DOCSLIB.ORG

Yu-Feng Hsu and Jerry A. Powell

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Yu-Feng Hsu and Jerry A. Powell Phylogenetic Relationships within Heliodinidae and Systematics of Moths Formerly Assigned to Heliodines Stainton (Lepidoptera: Yponomeutoidea) Yu-Feng Hsu and Jerry A. Powell Phylogenetic Relationships within Heliodinidae and Systematics of Moths Formerly Assigned to Heliodines Stainton (Lepidoptera: Yponomeutoidea) Yu-Feng Hsu and Jerry A. Powell UNIVERSITY OF CALIFORNIA PRESS Berkeley • Los Angeles • London UNIVERSITY OF CALIFORNIA PUBLICATIONS IN ENTOMOLGY Editorial Board: Penny Gullan, Bradford A. Hawkins, John Heraty, Lynn S. Kimsey, Serguei V. Triapitsyn, Philip S. Ward, Kipling Will Volume 124 UNIVERSITY OF CALIFORNIA PRESS BERKELEY AND LOS ANGELES, CALIFORNIA UNIVERSITY OF CALIFORNIA PRESS, LTD. LONDON, ENGLAND © 2005 BY THE REGENTS OF THE UNIVERSITY OF CALIFORNIA PRINTED IN THE UNITED STATES OF AMERICA Library of Congress Cataloging-in-Publication Data Hsu, Yu-Feng, 1963– Phylogenetic relationships within Heliodinidae and systematics of moths formerly assigned to Heliodines Stainton (Lepidoptera: Yponomeutoidea) / Yu-Feng Hsu and Jerry A. Powell. p. cm. Includes bibliographical references. ISBN 0-520-09847-1 (paper : alk. paper) — (University of California publications in entomology ; 124) 1. Heliodinidae—Classification. 2. Heliodinidae—Phylogeny. I. Title. II. Series. QL561.H44 H78 595.78 22—dc22 2004058800 Manufactured in the United States of America The paper used in this publication meets the minimum requirements of ANSI/NISO Z39.48-1992 (R 1997) (Permanence of Paper). Contents Acknowledgments, ix Abstract, xi Introduction ...................................................... 1 Problems in Systematics of Heliodinidae and a Historical Review ............ 4 Material and Methods ............................................ 6 Specimens and Depositories, 6 Dissections and Measurements, 7 Wing Venation Preparation, 7 Scanning Electron Microscope Preparation, 8 Species Discrimination and Description, 8 Larval Rearing Procedures, 8 Phylogenetic Methods, 9 Phylogeny of Heliodinidae ......................................... 12 Monophyly of Heliodinidae, 12 Characters and States Used in the Cladistic Analyses, 13 Results of Phylogenetic Analyses, 19 Systematic Account ................................................ 27 Family Heliodinidae Heinemann, 27 Key to the Genera of Heliodinidae and Species Groups of Aetole, 29 Heliodines assemblage, 31 Heliodines Stainton, 31 H. roesella (Linnaeus), 32 Epicroesa assemblage, 35 Epicroesa Meyrick, 35 Philocoristis Meyrick, 35 Lamprolophus assemblage, 36 Lamprolophus Busck, 36 Aetole assemblage, 36 ‘Heliodines’ princeps subassemblage, 36 ‘Heliodines’ princeps lineage, 36 ‘Heliodines’ princeps Meyrick, 37 Lithariapteryx subassemblage, 38 Lithariapteryx Chambers, 38 Neoheliodines Hsu, new genus, 38 Key to species of Neoheliodines, 40 N. nyctaginella (Gibson), new combination, 40 N. eurypterus Hsu, new species, 43 N. hodgesi Hsu, new species, 43 N. megostiellus Hsu, new species, 46 N. melanobasilarus Hsu, new species, 47 N. vernius Hsu, new species, 48 N. cliffordi (Harrison and Passoa), new combination, 51 N. arizonense Hsu, new species, 53 N. albidentus Hsu, new species, 55 Aetole subassemblage, 56 Embola Walsingham, 56 Key to species of Embola of North and Central America, 58 E. ionis (Clarke), new combination, 59 E. ciccella (Barnes and Busck), new combination, 61 E. cyanozostera Hsu, new species, 63 E. albaciliella (Busck), new combination, 64 E. friedlanderi Hsu, new species, 66 E. melanotela Hsu, new species, 67 E. autumnalis Hsu, new species, 68 E. sexpunctella (Walsingham), new combination, 69 E. dentifer Walsingham, 70 E. powelli Hsu, 71 E. xanthocephala Walsingham, 73 Heliogemma Hsu, new genus, 74 H. gigantea Hsu, new species, 75 H. grandis Hsu, new species, 76 H. preclara Hsu, new species, 77 Pseudastasia