World Catalogue of the Genus Carabus

Boleslav Brezina World Catalogue of the Genus Carabus October 2000 sample issue (best seen in 150% scale)

1 C A R A B U S Linnaeus, 1758

( D i g i t u l a t i ) CARABUS species: 7 LICHNOCARABUS 2 EUCARABUS 6 PARHOMOPTERUS 11 ARCHAEOCARABUS 37 ISIOCARABUS 10 OHOMOPTERUS 11

( L e p i d o s p i n u l a t i ) LIMNOCARABUS 1 EULEPTOCARABUS 1

( L i p a s t r i m o r p h i ) MORPHOCARABUS 20 TRACHYCARABUS 9 MIMOCARABUS 5 CRYPTOCARABUS 11 OPHIOCARABUS 14 CYCLOCARABUS 10 LIPASTER 1

( A r c h i c a r a b o m o r p h i ) ARCHICARABUS 9 ACROCARABUS 2 ISCHNOCARABUS 2 GNATHOCARABUS 1

( S p i n u l a t i ) APOTOMOPTERUS 122

( C r e n o l i m b i ) HEMICARABUS 4 HOMOEOCARABUS 1

( L a t i t a r s i ) NEOCARABUS 1 DIOCARABUS 12 TOMOCARABUS 7 STEPHANOCARABUS 1 SCAMBOCARABUS 6 SEMNOCARABUS 8 ULOCARABUS 2 PACHYSTUS 5 EURYCARABUS 2 NESAEOCARABUS 3 RHIPOCARABUS 1 TACHYPUS 4 MESOCARABUS 4 TANAOCARABUS 3 PACHYCARABUS 5 CAVAZZUTIOCARABUS 1 ORINOCARABUS 10 OREOCARABUS 7 CYTILOCARABUS 2 PIOCARABUS 1 TITANOCARABUS 2 QINLINGOCARABUS 5 HYPSOCARABUS 6 RHIGOIDOCARABUS 1 RHIGOCARABUS 30 AULONOCARABUS 12 LEPTOCARABUS 12 MEGANEBRIUS 21

( A r c i f e r i ) PLATYCARABUS 5 CHAETOCARABUS 2 HETEROCARABUS 1 HYGROCARABUS 1 CTENOCARABUS 2

( P r o c r u s t i m o r p h i ) CECHENOCHILUS 5 MICROTRIBAX 3 TRIBAX 17 NEOPLECTES 2 ARCHIPLECTES 18 MICROPLECTES 2

2 SPHODRISTOCARABUS 17 OXYCARABUS 1 PROCRUSTES 10 LAMPROSTUS 11 CATHOPLIUS 1 MACROTHORAX 3 RELICTOCARABUS 1 APOPLESIUS 4 INIOPACHYS 2 CHRYSOTRIBAX 2 CHRYSOCARABUS 5 PROCERUS 3 MEGODONTUS 18 IMAIBIUS 21 DEROPLECTES 5 AXINOCARABUS 1 GONIOCARABUS 7 EOTRIBAX 11 ALIPASTER 6 CECHENOTRIBAX 1 CRATOCARABUS 5 CRATOCEPHALUS 6 CRATOCECHENUS 3 CRATOPHYRTUS 5 PSEUDOTRIBAX 2 PANTOPHYRTUS 5 LEPTOPLESIUS 7 PSEUDOCRANION 22 SHUNICHIOCARABUS 1 CUPREOCARABUS 10 NEOPLESIUS 27 EOCECHENUS 10 CALOCARABUS 9 CRYPTOCECHENUS 2 ARISTOCARABUS 1 PAGOCARABUS 1 MEGODONTOIDES 4 PSEUDOCOPTOLABRUS 5 IMAIBIODES 1 ACATHAICUS 1 CATHAICUS 1 EUPACHYS 2 FULGENTICARABUS 1 TERATOCARABUS 1 CEPHALORNIS 1 CYCHROSTOMUS 2 ECCOPTOLABRUS 1 LASIOCOPTOLABRUS 1 COPTOLABRUS 14 COPTOLABRODES 1 SHENOCOPTOLABRUS 1 ACOPTOLABRUS 7 DAMASTER 1 subgenera: 115 species: 820

3 C A R A B U S Linnaeus, 1758 subgenera: 115 species: 81820 ( D i g i t u l a t i )

CARABUS Linnaeus, 1758; type species Carabus granulatus Linnaeus =Eutelocarabus Reitter, 1896; type species Carabus arcensis Herbst =Apocarabus Lapouge, 1930; type species Carabus stscheglowi Mannerheim =Nippocarabus Lapouge, 1930; type species Carabus vanvolxemi Putzeys ~Paracarabus Lapouge, 1930, nec Reitter, 1930; type species Carabus granulatus Linnaeus ~Neocarabus Hatch, 1949, nec Lapouge, 1930; type species Carabus granulatus Linnaeus species: 7

1 arcensis Herbst, 1784; cos Europe, Siberia, Far East, Japan ssp. arcensis Herbst, 1784; cos N,C-Europe:Slovenia to Scandinavia; W-Eur.Russia; "Pommern"(=NW-Poland) ~arvensis Paykull, 1790; iss of arcensis Herbst - =pomeranus Gmelin, 1788 "Pommern"(=currently NW-Poland) ~aereus Dejean, 1826, nec Dejean, 1826 "Alps of Steiermark" =seileri Heer, 1837 Switzerland:Schaffhausen ~cupreoaeneus Letzner, 1850; un ~marginatus Letzner, 1850; un ~niger Letzner, 1850; un ~nigrescens Letzner, 1850; un ~purpurescens Letzner, 1850; un ~versicolor Letzner, 1850; un ~viridiaeneus Letzner, 1850; un ~viridis Letzner, 1850; un ~ziegleri Géhin, 1876, nec Duftschmid, 1812; nom.pro aereus Dejean "Alps of Steiermark" ~cuprescens Dalla Torre, 1877; un ~fuscescens Dalla Torre, 1877; un ~virescens Dalla Torre, 1877; un ~nigrescens Westhoff, 1881; un ~nigrinus Westhoff, 1881, nec Motschulsky, 1865 ~aeratus Géhin, 1885, nec Quensel, 1806; nom.pro ziegleri Géhin "Alps of Steiermark" ~ruficrus Géhin, 1885; un ~nigrinopomeranus Rade, 1895; un =austriae Sokolar, 1907 ?Wienerwald ~detritus Lapouge, 1908, nec Drapiez, 1821 N-Germany:? ~simplex Lapouge, 1908; un "S-Germany & Austria" =noricus Sokolar, 1910 Austria:Alps:Wechsel =decipiens Lengerken, 1911 "Bayern" =germaniae Lengerken, 1911 Germany:? =borussicus Csiki, 1927; nom.pro detritus Lapouge N-Germany:? ~baldiai Mandl, 1963; un Austria:Vorarlberg:Hochfreschen, 1900m ~haideri Mandl, 1963; un Kärnten:Schleinitz, Lienzer Hütte env. ssp. conciliator Fischer, 1822 E-Siberia(Nerchinsk), Baykal Lake; N-Mongolia; Altays, Sayans =dahuricus Gebler, 1827 ?"Dauria" =kirbyi Fischer, 1827 Lena River, Taymyr Peninsula =vinculatus Gebler, 1830 Barnaul =kamtchaticus Motschulsky, 1846 Kamchatka(LT) =femoralis Motschulsky, 1865; nec ? "Amur"(LT) =amurensis Géhin, 1885; nom.pro femoralis Motschulsky "Amur"(LT) =concinnus Kraatz, 1886 ?"Amur" =viridiniger Kraatz, 1886 ?"Amur" ~borealis Poppius, 1906, nec Paykull, 1790 lower Lena River:"Shigansk" ~jelanensis Lapouge, 1908; un "Jelan"(=Baykal Lake env.) =poppiusi Csiki, 1927; nom.pro borealis Poppius lower Lena River:"Shigansk" =jurgitae Deuve, 1993 Altays Mts.:Aktash ssp. faldermanni Dejean, 1829 Far East, Korea; Shanxi, Shaanxi & NE-China; "Dauria" =bellus Kraatz, 1886 =nigerrimus Kraatz, 1886 =viridicoeruleus Kraatz, 1886 =provostii Fairmaire, 1888 "Mts.nr.Peking" =praesensum Cavazzuti, 1999 C-Shaanxi:Tongchuan:Shimen Shan, S-slope, 1300-1600m; 1m/1f ssp. hokkaidoensis Lapouge, 1924 Japan:Hokkaido("Teshio"); Kuriles; Sakhalin =sachalinensis Lapouge, 1906 "Sakhalin" ssp. baschkiricus Breuning, 1936 CE-Russia(Kazan’); C-Urals ~laticollis Motschulsky, 1850, nec Sturm, 1826 Baschkiria:Mt.Ilmen(=?Ilmen’ ca.20km NE of Zlatoust) ssp. sylvaticus Dejean, 1826 W-Europe:France(Seine-Marne:Ft.d’Eu), W-Switz., Belgium, Holland, W-Germany, Brit.Isles =schrickelli Dejean, 1826 LNF =alpicola Heer, 1837 ?Switzerland:Mt.Pilatus =anglicus Motschulsky, 1865 Windsore nr.London ~irregularis Beuthin, 1896, nec Fabricius, 1792 Haute-Loire =raetzeri Born, 1898 Switzerland:Neuenburger Jura ~costalis Lapouge, 1908; un France:Cévennes:Gerbier des Jonc =liebmanni Lengerken, 1911 W-France:Vosges =spencei Lengerken, 1911 "Pyrenees"(error) =sugintensis Brunier, 1922 W-France:Haut-Rhin:Sundgau(=area betw.Basel/Mulhouse) =subsylvaticus Perraudin, 1965 ?"Stoccarda" =jarrigei Colas, 1971 Cote d’Or:Chatillon; TNF =thebaudi Colas, 1971 Corréze:Les Monédiéres; TNF

