museo “Carmen Funes”, 8318 Plaza Huincul, Neuquén, Argentina. E-mail: [email protected] Abstract A new Late Cretaceous titanosaurian sauropod from the Sierra Barrosa locality (Anacleto Formation, Late Cretaceous, early Campanian) of Neuquén Province, Argentina, Barrosasaurus casamiquelai gen. et sp. nov., is described. The holotype of this species consists of three large and incomplete, although well-preserved, dorsal vertebrae (one probably the third, the next the seventh or eighth, and the last the ninth or tenth). The arrangement of neural arch laminae in the dorsal vertebrae of this titanosaur differs from that present in other genera in the following ways: the spinoprezygapophyseal laminae are well developed in the third vertebra, and relictual in the seventh or eighth vertebra; two spinodiapophyseal laminae are present in the seventh or eighth and in the ninth or tenth vertebrae, the anterior spinodiapophyseal lamina being more strongly developed than the posterior spinodiapophyseal lamina in the ninth or tenth vertebra. These characters, among others, allow the recognition of this individual as a new species of Titanosauria, which increases knowledge of the diversity of titanosaurs in the Late Cretaceous of Patagonia.
    [Show full text]
  • Paleontological Discoveries in the Chorrillo Formation (Upper Campanian-Lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina
    Rev. Mus. Argentino Cienc. Nat., n.s. 21(2): 217-293, 2019 ISSN 1514-5158 (impresa) ISSN 1853-0400 (en línea) Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina Fernando. E. NOVAS1,2, Federico. L. AGNOLIN1,2,3, Sebastián ROZADILLA1,2, Alexis M. ARANCIAGA-ROLANDO1,2, Federico BRISSON-EGLI1,2, Matias J. MOTTA1,2, Mauricio CERRONI1,2, Martín D. EZCURRA2,5, Agustín G. MARTINELLI2,5, Julia S. D´ANGELO1,2, Gerardo ALVAREZ-HERRERA1, Adriel R. GENTIL1,2, Sergio BOGAN3, Nicolás R. CHIMENTO1,2, Jordi A. GARCÍA-MARSÀ1,2, Gastón LO COCO1,2, Sergio E. MIQUEL2,4, Fátima F. BRITO4, Ezequiel I. VERA2,6, 7, Valeria S. PEREZ LOINAZE2,6 , Mariela S. FERNÁNDEZ8 & Leonardo SALGADO2,9 1 Laboratorio de Anatomía Comparada y Evolución de los Vertebrados. Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida Ángel Gallardo 470, Buenos Aires C1405DJR, Argentina - fernovas@yahoo. com.ar. 2 Consejo Nacional de Investigaciones Científicas y Técnicas, Argentina. 3 Fundación de Historia Natural “Felix de Azara”, Universidad Maimonides, Hidalgo 775, C1405BDB Buenos Aires, Argentina. 4 Laboratorio de Malacología terrestre. División Invertebrados Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida Ángel Gallardo 470, Buenos Aires C1405DJR, Argentina. 5 Sección Paleontología de Vertebrados. Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida Ángel Gallardo 470, Buenos Aires C1405DJR, Argentina. 6 División Paleobotánica. Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida Ángel Gallardo 470, Buenos Aires C1405DJR, Argentina. 7 Área de Paleontología. Departamento de Geología, Universidad de Buenos Aires, Pabellón 2, Ciudad Universitaria (C1428EGA) Buenos Aires, Argentina. 8 Instituto de Investigaciones en Biodiversidad y Medioambiente (CONICET-INIBIOMA), Quintral 1250, 8400 San Carlos de Bariloche, Río Negro, Argentina.