Walsingham, 78 Euheliodines Hsu, new genus, 79 E. chemsaki Hsu, new species, 79 E. jaliscella Hsu, new species, 81 Aetole Chambers, revised status, 82 Eximia Group, 83 A. eximia Hsu, new species, 84 A. insolita Hsu, new species, 86 Bella Group, 87 A. bella Chambers, 88 A. schulzella (Fabricius), new combination, 93 A. fulgida Hsu, new species, 95 A. prenticei Hsu, new species, 96 Tripunctella Group, 99 A. tripunctella (Walsingham), new combination, 101 A. unipunctella (Walsingham), new combination, 103 A. inusitata Hsu, new species, 107 A. aprica Hsu, new species, 108 A. cera Hsu, new species, 108 A. favonia Hsu, new species, 110 Extraneella Group, 111 A. extraneella (Walsingham), new combination, 112 Calcifer Group, 116 A. calcifer (Walsingham), new combination, 117 A. calciferoides Hsu, new species, 119 Galapagoensis Group, 121 A. galapagoensis (Heppner and B. Landry), new combination, 122 Copocentra Meyrick, 123 Scelorthus Busck, 124 Incertae Sedis, 124 ‘Heliodines’ aureoflamma Walsingham, 124 ‘H.’ marginata Walsingham, 125 ‘H.’ urichi Busck, 126 Biology ........................................................ 127 Larval Host Plant Relationships, 127 Life History, Voltinism, and Diapause, 129 Behavior, 129 Mating, 129 Hind Leg Posture, 130 Oviposition, 130 Larval Biology, 130 Pupal Biology, 131 Summary ...................................................... .133 Tables, 134 Literature Cited, 148 Figures 26–220, 159 Acknowledgments We are grateful to the following for assistance in field work, comments on the manuscript, the loan of specimens, and other contributions: John Brown, Systematic Entomology Laboratory, USDA, U.S. National Museum of Natural History, Washington, D.C.; John Chemsak, Essig Museum of Entomology, University of California, Berkeley; David Furth, formerly at the Museum of Comparative Zoology, Harvard University, Cambridge; Reinhard Gaeike, Deutsches Entomologisches Institut, Eberswalde; Terry Harrison, University of Illinois, Urbana; John Heppner, Florida State Collection of Arthropods, Gainesville; Ronald Hodges, Eugene, Oregon, formerly USDA, U.S. National Museum of Natural History, Washington, D.C.; Niels Kristensen, University of Copenhagen, Universitetsparken; Bernard Landry, Muséum d’histoire naturelle, Geneva; Jean-François Landry, Canadian National Collection, Ottawa; Martin Lodl, Naturhistorisches Museum Wien, Vienna, Austria; Carolina Martin, Museo Nacional de Ciencias Naturales, Madrid, Spain; Scott Miller, formerly of the Bishop Museum, Honolulu, Hawaii; Sigeru Moriuti and Toshiya Hirowatari, University of Osaka Prefecture, Sakai, Japan; Norman Penny, California Academy of Sciences, San Francisco; Michael Prentice, Jorge Santiago-Blay, and Robert Zuparko, University of California, Berkeley; Edward Riley, Texas A.&M. University, College Station; Michael Shaffer, Natural History Museum, London; Frederick Stehr, Michigan State University, East Lansing; David Wagner, University of Connecticut, Storrs; Shen-Horn Yen, Imperial College at Silwood Park, Ascot. Brown and Harrison provided detailed critical reviews of the manuscript that were exceptionally useful. Many plant identifications were made by Barbara Ertter, UC Herbarium, which retains the voucher specimens. YFH is especially indebted to John Doyen and Thomas Duncan, University of California, Berkeley, for many helpful comments and suggestions and Christopher Meacham and Brent Mishler for assistance with cladistic analysis. Finally, we are indebted to Gordon Nishida, who prepared the illustrations in electronic format, and provided other computer expertise enabling completion of the final copy. ix Abstract Heliodinidae traditionally have been characterized on the basis of forewing venation, color and scaling, and perch behavior, but none of these attributes defines monophyly. We identify four uniquely derived autapomorphies for the family: (1) M vein of forewing two-branched, presumably with M3 