4 ~bicolor (Keith, 1989); un ssp. carpathus Born, 1902; ?non priority Carpathians(C-Romania:Bucegi Mt.R., Mt.Negoi); E-Europe, S-Eur.Russia =?eremita Fischer, 1823; ?priority "Kaukasus"(error) ~romanus Marcu, 1937; un Romania:? =csikensis Csiki, 1942 CE-Romania:Haghimas Mts.nr.Balan =pseudonoricus Kenyery, 1983 Slovenia:"Bachergebirge:N-slope:1km S of Podvelka, 350m"(=ca.betw.Celje/Zagreb) ssp. venetianus Bernau, 1914 NE-Italy:Dolomiti, Veneto(Bosco di Cansiglio); Slovenia 2 deyrollei Gory, 1839 NW-Spain(Galicia:Sierra de Perache); N-Portugal ~luctuosus Géhin, 1885, nec Villers, 1789 N-Portugal:Braga ~chloromorphus Brañes, 1985; un ~cyaneomorphus Brañes, 1985; un ~goryi Brañes, 1985; un ~miniresplendens Brañes, 1985; un ~neotristis Brañes, 1985; un ~pecoudi Brañes, 1985; un ~splendidulus Brañes, 1985; un 3 granulatus Linnaeus, 1758 Europe, Brit.Isles, Balcans, Turkey, Siberia, Far East, Japan; introduced: Canada, USA ssp. granulatus Linnaeus, 1758 C,W,N-Europe, Brit.Isles, Balcans; S-Russia; introd.:SW-Canada:Brit.Columbia; ?LNF =chalybaeus Voet, 1776 =campestris Adams, 1817 Volga River Bank =confluens Fischer, 1827 Volga River =parallelus Faldermann, 1835 Volga River ~niger Letzner, 1850; un ~nigrescens Letzner, 1850; un ~rufofemoratus Letzner, 1850; un ~virescens Letzner, 1850; un ~rubripes Géhin, 1876, nec Duftschmid, 1812 ~cupreoaeneus Dalla Torre, 1877; un ~nigroaeneus Dalla Torre, 1877; un ~viridiaeneus Dalla Torre, 1877; un =forticostis Kraatz, 1878 =haematomerus Kraatz, 1878 =parvicollis Kraatz, 1878 ?"Steiermark" ~fulvipes Géhin, 1885, nec Fabricius, 1792 =wimmeli Schultz, 1900 Hamburg ~hudsonicus Lapouge, 1924, nec Motschulsky, 1865 ?Hudson Bay =granulatulus Mandl, 1955 ~crimeensis Breuning, 1933; un Krym Peninsula:?Jaila Mts. =hibernicus Lindroth, 1956 Ireland:Kerry Co.:Killarney; Hebrides; introduced:Quebec, N.Brunswick, N.Scotia ~atrocyanescens Lamy, 1973; un ssp. duarius Fischer, 1844; nom.pro elongatus Fischer Siberia, Altays, Sayans, Kazakhstan, Kyrgyzstan, N-Mongolia; "Dzhungaria" ~elongatus Fischer, 1842, nec Duftschmid, 1812 "Dzhungaria" =songaricus Motschulsky, 1850 "Des.Kirg.or."(LT) =dauricus Solsky, 1875 Irkutsk & Krasnoyarsk ~solskyanus Géhin, 1885; un =cupriculus Reitter, 1896 Siberia =expansus Lapouge, 1905 Selenga River =lenensis Poppius, 1906 lower Lena River ~luctuosus Lapouge, 1924, nec Villers, 1789 "Tunkun-Sayan"(=Siberia/Mongolia:Tunkunskiye Goltsy Mts.) =luctisonius Csiki, 1927; nom.pro luctuosus Lapouge "Tunkun-Sayan"(=Siberia/Mongolia:Tunkunskiye Goltsy Mts.) ssp. interstitialis Duftschmid, 1812 S-Austria(Sudkärnten); Italy, Corsica, S-Balcans; Turkey:Konya, Denizli =palustris Dejean, 1826 Roma env. =aetolicus Schaum, 1857 C-Greece:"Missolunghi"(=Mesolóngi) =debilicostis Kraatz, 1878 ?Piemonte =miriditus Apfelbeck, 1901 N-Albania:"Velipoja nr.Alessio"(=Lezhë) =paludicola Müller, 1903 Hercegovina:Narenta River nr.Metkovich =corsicus born, 1906 Corsica:Folelli ~ponticus Lapouge, 1924, nec Deyrolle, 1869 Turkey:? =ponti Csiki, 1927; nom.pro ponticus Lapouge Turkey:? ssp. corticalis Motschulsky, 1846, nec Fabricius, 1801; lsu S-Dagestan, NE-Georgia, N-Azerbaydzhan; "Georgia"; "Kachetia:Matany"(LT) =daghestanicus Lapouge, 1924 Dagestan ssp. leander Kraatz, 1878 C-Caucasus:N-macroslope:Teberda to W-Dagestan; ?LNF ssp. yezoensis Bates, 1883 Japan:Hokkaido("Sapporo & Junsai Lake nr.Hakodate"); Sakhalin ~sachalinensis Matsumura, 1911, nec Lapouge, 1906 Sakhalin ~praedator Lapouge, 1924; un "Yesso"(=Hokkaido) =karafutensis Csiki, 1927; nom.pro sachalinensis Matsumura Sakhalin ssp. telluris Bates, 1883 Japan:N,C-Honshu(Tonosawa); NE-China, Shanxi, Far East, Korea =pekinensis Fairmaire, 1887 "Peking" =ussuriensis Born, 1914 Sikhote-Alin Mt.R. ~gobiensis Lapouge, 1924; un "Inn-Shan & Kalgan"(=Hebei:Xianhua) 4 menetriesi Hummel, 1827 C-Europe, NE-Europe, N-Russia, Finland, Byelorussia, Latvia ssp. menetriesi Hummel, 1827 N-Europe, Finland, N-Russia(Novgorod env.), Byelorussia, Latvia; N-Slovakia:Oravská Lesná ssp. pacholei Sokolar, 1911 Austria:Oberöst.(Tanner Moor nr.Königswiesen); S,W,N-Bohemia: Šumava Mts. to Ústí n.Lab. ~bohemicus Tanzer, 1934; un S-Bohemia:"Mt.Schreiner foot:Schneider- & Oberschlag"(=Mt.Bobík:Krejčovice, Milešice) =knabli Mandl, 1951 E-Austria:"Nordtirol:Gem.Breitenwang bei Reutte"(=ca.60km WNW of Innsbruck); 1m =pseudogranulatus Nüssler, 1965 Germany:Sachsen:Gayer:Hermannsdorfer Wiese =witzgalli Reiser in Freude, 1976; ?nud ? 5 sculpturatus Ménétriés, 1832 SE-Azerbaydzhan:Lenkoran’, Talysh Mts.; N-Iran:Gilan, Elborz Mts., Gorgan env. =lenkoranus Reitter, 1896 ?"Lenkoran" =gilanicus Mandl, 1955 Gilan:?betw.Asalam/Heroabad =hyrcanus Heinz, 1977 Mazanderan:30km SE of Beshar 6 stscheglowi Mannerheim, 1827 E-Ukraine, E-Eur.Russia; C,S-Urals(Zlatoust betw.Ufa/Chelyabinsk) =zakharschewskii Motschulsky, 1845 Ukraine:"Charcov:Tschuguev"(=Kharkov:Chuguevka); "Rus.mer.:Svatova Luchka"(LT) 7 vanvolxemi Putzeys, 1875 Japan:Honshu(Chiuzenji nr.Nikko=Tochigi:N of Nikko); Sado Island =decurtatus Lapouge, 1922 Chiuzenji nr.Nikko

5 =noesskei Emden, 1932 Sado Island

LICHNOCARABUS Reitter, 1896; type species Carabus vinctus Weber species: 2

1 limbatus Say, 1825, nec Fabricius, 1776; lsu SE-Canada & NE-USA:Quebec-Virginia-Georgia-Illinois; Maryland =goryi Dejean, 1831 "North America" =clarkei Blumenthal, 1958 North Carolina:Mt.Clingman’s Dome; alpine form 2 vinctus Weber, 1801 SE-Canada & E-USA:Quebec-Indiana-Virginia-Florida; "N-America"(?Ithaca) =ligatus Germar, 1824 "North America" ~interruptus Say, 1825, nec Fabricius, 1775 ~carinatus Dejean, 1826, nec Duftschmid, 1812 Georgia =georgiae Csiki, 1927; nom.pro carinatus Dejean Georgia

EUCARABUS Géhin, 1885; type species Carabus ullrichi Germar =Loxocarabus Reitter, 1896; type species Carabus obsoletus Sturm =Tylocarabus Reitter, 1896; type species Carabus cumanus Fischer =Xystrocarabus Reitter, 1896; type species Carabus catenatus Panzer = Carabus catenulatus Scopoli species: 6

1 catenulatus Scopoli, 1763 Slovenia, Croatia, Bosnia, NE-Italy ssp. catenulatus Scopoli, 1763 Slovenia("Kranj"); Croatia, Bosnia; NE-Italy:Trieste env., SE-Alps; Switzerland:Lugano env. =catenatus Panzer, 1805 "Kranj" =herbsti Dejean, 1826 "Mts.of Croatia"(alpine form) =alternatus Haury, 1855 =subvirescens Motschulsky, 1865 "Idria" ~atropurpureus Dalla Torre, 1877; un ~atrovirescens Dalla Torre, 1877; un ~purpurescens Dalla Torre, 1877; un ~virescens Dalla Torre, 1877; un =korlevici Hoffmann, 1883 Croatia:"Lic plateau" =albanensis Géhin, 1885 ~subvirens Haury, 1885; iss of subvirescens Motschulsky - =troyeri Born, 1911 Croatia:Gospic ~aenescens Depoli, 1929; un =hosti Depoli, 1929 "Fiume env."(Rijeka) =rattii Mandl, 1966 Mt.Palanzone nr.Lago de Como(=ca.20km NE of Como) ssp. fontanellae Reitter, 1896 Croatia:Adriatic Sea coast & islands; "Zara"(=Zadar) =dinaricus Apfelbeck, 1904 Dinara Mts.:Troglav, Kamesnica =mosorensis Müller, 1930 "Mosor"(=nr.Split) 2 cumanus Fischer, 1823 C,W-Caucasus; S-Georgia; S-Eur.Russia:Stavropol; "Desertus Cumanus" =sobrinus Ménétriés, 1832 =submicans Reitter, 1888 "Circassia" =laetulus Reitter, 1896 "Circassia" ~lagodai Lutshnik, 1911; un Terek River 3 italicus Dejean, 1826 N,C-Italy, S-Switzerland ssp. italicus Dejean, 1826 N-Italy(Piemonte), S-Switzerland =halbherri Lapouge, 1898 =ronchettii Born, 1900 Lombardia:Val Sassina ~battonianus Mandl, 1966; un Liguria:Pra:Mt.Riondo ssp. rostagnoi Luigioni, 1904 C-Italy(Latium:Maccarese, Cisterno) ~rostagnonis Seidlitz, 1904; iss of rostagnoi Luigioni - 4 obsoletus Sturm, 1815, nec Rossi, 1790; lsu Carpathians ssp. obsoletus Sturm, 1815, nec Rossi, 1790; lsu Carpathians:Mor., Slov., SE-Poland, N-Hungary, E-Ukr., N,C-Rom.; "Österreich"(=?Moravia) =aureocupreus Reitter, 1880 "Trencsen"(=CW-Slovakia:Trenčín) =csikii Mallasz, 1901 "Kom.Bestercze-Naszod"(=NC-Romania:Bistrita-Nasaud) =prunneri Mallasz, 1901 "Kom.Also-Fehér"(=Alba Iulia) ~mallaszi Lapouge, 1908; un "N-Hungary"(=currently NW-Romania) ~uhligi Lapouge, 1908; un =uhligi Holdhaus, 1910 "Ceahlau, Caliman, Rareul, Nagy-Hagymas"(=Rarau, Haşmaşul Mare) =dudichi Csiki, 1927 ~bicoloratus Fleischer, 1930; un ~mallaszianus Breuning, 1932; un; nom.pro uhligi Holdhaus ~tesari Niedl, 1957; un M-Ukraine:Mukachevo ssp. carpathicus Palliardi, 1825 C,SW-Romania:Fagaraş Mt.R. to Banat; "Transsylvanian Alps" =euchromus Palliardi, 1825 "Banat" ~sacheri Thomson, 1875; un =bielzi Birthler, 1886 C-Romania:"Çibin"(Çindrel Mt.R.) =deubelianus Fleischer, 1911 "Kapellenberg nr.Kronstadt"(=Braşov) ssp. nagyagensis Seidlitz, 1888; nom.pro procerus Bielz W-Romania:Bihor Mts.; Cluj; Hunedoara & Alba:("Nagyag, Zalatna"=Sacarimb, Zlatna) ~carpathicusnagyagensis Birthler, 1886; un "Nagyag, Zalatna"(=Sacarimb, Zlatna) ~procerus Bielz, 1887, nec Reitt.; nom.pro carpathicusnagyagensis Birth. "Nagyag, Zalatna"(=Sacarimb, Zlatna) =fossulifer Fleischer, 1893 "Kom.Bihar:Hadymadfalva" ~cupreus Fleischer, 1930; un ~viridis Fleischer, 1930; un =tippmanni Breuning, 1936 "Zalatna"(=Alba:Zlatna) ~paranagyagensis Lie, 1990; un Arad/Hunedoara:Savirsin, Pojoga 5 parreyssi Palliardi, 1825; ?iss of paraysi Palliardi Croatia, Bosnia; Montenegro ssp. parreyssi Palliardi, 1825; ?iss of paraysi Palliardi Croatia, Bosnia; Montenegro:Volujak, Durmitor Massif ~paraysi Palliardi, 1825; ?cos of parreyssi Palliardi - =gattereri Géhin, 1882 "Steiermark"(error) =purpurifer Reitter, 1885 =dobiaschi Beuthin, 1894 "Bosnia"

