40 5.2.1 Terrestrial Palynomorphs Laevigate Trilete Spores
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5.2.1 Terrestrial Palynomorphs Laevigate trilete spores Genus: Aulisporites LESCHIK 1954 Aulisporites astigmosus (LESCHIK 1956a) KLAUS 1960 1956 Calamospora astigmosa sp. nov . – LESCHIK, p. 22, Plate 2 Fig. 17 1960 Aulisporites astigmosus (Leschik) nov. Comb . – K LAUS , p. 119 - 120, Plate 28, fig. 2. Genus: Calamospora SCHOPF , WILSON & B ENTALL 1944 Calamospora tener (LESCHIK 1955) DE JERSEY 1962 1955 Laevigatisporites tener sp. nov. – LESCHIK , p. 13, Plate 1., Fig. 20. 1955 Punctatisporites flavus – LESCHIK , p. 31, Plate 4, Fig. 2. 1958 Calamospora mesozoicus – COUPER , p. 132, Plate 15/3+4. 1960 Calamospora nathorstii – KLAUS , p. 116, Plate 28, Fig. 1. 1962 Calamaospora tener (L ESCHIK ) n. comb. - de Jersey, p. 3-4, Plate 1, fig 9 –10. 1964 Calamospora tener (LESCHIK 1955) n. comb. – MÄDLER (a), p. 92, Plate 8, Fig. 2. Genus: Cingulizonates DYBOVA & J ACHOWICS 1957 Cingulizonates rhaeticus (R HEINHARDT ) S CHULZ 1967 1962 Cingulatizonates rhaeticus sp. nov. – RHEINHARDT , P. 702, P LATE 2 F IG . 3 1964 Anulatisporites drawehni MAEDLER , P. 177, P LATE 2, F IGS . 1 – 2 1966 Cingulatizonates delicatus ORLOWSKA – Z WOLINSKAP I 014, PLATE 7 FIGS 36 - 38 1967 Cingulatizonates rhaeticus – SCHULZ P. 584, PLATE .13, F IG . 6- 7 Genus: Concavisporites THOMSON & P FLUG 1953 p. 49, Plate 1, Fig. 19 1953 Concavisporites gen. nov. – THOMSON & P FLUG , p. 49. 1959 Toroisporites gen. nov. – KRUTZSCH ,, p. 90. Concavisporites crassexinius NILSSON 1958 p.35, Plate 1, Fig. d 1958 Concavisporites crassexinius sp. nov. – NILSSON , p . 35, Plate 1, Fig. 11. Concavisporites mesozoicus sensu BÓNA Comment: Sporomorphs described by Bóna as C. mesozoicus are comparable to Concavisporites variverrucatus described by COUPER 1958. Concavisporites parvulus sensu BONA The spores, described by Bóna as C. parvulus with a diameter of 20-24 µm are smaller as the other species of this genus. 40 Concavisporites rhaetoliassicus ACHILLES 1981 1981 Concavisporites rhaetoliassicus sp. nov. – ACHILLES , p. 13 –14 , Plate 1, Fig. -13 - 15. Genus: Cornutisporites SCHULZ 1962 p. 310, plate 1, Fig. 7 - 8 Cornutisporites seebergensis SCHULZ 1962 1967 Cornutisporites seebergensis SCHULZ , p. 310, Plate 1, Fig. 7 - 8. Genus: Cyathidites COUPER 1953 p. 27, Plate 2, Fig. 11, 12. Cyathidites australis COUPER 1953 1953 Lygodiosporites adriennis POTONIE & G ELLETICH f. mesozoicus Thiergart; COOKSON , p. 470, Plate 2, Fig. 29. 1953 Cyathidites australis COUPER , p. 27, Plate 2, Fig. 11, 12. 1961 Lygodiumsporites adriennis POTONIE & G ELLETICH ; B OLKOVITINA , p. 104-105, Plate 31, Fig. 3e (illustration of specimen figured by COOKSON 1953). 1961 Cyathidites australis COUPER ; Dev, p. 43, Plate 1, Fig. 1. 1963 Leiotriletes sp. Type A.; SAAD , p. 120, Plate 33, Fig. 1-5. 1963 Cyathidites australis COUPER ; DETTMANN , p. 22, Plate 1, Fig. 1-3. 1965 Cyathidites cf. australis COUPER ; DORING , p. 19, Plate 3, Fig. 1-3. 1966 Cyathidites australis COUPER ; BURGER , p. 237, Plate 5, Fig. 2. 1968 Deltoidospora australis ; CHALONER & MUIR , Plate 16, Fig. a. 1969 Cyathidites australis COUPER ; NORRIS , p. 582, Plate 102, Fig. 1. 