lost; (2) tegumen greatly expanded posteriorly, forming a sclerotized, hollow tube; (3) ventral branches of apophyses anteriores originating from a fused transverse bridge; and (4) pupa with long, stiff dorsal and lateral setae. Phylogenetic relationships among genera and species groups of world Heliodinidae are constructed using parsimony and character compatibility as optimality criteria, with representatives of six other families of Yponomeutoidea as outgroups. Results of the analyses show Heliodines Stainton, as formerly recognized (i.e., all the species with conspicuous red markings on the forewings), to be a polyphyletic assemblage. To accommodate the New World fauna, two old names, Aetole Chambers and Embola Walsingham, have been resurrected from synonymy, and three new genera are described: Neoheliodines Hsu (Type species: Heliodines nyctaginella Gibson, 1914), Heliogemma Hsu (Type species: H. gigantea Hsu), and Euheliodines Hsu (Type species: E. chemsaki Hsu). The South American genus Crembalastis Meyrick is synonymized with Embola. A descriptive taxonomy is provided for North and Central American and Caribbean species formerly assigned to Heliodines; 45 species are treated, 25 of which are described as new: Aetole fulgida (TL: Sinaloa, Mexico), A. prenticei (Calif.), A. eximia (Baja Calif., Mexico), A. insolita (El Salvador), A. cera (Calif.), A. favonia (Calif.), A. inusitata (Baja Calif., Mexico), A. aprica (Texas), A. calciferoides (Veracruz, Mexico); Embola autumnalis (Ariz.), E. cyanozostera (Nevada), E. friedlanderi (San Luis Potosí, Mexico), E. melanotela (Haiti); Euheliodines chemsaki (S. L. P., Mexico), E. jaliscella (Jalisco, Mexico); Heliogemma gigantea (Jalisco, Mexico), H. grandis (Tamaulipas, Mexico), H. preclara (Jalisco, Mexico); Neoheliodines albidentus (Ariz.), N. arizonense
Load more

Recommended publications
  • SYSTEMATICS of the MEGADIVERSE SUPERFAMILY GELECHIOIDEA (INSECTA: LEPIDOPTEA) DISSERTATION Presented in Partial Fulfillment of T
    SYSTEMATICS OF THE MEGADIVERSE SUPERFAMILY GELECHIOIDEA (INSECTA: LEPIDOPTEA) DISSERTATION Presented in Partial Fulfillment of the Requirements for The Degree of Doctor of Philosophy in the Graduate School of The Ohio State University By Sibyl Rae Bucheli, M.S. ***** The Ohio State University 2005 Dissertation Committee: Approved by Dr. John W. Wenzel, Advisor Dr. Daniel Herms Dr. Hans Klompen _________________________________ Dr. Steven C. Passoa Advisor Graduate Program in Entomology ABSTRACT The phylogenetics, systematics, taxonomy, and biology of Gelechioidea (Insecta: Lepidoptera) are investigated. This superfamily is probably the second largest in all of Lepidoptera, and it remains one of the least well known. Taxonomy of Gelechioidea has been unstable historically, and definitions vary at the family and subfamily levels. In Chapters Two and Three, I review the taxonomy of Gelechioidea and characters that have been important, with attention to what characters or terms were used by different authors. I revise the coding of characters that are already in the literature, and provide new data as well. Chapter Four provides the first phylogenetic analysis of Gelechioidea to include molecular data. I combine novel DNA sequence data from Cytochrome oxidase I and II with morphological matrices for exemplar species. The results challenge current concepts of Gelechioidea, suggesting that traditional morphological characters that have united taxa may not be homologous structures and are in need of further investigation. Resolution of this problem will require more detailed analysis and more thorough characterization of certain lineages. To begin this task, I conduct in Chapter Five an in- depth study of morphological evolution, host-plant selection, and geographical distribution of a medium-sized genus Depressaria Haworth (Depressariinae), larvae of ii which generally feed on plants in the families Asteraceae and Apiaceae.