6 =padewiethi Csiki, 1906 S-Velebit Mts.:Stirovaca ~ganglbaueri Apfelbeck, 1890, nec Reitter, 1888 S-Bosnia:? =planinaecola Csiki, 1906; nom.pro ganglbaueri Apfelbeck S-Bosnia:? =squamisculptus Reitter, 1908; nom.pro ganglbaueri Apfelbeck S-Bosnia:? ssp. plassensis Born, 1907 Bosnia:Adriatic Sea coast(Plasa); E-Serbia:Frushka Gora =strauchi Born, 1908 N-Bosnia:Bihac env. =tippmannianus Breuning, 1943 E-Serbia:Frushka Gora 6 ullrichi Germar, 1824; ?iss of ulrichii Germar C,E-Europe, Hungary, Ukraine, Romania, Bulgaria, N-Yugoslavia ssp. ullrichi Germar, 1824; ?iss of ulrichii Germar CE-Germany, S-Poland, Bohemia to Slovakia, NE-Hung., Ukraine; N,C-Romania; "Schlesien" ~ulrichii Germar, 1824; ?cos of ullrichi Germar - =leuckarti Petri, 1885 "Siebenburgen:Schässburg"(=Sighişoara) =podolicus Semenov, 1898 "Vinnitza"(=W-Ukraine:Vinnitsa) =comanensis Born, 1902 SE-Romania:Dobrogea lowland:Comana, Braila; NE-Bulgaria:Razgrad =pavlitscheki Born, 1907 "Bukowina" =tigancanus Born, 1908 SE-Romania:Dobrogea lowland:"Tiganca" =intercessor Sokolar, 1912 NC-Hungary:Nógrád:Rétság ~kardaschi Fleischer, 1926; un "Vlarapass"(=Moravia:Vlárský průsmyk) =volhynicus Lutshnik, 1927 "Wolhynia" =planitiae Csiki, 1929 E-Hungary(Alföld region:?Debrecen, ?Gyula) =donovalensis Smetana, 1951 C-Slovakia:Nízké Tatry Mt.R.:Donovaly ssp. rhilensis Kraatz, 1876 Austria, Belg., Holl., Germany, W-Bohemia, Hung., CW-Romania, N-Yug.; Bulgaria(Rila Mts.) ~cupreoaeneus Dalla Torre, 1877; un "Oberösterreich" ~picipes Dalla Torre, 1877; un ~viridiaeneus Dalla Torre, 1877; un =viridulus Kraatz, 1878 ~parvus Géhin, 1885; un "Steiermark" ~stussineri Géhin, 1885; un Slovenia:"Krain" =jaroslawi Fleischer, 1898 "Kom.Bihar" =werneri Reitter, 1899 Bosnia:Vradac =papukensis Koca, 1900 Serbia:Slavonia:Papuk Mts. =pernix Csiki, 1904 =slivensis Apfelbeck, 1904 CE-Bulgaria:Sliven =sokolari Born, 1904 S-Austria:Leitha Mts. =baranyensis Sokolar, 1908 SC-Hungary:"Kom.Baranya" ~germanicus Sokolar, 1908, nec Semenov, 1898 "Germany, Nieder- & Oberösterreich" =alamannicus Csiki, 1927; nom.pro germanicus Sokolar "Germany, Nieder- & Oberösterreich" ~stegmanni Mandl, 1969; un Niederbayern:Straubing ~serbomacedonicus Kenyery, 1970; un S-Serbia:? ~kucajensis Pavićević & Toševski, 1988; un NE-Serbia:Kučaj Mts.:Babina Glava:above Resavska pečina Cave ssp. fastuosus Palliardi, 1825 SW-Romania:Banat(Mehadia) ~cupreonitens Kraatz, 1878, nec Chevrolat, 1861 ~parallelus Kraatz, 1878, nec Faldermann, 1835 "Banat:Nemet Bogsan"(=Bocsa nr.Reşita) ~robustus Kraatz, 1878, nec Deyrolle, 1869 "Banat" ~superbus Kraatz, 1878, nec Schaufuss, 1872 "Banat:Moldowa"(=Moldova Noua) ~viridilimbatus Kraatz, 1878, nec Motschulsky, 1859 "Banat:Nemet Bogsan"(=Bocsa nr.Reşita) =glaucus Haury, 1878 Banat:Bazias ~cyaneus Bodemeyer, 1881, nec Fabricius, 1776 Banat:Reşita ~cuprinus Lapouge, 1902; un Banat:Bazias ~fabrizioi Battoni & Breuning, 1970; un E-Serbia:Negotin ~transdiernae Pavićević & Toševski, 1988; un NE-Serbia:Tekija(=Dunav right bank opposite to Orşova) ~alionidiernae Lie, 1989; ?nud Banat:Alion nr.Orşova ~superboides Lie, 1989; ?nud Banat:"sanatoriul Marilla de linga Oravita"(=Oravita env.) ~alioni Lie, 1990; un; ?nom.pro alionidiernae Lie Banat:Alion nr.Orşova ~ciutagrosi Lie, 1990; un Banat:dealul Ciuta nr.Groşi(=ca.15km NE of Faget) ~feregarii Lie, 1990; un Banat:Feregari/Domogled valley nr.Baile Herculane ~subfastuosus Lie, 1990; un Banat: betw.Cavaran/Tincova(=ca.30km SE of Lugoj) ~subsuperbus Lie, 1990; un; ?nom.pro superboides Lie Banat:"sanatoriul Marilla de linga Oravita"(=Oravita env.) ~topeniae Savulescu, 1992; un Banat:Topenia valley(=NE of Baile Herculane) =subfastuosus Lie, 1996 Banat: betw.Cavaran/Tincova(=ca.30km SE of Lugoj) ~subsuperbussuperboides Lie, 1996; un Banat:"sanatoriul Marilla de linga Oravita"(=Oravita env.) ~umbrosus Lie, 1996; un Banat:Poiana Rusca Mts.:Nadrag & Borlova env. ssp. arrogans Schaum, 1858 NE-Serbia:Majdanpek, Donji Milanovac; "Serbia" ~ceremosnjensis Pavićević & Toševski, 1988; un NE-Serbia:Homolje Mts.:Ceremošnja ~gornjakensis Pavićević & Toševski, 1988; un NE-Serbia:Šetonje:Gornjak monastery env. ~pseudoarrogans Pavićević & Toševski, 1988; un NE-Serbia:Donji Milanovac

PARHOMOPTERUS Lapouge, 1931; type species Carabus sternbergi Roeschke =Orientocarabus Kwon & Lee, 1984; type species Carabus billbergi Mannerheim species: 11

1 angustus Roeschke, 1898 CE-Korea:"Gensan"(=Wonsan); 1f 2 billbergi Mannerheim, 1827 Russia:Far East, Korea, NE-China ssp. billbergi Mannerheim, 1827 Russia:Far East("Dauria"); N-Korea; NE-China:Heilongjiang, Jilin =sedakovi Fischer, 1844 =stschukini Fischer, 1844 =aequatus Motschulsky, 1845 =cupricollis Kraatz, 1886 ~niger Kraatz, 1886, nec Linnaeus, 1761 =pumilus Kraatz, 1886 "Amur River" ~viridicollis Kraatz, 1886, nec Kraatz, 1882 =charbinensis Breuning, 1961 NE-China:Heilongjiang:Harbin ~mandschuricus Breuning, 1961, nec Semenov, 1898 =quelpartensis Breuning, 1961 S-Korea:?"Quelpart Island"(=Jeju-do); ?error =maoershanensis Li, 1992 NE-China:Heilongjiang:Shangzhi:Mao’er Shan