    [Show full text]
  • Lyons SCIENCE 2021 the Influence of Juvenile Dinosaurs SUPPL.Pdf
    science.sciencemag.org/content/371/6532/941/suppl/DC1 Supplementary Materials for The influence of juvenile dinosaurs on community structure and diversity Katlin Schroeder*, S. Kathleen Lyons, Felisa A. Smith *Corresponding author. Email: [email protected] Published 26 February 2021, Science 371, 941 (2021) DOI: 10.1126/science.abd9220 This PDF file includes: Materials and Methods Supplementary Text Figs. S1 and S2 Tables S1 to S7 References Other Supplementary Material for this manuscript includes the following: (available at science.sciencemag.org/content/371/6532/941/suppl/DC1) MDAR Reproducibility Checklist (PDF) Materials and Methods Data Dinosaur assemblages were identified by downloading all vertebrate occurrences known to species or genus level between 200Ma and 65MA from the Paleobiology Database (PaleoDB 30 https://paleobiodb.org/#/ download 6 August, 2018). Using associated depositional environment and taxonomic information, the vertebrate database was limited to only terrestrial organisms, excluding amphibians, pseudosuchians, champsosaurs and ichnotaxa. Taxa present in formations were confirmed against the most recent available literature, as of November, 2020. Synonymous taxa or otherwise duplicated taxa were removed. Taxa that could not be identified to genus level 35 were included as “Taxon X”. GPS locality data for all formations between 200MA and 65MA was downloaded from PaleoDB to create a minimally convex polygon for each possible formation. Any attempt to recreate local assemblages must include all potentially interacting species, while excluding those that would have been separated by either space or time. We argue it is 40 acceptable to substitute formation for home range in the case of non-avian dinosaurs, as range increases with body size.
    [Show full text]
  • Titanosaur Trackways from the Late Cretaceous El Molino Formation of Bolivia (Cal Orck’O, Sucre)
    Annales Societatis Geologorum Poloniae (2018), vol. 88: 223 – 241. doi: https://doi.org/10.14241/asgp.2018.014 TITANOSAUR TRACKWAYS FROM THE LATE CRETACEOUS EL MOLINO FORMATION OF BOLIVIA (CAL ORCK’O, SUCRE) Christian A. MEYER1, Daniel MARTY2 & Matteo BELVEDERE3 1 Department of Environmental Sciences, University of Basel, Bernoullistrasse 32, CH-4056 Basel, Switzerland; e-mail: [email protected] 2 Museum of Natural History Basel, Augustinergasse 2, CH- 4000 Basel, Switzerland; e-mail: [email protected] 3 Office de la culture, Paléontologie A16, Hôtel des Halles, P.O. Box 64, CH-2900 Porrentruy 2, Switzerland; e-mail: [email protected] Meyer, C. A., Marty, D. & Belvedere, M., 2018. Titanosaur trackways from the Late Cretaceous El Mo- lino Formation of Bolivia (Cal Orck’o, Sucre). Annales Societatis Geologorum Poloniae, 88: 223 – 241. Abstract: The Cal Orck’o tracksite is exposed in a quarry wall, approximately 4.4 km NW of Sucre (Department Chuquisaca, Bolivia) in the Altiplano/Cordillera Oriental, in the El Molino Formation (Middle Maastrichtian). Fossiliferous oolitic limestones, associated with large, freshwater stromatolites and nine levels of dinosaur tracks in the El Molino Formation document an open lacustrine environment. The main track-bearing level is almost vertical with a surface area of ~ 65,000 m2. The high-resolution mapping of the site from 1998 to 2015 revealed a total of 12,092 individual dinosaur tracks in 465 trackways. Nine different morphotypes of dinosaur tracks have been documented. Amongst them are several trackways of theropods, orni- thopods, ankylosaurs and sauropods, with the latter group accounting for 26% of the trackways.
    [Show full text]
  • Abelisauroidea and Carchardontosauridae (Theropoda, Dinosauria) in the Cretaceous of South America
    Abelisauroidea and carchardontosauridae (theropoda, dinosauria) in the cretaceous of south america. Paleogeographical and geocronological implications Carlos Roberto dos Anjos Candeiro, Agustín Guillermo Martinelli ABELISAUROIDEA AND CARCHARDONTOSAURIDAE (THEROPODA, DINOSAURIA) IN THE CRETACEOUS OF SOUTH AMERICA. PALEOGEOGRAPHICAL AND GEOCRONOLOGICAL IMPLICATIONS Abelisauroidea e Carcharodontosauridae (Theropoda, Dinosauria) na América do Sul durante do Cretáceo. Implicações paleogeograficas e geocronologicas Carlos Roberto dos Anjos Candeiro Depart. Geologia, Universidade Federal do Rio de Janeiro [email protected], [email protected] Agustín Guillermo Martinelli Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Av. Ángel Gallardo 470, C1405DJR, Buenos Aires, Argentina [email protected] Artigo recebido em 18/10/2004 e aceito para publicação em 31/08/2005 ABSTRACT: In this contribution an up-to-date list of abelisauroid ceratosaurians and carcharodontosaurid allosaurians recognized in South America is presented. Abelisauroids and carcharodontosaurids in South America show rich species diversity and a wide range of temporal and geographical distribution. At least eight formally described species of Abelisauroidea are recognized in Argen- tina and only one in Brazil; in contrast, only one species of Carcharodontosauridae is known for all South America. The fossil record of abelisauroids and carcharodontosaurids in South America shows a dominance of abelisauroids in the upper late Cretaceous, while the dominance of carcharodontosaurids as large predators was during the Cenomanian-Turonian. Although knowl- edge of the evolution of Abelisauroidea and Carcharodontasauridae in South America, as well as in the rest of Gondwana is still far for being complete, intensive explorations in recent years have provided greater insight into the composition of theropod faunas in the Cretaceous of Gondwana.
