American Upper Cretaceous Echinoidea By C. WYTHE COOKE A SHORTER CONTRIBUTION TO GENERAL GEOLOGY GEOLOGICAL SURVEY PROFESSIONAL PAPER 254-A Descriptions and illustrations of. Upper Cretaceous echinoids from the Coastal Plain and western interior of the Un!·ted States andfrom Guatemala UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1953 UNITED STATES DEPARTMENT OF THE INTERIOR Douglas McKay, Secretary GEOLOGICAL SURVEY W. E. Wrather, Director For sale by the Superintendent of Documents, U. S. Government Printing Office Washington 25, D. C. -Price $1.00 (paper cover) CONTENTS Page Page Abstract __________________________________________ _ 1 Systematic descriptions _______ ---- ____________ ------_ 4 Introduction ______________________________________ _ 1 References----------------------------------------­ 38 Key to the genera of American Upper Cretaceous IndeX--------------------------------------------- 41 Echinoida _______________________________________ _ 4 ILLUSTRATIONS [Plates 1-16 follow index] PLATE 1. Cidaris, Salenia, Porosoma, Codiopsis, and Rachiosoma. 2. Pedinopsis. 3. Lanieria, Rachiosoma, Phymosoma, and Conulus. 4. Globator, Nucleopygns, Lefortia, Faujasia, and Pygurostoma. 5. Catopygus, Phyllobrissus, Clarkiella, and H ardouinia. 6-8. H ardouinia. 9. H olaster and Pseudananchys. 10. Echinocorys, Cardiaster, and H olaster. 11. Cardiaster, I somicraster, and Echinocorys. 12, 13. Hemiaster. 14. Cardiaster, H emiaster, and Linthia. 15. Proraster and Micraster. 16. Hemiaster, Barnumia, and Pygurostoma. Page -FIGURE 1. Stratigraphic location of some echinoid-bearing formations of the Coastal Plain_------------------------------­ 2 IJ',[ AMERICAN UPPER CRETACEOUS ECHINOIDEA By C. WYTHE COOKE ABSTRACT Alabama Museum of Natural History at Tuscaloosa, Most of the 58 species (in 8 genera) of known Upper the Paleontological Research Laboratories at States-. Cretaceous echinoids are found in the Gulf series in the ville, North Carolina, and the University of Alberta at States from Texas to North Carolina. Of six species Edmonton. For this additional material I am indebted that came from the western interior United States only to Dr. Horace G. Richards, Miss Winnie McGlamery, Hemiaster humphreysanus occurs also in the Gulf series. Dr. Richard L. Casanova, and Prof. P. S. Warren. Ten species from New Jersey, Delaware, or Maryland Especial thanks are due to L. W. Stephenson, John have not been recognized elsewhere. Three species B. Reeside, Jr., and William A. Cobban, all of the from Guatemala are included. United States Geological Survey, who contributed Descriptions of the genera herein given Include refer­ important data on the stratigraphic ranges of several ences to the type species and mention .their character­ species. istic features. One new genus, Barnumia, is described. The scope of this report is less inclusive than might Also included are a dichotomous key to the genera, be inferred from the title, which was selected for brevity. keys to the species of Hardouinia and Hemiaster, and a The word American as used here applies primarily to correlation table of echinoid-bearing formations. the United States, though a few of the species described The following species are described as new: Salenia occur also in Mexico, Brazil, Guatemal~, or in Canada. hondoensis, Porosoma reesidei, Codiopsis stephensoni, Only the post-Cenomanian Upper Cretaceous echinoids Rachiosoma hondoensis, Lanieria uvaldana, Conulus are described. This excludes the species from the stephensoni, Globator vaughani, Faujasia chelonium, Upper Cretaceous Buda limestone of Texas, which, Catopygus mississippiensis, Lejortia trojana, Hardouinia though of early Cenomanian age, is classified as part mcglameryae, Pseudananchys stephensoni, Holaster fer­ of t.he Comanche series, most of which is of early alis, Cardiaster deciper, Isomicraster rossi, Isomicraster Cretaceous age. The echinoid fauna of the Buda has danei, Barnumia browni, Hemi.aster arcolensis, Hemiaster been described by Whitney (1916), and its species sabinal, Hemiaster amelianus, and Micraster uddeni. were included in my paper on the Comanche echinoids Transfers of species to a different genus include (Cooke, 1946). Cyphosoma mortoni De Loriol to Rachiosoma; Pseudo­ Also excluded from the report are the Cretaceous diadema hilli Clark to Phymosoma; Echinobrissus echinoids of the Pacific coastal States. These were texanus Clark to Nucleopygus; Clypeaster geomet1·icus described by Kew (1920), who lists the following Morton to Pygurostoma; Echinobrissus cubensis W eisbord Cretaceous species: Cidaris tehemaensis Clark, Oatopy­ to Phyllobrissus; Cassidulus hemisphericus Slocom to gus californicus Kew, Hemiaster cholamensis Kew, Clarkiella; and Cassidulus emmonsi Stephenson, Cassi­ Hemiaster californicus Clark, Hemiaster alamedensis dulus keltumi Stephenson, Cassidulus micrococcus Gabb, Kew, Hemiaster oregonensis Kew, and Epiaster depressus Cassidulus aequoreus Morton, Clypeaster florealis Mor­ Kew. ton, Cassidulus stantoni Clark, and Cassidulus taylori The greater number of species described herein were Warren to Hardouinia. derived from the Gulf series in the states from Texas to North Carolina. Also all the known species from INTRODUCTION the western interior are included. These are Porosoma. This report is based chiefly on collections in the reesidei Cooke, n. sp.; Hardouinia stantoni (Clark); United States National Museum. Besides these, which Hardouinia taylori (Warren); Hemiaster beecheri Clark; include most of the material studied by William Bullock Hemiaster humphreysanus Meek and Hayden; Gardi­ Clark (1891, 1893, 1915) and Lloyd W. Stephenson aster curtus Clark; and Holaster feralis Cooke, n. sp. (1927, 1941), some specimens, including types, were The type of Scutellaster cretaceus Cragin, found at loaned by the Academy of National Sciences of Phila­ Colorado Springs, Colorado, and heretofore supposed delphia, the New Jersey State Museum at Trenton, the to be Cretaceous, is probably a curio from the west 1 2 SHORTER CONTRIBUTIONS TO GENERAL GEOLOGY Coast discarded by some tourist. It is certainly not to the standard Upper Cretaceous time scale of Europe. Cretaceous. It has been identified by J. Wyatt Durham Only the echinoid-bearing formations are listed in that {1953, pp. 147-148) as the common Pliocene species table. For the complete sequence in the Coastal Plain Anorthoscutum interlineatum (Stimpson). the reader is referred to tables by Stephenson, King, Eight species from New Jersey and Delaware are Monroe, and Imlay (1942) and by Monroe (1946); represented for the most part by casts of the interior. for the western interior of the United States to tables They are Pygurostoma geometricum (Morton), Catopygus by Cobban and Reeside (1952). The division of the pusillus Clark, Catopygus ( Oolopygus) williamsi Clark, Maestrichtian into three parts is arbitrary. It signifies Hardouinia florealis (Morton), "Hemiaster" kummeli merely relative position. Clark, Hemiaster welleri Clark, Hemiaster delawarensis The distribution of species according to their geologic Clark, and Cardiaster smocki Clark. These species horizon is shown -in the accompanying table. It will have not been recognized elsewhere, and the state of be noted that the geologic range of most species is preservation of most of them is such as to make recogni­ limited to one stage. For that reason, most echinoids tion uncertain. One species, Cardiaster marylandicus are excellent horizon markers. Clark, comes from the Monmouth formation of Mary­ land. The initials U.S.G.S. followed by a serial number The stratigraphic location of most of the geologic indicates an entry in the register of Mesozoic localities formations from which the American Upper Cretaceous of the Paleontology and Stratigraphy Branch of the echinoids have been derived is indicated in the correla­ United States Geological Survey. The initials U.S.N.M. tion table, Figure 1, in which the formations are referred stand for the United States National Museum. NORTH CAROLINA, TEXAS ARKANSAS MISSISSIPPI, ALABAMA NEW JERSEY SOUTH CAROLINA Q) Escondido formation Kemp clay Providence sand ~ @ ,..:) §' Corsicana marl Prairie Bluff chalk ~ ~ 2~--------------------~ .~ :0 ~ >.. "0 >.. Nacatoch sand Ripley formation Peedee formation Navesink marl ti :E ~ ~-- > ~----------------------r-----------+-----------------~------------~r----------------1 ~ ~ ro >.. Z ro Neylandville marl Saratoga chalk Mount Laurel sand [:4 Marshalltown formation c ro Anacacho limestone and ·a Taylor marl Annona chalk c0 Woodbury clay Q) ~r---~------------------------~----------~----------------~c ro ·a Arcola sandstone member .£ Merchantville clay @ of the Mooreville chalk ~ - Austin chalk :::: ~-------?--------~ .~ (.) C1j Tombigbee sand member ·a of the Eutaw formation 0 u Eagle Ford shale Buda limestone FIGURE I.-Stratigraphic location of some echinoid-bearing formations of the Coastal Plain. AMERICAN UPPER CRETACEOUS ECHINOIDEA 3 Stratigraphic range of American Upper Cretaceous echinoids Maestrichtian Turo­ Coni­ San­ Campa- l----------------~-----­ nian acian tonian nian Lower Middle Upper Cidaris wahalakensis De LorioL _____________________________________________________________ - _----- -------- X C. texanus Clark__________________________________________________________________ X -------- -------- -------- Salenia hondoensis Cooke, n. sp_____________________________ ________ ________ ________ X -------- -------- -------- ~~d~~~~~;~~~~~i 2~~~~~~~-s~--~~=========================== ========