    [Show full text]
  • Colorado Wildlife Action Plan: Proposed Rare Plant Addendum
    Colorado Wildlife Action Plan: Proposed Rare Plant Addendum By Colorado Natural Heritage Program For The Colorado Rare Plant Conservation Initiative June 2011 Colorado Wildlife Action Plan: Proposed Rare Plant Addendum Colorado Rare Plant Conservation Initiative Members David Anderson, Colorado Natural Heritage Program (CNHP) Rob Billerbeck, Colorado Natural Areas Program (CNAP) Leo P. Bruederle, University of Colorado Denver (UCD) Lynn Cleveland, Colorado Federation of Garden Clubs (CFGC) Carol Dawson, Bureau of Land Management (BLM) Michelle DePrenger-Levin, Denver Botanic Gardens (DBG) Brian Elliott, Environmental Consulting Mo Ewing, Colorado Open Lands (COL) Tom Grant, Colorado State University (CSU) Jill Handwerk, Colorado Natural Heritage Program (CNHP) Tim Hogan, University of Colorado Herbarium (COLO) Steve Kettler, U.S. Fish and Wildlife Service (USFWS) Andrew Kratz, U.S. Forest Service (USFS) Sarada Krishnan, Colorado Native Plant Society (CoNPS), Denver Botanic Gardens Brian Kurzel, Colorado Natural Areas Program Eric Lane, Colorado Department of Agriculture (CDA) Paige Lewis, The Nature Conservancy (TNC) Ellen Mayo, U.S. Fish and Wildlife Service Mitchell McGlaughlin, University of Northern Colorado (UNC) Jennifer Neale, Denver Botanic Gardens Betsy Neely, The Nature Conservancy Ann Oliver, The Nature Conservancy Steve Olson, U.S. Forest Service Susan Spackman Panjabi, Colorado Natural Heritage Program Jeff Peterson, Colorado Department of Transportation (CDOT) Josh Pollock, Center for Native Ecosystems (CNE) Nicola Ripley,
    [Show full text]
  • Pima County Plant List (2020) Common Name Exotic? Source
    Pima County Plant List (2020) Common Name Exotic? Source McLaughlin, S. (1992); Van Abies concolor var. concolor White fir Devender, T. R. (2005) McLaughlin, S. (1992); Van Abies lasiocarpa var. arizonica Corkbark fir Devender, T. R. (2005) Abronia villosa Hariy sand verbena McLaughlin, S. (1992) McLaughlin, S. (1992); Van Abutilon abutiloides Shrubby Indian mallow Devender, T. R. (2005) Abutilon berlandieri Berlandier Indian mallow McLaughlin, S. (1992) Abutilon incanum Indian mallow McLaughlin, S. (1992) McLaughlin, S. (1992); Van Abutilon malacum Yellow Indian mallow Devender, T. R. (2005) Abutilon mollicomum Sonoran Indian mallow McLaughlin, S. (1992) Abutilon palmeri Palmer Indian mallow McLaughlin, S. (1992) Abutilon parishii Pima Indian mallow McLaughlin, S. (1992) McLaughlin, S. (1992); UA Abutilon parvulum Dwarf Indian mallow Herbarium; ASU Vascular Plant Herbarium Abutilon pringlei McLaughlin, S. (1992) McLaughlin, S. (1992); UA Abutilon reventum Yellow flower Indian mallow Herbarium; ASU Vascular Plant Herbarium McLaughlin, S. (1992); Van Acacia angustissima Whiteball acacia Devender, T. R. (2005); DBGH McLaughlin, S. (1992); Van Acacia constricta Whitethorn acacia Devender, T. R. (2005) McLaughlin, S. (1992); Van Acacia greggii Catclaw acacia Devender, T. R. (2005) Acacia millefolia Santa Rita acacia McLaughlin, S. (1992) McLaughlin, S. (1992); Van Acacia neovernicosa Chihuahuan whitethorn acacia Devender, T. R. (2005) McLaughlin, S. (1992); UA Acalypha lindheimeri Shrubby copperleaf Herbarium Acalypha neomexicana New Mexico copperleaf McLaughlin, S. (1992); DBGH Acalypha ostryaefolia McLaughlin, S. (1992) Acalypha pringlei McLaughlin, S. (1992) Acamptopappus McLaughlin, S. (1992); UA Rayless goldenhead sphaerocephalus Herbarium Acer glabrum Douglas maple McLaughlin, S. (1992); DBGH Acer grandidentatum Sugar maple McLaughlin, S. (1992); DBGH Acer negundo Ashleaf maple McLaughlin, S.