    [Show full text]
  • Analysis of Titanopodus Tracks from the Late Cretaceous of Mendoza, Argentina
    “main” — 2011/1/12 — 13:08 — page 1 — #1 Anais da Academia Brasileira de Ciências (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 www.scielo.br/aabc Speeds and stance of titanosaur sauropods: analysis of Titanopodus tracks from the Late Cretaceous of Mendoza, Argentina BERNARDO J. GONZÁLEZ RIGA Departamento de Paleontología, IANIGLA, CCT-CONICET-Mendoza, Avda. Ruiz Leal s/n, Parque Gral. San Martín (5500) Mendoza, Argentina/Instituto de Ciencias Básicas, Universidad Nacional de Cuyo Manuscript received on November 13, 2009; accepted for publication on June 21, 2010 ABSTRACT Speed estimations from trackways of Titanopodus mendozensis González Riga and Calvo provide information about the locomotion of titanosaurian sauropods that lived in South America during the Late Cretaceous. Titanopodus ichnites were found at Agua del Choique, a newly discovered track site in the Loncoche Formation, Late Campanian-Early Maastrichtian of Mendoza, Argentina. This speed study follows the hypothesis of dynamic similarity proposed by Alexander. As a refinement of this method, a complementary equation is presented here based on an articulated titanosaurian specimen collected in strata that are regarded as correlative to those that have yielded Titanopodus tracks (Allen Formation, Neuquén Basin). This analysis indicates that hip height can be estimated as 4.586 times the length of the pes track in derived titanosaurs. With an estimation of the hip height and the stride measurements, the speed is calculated. The study of two wide-gauge trackways indicates that Titanopodus ichnites were produced by medium- sized titanosaurs (hip height of 211-229 cm) that walked at 4.7-4.9 km/h towards the south and southwest, following, in part, a sinuous pathway.
    [Show full text]
  • Dinosaurs (Reptilia, Archosauria) at Museo De La Plata, Argentina: Annotated Catalogue of the Type Material and Antarctic Specimens
    Palaeontologia Electronica palaeo-electronica.org Dinosaurs (Reptilia, Archosauria) at Museo de La Plata, Argentina: annotated catalogue of the type material and Antarctic specimens Alejandro Otero and Marcelo Reguero ABSTRACT A commented-illustrated catalogue of non-avian dinosaurs housed at Museo de La Plata, Argentina is presented. This represents the first commented catalogue of the La Plata Museum dinosaurs to be published. This includes the type material as well as Antarctic specimens. The arrangement of the material was made in a phylogenetic fashion, including systematic rank, type material, referred specimens, geographic and stratigraphic location, and comments/remarks, when necessary. A total of 13 type specimens of non-avian dinosaurs are housed at the collection of Museo de La Plata, including eight sauropods, one theropod, one ornithischian, and three ichnotaxa. There are four Antarctic specimens, one of which is a holotype, whereas other corresponds to the first sauropod dinosaur registered for that continent. Alejandro Otero. División Paleontología de Vertebrados, Museo de La Plata. Paseo del Bosque s/n (1900), La Plata, Argentina. [email protected] Marcelo Reguero. División Paleontología de Vertebrados, Museo de La Plata. Paseo del Bosque s/n (1900), La Plata, Argentina. [email protected] Keywords: Saurischia; Ornithischia; Holotype; Lectotype; Patagonia; Antarctica INTRODUCTION housed at the DPV collection, reaching more than 600 individual bones. These include 13 type mate- The Museo de La Plata (MLP), constructed in rial and otherwise important dinosaur specimens, 1884, is one of the oldest museums of natural his- which include ornithischians (e.g., ornithopods, tory of South America. The collection of the ankylosaurs), saurischians (e.g., theropods, sau- División Paleontología de Vertebrados (DPV) was ropodomorphs), as well as three ichnospecies created in 1877 when the museum’s future Direc- included (see Table 1 for a summary of the taxa tor, Dr.
