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Phenological Responses to Climate in the Alberta Native Flora: Herbarium Specimens Reveal Differential Responsiveness Between Species in Mesic and Xeric Habitats
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2019-03-01 Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats Porto, Cassiano Porto, C. (2019). Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats (Unpublished master's thesis). University of Calgary, Calgary, AB. http://hdl.handle.net/1880/109929 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats by Cassiano Porto A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE GRADUATE PROGRAM IN BIOLOGICAL SCIENCES CALGARY, ALBERTA MARCH, 2019 © Cassiano Porto 2019 UNIVERSITY OF CALGARY Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats by Cassiano Porto A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE GRADUATE PROGRAM IN BIOLOGICAL SCIENCES Research Supervisor: Dr. -
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
Native Plants for Conservation, Restoration & Landscaping
ABOUT THE NATIVE PLANTS FOR CONSERVATION, WHAT ARE NATIVES? For more information, refer to field guides and publications RESTORATION AND LANDSCAPING PROJECT Native species evolved within specific regions and dispersed on local natural history for color, shape, height, bloom times This project is a collaboration between the Virginia Depart- throughout their range without known human involvement. and specific wildlife value of the plants that grow in your ment of Conservation and Recreation and the Virginia Native They form the primary component of the living landscape region. Visit a nearby park, natural area preserve, forest or Plant Society. VNPS chapters across the state helped to fund and provide food and shelter for wildlife management area to learn about common plant the 2011 update to this brochure. native animal species. Native associations, spatial groupings and habitat conditions. For The following partners have provided valuable assistance plants co-evolved with specific recommendations and advice about project design, throughout the life of this project: native animals over many consult a landscape or garden design specialist with thousands to millions of experience in native plants. TheNatureConservancy–VirginiaChapter•Virginia years and have formed TechDepartmentofHorticulture•VirginiaDepartmentof complex and interdependent WHAT ARE NON-NATIVE PLANTS? AgricultureandConsumerServices•VirginiaDepartment relationships. Our native Sometimes referred to as “exotic,” “alien,” or “non- of Environmental Quality, Coastal Zone Management fauna depend on native indigenous,” non-native plants are species introduced, Program•VirginiaDepartmentofForestry•Virginia flora to provide food and DepartmentofGameandInlandFisheries•Virginia Native intentionally or accidentally, into a new region by cover. -
Swing Through
Swing Through 20m Swing Through is an interactive agility garden that connects the user to Canada’s diverse landscape, as well as its major economic industry. The garden is a series of thirteen finished lumber posts that dangle from a large steel structure, creating “tree swings”. On the swings are climbing holds where visitors can use the holds to climb up and across the tree swings. Directly under the tree swings are thirteen colour-coordinated stumps that give the user an extra boost, if needed. The thirteen timber tree swings represent Canada’s ten provinces and three territories by using wood from the official provincial and territorial trees. Surrounding this structure of Canadian trees is a garden divided into thirteen sections displaying the native plants of each province and territory. This representative regional plantings encompassing the swings, creating a soft edge. 10m Swing Through allows visitors to touch, smell, and play with the various YT NT NU BC AB SK MB ON QC NL NB PE NS natural elements that make our country so green, prosperous and beautiful. PLAN | 1:75 Yukon Nunavut Alberta Manitoba Quebec New Brunswick Nova Scotia Tree: Subapline fir, Abies lasiocarpa Tree: Balsam Poplar, Populus balsamifera Tree: Lodgepole pine, Pinus contorta Tree: Balsam fir, Abies balsamea Tree: Yellow birch, Betula alleghaniensis Tree: Balsam fir, Abies balsamea Tree: Red spruce, Picea rubens Plants: Epilobium angustifolium, Plants: Saxifraga oppositifolia, Rubus Plants: Rosa acicularis Prunus virginiana, Plants: Pulsatilla ludoviciana, -
Key to Table Abbreviations