7 =olliveirianus Deuve, 1999 "Heilongjiang:Yichang, Hongxing, Linchang"(=?ca.60km of NE of Yichun:Hongxing) =jilinicola Deuve & Li, 1999 Jilin:"Hunjiang, Linjiang, Mayihe" ssp. coreicola Deuve, 1989 N-Korea:Machon Riong, Tachon env.; ?Pukde-chon; ?Thomak-tog nr.Chongjin ssp. lupisylvaticus Imura, 1993 NC-Korea:Nangnim Mts. 3 cartereti Deuve, 1982 C-Korea ssp. cartereti Deuve, 1982 CE-Korea:C-Taebaek-san Mt.R.:Mt.Sorak Massif:Bi-Sum-Dae ssp. ronkayi Deuve, 1992 CE-Korea:N-Taebaek-san Mt.R.:Kumgang-san Massif:Mt.Mammul-san; 1m ssp. omisanensis Lassalle, 1999 CE-Korea:S-Taebaek-san Mt.R.:Omisan 4 manifestus Kraatz, 1881 NE-China, Korea ssp. manifestus Kraatz, 1881 NE-China:Shanxi, Jilin, Hebei, Beijing; "Mts.of East China" =moupinensis Breuning, 1967 C-China:Sichuan:"Moupin"(=Baoxing); error ssp. guanmenshanus Imura, 1991 NE-China:E-Liaoning:Benxi env.(Guanmen Shan); ?C-Korea:Soul =paeninsularis Deuve, 1992 C-Korea:"Séoul"(=Soul) ssp. pyonganicola Deuve & Li, 1998 N-Korea:Pyongan-pukto:NE of Sinuiju:Uiju:Chunsan-ri 5 mianyangensis Deuve & Li, 1998 NE-China:SE-Liaoning:Gaizhou(=Gai Xian):Mianyangdingzhi Shan 6 namhaedoensis Kwon & Lee, 1984 S-Korea:Namhae-do Island(Mt.Kumsan) 7 nitididorsus Ishikawa & Kim, 1983 S-Korea:S-part ssp. nitididorsus Ishikawa & Kim, 1983 SC-Korea:Kyongsang-namdo:Chirisan Massif("Mt.Jirisan") =chirisanensis Kwon & Lee, 1984 SC-Korea:Mt.Chirisan ssp. paegunsanus Imura, 1997 SE-Korea:Chollanam-doo:E-part:Kwang-yang-gun:Mt.Paegun-san 8 pawlowskianus Deuve, 1989 N-Korea ssp. pawlowskianus Deuve, 1989 NW-Korea:Myohyang-san Mt.R.:Hyangsan-Chon, Sangwon-am; Mt.Chonma-san ssp. peregrinus Cavazzuti & Ratti, 1998 N-Korea:"S.Hamgyong:Puksubaek Shan"(=Hangyong-namdo:Puksubaek-san Massif); 1m 9 sternbergi Roeschke, 1898 Korea, E-China ssp. sternbergi Roeschke, 1898 S,C-Korea("Gensan"=Wonsan) =touzalini Lapouge, 1911 S-Korea:"Quelpart Island"(=Jeju-do) ~chicuanus Breuning, 1928; un "Chi-kuan-shan"(=E-Liaoning:Mts.nr.Fengcheng) =longior Breuning, 1974 C-Korea:"Hpoung-Syou-Ouen" =huchangensis Kwon & Lee, 1984 NC-Korea:Pyongan-pukto:Huch’ang =palgongsanensis Kwon & Lee, 1984 SE-Korea:Kyongsang-pukto:SE of Kumi:Mt.Palgong-san =sobaeksanensis Kwon & Lee, 1984 SE-Korea:Kyongsang-pukto:Mt.Sobaek-san =honamensis Kwon & Lee, 1984 SE-Korea:Cholla-namdo:NW of Sunch’on:Mt.Chogye-san, Mt.Turyun-san ssp. fenghuangshanus Imura, 1991 NE-China:E-Liaoning:Fenghuang Shan nr.Fengcheng ssp. schnelli Lassalle, 1999 S-Korea:Kyongsang-namdo:Songgyesa ssp. lauschanensis Born, 1913 CE-China:Shandong:"Laushan nr.Kiaochow"(=Lao Shan NE of Qingdao) ssp. kojedoensis Kwon & Lee, 1984 S-Korea:Koje-do Island ssp. tongdosanus Kwon & Lee, 1984 SE-Korea:Kyongsang-namdo:Pusan:Seung Lak Mts:"Mt.Yongch’wisan"(=Yangsan N of Pusan) 10 szeli Deuve, 1994 NE-Korea:Hamgyong-pukto:Yonsa(=?Yonsu); 1m 11 xiuyanensis Deuve & Li, 1998 NE-China:SE-Liaoning:Xiuyan Xian:Dayingzhi:Shajinchun:Laoping Ding

ARCHAEOCARABUS Semenov, 1898; type species Carabus relictus Semenov =Acoptopterus Lapouge, 1927; type species Carabus vigilax Bates species: 37

1 agamemnon Breuning, 1943 C-Sichuan:Emei Shan; 1m 2 anhuinus Imura, 1996 SW-Anhui:Dabie Shan:Qianshan Xian(Tianzhu Mts.), Yuexi Xian 3 baimanorum Deuve, 1999 N-Sichuan:Pingwu Xian:Wanbacui, 2500m 4 battoniensis Deuve, 1991 Gansu, Sichuan ssp. battoniensis Deuve, 1991 S-Gansu:"Minchow"(=Min Xian); 1m ssp. nanpingensis Deuve, 1992 N-Sichuan:Nanping Xian, 2600m ssp. ludingensis Deuve & Vigna Taglianti, 1992 W-Sichuan:Luding Xian:Xin Shin 5 bornianus Hauser, 1922 Yunnan:"Tali-sen-fu"(=Dali; LT),"Kuh-tsin-fu, Kouang-hsien, Pe-yen-tsin, Yunnan-fu" ~albrechti: Maindron, 1906, non Morawitz, 1862 - =cecileae Beheim & Breuning, 1943 Yunnan:"Pe-yen-tsin"(=Yanfeng nr.Dayao, betw.Dali/Kunming) 6 camilloi Cavazzuti & Ratti, 1998 NE-Sichuan:Ping ba Zhen, Taiping(=Wanyuan) 7 claricollis Cavazzuti & Ratti, 1998 N-Jiangxi:Lu Shan, 1300m; 1m 8 daiyunshan Kleinfeld, 1998 Fujian:Daiyun Shan:Dehua, 25:41N/118:12E/1500m; 1m 9 dongchuanicus Deuve, 1994 N-Yunnan, S-Sichuan ssp. dongchuanicus Deuve, 1994 NE-Yunnan:Dongchuan ssp. mianshanicus Cavazzuti, 1997 S-Sichuan:Mianshan, 2600-2800m(=ca.50km N of Xichang) ssp. luojicus Cavazzuti, 1998 S-Sichuan:Lunan Shan:Xichang:Luoji, 3200m ssp. meiguensis Imura, 1998 S-Sichuan:Daliang Shan:Meigu Xian:Mt.Dafengding, 28:40N/103:07E; Meigu env. 10 dreuxioides Deuve, 1998 NW-Hunan:Sangzhi; 1m 11 emanuelei Imura, 1997 SW-Hunan:S of Suining:Huangshan 12 fontellus Deuve, 1997 Guizhou; ?NW-Hunan ssp. fontellus Deuve, 1997 "Guizhou"; 1m ssp. semifuscus Cavazzuti & Ratti, 1999 NE-Guizhou:Tongren:betw.Wanshan/Tongren:Chadian ssp. attrepidans Deuve, 2000 ?NW-Hunan:Dayong env.; "Gansu:Wenxian"(probably incorrect) 13 fuscomarginatus Kleinfeld, 1997 Fujian, Guangdong; =?gressittianus ssp. ssp. fuscomarginatus Kleinfeld, 1997 C-Fujian: SE of Yong ’an: Mt.Tianbaoyan, 25:58N/117:22E/1100m ssp. shanminicus Deuve, 1999 "Fujian:Shanmin"; 1m ssp. bambousicola Deuve & Tian, 1999 NC-Guangdong:Ruyuan Xian:Nanling Reserve:Mt.Shikengkong ssp. daanensis Deuve, 1999 Fujian:Wuyi Shan:Da’an 14 koidei Imura, 1996 SW-Sichuan:Muli, 3240m; 3m 15 kucerai Deuve, 1997 S-Shaanxi:Lüeyang; 1f 16 kweitshauensis Mandl, 1975 Guizhou:"Shih-men-kan"; 1f 17 latens Cavazzuti & Ratti, 1998 SW-Hunan:Suijin:Huang Shan, 1400m 18 latipennis Breuning, 1932 Sichuan; 2f 19 mianningensis Kleinfeld, 2000 S-Sichuan:Mianning, 28:33N/102:09 20 morphocaraboides Deuve, 1989 N-Yunnan, S-Sichuan ssp. morphocaraboides Deuve, 1989 NE-Yunnan:Dongchuan env.(Dongchuan City:NW-part:Laoxue He valley, 3200m) ssp. pigmentatus Cavazzuti, 1998 N-Yunnan:Mts.betw.Dayao/Yangpengzi, 3000-3200m ssp. yao Imura, 1999 NE-Yunnan:Qiaojia Xian:Yao Shan, 27:11N/103:01E, 3300-3800m ssp. zhaojueanus Imura, 1999 S-Sichuan:Zhaojue Xian:between Xichang/Meigu; 1f

