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Noorntaztesa AMERICAN MUSEUM PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET NEW YORK, N.Y

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Noorntaztesa AMERICAN MUSEUM PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET NEW YORK, N.Y Noorntaztesa AMERICAN MUSEUM PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET NEW YORK, N.Y. 10024 U.S.A. NUMBER 2673 MARCH 23, 1979 FIREDERICK H. RINDGE A Revision of the North American Moths of the Genus Lomographa (Lepidoptera, Geometridae) AMERICAN MUSEUM No iltates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 2673, pp.1-18, figs. 1-20, maps 1-3 March 23, 1979 A Revision of the North American Moths of the Genus Lomographa (Lepidoptera, Geometridae) FREDERICK H. RINDGE' ABSTRACT The members of this genus have been referred to keys are given for them. Each one is redescribed; the name Bapta for many years; however, the correct both the adults and their genitalia are illustrated. generic assignment is Lomographa Hubner. Both Distributional maps show the known range for each genera have the same type species; Lomographa was species. One change in synonymy is made: virginalis published four years before Bapta. The four North Cassino and Swett is placed as a synonym of American species are revised for the first time, and glomneraria Grote. INTRODUCTION For more than 60 years the North American two generic terms was incorrect. Bapta has members of this genus have been placed under been utilized by such workers as Barnes and the name Bapta. However, Lomographa was McDunnough (1917, p. 111); McDunnough published before Bapta and is the correct ge- (1938, p. 156) and Forbes (1948, p. 274) in neric name. The four North American species North America; Inoue (1956, p. 303) in Japan; have never been revised; their genitalia have Bleszynski (1966, p. 30) in Poland; Silva Cruz neither been described nor figured. In addition, and Gongalves (1977, p. 17) in Portugal; and the relationships of this genus have never been Forster and Wohlfahrt ([1978], p. 200) in Eu- satisfactorily explored. These considerations led rope. Kloet and Hincks (1972, p. 67) gave the me to undertake the present revisionary study. correct usage and synonymy, namely that One of the major problems with this group Bapta is a synonym of Lomographa because has been the proper application of the correct both generic names have the same type species generic name. For many years this assemblage and the former was published four years after in both the Old World and the New World was Lomographa. Now that this typology has been known primarily as Bapta, with Lomographa established it will be necessary to review all being applied to an entirely different group. species described under both these generic Much of this usage probably followed Prout names to establish their proper placement. The (1909, p. 280; 1915, p. 314); unfortunately, he present paper is a revision of the four species did not have the correct type species for either found in North America, north of Mexico. Bapta or Lomographa, and so his use of the Another problem has been the misidentifi- 'Curator, Department of Entomology, the American Museum of Natural History. Copyright (©) The American Museum of Natural History 1979 ISSN 0003-0082 / Price $1.55 2 AMERICAN MUSEUM NOVITATES NO. 2673 cation or incorrect use of other generic names. one another. Very little or nothing is known The North American species have been vari- about the early stages of Palyas and Phry- ously placed, incorrectly, in Corycia, gionis, so comparisons of the immatures with Orthofidonia, Deilinia, Cabera, and Macaria Lomographa are not possible at this time; this as well as in Bapta-but never, until now, in should be done when material becomes avail- Lomographa. Three of our species were de- able. Based on the material at hand, I am scribed from females, with the males not being accepting Ferguson's placement of these genera known until some time later. This, in itself, in the Baptini. complicated the placing of these taxa in the The North American species usually are on proper genus, as males have more characters of the wing in late spring and early summer, with generic value than do females. Barnes and at least some of them being partial or primarily McDunnough (1917, p. 111) were the first to day flying. In the northern portion of their correctly group the four species under a single range, they are apparently basically single generic heading. brooded; in the south it is possible that more Morphologically, the species form a com- than one generation per year is produced. Very pact and easily recognized group, as the adult little seasonal variation is present in the species characters and male genitalia are quite similar of Lomographa. This also applies to individual to one another. This is also true for the female and geographic variability, with the exception genitalia of three of the four