    [Show full text]
  • Theropod Dinosaurs from Argentina
    139 Theropod dinosaurs from Argentina Martín D. EZCURRA1 & Fernando E. NOVAS2 1CONICET, Sección Paleontología Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Av. Angel Gallardo 470, Buenos Aires, C1405DJR, Argentina. [email protected], Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Av. Ángel Gallardo 470, Buenos Aires, C1405DJR, Argentina. [email protected] Abstract. Theropoda includes all the dinosaurs more closely related to birds than to sauropodomorphs (long-necked dinosaurs) and ornithischians (bird-hipped dinosaurs). The oldest members of the group are early Late Triassic in age, and non-avian theropods flourished during the rest of the Mesozoic until they vanished in the Cretaceous-Palaeogene mass extinction. Theropods radiated into two main lineages, Ceratosauria and Tetanurae, which are well represented in Cretaceous rocks from Argentina. Ceratosaurians are the most taxonomically diverse South American non-avian theropods, including small to large-sized species, such as the iconic horned dinosaur Carnotaurus. Argentinean tetanurans are represented by multiple lineages that include some of the largest carnivorous dinosaurs known worldwide (carcharodontosaurids), the enigmatic large-clawed megaraptorans, and small to medium-sized species very closely related to avialans (e.g. unenlagiids). The Argentinean non-avian theropod record has been and is crucial to understand the evolutionary and palaeobiogeographical
    [Show full text]
  • A New Titanosaur Sauropod Dinosaur from the Upper Cretaceous of North Patagonia, Argentina
    A new titanosaur sauropod dinosaur from the Upper Cretaceous of North Patagonia, Argentina LEONARDO S. FILIPPI, RODOLFO A. GARCÍA, and ALBERTO C. GARRIDO Filippi, L.S., García, R.A., and Garrido, A.C. 2011. A new titanosaur sauropod dinosaur from the Upper Cretaceous of North Patagonia, Argentina. Acta Palaeontologica Polonica 56 (3): 505–520. A new sauropod titanosaur from the Upper Cretaceous Anacleto Formation is described. Narambuenatitan palomoi gen. et sp. nov., is diagnosed by cranial and axial autapomorphies. The holotype, which represent a subadult individual, con− sists of the left premaxilla and maxilla, braincase, both quadrates, one cervical vertebrae, one dorsal vertebra, fragments of cervical and dorsal ribs, seventeen caudal vertebrae, caudal transverse processes, fragments of haemal arches, left ster− nal plate, right coracoid, left humerus, left ulnae, both pubes, iliac pedicel, proximal fragment of right ischia, and an in− complete left femur. The phylogenetic analysis indicates that Narambuenatitan is a non−eutitanosaurian lithostrotian, and that it shares with Epachthosaurus a neural spine in middle caudal vertebrae which are laminar and posteriorly elongated. Key words: Sauropoda, Titanosauria, phylogeny, Campanian, Cretaceous, Anacleto Formation, Neuquén Basin, Ar− gentina. Leonardo S. Filippi [[email protected]], Museo “Carmen Funes”, Av. Córdoba 55, (8318) Plaza Huincul, Neuquén, Argentina; Rodolfo A. García [[email protected]], Inibioma – Museo de Geología y Paleontología, Universidad Nacional del Comahue, Buenos Aires 1400, (8300) Neuquén, Argentina; Alberto C. Garrido [[email protected]], Museo Provincial “Prof. Dr. Juan Olsacher”, Dirección General de Minería, Elena de Vega 472, (8340) Zapala, Neuquén, Argentina. Received 7 February 2010, accepted 23 December 2010, available online 29 December 2010.
    [Show full text]