8 21 nanosomus Hauser, 1931 Sichuan, Guangxi, Guizhou, Hubei ssp. nanosomus Hauser, 1931 SE-Sichuan:"Kin-fu Shan:Sung-Kanho River, 2000m"(=Jinfo Shan; LT) ssp. cordithoracicus Deuve, 1989 NE-Guangxi:N of Guilin:Longsheng Xian:Huaping; Miao’er Shan; Qianjiadong ssp. maotaiophilus Deuve, 1996 SE-Guizhou:Leigong Shan nr.Kaili ssp. wufengensis Kleinfeld, 1998 SW-Hubei:Yizhuxiang:Wufeng, 30:13N/110:04E 22 nestor Breuning, 1934 Sichuan, Hubei, C-Guizhou ssp. nestor Breuning, 1934 SE-Sichuan:"Kin-fu Shan"(=Jinfo Shan); W-Hubei:Hefeng Xian ssp. xianfengicus Deuve & Li, 2000 SW-Hubei:Xianfeng Xian:Laoyan ssp. emeicola Deuve & Imura, 1992 Sichuan:Emei Shan ssp. barbulatus Cavazzuti & Ratti, 1999 C-Guizhou:Mt.Yunwu; 1f 23 niobe Kleinfeld, 2000 W-Yunnan:Xuebang Shan:Lujing River, Lanping, 26:27N/ 99:23E, 4000m 24 oblongior Deuve, 1992 N-Yunnan ssp. oblongior Deuve, 1992 N-Yunnan:Lijiang, 2500-3000m:Ganhaizi Pass ssp. yunlingicus Deuve, 1994 NW-Yunnan:betw.Zhongdian/Weixi:Yunling Shan Mt.R. 25 paris Breuning, 1932 C-Sichuan:"Mts.NW of Ya-chou-fu"(=Ya’an); Wolong env. ~modestus Oberthür, 1884, nec Fabricius, 1801 LNF(illustration only) 26 pseudolatipennis Deuve, 1991 Shaanxi, Gansu, Sichuan, Hubei ssp. pseudolatipennis Deuve, 1991 S-Shaanxi:Long Xian(left Wei He River bank); S-Gansu:Tianshui, ?Zhugqu ssp. wenxianensis Deuve, 1992 S-Gansu:Wenxian env. ssp. bashanensis Imura, 1995 NE-Sichuan:Dabashan Mts.:Chengkou Xian:Bashan env. ssp. shennongjianus Imura, 1996 W-Hubei:Shennongjia env.(Yanzi Pass ca.40km NNE of Mt.Dashennongjia) ssp. qinlingicus Deuve, 1998 S-Shaanxi:Qinling Shan Mts.:15km N of Xunyuangba, 1700m; Zhiuzhi Xian:Houzhenzi ssp. shansutiaensis Lassalle & Remond, 1998 N-Sichuan:"Shansutia, 2700m"(=ca.80kmNW of Pingwu:32:51N/104:11E) ssp. huayangzhen Cavazzuti, 1999 S-Shaanxi:80km N of Yang Xian:Huayang Zhen, 1000-1400m ssp. lantianensis Cavazzuti, 1999 SE-Shaanxi:pass betw. Lantian/Shang Xian, 1300m ssp. yingpanjienus Cavazzuti, 1999 S-Shaanxi:betw.Shagonjie/Yingpanjie, 1500m ssp. yangxianensis Deuve, 1999 S-Shaanxi:Yang Xian ssp. gracilescens Deuve, 1999 N-Sichuan:Pingwu Xian:"Cheutiapao, 2300m" 27 relictus Semenov, 1898 C-Sichuan:"betw.Sing-pu-guan/Tan-chua"(=Tonghua W of Wenchuan); 1f 28 shamaevi Imura, 1996 C-Sichuan:nr.Wolong, 1700-2300m 29 tianbaoshan Kleinfeld, 1998 C-Fujian: SE of Yong ’an :Mt.Tianbaoyan, 25:58N/117:22E/1100m; 1f 30 tieguanzi Imura, 1990 C-Sichuan:Emei Shan 31 vigil Semenov, 1898; nom.pro striatus Semenov Gansu, Sichuan, Hubei, Shaanxi ssp. vigil Semenov, 1898; nom.pro striatus Semenov S-Gansu:"Minchow"(=Min Xian) ~striatus Semenov, 1887, nec DeGeer, 1781 S-Gansu:"Minchow"(=Min Xian) ssp. pseudoparis Deuve, 1992 W-Shaanxi:Long Xian, Taibai Shan Massif ssp. dabashanus Imura, 1995 NE-Sichuan:Dabashan Mts.:Chengkou Xian:Bashan env. ssp. nigrominimus Deuve & Mourzine, 1997 S-Gansu:betw.Wenxian/Wudu:Toqizi, 2300-2500m(=nr.Linjiang) ssp. pseudotosanus Deuve & Mourzine, 1997 S-Gansu:E of Wudu:Wangziguan, 2000m; 5f ssp. pyrochromus Deuve, 1997 S-Gansu:Tianshui env.; 1f ssp. quercinus Cavazzuti, 1999 C-Shaanxi:Tongchuan:Shimen Shan, S-slope, 1300-1600m; 1m/1f ssp. cordulatus Cavazzuti, 1999 S-Shaanxi:80km N of Yang Xian:Huayang Zhen, 1000-1400m ssp. guangwushanus Imura & Su, 2000 NE-Sichuan:Nanjiang Xian:pass betw.Shanliangqu/Taoyuan:Mt.Guangwu Shan ssp. ohshimaorum Deuve, 1991 W-Hubei:Shennongjia ssp. giacomazzoi Deuve, 1992 W-Sichuan:180km SW of Chengdu:Xin Gou 32 vigilax Bates, 1890 C-Sichuan:"Chia-ting-fu"(=Leshan); "Wa-shan"(=Mts.betw.Fulin/Ya’an), Emei Shan 33 wuangxianus Kleinfeld, 1998 E-Sichuan:W-Wuangxian:Zhuitoushi Mt., 30:40N/108:20E 34 wushanling Kleinfeld, 1998 CW-Hubei:Changyang:Wushanling Mt., 30:30N/111:12E 35 xingshanensis Imura, 1996 CW-Hubei:W of Xingshan; ?W-Henan:Lushi env. 36 xupuensis Kleinfeld, 1998 W-Hunan:S of Xupu:Liangfengjie, 27:54N/110:36E 37 yunnanus Fairmaire, 1886 Yunnan, SW-Sichuan ssp. yunnanus Fairmaire, 1886 Yunnan:Dali env.; Lijiang env.:Ganhaizi Pass, Yulongxue Shan ssp. guerryanus Breuning, 1961 Yunnan:"Hang-kia-pin"(=betw.Dali/Lijiang:N of Jizu Shan) ssp. pseudoyunnanus Deuve, 1994 Yunnan:"Ta-pin-tze"(=betw.Dali/Lijiang:N of Jizu Shan); Lijiang env. ssp. yanyuanicus Cavazzuti, 1996 SW-Sichuan:Yanyuan, 2650m ssp. hei Cavazzuti, 1996 SW-Sichuan/NW-Yunnan:Yongning env.:W-bank of Lugu Lake; 1m

ISIOCARABUS Reitter, 1896; type species Carabus fiduciarius Thomson species: 10

1 dargei Deuve, 1987 N-Jiangxi:Lu Shan Mts.(Lu Shan, 1000m) =xingziensis Kleinfeld, 1996 N-Jiangxi:Lu Shan Mts.:ca.29:26N/115:57E(=ca.12km NW of Xingzi) =kulingensis Kleinfeld & Schütze, 1997 N-Jiangxi:"Ku-ling"(=Guling:Lu Shan Mts.); 1m 2 dreuxi Deuve, 1994 Guizhou, Guangxi, Hunan, Guangdong ssp. dreuxi Deuve, 1994 SE-Guizhou:Leigong Shan nr.Kaili ssp. dyseimatus Cavazzuti, 1995 NE-Guizhou:Tongren; 1m/1f ssp. jinxiuensis Imura, 1995 CE-Guangxi:Dayao Shan:Jinxiu Xian:Mt.Lao Shan; SW-Guandong:Mt.Datian Ding =xinyingensis Kleinfeld, 1998 SW-Guangdong:Xinyin:Yunwu Shan, Datian Ding, 22:16N/111:15E; Luoding, 22:46N/111:33E ssp. yangmingensis Deuve, 1995 S-Hunan:E of Youngzhou:Yangming Shan; NE-Guangxi:Guanyang:Qianjiadong ssp. liangshanicus Kleinfeld, 1998 W-Hunan:Liang Shan:Huaihua, 27:33N/109:57E; Xuefeng Shan:Dongkou; Tianzhi Shan ssp. loccaianicolor Deuve & Tian, 1999 E-Guangxi:Nanxiang:Huashuichong Reserve (nr. Hezhou) ssp. guandongicola Deuve & Tian, 2000 NC-Guangdong:Ruyuan Xian:Nanling Reserve:Mt.Shikengkong, 1350-1700m; 1m 3 fiduciarius Thomson, 1856 E-China, Korea ssp. fiduciarius Thomson, 1856 Hubei, Jiangxi, Anhui, Jiangsu; ?Sichuan; "Nordchina" ssp. chusanensis Nicolas, 1924 Zhejiang:Zhousan Archipelago("Chusan"=Zhousan Island) ssp. kirinicus Csiki, 1927; nom.pro auricollis Born N,C-Korea; NE-China:Jilin, ?Liaoning; "Korea" ~auricollis Born, 1922, nec Waterhouse, 1867 "Korea" ssp. saishutoicus Csiki, 1927; nom.pro insularis Lapouge S-Korea:Jeju-do Island; "Quelpart Island"(=Jeju-do) ~insularis Lapouge, 1911, nec Hope, 1837 "Quelpart Island"(=Jeju-do) ssp. tim Kleinfeld, 1999 E-Sichuan:Daba Shan:Wanyuan env., 32:44N/108:03E, 1500m 4 gressittianus Mandl, 1975 Jiangxi/Guangdong/Fujian borders; Hunan ssp. gressittianus Mandl, 1975 Jiangxi/Guangdong/Fujian:"Hong-san"(=Hong Shan, 25:00N/115:50E); Fujian:Daiyun Shan ssp. hunanicola Deuve & Yu, 1992 Hunan:Jiangyong, 27:20N/111:20E; 1f 5 hienfoungi Thomson, 1857 Jiangxi, Anhui, Jiangsu, Shanghai, Zhejiang

9 ssp. hienfoungi Thomson, 1857 Jiangxi, Anhui, Jiangsu, Shanghai; "Nordchina" ssp. chinensis Breuning, 1927 Zhejiang:"Chu-chow"(=Lishui); Jiangxi:"Kiukiang"(=Jiujiang) 6 kiukiangensis Bates, 1888 Jiangxi, Zhejiang, Anhui, Guangxi ssp. kiukiangensis Bates, 1888 Jiangxi("Kiukiang"=Jiujiang), Zhejiang =suensonianus Mandl, 1979 NC-Jiangxi:"Yao-chow"(=Poyang Lake E-coast:Boyang); 1f =heterodynamus Deuve, 1991 Jiangxi:Luofang(=betw.Nanchang/Pingxiang:ca.20kmNE of Xinyu); 1m ssp. alzonai Deuve, 1990 "Sichuan:Ta-zien-lu"(=Kangding; ?error); 1f ssp. tianzhuicus Imura, 1996 SW-Anhui:E-Dabieshan:Qianshan Xian:Tianzhu Mts.; 2f ssp. ronganensis Deuve, 1998 Guangxi:Rong’an Xian:Mt.Yuanbao Shan; 5f ssp. pseudoronganensis Deuve & Tian, 2000 NE-Guangxi:Longsheng Xian; 1f 7 miaorum Lassalle & Prunier, 1993 Guangxi:Longsheng Xian:Miao’er Shan 8 orphniopterus Hauser, 1920; nom.pro castanopterus Hauser Fujian, Zhejiang, Jiangxi ssp. orphniopterus Hauser, 1920; nom.pro castanopterus Hauser Fujian("Yun-ling-shan, Koaleng"=NE of Chong’an; LT); Zhejiang ~castanopterus Hauser, 1920, nec A.& J.B.Villa, 1833 Fujian:"Yun-ling-shan, Koaleng"(=NE of Chong’an; LT) ssp. klapperichi Beheim & Breuning, 1943 Fujian:"Kuatun"; C-Jiangxi:Chongren ssp. fulvoligatus Cavazzuti & Ratti, 1998 C-Fujian:Tianbaoyan(=Mts.SE of Yong’an) 9 pustululatus Deuve, 1993 NE-Hubei:Dabie Shan:Jiuzihe; ?C-Jiangxi:Chongren(?error); "Yunnan"(error) 10 strandiellus Breuning, 1934 Jiangxi("Kien-chang"=Nancheng ca.150km SE of Nanchang); N-Hunan =dayongensis Kleinfeld, 1996 NW-Hunan:NW of Dayong:Zhangjiajie, 29:08N/110:29E =microgranum Cavazzuti & Ratti, 1999 NW-Hunan:Guzhang:Gaowangjie; NE-Guizhou:betw. Wanshan/Tongren:Chadian