    [Show full text]
  • US Fish and Wildlife Service
    BARNEBY REED-MUSTARD (S. barnebyi ) CLAY REED-MUSTARD SHRUBBY REED-MUSTARD (S,arguillacea) (S. suffrutescens) .-~ U.S. Fish and Wildlife Service UTAH REED—MUSTARDS: CLAY REED-MUSTARD (SCHOENOCRAMBE ARGILLACEA) BARNEBY REED—MUSTARD (SCHOENOCRAMBE BARNEBYI) SI-IRUBBY REED-MUSTARD (SCHOENOCRAMBE SUFFRUTESCENS) RECOVERY PLAN Prepared by Region 6, U.S. Fish and Wildlife Service Approved: Date: (~19~- Recovery plans delineate reasonable actions which are believed to be required to recover and/or protect the species. Plans are prepared by the U.S. Fish and Wildlife Service, sometimes with the assistance of recovery teams, contractors, State agencies, and others. Objectives will only be attained and funds expended contingent upon appropriations, priorities, and other budgetary constraints. Recovery plans do not necessarily represent the views or the official positions or approvals of any individuals or agencies, other than the U.S. Fish and Wildlife Service, involved in the plan formulation. They represent the official position of the U.S. Fish and Wildlife Service only after they have been signed by the Regional Director or Director as an~roved Approved recovery plans are subject to modification as dictated by new findings, changes in species status, and the completion of recovery tasks. Literature Citation should read as follows: U.S. Fish and Wildlife Service. 1994. Utah reed—mustards: clay reed—mustard (Schoenocrambe argillacea), Barneby reed-mustard (Schoenocrambe barnebyl), shrubby reed—mustard (Schoenacranibe suffrutescens) recovery plan. Denver, Colorado. 22 pp. Additional copies may be purchased from: Fish and Wildlife Reference Service 5430 Grosvenor Lane, Suite 110 Bethesda, Maryland 20814 Telephone: 301/492—6403 or 1—800—582—3421 The fee for the plan varies depending on the number of pages of the plan.
    [Show full text]
  • Nota Lepidopterologica
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Nota lepidopterologica Jahr/Year: 1994 Band/Volume: 17 Autor(en)/Author(s): Riedl Tadeusz Artikel/Article: Une nouvelle espèce européenne du genre Pancalia Stephens (Lepidoptera : Cosmopterigidae, Antequerinae) 87-91 ©Societas Europaea Lepidopterologica; download unter http://www.biodiversitylibrary.org/ und www.zobodat.at Nota lepid. 17 (1/2) : 87-91 ; 30.XI.1994 ISSN 0342-7536 Une nouvelle espèce européenne du genre Pancalia Stephens (Lepidoptera : Cosmopterigidae, Antequerinae) Tadeusz Riedl Witosa 5/5, 80-809 Gdansk, Pologne Summary A new European species of the genus Pancalia Stephens (Lepidoptera, Cosmo- pterigidae, Antequerinae) — Pancalia baldizzonella sp. n. is described from southern Italy. Adults and genitalia of both sexes are illustrated. The new species is most closely related to P. latreillella Curtis and P. nodosella (Bruand), from which it can be distinguished by the wing markings and genitalic cha- racters. Résumé Description de Pancalia baldizzonella sp. n. d'Italie méridionale. Les adultes et les armures génitales des deux sexes de ce taxon sont figurées. Zusammenfassung Pancalia baldizzonella sp. n. wird aus Süditalien beschrieben. Die Imagines und die Genitalien beider Geschlechter werden abgebildet. Die neue Art steht P latreillella Curtis und P nodosella (Bruand) am nächsten, von denen sie sich durch die Flügelzeichnung und Genitalmerkmale unterscheidet. Introduction Depuis la parution d'un article de Gaedike (1967) consacré aux espèces de Pancalia Stephens, 1829, ce genre et ses espèces ont été étudiés à plusieurs réprises (Leraut, 1984 ; Riedl, 1984 ; Sinev, 1985). Ces études ont permis d'expliquer la position systématique du genre en question et de confirmer, après deux révisions, l'existence de 9 espèces nomi- nales habitant uniquement la zone paléarctique, à savoir P.