Key to Abbreviations Height Soil Type Height of plant, in feet1 - varies from state to state, A Average moisture, soil type soil, growing season, other factors a mixture of loam, humis D Dry soil, average type Cl Clay soil H1 Smallest average height M Moist soil, average type H2 Largest average height W Wet soil, average type .08' = 1" Wa Water's edge or in water .16' = 2" Wd Well-drained soil .25' = 3" .33' = 4" Tolerance .5' = 6" DT Drought Tolerant .66' = 8" WT Wet Tolerant .75' = 9" .83' = 10" Color Spread A All colors, see description Diameter of full grown plant, B Blue in feet, (see above) - Br Brown varies from state to state, soil, G Green growing season, other factors O Orange Determines spacing needs. Pn Pink Spr1 Smallest average spread Pr Purple Spr2 Largest average spread R Red Y Yellow Sunlight W White Amount of sunlight preferred by plant. Usually has range of Attraction sunlight requirements. Follow B Attracts birds requirements carefully. Usually H Attracts hummingbirds Partial Sun plants prefer afternoon Bt Y, attracts butterflies sun, while Partial Shade plants Y1, attracts moths prefer morning sun. GW Y, attracts general wildlife FS Full Sun N, may repel wildlife PS Partial Sun PSh Partial Shade FSh Full Shade Bloomtime Usual time plant blooms, may vary slightly from state to state Bloomtime listed by Month Page 1 TABLE 1 Partial Listing of Kentucky Wildflowers by Common Name See KEY for abbreviation explanation Common Name Scientific Name H1 H2 Spr1 Spr2 S1 S2 Bl1 Bl2 So1 So2 C1 C2 C3 B H Bt GW Aloe, American, or -
THE RED LILY, Lilium Philadelphicum, in SASKATCHEWAN
NATURE LIBRARY PRAIRIE PHOENIX: THE RED LILY, Lilium philadelphicum, IN SASKATCHEWAN Bonnie J. Lawrence and Anna L. Leighton. 2005. Nature Saskatchewan Special Publication No. 25. 139 pp. 24.95 Can. Softcover 21.5 x 25.3 cm, (8.5 x 10 inches). As the culmination of more than a decade of direct study and the accumulation of in excess of a century of anecdotal and scientific literature, the book is an ecological treatise. It takes the reader from details of anatomical structure through phases of growth, and provides information on range, habitats, associated species and interactions with other components of the lily’s ecosystem. It is a treat to learn not only about the hidden secrets of the lily’s life history, but also about the habits of the animal life with which it shares space, both underneath the ground and above it. The photographs, illustrations and figures throughout the book are At first glance, the large format and superb, with the names of many of our plethora of photographs in this book well-known naturalists represented in lead one to believe that it might be what the photo credits, supporting and is commonly referred to as a “coffee enhancing the excellent material table” book, the kind that you can flip obtained by the authors. Is it any through idly as you converse with wonder that we can never tire of friends or watch the TV news. But seeking out the lilies every summer those books favor titillating the visual just to absorb a little of their beauty sense over providing an educational into our own souls? experience and this book definitely does not fit that category. -
Interpretation & Conservation Education Plan
United States Department of Daniel Boone National Forest Agriculture Forest Service Region 8 Daniel Boone DInterpretation & National Forest B October 2008 Conservation Education Plan Completed with assistance from the Center for Design & Interpretation The U.S. Department of Agriculture (USDA) prohibits discrimination in all its programs and activities on the basis of race, color, national origin, age, disability, and where applicable, sex, marital status, familial status, parental status, religion, sexual orientation, genetic information, political beliefs, reprisal, or because all or part of an individual’s income is derived from any public assistance program. (Not all prohibited bases apply to all programs.) Persons with disabilities who require alternative means for communication of program information (Braille, large print, audiotape, etc.) should contact USDA’s TARGET Center at (202) 720-2600 (voice and TDD). To file a complaint of discrimination, write to USDA, Director, Office of Civil Rights, 1400 Independence Avenue, S.W., Washington, D.C. 20250-9410, or call (800) 795-3272 (voice) or (202) 720-6382 (TDD). USDA is an equal opportunity provider and employer. i Acknowledgments The Daniel Boone NF Interpretive and Conservation Education Plan was made possible through the dedicated effort of a Core Team of inspired employees: Kimberly Morgan, Public Affairs Specialist and Core Team Leader Myra Williamson, Recreation Program Manager AMason Miller, Recreation, Engineering, Lands, and Minerals Staff Officer John Kinney, Redbird District -