OHOMOPTERUS Reitter, 1896; type species Carabus dehanii Chaudoir species: 11

1 albrechti Morawitz, 1862 Hokkaido, Honshu ssp. albrechti Morawitz, 1862 S,C-Hokkaido(Oshima:Hakodate; LT) =multistriatus Motschulsky, 1865 "Japan" ssp. freyi van Emden, 1932 Sado Island ssp. esakianus Nakane, 1961 CE-Honshu:Chiba, Kanagawa, Tochigi; ?LNF ssp. okumurai (Ishikawa, 1966) C-Honshu:Yamanashi(Mt.Daibosatzu), Nagano ssp. tohokuensis (Ishikawa, 1984) N-Honshu:Tohoku district:Fukushima to Aomori(Iwate:Morioka:Matsuzono) ssp. awashimae Ishikawa & Takami, 1996 Niigata:Awashima Island ssp. echigo Ishikawa & Takami, 1996 Honshu:Niigata(Yunotani-mura:Mugurazawa) & adj.regions of Nagano & Fukushima Pref. ssp. hidakanus Ishikawa & Takami, 1996 C-Hokkaido:Sapporo env.to Hidaka Province(Biratori) ssp. itoi Ishikawa & Takami, 1996 NE-Hokkaido:Tokachi(Urahoro:Rushin) to Nemuro:Shiretoko Peninsula 2 daisen (Nakane, 1953) S-Honshu ssp. daisen (Nakane, 1953) SW-Honshu:Tottori:Mt.Daisen ssp. okianus (Nakane, 1961) Shimane:Oki Islands:Dogo Island 3 dehanii Chaudoir, 1848 Honshu, Kyushu, Shikoku ssp. dehanii Chaudoir, 1848 S,C-Honshu; N,C-Kyushu; "Japan" =punctatostriatus Bates, 1873 ~strenuus Breuning, 1932, nec Panzer, 1797 Kyushu:Konose =imafukui (Nakane, 1962) ssp. ishidai Imura & Mizusawa, 1995 Kumamoto:Amakusa-gun:Gosho-no-ura-jima Island:Karasu-ga-togé Pass ssp. katsumai Imura & Mizusawa, 1995 NE-Shikoku:Tokushima(Tokushima-shi:Mt.Bi-zan), Kagawa ssp. koshikicola Imura & Mizusawa, 1995 Kagoshima:Kami-koshiki-jima Island:Mt.Miné-no-yama:E-slope ssp. kumaso Imura & Mizusawa, 1995 S-Kyushu:Kumamoto/Kagoshima/Miyazaki(Ebino-shi:Okobira) borders region ssp. tosanus (Nakane, Iga & Ueno, 1953) S-Shikoku:Ehime, Kochi ssp. ishizuchianus (Nakane, 1960) SW-Shikoku:Ehime:Mt.Ishizuchi ssp. kawanoi (Kamiyoshi & Mizoguchi, 1960) SE-Shikoku:S-Tokushima(Fukuwara:Mt.Tsurugi), Ehime, Kochi =hiraii (Nakane, 1960) ssp. botchan Imura & Mizusawa, 1995 SW-Shikoku:Takanawa-hantó Peninsula(Matsuyama-shi:Mt.Inoko-yama) 4 esakii Csiki, 1927; nom.pro lewisi Géhin & Haury CE-Honshu:Shizuoka, Kanagawa; Yamanashi:S-part; ?Chiba:Sawara ~lewisi Géhin & Haury, 1885, nec Rye, 1872 ?Chiba:Sawara =shizuokaensis (Nakane, 1953) Shizuoka:? 5 insulicola Chaudoir, 1869 Honshu, Sado Island ssp. insulicola Chaudoir, 1869 N,C-Honshu, Sado Island; "Japan" =kaempferi Thomson, 1875 ssp. nishikawai (Ishikawa, 1966) C-Honshu:Chiba:Boso Peninsula:S-part(Awa env.:Mt.Tomisan) ssp. pseudinsulicola (Ishikawa, 1966) C-Honshu:Nagano:upper Tenryu River:left bank(Ina env.:Nosoko) ssp. arrowianus Breuning, 1934 C-Honshu:Aichi, Shizuoka; "Hyogo:Kobe"(LT; error) ~nagoyaensis (Nakane, 1961); un C-Honshu:Aichi:Nagoya ssp. nakamurai (Ishikawa, 1966) C-Honshu:Nagano(Mt.Jimbagatayama) & Shizuoka:middle Tenryu River:left bank ssp. komiyai (Ishikawa, 1966) C-Honshu:Shizuoka:betw.Oi/Tenryu Rivers(Ogasa:Mt.Ogasayama) ssp. murakii Ishikawa & Kubota, 1984 C-Honshu:Mie:N-part of Shima Peninsula:(Watarai-gun:Futami-cho:Sho) ssp. hidaosa Ishikawa & Kubota, 1995 C-Honshu:Gifu:betw.Nagara/Hida/Jintsu Rivers(Takayama-shi:Ohbora-cho) ssp. minoensis Ishikawa & Kubota, 1995 C-Honshu:Gifu:betw.Kiso/Nagara Rivers(Gifu-shi:Mt.Kinkazan) ssp. maiyasanus Bates, 1873 C-Honshu:Hyogo(Kobe:Moon Temple-Maiyasan), Kyoto, Tottori, Fukui, Shiga, Gifu, Mie ssp. hokurikuensis Ishikawa & Kubota, 1995 C-Honshu:Toyama(Asahi-machi:Mt.Shiroyama nr.Miyazaki), Niigata, Ishikawa ssp. yoroensis Ishikawa & Kubota, 1995 C-Honshu:Gifu & Mie:Yoro Mt.R.(Mie:Kuwana-gun:Tado-cho:Mt.Tado-yama) ssp. ohkawai (Nakane, 1968) C-Honshu:Mie:Shima Peninsula(Ugata) ssp. shigaraki (Hiura & Katsura, 1971) C-Honshu:Shiga(Kurita-gun:Mt.Aboshiyama:W-slope); Mie ssp. takiharensis Katsura & Tominaga, 1978 C-Honshu:Kii Peninsula:Nara, Mie(Taki-gun:Ohdai-cho:Shimosuga) ssp. suzukanus Ishikawa & Kubota, 1995 C-Honshu:Mie:SE-foot of Suzuka Mt.R.(Suzuka-shi:Nagasawa-shinden) 6 japonicus Motschulsky, 1857 Kyushu, Shikoku, S-Honshu ssp. japonicus Motschulsky, 1857 Kyushu, Shikoku; "Japan" =corvinus Motschulsky, 1865 "Japan" ~striatus Chaudoir, 1869, nec DeGeer, 1781 "Nordchina"(error) =gisellae Csiki, 1927; nom.pro striatus Chaudoir "Nordchina"(error) ~karatsuanus (Nakane, 1961); un Kyushu:Karatsu ssp. tsushimae Breuning, 1932; emend.pro tsuchimae Breuning, 1932 Nagasaki:Tsushima Island ~tsuchimae Breuning, 1932; un; ios - ssp. chugokuensis (Nakane, 1961) S-Honshu:Shimane(Mt.Takashiro), Hiroshima, Yamaguchi, Okayama, Hyogo ssp. ikiensis (Nakane, 1968) Nagasaki:Iki Island

10 ssp. chotaroi Imura, Dejima & Mizusawa, 1993 Nagasaki:Ikitsuki Island ssp. hiradonis Imura, Dejima & Mizusawa, 1993 Nagasaki:Hirado Island:Mt.Yasuman-dake ssp. onodai Imura, Dejima & Mizusawa, 1993 Kagoshima:Shimo-koshiki Island:Mt.Kuchi-dake ssp. okinoshimanus Imura, Dejima & Mizusawa, 1993 Kochi:Oki-no Island:Moshima, Mt.Imose-yama ssp. awajiensis Imura, Dejima & Mizusawa, 1993 Hyogo:Awaji Island:Mt.Sen-zan ssp. seizaburoi Imura, Dejima & Mizusawa, 1993 Kagawa:Shodo Island:Kankakei valley, Mt.Hoshigajo ssp. nozakicola Imura & Mizusawa, 1994 Nagasaki:Nozaki Island ssp. wakamatsuensis Imura & Mizusawa, 1994 Nagasaki:Wakamatsu Island ssp. mochizukii Imura & Mizusawa, 1994 Ehime:Oshima Island:Mt.Nenbutsu-yama ssp. umekii Imura & Mizusawa, 1999 Yamaguchi:Iwakuni env., Yashiro-jima Isl.(Mt.Kano-zan); Hiroshima: Kami-kamagari-jima Isl. ssp. yoshiyukii Imura & Mizusawa, 1999 Shikoku:Kochi:S of Kochi City:Washio Hills(Mt.Washio-yama) 7 kimurai (Ishikawa, 1966) C-Honshu:Shizuoka(Mt.Ryuzosan nr.Shizuoka City); Yamanashi:S-part 8 lewisianus Breuning, 1932 C-Honshu ssp. lewisianus Breuning, 1932 C-Honshu:Kanagawa(Hakone), Yamanashi, Shizuoka ssp. awakazusanus Ishikawa, 1981 C-Honshu:Chiba:Boso Peninsula:S-part(Awa-gun:Mt.Kiyosumiyama) 9 uenoi (Ishikawa, 1960) C-Honshu:Nara(Yamato env.:Mt.Kongo-zan), Osaka 10 yaconinus Bates, 1873 Honshu, Shikoku ssp. yaconinus Bates, 1873 S-Honshu:Hyogo(LT), Kyoto, Okayama, Hiroshima; Awajishima Island ssp. blairi Breuning, 1934 C-Honshu:Toyama(Fushiki), Ishikawa ssp. cupidicornis Ishikawa & Kubota, 1994 S-Honshu:Mie(Ureshino-cho:Shaden), Nara, Osaka, Wakayama ssp. maetai Ishikawa & Kubota, 1994 S-Honshu:Tottori(Aoya-cho:Ikenotani), Shimane, Hiroshima ssp. oki Ishikawa & Kubota, 1994 Shimane:Oki Islands:Dogo Island(Minamidami:Fuse-mura) ssp. seto Ishikawa & Kubota, 1994 Shikoku:Ehime:Nuwashima Island, Futagamijima Island ssp. sotai Ishikawa & Kubota, 1994 S-Honshu:Kyoto(Kitashirakawa Botanical Garden), Shiga, Mie, Gifu ssp. yamaokai Ishikawa & Kubota, 1994 N-Shikoku(Ehime:Matsuyama:Yoshifuji); Nakajima & Gogoshima Isl.; Honshu:Wakayama ssp. iwawakianus (Nakane, 1953) C-Honshu:Osaka/Nara:Kongo & Izumi Mt.R.); Wakayama; (LNF; cotypes from various loc.) ssp. kiiensis (Nakane & Iga, 1953) C-Honshu:Kii Peninsula:Wakayama, Nara:S of Kinokawa River (LNF; cotypes from Kii Pen.) ssp. muro Ishikawa & Kubota, 1996 C-Honshu:Mie:Muro Mts.(Nabari-shi:Shorenji) ssp. narukawai Ishikawa & Kubota, 1996 C-Honshu(Mie:Kaozaka-cho:Azaka-jinja); Shiga:Suzuka & Nunobiki Mt.R., Hakusan-cho ssp. shima Ishikawa & Kubota, 1996 C-Honshu:Mie:Shima Peninsula:S of Miya River(Toba-shi:Toinoyama:Mt.Asamagatake) 11 yamato (Nakane, 1953) C-Honshu:Nara(Yamato), Osaka, Kyoto, Shiga, Mie, Aichi, ?Gifu ( L e p i d o s p i n u l a t i )