2003 Cyathidites australis COUPER ; CHEN & Z HANG , p. 305, Plate 1, Fig. 6-8. 2003 Deltoidospora australis (COUPER ) POCOCK ; QUATTROCCHIO , et al. p. 66, Plate 1, Fig. 1. 2004 Cyathidites australis COUPER ; CHEN & Z HANG , p. 205, Plate 1, Fig. 7, 8. 2006 Deltoidospora australis (COUPER ) POCOCK ; QUATTROCCHIO , et al. p. 594, Fig. 6E. 2006 Cyathidites australis COUPER ; MACPHAIL & CANTRILL , p. 619, Plate 1, Fig. 11. Natural affinity: Filicopsida; various genera and families; Cyatheaceae ( Cyathea ), Dicksoniaceae, Schizaeaceae (Lygodium ). Cycathidites minor COUPER , 1953 1953 Cyathidites australis COUPER , p. 28, Plate 2, Fig. 13 Genus: Deltoidospora MINER 1935 p. 613 plate 24 fig. 7 Deltoidospora sp. sensu BONA Description: Trilete spores, concavely triangular to subcircular; Y mark distinct, rays at least 2/3 radius; exine two-layered, smooth or infrapunctate, with or without exinal folds (kyrtome or less) along the Y mark; 25-80 µm. 41 Genus : Densosporites BERRY 1937 p. 157 Densosporites fissus ( REINHARDT 1964) S CHULZ 1967 1964 Densoisporites fissus sp.nov. REINHARDT p. 54 plate 2 fig 1 – 3 1964 Cingulatizonites rhaeticus MAEDLER 1964 p. 184 plate 2 fig. 18 - 19 1967 Densosporites fissus (R HEINHARDT ) comb. nov. SCHULZ 582, PLATE 12, FIG . 5 - 6 Densoisporites sp. Plate 1, Fig. c Genus : Dictyophyllidites COUPER , 1958 p. 140 plate 21 fig. 5 - 6 Dictyophyllidites harrisii COUPER , 1958 1958 Dictyophyllidites harrisii COUPER , p. 140 plate 21 fig. 5 - 6 Original diagnosis: Trilete, laesurae long, distinct, commissures clearly raised, bordered by a distinct margo; equatorial contour triangular, distal surface markedly convex, proximal less so; exine smooth and comparatively thin. The genus is intended for dispersed spores of the type met within the Jurassic fern Dictyophyllum . Genus: Leiotriletes NAUMOVA 1939 p. 355 emend. POTONIÉ & K REMP 1955 Leiotriletes sp. Genus: Stereisporites THOMSON & P FLUG 1953 p. 53 Stereisporites spp. DE JERSEY & R AINE 1990 1990 Stereisporites sp.; DE JERSEY & RAINE , p. 22, Plate 1, Fig. L. Natural affinity: Bryophyta. Genus Todisporites COUPER 1958 P. 134 Todisporites major COUPER 1958 1958 Todisporites major COUPER , p. 134, Plate 16, Fig. 6-8. 1964 Punctatisporites major (COUPER ) KEDVES & S IMONCSICS , p. 13, Plate 3, Fig. 1, 2. 1966 Todisporites major COUPER ; Helal, p. 86, Plate 31, Fig. 5. 1977 Punctatisporites major (COUPER ) DORHOFER , pp. 20-21, Plate 2, Fig. 10-12 (in part). (includes T. minor COUPER in synonymy) 1978 Todisporites major COUPER ; GUY -OHLSON , pp. 40-41, Plate 6, Fig. 58. 1988 Punctatisporites major (COUPER ) DORHOFER ; PONS , p. 82, Plate 17, Fig.7. 1989 Todisporites cf. major COUPER ; PLANDEROVA , p. 40, Plate 11, Fig. 6. 1992 Puncatisporites sp. cf. P. major (COUPER ) DORHOFER ; BRENNER & Bickoff, p. 164, Plate 5, Fig. 11. 1993 Todisporites major COUPER ; MANDAOKAR , p. 134-135, Plate 2, Fig. 11, 14, 18. 1995 Punctatosporites COUPER ; RAVN , p. 64, Plate 8, Fig. 21. 42 1996 Todisporites major COUPER ; PRAMPARO et al., p. 398. Natural affinity: Filicopsida; Osmundaceae?. Todisporites minor COUPER 1958 p. 135 Plate 16, Fig. 9 -10 See 1955 Laevigatosporites globosus LESCHIK , p. 11, Plate 1, Fig. 5. 