Renny Grilz Box 382 Aberdeen, SK S0K 0A0 (306)253-‐4219 Seeds@Growwildflowers.Com Blazingstarwildflowers @Growwildflowers
WILD HARVESTING OF NATIVE SEED Renny W. Grilz, P.Ag. Blazing Star Wildflower Seed Company Aberdeen, SK www.GrowWildflowers.com seeds@GrowWildflowers.com @GrowWildflowers BlazingStarWildflowers About Blazing Star Wildflower Seed Company Wild Harvest vs Nursery Knowing the Species How common is the species? Identification – vegetative, flowering, and seed stages Life Cycle – Annual, biennial, perennial Propagation Method – Seeds, rhizomes, stolons, bulblets Knowing the Species Approximate flowering and seed set cycle Habitat types Slope location Sensitivity of the species to harvesting Flowering triggers Response to site management Proper collection and storage techniques Knowing the Species Morgan et al., 1995 Knowing the Species Prairie Crocus – Anemone patens Prairie Crocus Three-‐flowered Avens – Geum triflorum Western Red Lily – Lilium philadelphicum Milkweed – Asclepias sp. Showy Milkweed Dwarf Milkweed Plains Rough Fescue – Festuca altaica hallii Where To Harvest Obtain permits to collect on crown lands Obtain permission to collect on private lands Avoid rare or fragile habitats Highway and road ditches – can be dangerous Avoid land infested with exo�c and weedy species Avoid areas where there is high human traffic Consider the terrain and the capabili�es of your equipment Site Management When To Harvest Collect when sites are least suscep�ble to damage Harves�ng seed that has not fully ripened may result in reduced viability Collect ripe seeds (firm and dry) Seeds are usually ripe 6 to 8 weeks a�er flowering When To Harvest Indicators of ripe seed: – Stem will be brown and dry – Seeds or seed head will begin to dry and change colour as it ripens – Seed strips off easily by hand – Tapping seed heads gently into your palm produces numbers of seeds – Colour -‐ the seeds o�en turning white, black, brown or grey Seeds that are difficult to remove from the plant and are s�ll green are not ripe Harves�ng Na�ve Seed 1. -
Threatened Lilies Reach for the Sky
Threatened Lilies Reach for the Sky I was thinking the other day about an old friend, conservationist Rusty Walton, who passed away last year. To say he was quiet would be an understatement. I could rarely make a smile bend on his face no matter how hard I tried. However, as I think back on the times I spent with him it always makes me smile. He was a dedicated naturalist and an accomplished ecologist. He introduced me to my first wild patch of wood lily on the Vineyard almost ten years ago. Recently I went back to the same spot, and surprisingly the lilies, like Rusty were gone. The wood lily (Lilium philadelphicum) has been drastically reduced on the Vineyard by large populations of deer. Wood lily has a natural distribution range across thirty seven US states. However similar to here on the Vineyard, they are losing ground across North American due to the over population of deer. Frequently I am asked about what it means when a plant goes extinct. On the Vineyard I am talking about “local” extinction, not the complete loss of a species from our planet. The underlying tragedy of local species extinction is the loss of the co-dependent species that have formed a mutualistic (beneficial) relationship with plants. These species include its insect pollinators that use it as a nectar source, and insects that may also feed on parts of the plant or lay their eggs on it to support their lifecycle. The dependence goes further with the birds that feed on those insects. -
Fire in the Southeastern Grasslands, By