    [Show full text]
  • MASTER PLANT LIST for WOODLAND WATER-WISE MOW
    MASTER PLANT LIST for WOODLAND WATEWATERR ‐WISE MOMOWW STRIPSTRIPSS Plant species included below are recommended for use in the Woodland Water‐Wise Mow Strips. See individual planting plans for design layouts, site preparation, installation and maintenance tips. SHRUBS COMMON NAME Height Width Exposure Description Botanical Name AUTUMN SAGE 3' 3' sun/part shade Small shrub with showy flowers that attract hummingbirds and beneficial insects. Many color Salvia greggii varieties flowers profusely in the spring and fall BLUE BLOSSOM (N) 3' 3' sun/part shade Best small ceanothus for Central Valley gardens; clusters of dark-violet flowers bloom in spring; Ceanothus maritimus attracts beneficial insects. Little or no pruning 'Valley Violet' required. Drought tolerant. CLEVELAND SAGE (N) 3' 3' sun/part shade Evergreen shrub produces maroon-stemmed, blue-violet flowers in spring; attracts Salvia clevelandii hummingbirds, butterflies and beneficial insects. ''WinnifredWinnifred GilmanGilman'' RemoveRemove ooldld flflowerower stastalkslks iinn summer; prune to maintain compact form. Very drought tolerant. COMPACT OREGON GRAPE 1‐3' 2‐3' part shade/shade Dark, grape-like fruits provide food for native birds; tough plant that tolerates a variety of Mahonia aquifolium garden conditions; attracts beneficial insects 'Compacta' (N) and birds. Drought tolerant. GOODWIN CREEK LAVENDER 3' 3' sun More heat resistant than English lavenders; long springi andd summer blbloom; attracts Lavendula x ginginsii hummingbirds and beneficial insects; cut back 'Goodwin Creek Grey' after flowering; drought tolerant. SPANISH LAVENDER 1.5‐3' 2‐3' sun Showiest of all the lavenders; blooming in spring; cut back to removed old flowers; attracts Lavandula stoechas butterflies and beneficial insects; drought tolerant. RED YUCCA ((N)N) 3‐4' 4 3‐ 44'sun Attractive spiky-lookingpy g leaves; ; blooms all summer long; attracts hummingbirds; very heat Hesperaloe parviflora and drought tolerant.
    [Show full text]
  • Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument
    Schmidt, Drost, Halvorson In Cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument Plant and Vertebrate Vascular U.S. Geological Survey Southwest Biological Science Center 2255 N. Gemini Drive Flagstaff, AZ 86001 Open-File Report 2006-1163 Southwest Biological Science Center Open-File Report 2006-1163 November 2006 U.S. Department of the Interior U.S. Geological Survey National Park Service In cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument By Cecilia A. Schmidt, Charles A. Drost, and William L. Halvorson Open-File Report 2006-1163 November, 2006 USGS Southwest Biological Science Center Sonoran Desert Research Station University of Arizona U.S. Department of the Interior School of Natural Resources U.S. Geological Survey 125 Biological Sciences East National Park Service Tucson, Arizona 85721 U.S. Department of the Interior Dirk Kempthorne, Secretary U.S. Geological Survey Mark Myers, Director U.S. Geological Survey, Reston, Virginia: 2006 Note: This document contains information of a preliminary nature and was prepared primarily for internal use in the U.S. Geological Survey. This information is NOT intended for use in open literature prior to publication by the investigators named unless permission is obtained in writing from the investigators named and from the Station Leader. Suggested Citation Schmidt, C. A., C. A. Drost, and W. L. Halvorson 2006. Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument. USGS Open-File Report 2006-1163.