1958 Todisporites minor COUPER , p. 135, Plate 16, Fig. 9, 10. 1968 Todisporites sp. A.; JAIN , p. 9, Plate 1, Fig. 5. 1969 Todisporites minor COUPER ; VAGVOLGYI & H ILLS , p. 160, Plate 1, Fig. 13. 1973 Todisporites cf. T. minor COUPER ; STONE , p. 64, Plate 10, Fig. 51. 1974 Todisporites minor COUPER ; RAMANUJAM & S RISAILAM , p. 73, 75, Plate 2, Fig. 11. 1975 Todisporites cf. minor COUPER ; VOLKHEIMER & QUATTROCCHIO , p. 204-205, Plate 1, Fig. 2. 1976 Todisporites minor COUPER ; ROGALSKA , Plate 10, Fig. 153-156. 1977 Punctatisporites globosus (LESCHIK ) LUND , p. 52, Plate 1, Fig. 11. 1977 Punctatisporites major (COUPER ) DORHOFER ,p. 20-21 (in part). 1977 Todisporites minor COUPER ; STAPLETON & B EER , pp. 6-7, Plate 2, Fig. 9. 1985 Todisporites rotundiformis (MALJAVKINA ) POCOCK ; Yu et al., p. 65-66, Plate 12, Fig. 22. 1986 Todisporites minor COUPER ; ASHRAF & E RBEN , p. 128, Plate 2, Fig. 10. 1986 Punctatisporites minor (COUPER ) BRENNER , p. 143, Fig. 6.10. 1086 Todisporites minor COUPER ; Nanjing Institute etc., p. 200, Plate 1, Fig. 18, 39, 40, 46. 1991 Punctatisporites globosus (LESCHIK ) LUND ; DYBKJAER , p. 19, Plate 1, Fig. 8. 1991 Punctatisporites minor COUPER ; ZHANG & ZHAN , p. 76, Plate 3, Fig. 18. 1992 Todisporites cf. T. minor COUPER ; KUMAR , p. 84, Plate 1, Fig. 8. 1992 Todisporites minor COUPER ; ELA & M AHROUS , p. 607, Fig. 5.1. 1998 Todisporites cf. minor COUPER ; SONG , p. 344, Plate 1, Fig. 23, 24. 1998 Todisporites minor COUPER ; SHANG , p. 444, Plate 3, Fig. 2. Natural affinity: Filicopsida; ?Osmundaceae. Genus: Uvaesporites DOERING 1965 P. 39 Uvaesporites argenteaeformis (BOLKOVITINA 1953) SCHULZ 1967 1953 Stenozonatriletes argenteaeformis BOLKOVITINA , p. 51, Plate 7, Fig. 9. 1962 Triletes reissingeri RHEINHARDT , p. 707, plate 2 fig. 1-2 1967 Uvaesporites argenteaeformis (BOLKOVITINA ) SCHULZ , p. 560, Plate 2, Fig. 10, 11. 1997 Uvaesporites argenteaeformis (BOLKOVITINA ) SCHULZ ; ZHANG & GRANT -MACKIE , p. 18, Plate 2, Fig. 7; Plate 4, Fig. 11, 12. 2000 Uvaesporites argenteaeformis (BOLKOVITINA ) SCHULZ ; GAO et al., p. 222, Plate 8, Fig. 16. 2001 Uvaesporites argenteaeformis (BOLKOVITINA ) SCHULZ ; VAJDA , pp. 417, 421, Fig. 10I, 15H. Natural affinity: Lycopsida. Sculptured trilete Spores Genus: Acanthotriletes (Naumova 1939) p. 355 POTONIÉ & K REMP 1954 Acanthotriletes varius NILSSON 1958 page 42 Plate 2, Fig. 10 1958 Acanthotriletes varius sp. nov. – NILSSON , p. 42, Plate 2, Fig. 10. 43 1964 Anemiidites spinosus sp. nov. – MÄDLER (b), p. 180, Plate 2, Fig. 11. Genus: Baculatisporites PFLUG & T HOMSON 1953 p. 56 Baculatisporites sp. 1934 Baculatisporltes primarius WOLFF 1953 PFLUG & THOMSON in THOMSON & PFLUG , 1953, p. 56. Comment: Krutzsch 1967 after having studied spores of all recent species of Osmunda , proposes to maintain Verrucosisporites for Paleozoic spores of this affinity and morphology, and to broaden the diagnosis of Baculatisporites so it will encompass all Mesozoic spores with osmundoid character of ornamentation, which includes ruguiate and baculate sculpture. Similar sculpture can be found, amongst others, in the genera Todea and Lepidopteris of the Osmundaceae . Krutzsch states that at present only few fossil spores can be identified