Fire in the Southeastern Grasslands RICHARD J. VOGL Department of Biology California State University Los Angeles, CA 90032 INTRODUCTION ~ERE has been more research on the effects of fire in the southeastern United States than in any region of North America. Most studies have been concerned with the effects of fire on the trees, including the role of fire in controlling hardwood suc cession, fire damage to trees, the effects of fire on soils and litter, the influence of fire on conifer growth and reproduction, and the relationships of fire to tree diseases (Garren 1943; Ahlgren and Ahlgren 1960; Cushwa 1968). A lesser, but stilI substantial number of studies have been focused on the effects of fire on forage yields and livestock production (Wahlenberg et al. 1939), and the use of fire in wildlife management in the Southeast. But academic or phy tosociological studies of the vegetational composition and of the effects of fire on the understory vegetation are generally lacking. Except for some range and wildlife research and several general studies (Wells and Shunk 1931; Leukel and St<Jkes 1939; Biswell and Lemon 1943; Burton 1944; Lemon 1949, 1967; Campbell 1955; Biswell1958; Hodgkins 1958; Arata 1959; Cushwa et al. 1966, 1970; Wolters 1972) , most investigators have ignored the herbaceous cover or grassland vegetation under southeastern trees. Even early botanists often became more interested in the unusual botanical features such as the southern extent of Appalachian tree species (Harper 1943, 1952), the description of the silaceous dunes of the 175 RICHARD J. VOGL Gulf Coast (Kurz 1942), the habits of eastern red cedar (Harper 1912), the vegetation of the Okefenokee Swamp (Wright and Wright 1932), or why the Black Belt Prairie of Alabama was treeless (Ranking and Davis 1971), thereby neglecting the widespread and common grassland vegetation and its relationship to fire. -
Cumberland Plateau Geological History
National Park Service U.S. Department of the Interior Big South Fork National River and Recreation Area Oneida, Tennessee Geology and History of the Cumberland Plateau Geological History Rising over 1000 feet above the region around it, the Cumberland Plateau is a large, flat-topped tableland. Deceptively rugged, the Plateau has often acted as a barrier to man and nature’s attempts to overcome it. The Plateau is characterized by rugged terrain, a moderate climate, and abundant rainfall. Although the soils are typically thin and infertile, the area was once covered by a dense hardwood forest equal to that of the Appalachians less than sixty miles to the east. As a landform, this great plateau reaches from north-central Alabama through Tennessee and Kentucky and Pennsylvania to the western New York border. Geographers call this landform the Appalachian Plateau, although it is known by various names as it passes through the differ ent regions. In Tennessee and Kentucky, it is called the Cumberland Plateau. Within this region, the Cumberland River and its tributaries are formed. A view from any over- look quickly confirms that the area is indeed a plateau. The adjoining ridges are all the same height, presenting a flat horizon. The River Systems The Clear Fork River and the New River come together to form the Big South Fork of the Cumberland River, the third largest tributary to the Cumberland. The Big South Fork watershed drains an area of 1382 square Leatherwood Ford in the evening sun miles primarily in Scott, Fentress, and Morgan counties in Tennessee and Wayne and Overlooks McCreary counties in Kentucky. -
Chapter 4 Native Plants for Landscape Use in Kentucky
Chapter 4 Native Plants for Landscape Use In Kentucky A publication of the Louisville Water Company Wellhead Protection Plan, Phase III Source Reduction Grant # X9-96479407-0 Chapter 4 Native Plants for Landscape Use in Kentucky Native Wildflowers and Ferns The U. S. Department of Transportation, (US DOT), has developed a listing of native plants, (ferns, annuals, perennials, shrubs, and trees), that may be used in landscaping in the State of Kentucky. Other agencies have also developed listings of native plants, which have been integrated into the list within this guidebook. While this list is, by no means, a complete report of the native species that may be found in Kentucky, it offers a starting point for additional research, should the homeowner wish to find additional KY native plants for use in a landscape design, or to check if a plant is native to the State. A reference book titled Wildflowers and Ferns of Kentucky, which was recommended by personnel at the Salato Wildlife Center as an excellent reference for native plants, was also used to develop the list. (A full bibliography is listed at the end of this chapter.) While many horticultural and botanical experts may dispute the inclusion of specific plants on the listing, or wish to add more plants, the list represents the latest information available for research, by the amateur, at the time. The information listed within the list was taken at face value, and no judgment calls were made about the suitability of plants for the list. The author makes no claims as to the completeness, accuracy, or timeliness of this list.