LIMNOCARABUS Géhin, 1876; type species Carabus clathratus Linnaeus species: 1

1 clathratus Linnaeus, 1761 N,C-Europe, Brit.Isles, Balcans, Turkey, Russia, Central Asia, Siberia, Far East, Japan ssp. clathratus Linnaeus, 1761 N,C-Europe, British Isles, Russia, Central Asia, Siberia; LNF(?N-Germany) =aeneopunctatus DeGeer, 1774 =longus Voet, 1778 =adspersus Gmelin, 1788 ~eversmanni Motschulsky, 1846, nec Fischer, 1832 "Ichym" ~detritus Letzner, 1850; un "Silesia" ~cupreus Schilsky, 1888, nec Linnaeus, 1758 N-Germany:Borkum Island, Warnemünde, Zingst =jansoni Kraatz, 1890 Scottland ~multipunctatus Kraatz, 1890, nec Linneaus, 1758 "Westpreussen" ~borealis Born, 1908, nec Paykull, 1790 =foveolatoseriatus Reitter, 1896 Altays =boreus Csiki, 1927; nom.pro borealis Born ~aeratus Culot, 1986; un ssp. stygius Ganglbauer, 1890 Azerbaydzhan:Lenkoran env.; N-Iran ssp. auraniensis Müller, 1902 Austria, S-Moravia, Hung., SE-France; Italy; Balcans(?"Fiume"=Rijeka); Cauc., Armen., Turkey ~laccophilus Reitter, 1896; un "Caucasus:Mahmutli" =arelatensis Lapouge, 1903 SE-France:Camargue, Montpellier =antonellii Luigioni, 1921 C-Italy:Latium:Maccarese,Fiumicino, Roma env. =epirensis Purkyně, 1928 NW-Greece:Ipeíros:"Janina"(=Ioánnina) ssp. maacki Morawitz, 1862 Far East(Hanka Lake):Ussuri, Sikhote-Alin; Korea; Japan:N,C-Honshu =aquatilis Bates, 1883 Japan:Honshu:Shimonosawa Lake(=?Nagano:Suwa-ko Lake) =viridipunctus Kraatz, 1886 Far East:Suyfun River(=W of Vladivostok)

EULEPTOCARABUS Nakane, 1956; type species Carabus porrecticollis Bates species: 1

1 porrecticollis Bates, 1883 Honshu ssp. porrecticollis Bates, 1883 N,CE-Honshu; "Honshu, Urasa"(=Niigata:Minami-Uwonuma Co.:Yamato Town) ssp. kansaiensis Nakane, 1961 S,CW-Honshu; Osaka:Mt.Iwawaki ( L i p a s t r i m o r p h i )

MORPHOCARABUS Géhin, 1885; type species Carabus monilis Fabricius =Ancylocarabus Reitter, 1896; type species Carabus tarbagataicus Kraatz =Apostocarabus Reitter, 1896; type species Carabus odoratus Motschulsky =Leptinocarabus Reitter, 1896; Carabus acutesculptus Chaudoir = Carabus venustus Morawitz =Pancarabus Reitter, 1896; type species Carabus aeruginosus Fischer =Promorphocarabus Reitter, 1896; type species Carabus gebleri Fischer =Gigantocarabus Semenov, 1898; type species Carabus gebleri Fischer =Sajanocarabus Lutshnik, 1924; type species Carabus kozhantschikowi Lutshnik =Amorphocarabus Lapouge, 1930; type species Carabus henningi Fischer =Basilicocarabus Lapouge, 1930; type species Carabus regalis Fischer species: 20

1 aeruginosus Fischer, 1822 E-Eur.Russia, Siberia, Altays, Sayans ssp. aeruginosus Fischer, 1822 E-Eur.Russia, Siberia, Altays("Ridders"=Leninogorsk; ?by error), Sayans

11 =aereus Dejean, 1826 =gmelini Fischer, 1827 =incertus Motschulsky, 1846 "Sib.or."(LT) =subcostatus Motschulsky, 1850 "Sib.or." ~capucinus Géhin, 1885; un =cereus Lapouge, 1912 "Transbaykalia" ssp. hermani Mannerheim, 1827 Urals(Zlatoust), W-Siberia ssp. salechardensis Obydov, 1999 NW-Siberia(Salekhard env.) & NE-Eur.Russia; ?NC-Siberia ssp. eschscholtzi Mannerheim, 1827 Altays, Sayans, N-Mongolia; "Sibiria" ssp. tulatensis O.Berlov & Shilenkov, 1996 Altays:Tigiretsky Mt.R.(Tulata env.) ssp. zyrjanovskianus Shilenkov & O.Berlov, 1996 W-Altays:Bukhtarma River basin(Zyryanovsk) ssp. verzhutzkii O.Berlov & Shilenkov, 1996 NW-Altays:Anuysky & Seminsky Mt.R.:Ch’ornyy Anuy env., Shebalino env. ssp. kolyvanus Shilenkov & O.Berlov, 1996 Russia:NW-Altays:Kolyvansky Mt.R.:Kolyvan env.:Kamenka village 2 chaudoiri Gebler, 1847 C, NE-Siberia, ?N-Mongolia ssp. chaudoiri Gebler, 1847 Mts.nr.Kuznetsk(=Kuznetsky Alatau) & Kan River(=E of Krasnoyarsk); "Altay"(LT) ssp. mongolorum Csiki, 1927; nom.pro neglectus Lapouge, 1913 N-Mongolia; "Tunkun, Sajan"(=Siberia/Mongolia:Tunkunskye Goltsy Mts.; LT) ~neglectus Lapouge, 1913, nec Kraatz, 1887 "Tunkun, Sajan"(=Siberia/Mongolia:Tunkunskye Goltsy Mts.; LT) ssp. czadanicus Obydov, 1997 W-Tuva:Chadan(error; in fact:C-Tuva:Khadyn ca.50km NW of Kyzyl) ssp. marusiki Obydov, 1997 SE-Tuva:Sangilen Mts.:Kargy River valley:50:31N/97:01E ssp. zaikai Obydov, 1999 C-Tuva:Akademika Obrucheva Mt.R.:Sainak Pass, 52:00N/95:34E ssp. shilenkovi O.Berlov, 1989 Tuva to NE-Siberia(Chukot Peninsula:Bilibinsk:Omolon vill.); Yakutsk:Ust’-Yansk 3 excellens Fabricius, 1798 NE-Romania, Moldavia, Ukraine; W-Russia: Kursk env.; Byelorussia =goldeggi Duftschmid, 1812 "Podolia" =erythromerus Dejean, 1826 Moldavia:Bendery =erythrocnemus Fischer, 1827 =erythrodes Fischer, 1827 ~chrysitis Letzner, 1850; un ~polonicus Lomnicki, 1886, nec Motschulsky, 1850 "Galicia:Lubaczow"(=SE-Poland:Lubaczow E of Rzeszów) =frivaldskyi Kraatz, 1887 "Galicia" =lomnitzkii Reitter, 1896; nom.pro polonicus Lomnicki "Galicia:Lubaczow"(=SE-Poland:Lubaczow E of Rzeszów) =sturmi Schilsky, 1899 =moldaviensis Born, 1903 S-Moldavia =jasilkowskii Born, 1905 "Moldavia:Bojan" ~pulchellus Csiki, 1906, nec Brüggemann, 1873 "Tavarnók"(=SW-Slovakia:Tovarníky; error) ~indiconotus Lapouge, 1906; un ~paradoxus Lapouge, 1906; un ~rubicundus Lapouge, 1906; un ~pseudomoldaviensis Hormuzaki, 1907; un NE-Romania:"Jassy"(=Iaşi) =pyretanus Hormuzaki, 1907 N-Moldavia =rareulensis Born, 1907 NE-Romania:Rarau Mts.(alpine form) ~paradoxalis Lapouge, 1909; un =feketei Csiki, 1926; nom.pro pulchellus Csiki "Tavarnók"(=SW-Slovakia:Tovarníky; error) ~botezati Marcu, 1934; un ~striatulus Marcu, 1934; un 4 gebleri Fischer, 1817 Altays Mts.:Zmeinogorsk, Zyryanovsk env. ssp. gebleri Fischer, 1817 Altays Mts.:Zmeinogorsk env. ssp. ultimus Obydov, 1999 Altays Mts.:E-Kazakhstan:Bukhtarma River valley(Zyryanovsk env.) 5 henningi Fischer, 1817 E-Eur.Russia, Urals; C,W-Siberia, Altays, Sayans; ?N-Mongolia ssp. henningi Fischer, 1817 E-Eur.Russia, Urals, W-Siberia(Barnaul), Altays, Sayans =sahlbergi Dejean, 1826 Altays, Sayans, N-Mongolia, Baykal Lake env.; "Siberia" ~ruficornis Kraatz, 1882, nec DeGeer, 1774 Altays =spurius Kraatz, 1882 ~unicolor Kraatz, 1882, nec Fabricius, 1787 =continuus Lapouge, 1906 "Jelan"(=Baykal Lake env.) =roeschkei Csiki, 1901 Yenisey River =sajanicus Csiki, 1901 Sayans =barnaulanus Born, 1922 Barnaul =uralicus Born, 1922 Urals ssp. oviformis Beheim & Breuning, 1943 arctic W-Siberia("Kantaika"=Khantayka SW of Noril’sk) ssp. hamarensis Shilenkov, 1996 Russia:Hamar-Daban Mt.R.:Snezhnaya River valley ssp. melyachi Shilenkov & Menstshikov, 1996 Russia:W-Altays:(Kolyvansky Mt.R.:Kolyvan env.:Kamenka village) ssp. saldaitisi Obydov, 1997 SE-Tuva:Sangilen Mts.:Balyktyk-Khem River source:50:18N/96:24E, 2400m 6 hummeli Fischer, 1823 Siberia; Altays, Sayans, Mongolia, Far East, Korea, NE-China ssp. hummeli Fischer, 1823 C,W,E-Siberia(Nerchinsk); Altays, Sayans, Mongolia =burnaschewi Dejean, 1826 "Siberia" =cyaneoviolaceus Motschulsky, 1844 "Dahuria, Irkutsk" =obversus Motschulsky, 1846 Nerchinsk(LT) =ochoticus Mannerheim, 1852 Far East:"Port Ajan" ~multistriatus Lapouge, 1924, nec Motschulsky, 1865 =densestriatus Csiki, 1927; nom.pro multistriatus Lapouge ssp. middendorfi Ménétriés, 1851 Russia:Far East:Bolshoy Shantar Island ssp. stolidus Lapouge, 1924 ?SE-Siberia:Hamar-Daban Mt.R.; ?Sayans ssp. jurgae Obydov, 1995 Yakutsk:Suntar-Khayata Mt.R.:Khandyga R.(error; in fact: tundra 300km N of Khandyga R.) ssp. nevelskii Shilenkov, 1996 S-Sakhalin(Kril’on Peninsula:Kirilovo); C-Sakhalin ssp. smaragdulus Kraatz, 1878, nec Fabricius; 1798; lsu Russia:Far East(middle Amur River), N-Korea, NE-China:Jilin =chrysothorax Kraatz, 1886 Suyfun River(=W of Vladivostok) ~decoloratus Kraatz, 1886, nec Reitter, 1884 Suyfun River(=W of Vladivostok) =purpuricollis Kraatz, 1886 Suyfun River(=W of Vladivostok) =purpuripennis Kraatz, 1886 "middle Amur River" =subaeneus Kraatz, 1886 Suyfun River(=W of Vladivostok) =subauratus Kraatz, 1886 Suyfun River(=W of Vladivostok) =viridimarginatus Kraatz, 1886 Suyfun River(=W of Vladivostok) ~viridipennis Kraatz, 1886, nec Lewis, 1880 "Amur" =buriatus Lapouge, 1924 "middle Amur River" =amurlandicus Shilenkov, 1994; nom.pro viridimarginatus Kraatz Suyfun River(=W of Vladivostok)