    [Show full text]
  • Vascular Plants Ventana Double Cone Trail
    CALIFORNIA NATIVE PLANT SOCIETY – VASCULAR PLANTS VENTANA DOUBLE CONE TRAIL Acer macrophyllum - big-leaved maple Epilobium minutum - minute willow-herb Acmispon argophyllus - silver-leaved lotus Ericameria nauseosus var. speciosa - common rabbit-brush Acmispon glaber - deerweed Erigeron petrophilus - rock daisy Acmispon grandiflorus - large-flowered lotus Eriodictyon californicum - yerba santa Acmispon parviflorus - small-flowered lotus Eriogonum fasciculatum var. foliolosum - California buckwheat Adenostoma fasciculatum - chamise Eriogonum nudum var. pubiflorum - naked eriogonum/tibinagua Agoseris grandiflora - large-flowered agoseris Eriogonum saxatile - rock buckwheat Allophyllum gilioides - straggling gilia Eriophyllum confertiflorum - golden yarrow Antirrhinum multiflorum - sticky snapdragon Festuca microstachys - Nuttall's fescue Arbutus menziesii - madrone Festuca myuros - rattail fescue Arceuthobium campylopodum - western dwarf mistletoe Frangula californica - California coffeeberry Arctostaphylos glandulosa - Eastwood's manzanita Galium angustifolium - narrow-leaved bedstraw Boechera breweri - Brewer's rock cress Galium aparine - goose-grass Bromus carinatus var. carinatus - California brome Galium californicum ssp. flaccidum - California bedstraw Bromus diandrus - ripgut grass Galium californicum ssp. luciense - Lucia bedstraw Bromus grandis - tall brome Galium clementis - Santa Lucia bedstraw Bromus laevipes - woodland brome Galium porrigens - climbing bedstraw Bromus madritensis ssp. rubens - red brome Garrya flavescens - ashy
    [Show full text]
  • Field Bindweed in California
    ers, seeds, or seed capsules. The most Field bindweed in California destructive organisms are the seed bee- tles (Bruchidae), especially Sperrnopha- The outlook for biological control gus sericeus (Geoffroy). In Greece, smut caused by the fungus Thecophora se- minis-convolvuli (Desmaz.) Lioro infects Sara S. Rosenthal 0 Lloyd A. Andres 0 Carl B. Huffaker up to 17 percent of the seeds. Stem and root feeders. The remain- ing Mediterranean natural enemies at- tack the stems and roots. The agromyzid fly, Melanagromyza albocilia Hendel, is Biological control is seldom attempt- fornia, we have found a pattern that the most widespread stem borer. The ed against weeds on cultivated lands, may be used to help select the most pyralid moth, Noctuelia floralis because herbicides generally provide suitable European organisms for bio- (Huebner), attacks stems and roots ex- economical, efficient control, and farm- logical control. ternally. The most common root feeder ing practices may interfere with the Our discussion focuses on such spe- is the larva of the flea beetle, Longitar- biological agents. Field bindweed, how- cies and excludes many insects attract- sus pellucidus Foudras (or very near). ever, reproduces from seeds that may ed to the flowers for pollen or nectar as The adult feeds on the leaves. remain alive in the soil for more than 40 well as many crop pests for which field years and from an extensive perennial bindweed is an alternate host. For ex- California plant feeders root system. Control by cultivation or ample, we did not include the spider Foliage feeders. As might be expect- chemicals is difficult, at best.
    [Show full text]
  • Biodiversity and Ecology of Critically Endangered, Rûens Silcrete Renosterveld in the Buffeljagsrivier Area, Swellendam
    Biodiversity and Ecology of Critically Endangered, Rûens Silcrete Renosterveld in the Buffeljagsrivier area, Swellendam by Johannes Philippus Groenewald Thesis presented in fulfilment of the requirements for the degree of Masters in Science in Conservation Ecology in the Faculty of AgriSciences at Stellenbosch University Supervisor: Prof. Michael J. Samways Co-supervisor: Dr. Ruan Veldtman December 2014 Stellenbosch University http://scholar.sun.ac.za Declaration I hereby declare that the work contained in this thesis, for the degree of Master of Science in Conservation Ecology, is my own work that have not been previously published in full or in part at any other University. All work that are not my own, are acknowledge in the thesis. ___________________ Date: ____________ Groenewald J.P. Copyright © 2014 Stellenbosch University All rights reserved ii Stellenbosch University http://scholar.sun.ac.za Acknowledgements Firstly I want to thank my supervisor Prof. M. J. Samways for his guidance and patience through the years and my co-supervisor Dr. R. Veldtman for his help the past few years. This project would not have been possible without the help of Prof. H. Geertsema, who helped me with the identification of the Lepidoptera and other insect caught in the study area. Also want to thank Dr. K. Oberlander for the help with the identification of the Oxalis species found in the study area and Flora Cameron from CREW with the identification of some of the special plants growing in the area. I further express my gratitude to Dr. Odette Curtis from the Overberg Renosterveld Project, who helped with the identification of the rare species found in the study area as well as information about grazing and burning of Renosterveld.