species. The three of glomeraria; this problem is discussed under that occur in eastern North America have the that species. papillae anales with slender tapering setae and As the included species are relatively easy to with the apophyses attached anteriorly; the sin- recognize, I am accepting most of the refer- gle western species (elsinora) has larger pa- ences in the literature as being valid. The ma- pillae anales with capitate setae and with the jority of citations refer to geographic apophyses attached medially. The former repre- distribution; these I have listed chronologically sents the plesiomorphic condition, as it is found under Distribution for each species. My distri- in the two Palearctic species (bimaculata and bution maps are based entirely on specimens I temnerata) examined, as well as in practically have examined, and do not include literature all of the other species of Lomographa I have references. dissected from Middle and South America and During the course of this study, I have ex- from New Guinea, and in the members of the amined 1807 specimens (1153 males, 654 fe- two other genera of the Baptini found in North males) and 57 genitalic preparations (30 males, America. Only one other species with ap- 27 females) of North American species. This omorphic female genitalia has been studied in includes the four primary types in North Amer- addition to elsinora; the former occurs in south- ica (in AMNH, MCZ, and USNM). The three em Mexico, and has large papillae anales with in the British Museum (Natural History) were a single row of thick inwardly curving setae on not personally examined; color transparencies each side, and with the apophyses attached me- of them were kindly furnished me by D. S. dially. Fletcher of that institution. In addition, I have The relationships of the Lomographa within studied a number of Neotropical and Old World the Baptini, at least in the New World, are specimens and their genitalia. The majority of poorly known; this is due in large part to an the dissections were prepared by me; in addi- almost complete lack of study of the Neotropi- tion, slide mounts of the antennae and legs cal Ennominae. Ferguson (personal commun.) were made of the North American and two has suggested that Palyas and Phrygionis Palearctic species. All the North American should be placed in the Baptini. Superficially, specimens studied by me at the American Mu- the members of these primarily tropical genera seum of Natural History have had identification do not look like the species of Lomographa; labels placed on their pins (with the exception morphologically the adults are quite similar to of primary types). The majority of specimens 1979 RINDGE: LOMOGRAPHA 3 (1104) and slides (44) are in the collection of ring north of Mexico by the simple antennae of that institution. the males, by the much smaller socius in the The following abbreviations have been used: male genitalia, and by the more elongate fe- AMNH, the American Museum of Natural History male genitalia. CNC, the Canadian National Collection ADULT: Head, eyes large, round; front flat MCZ, Museum of Comparative Zoology, Harvard or slightly swollen, extending beyond eyes for University short distance, dorsal margin rounded or angu- USNM, National Museum of Natural History, late; tongue elongate; palpi short, not quite Smithsonian Institution reaching front; antennae of both sexes simple, with from about 45 to 60 segments. Thorax ACKNOWLEDGMENTS slender; patagia with numerous hairlike scales; I acknowledge with thanks the cooperation fore tibia unarmed, process one-third to one- and aid of the following colleagues who have half length of segment in male, slightly shorter me to study types and specimens in in female; hind tibia with two pairs of elongate allowed spurs, not dilated and without hair pencil in their charge, who have furnished me with spec- male. Abdomen slender, without tufts and imens and photographs, and who have been without row of setae on third segment of male, kind enough to answer many questions: Mr. and eighth segment unmodified. Forewings Andre Blanchard, Houston, Texas; Mr. D. S. for the Department of Entomology, broadly triangular, with 12 veins; with or with- Fletcher out areole; R1 shortly united with Sc, then ei- British Museum (Natural History); Mr. K. B. ther free to costa or shortly united with R2, Bolte for the Canadian National Collection; Mr. R5 from stalk before R3+4; M1 from upper an- Julian P. Donahue for the Natural History Mu- gle; udc and mdc straight, ldc weakly curved seum of Los Angeles County; Dr. D. C. Fergu- anteriorly; Cu1 from below lower angle; fovea son for the National Museum of Natural absent. Hind wings broad; frenulum strong; Sc History, Smithsonian Institution; Dr. Alfred F. approximate to R in basal portion of cell; R Newton, Jr., for the Museum of Comparative and M1 from before upper angle; M3 from Zoology; and Dr.
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