    [Show full text]
  • Beobachtungen an Einer Südosthessischen Population Von Chysoesthia Drurella (Fabricius, 1775) (Lepidoptera: Gelechiidae, Gelechiinae) 181-184 Nachr
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Nachrichten des Entomologischen Vereins Apollo Jahr/Year: 2011 Band/Volume: 32 Autor(en)/Author(s): Weyh Rolf E. Artikel/Article: Beobachtungen an einer südosthessischen Population von Chysoesthia drurella (Fabricius, 1775) (Lepidoptera: Gelechiidae, Gelechiinae) 181-184 Nachr. entomol. Ver. Apollo, N. F. 32 (3/4): 181–184 (2012) 181 Beobachtungen an einer südosthessischen Population von Chysoesthia drurella (Fabricius, 1775) (Lepidoptera: Gelechiidae, Gelechiinae) Rolf E. Weyh Dipl.-Biol. Rolf E. Weyh, Somborner Straße 14, D-63579 Freigericht, Deutschland; [email protected] Zusammenfassung: Beobachtungen zu Larvalbiologie, Ver- Erscheinungszeiten und Generationenzahl puppungsweise, Überwinterungsstadium und Ge ne ra tio- nen zahl sowie zur Verbreitung der in Hessen erstmals nach- Die ersten Minen sowie einzelne fliegende Falter von C. ge wiesenen Palpenmotte Chysoesthia drurella (Fa bri ci us, drurella konnten in den Jahren 2008 bis 2011 jeweils ab 1775) (Gelechiidae) werden beschrieben, Falter und Prä ima- der dritten Maidekade festgestellt werden, das heißt, die ginalstadien farbig abgebildet. Flugzeit der ersten Faltergeneration, die bisher leider nicht direkt beobachtet werden konnte, dürfte im Mai Observations on a population of Chysoesthia drurella (Fabricius, 1775) in southeastern Hesse, Germany stattfinden. In den folgenden etwa 20 Tagen konn ten (Lepido ptera: Gelechiidae, Gelechiinae) sehr selten einzelne Falter gesichtet werden. Abstract: Information on larval biology, pupation, over win- Ab der drit ten Junidekade bis Mitte Juli traten dann te ring and number of generations as well as to distribution wie der in gro ßer Anzahl fliegende und sitzende Falter of the palpmoth Chysoesthia drurella (Fa bri ci us, 1775) auf.
    [Show full text]
  • Sweetpotato Leaf Miner (341)
    Pacific Pests and Pathogens - Fact Sheets https://apps.lucidcentral.org/ppp/ Sweetpotato leaf miner (341) Photo 1. Larval mines of the sweetpotato leafminer, Photo 1. Adult sweetpotato leafminer, Bedellia Bedellia somnulentella. somnulentella. Photo 3. Adult sweetpotato leafminer, Bedellia somnulentella, showing the fringed fore and hindwings. Common Name Sweetpotato leaf miner, morning glory leafminer Scientific Name Bedellia somnulentella; previously known as Bedellia ipomoeae. Distribution Worldwide. Asia, Africa, North America, Europe, Oceania. It is present in Australia, Fiji1, New Zealand, and Papua New Guinea. Hosts Sweet potato, kangkong (Ipomoea aquatica), and bindweeds (Calstegia and Convolvulus species). Symptoms & Life Cycle The damage is done by the larvae. Mines are at first serpentine (snake-like), and filled with frass; they become yellow-brown and later grow in width, forming blotches (Photo 1). At this stage, the larvae protrude from the blotch to defaecate. Later still, holes are produced in the leaves as the blotches are destroyed. The eggs are laid on the leaves, usually on the underside near the midrib, veins or base of the leaf blade. The larvae are yellowish with pink spots along the back, developing into red rounded projections on all segments. When mature, the larvae are 5.5 mm long. They exit the mines and make a network of silk threads on the underside of the leaf. When not feeding, they move around inside the threads, above the leaf surface. Frass often catches in the web of threads. The larvae pupate there. Adults are greyish brown, 3.5-4 mm long, with a tuft of light yellow to brown hairs on the head, and wingspan of 8-10 mm (Photos 2&3).
    [Show full text]