12 ssp. tristiculus Kraatz, 1879 Far East(Suyfun River mouth W of Vladivostok) =gracilentus Kraatz, 1881 "Innshan Mts.W of Peking" ssp. pusongensis Imura, 1993 NC-Korea:Nangnim Mts.:Pusong, Paektu-san Mass.; NE-China:Jilin; Far East:Vladivostok env. =gustavi Shilenkov, 1996 Far East:Vladivostok env.:Chornye Gory Mts.:Kedrovaya Pad’ =wangxingensis Deuve & Li, 1998 E-Jilin:"Wangxing Xian:Daxinggou"(correctly:Wangqing Xian) ssp. suensoni Mandl, 1977 NE-Shanxi:NE of Xin Xian:Wutai Shan Mts.; Fang Shan W of Taiyuan ~fuscipennis Mandl, 1979; un NE-Shanxi:NE of Xin Xian:Wutai Shan Mts. 7 karpinskii Kryzhanovskij & Matveev, 1993 SW-Siberia:South Urals:Mt.Iremel, Yamantau, Zigalga 8 kozhantschikovi Lutshnik, 1924 West Sayans:Kazyr Suk River =martjanowi Lutshnik, 1924 West Sayans:Kazyr Suk River 9 mestscherjakovi Lutshnik, 1924 West Sayans, East Sayans ssp. mestscherjakovi Lutshnik, 1924 West Sayans(Buyba Lake), East Sayans ssp. sambylensis Shilenkov, 1996 West Sayans:Dzhembashsky Mt.R.(Abakan River basin:Sambyl Pass) 10 michailovi Kabak, 1992 NE-Kazakhstan:Sarym-Sakty Mt.R.:45km WSW of Katon-Karagay:E of Ornek 11 monilis Fabricius, 1792 France, Belgium, Holland, Switzerland, Germany(Halle), Brit.Isles, Ireland ~affinis Panzer, 1798, nec Schrank, 1781 Germany:Würzburg =consitus Panzer, 1809 Bern =kronii Hoppe & Hornschuch, 1825 Salzburg(error) & Mühlhausen =amoenus Baudet-Lafarge, 1836 Puy-de-Dome:Mt.Doré =?helveticus Heer, 1837; ?hyb Switzerland:?"Guttannen, Herzogenbuchsee"; ?hyb monilis/cancellatus =schartowi Heer, 1838 "Jura" =gracilis Küster, 1846 "Bayern" ~regularis Wissmann, 1848, nec Fischer, 1823 "Thüringen" =scolecaster Gistl, 1857 "Helvetia" =veris Gistl, 1857 "Germany" =rufofemoratus Motschulsky, 1865 "Helvetia" =taunicus Heyden, 1871 W-Germany:"Mt.Taunus & Vogelsberg"(alpine form) ~femoralis Géhin, 1876, nec Motschulsky, 1865 ~azurescens Dalla Torre, 1877; un ~viridescens Dalla Torre, 1877; un =alticola Bellier, 1880 Alpes-Hte-Provence:Digne env. =anomalus Géhin, 1880 Switzerland:Mt.Pilatus =femoratus Géhin, 1880 =interpositus Géhin, 1880 "Parigi" =rugatinus Géhin, 1880 =sabaudus Géhin, 1880 "Jura" ~nigritulus Kraatz, 1881, nec Kraatz, 1876 "Grande Chartreuse" ~rubricrus Géhin, 1885; un; nom.pro rufofemoratus Motschulsky "Helvetia" ~trilineatus Haller, 1885, nec Fabricius, 1792 "Schweizer Jura" ~varicolor Joerin-Gerber, 1888; un "Jura" =bavaricus Beuthin, 1896 Bayern:Lechfeld ~interruptus Beuthin, 1896, nec Fabricius, 1775 =moestulus Beuthin, 1896 =sengstacki Beuthin, 1896 =?subschartowi Lapouge, 1898; ?hyb Haute-Garonne:Luchon; ?hyb monilis/cancellatus =meridionalis Born, 1905 S-France ~anglicus Lapouge, 1906; un England:? ~externus Lapouge, 1906; un ~gaultieri Lapouge, 1906; un Loire-Atlantique:Nantes =tugenus Lapouge, 1906 "Kanton Appenzell" =perreti Born, 1907 Isere:Notre-Dame-la-Salette(alpine form) ~arvernus Barthe, 1908; un Puy-de-Dome:Mt.Doré ~insularis Born, 1908, nec Hope, 1837 England:? ~bavariensis Lapouge, 1909; iss of bavaricus Beuthin - =pustulatus Lapouge, 1909 Puy-de-Dome =tigurinus Lapouge, 1909 ~kellneri Hubenthal, 1910; un "Thüringen" =nicodii Pic, 1910 =rhodanicus Lapouge, 1916 S-France:? =sequanus Lapouge, 1916 Cote-d’Or:Brazay en Morvan =succisus Lapouge, 1916 France:St.Gaudens =subpyrenaeus Barthe, 1924 Haute-Garonne:St.Gaudens, Col d’Aspin ~charbonieri Sirguey, 1931; un Puy-de-Dome:Mt.Doré ~cylindricus Sirguey, 1931, nec Herbst, 1784 Puy-de-Dome ~cylindricusniger Sirguey, 1931; un Puy-de-Dome ~cylindricusviolaceus Sirguey, 1931; un Puy-de-Dome ~femoratusviolaceus Sirguey, 1931; un =incostatus Sirguey, 1931 Puy-de-Dome ~incostatusfemoralis Sirguey, 1931; un Puy-de-Dome ~incostatusniger Sirguey, 1931; un Puy-de-Dome ~incostatusviolaceus Sirguey, 1931; un Puy-de-Dome ~interposituscyaneus Sirguey, 1931; un ~interpositusniger Sirguey, 1931; un ~interpositusviolaceus Sirguey, 1931; un =lavagnei Sirguey, 1931 Puy-de-Dome:Mt.Doré ~marcoti Sirguey, 1931; un ~pustulatusniger Sirguey, 1931; un ?Puy-de-Dome ~pustulatusviolaceus Sirguey, 1931; un ?Puy-de-Dome ~rhodanicusviolaceus Sirguey, 1931; un ~sequanusniger Sirguey, 1931; un S-France:? ~sequanusviolaceus Sirguey, 1931; un S-France:? ~theroudi Sirguey, 1931; un =theryi Sirguey, 1931 ~trapeti Sirguey, 1931, nec Bleuse, 1885 Puy-de-Dome:Mt.Doré ~trilineatusniger Sirguey, 1931; un ?"Schweizer Jura"

13 ~trilineatusviolaceus Sirguey, 1931; un ?"Schweizer Jura" ~saouensis Mandl, 1967; un Drôme:Ft.de Saou =daniellae (Bellon & Tarrier, 1972) ?Alpes-Maritimes:Col de Champs =ventusicus (Bellon & Tarrier, 1972) =montichares (Hansen & Gaskowiak, 1973) Ariége:Pailhéres =dupeuxi Deuve, 1974 Charente:Foret de Braconne =benoiti (Devecis, 1975) Corréze:Albussac ~meneensis Grotz & Grotz, 1975; un Drôme:Col de Menée =norensis Pham, 1977 Aude:Pic de Nore =rosayanus Dufour, 1978 Cote-d’Or:Rosay =fagnensis Grotz, 1978 ~nepos Ivanovs, 1981; un N-Germany:Niedersachsen:Verden an der Aller(=SE of Bremen); =?excellens Fabricius =alpigradus Sermet, 1985 ~vincentianus Devecis, 1991; un Cantal:Col d’Aulac 12 odoratus Motschulsky, 1844; ?non priority Siberia, Altays, Sayans; ?N-Mongolia ssp. odoratus Motschulsky, 1844 Russia:Baykal Lake:SE-coast; Hamar-Daban Mts.(LT) ~vulneratus Lapouge, 1915; un =batesianus Deuve, 1995 "Baikal"; 1m ssp. viridilimbatus Motschulsky, 1845 SE-Siberia & Far East:Barguzin Mt.R., Amur; CE-Siberia:S of Yakutsk:Amga; "Amur bor."(LT) ssp. irkoutskensis Lapouge, 1915 Irkutsk env. ssp. dohrni Gebler, 1847; ?non priority C-Altays; "Siberia, Altay"(LT) =?microchondrus Fischer, 1842; inc; ?priority "Songoria:Kreis Ajagus"(=?Ayaguz 300 km S of Semipalatinsk) ~interstitialis Motschulsky, 1845; nud; nom.pro alysidotus: Fischer - (Siberia:Irtysh River; invalid LT) ~irkoutskensis Géhin, 1885; nud; nom.pro interstitialis Motschulsky - ssp. baeri Ménétriés, 1851 NC,NE-Siberia:Yenisey to Kolyma River; "Boganida"(=S-part of Taymyr Peninsula) =tolli Poppius, 1910 Chara-Ullah Mts.betw.Lena/Jana River(=Verkhoyansky Mt.R.) ssp. chaffanjoni Lesne, 1897 arctic E-Siberia:Bulun nr.Lena River mouth ssp. melleus Lapouge, 1909; ?un East Sayans:Mondy; ?NC-Mongolia ssp. septentrionalis Breuning, 1932 NW,NC-Siberia:arctic Urals("Obdorsk"=Salekhard) to Yenisey River ssp. kamtschatensis Breuning, 1942 NE-Siberia:Kamchatka:Penzhinskaya Guba:Shestakovo ssp. altyntuanus Shilenkov, 1996 Russia:E-Altays:Chulyshman River:Mt.Altyntu, ?Mt.Korumbu; ?alpine form ssp. antropovi Shilenkov, 1996 Russia:East & West Tannu-Ola Mt.R.(East Tannu-Ola:Shuurmak env.) ssp. bargusinicus Shilenkov, 1996 Russia:Barguzinsky Mt.R.(Davshe River valley) ssp. dabanensis Shilenkov, 1996 Russia:Irkutsk:Hamar-Daban Mt.R.:S-slope(Snezhnaya River valley) ssp. shaman Shilenkov, 1996 Russia:West Sayans:Mt.Shaman ssp. telezkianus Shilenkov, 1996 Russia:E-Altays:Teletskoe Lake env.:(SW-bank:Mt.Altyntu); Chulyshman Mts. ssp. krugeri Obydov, 1999 C-Tuva:Akad.Obrucheva Mt.R.:Koptu Pass, 51:56N/95:32E; Sainak P., 52:00N/95:34E ssp. magadanicus Obydov, 1999 E-Siberia:Magadan env.(Magadan) ssp. martjanovianus Obydov, 1999 Russia:West Sayans:Sayanogorsk 13 regalis Fischer, 1822 E-Eur.Russia; C,W-Siberia, Altays, Sayans; N-Mongolia ssp. regalis Fischer, 1822 E-Eur.Russia; W-Siberia(Barnaul) =cuprinus Fischer, 1822 =cyanicollis Dejean, 1826 =pasianax Fischer, 1827 =interstinctus Motschulsky, 1865 Irtysh River ~viridicinctus Kraatz, 1882, nec Schaufuss, 1872 =viridicollis Kraatz, 1882 =jureceki Born, 1922 "Mts.nr.Simbirsk"(=Ulyanovsk) =volgensis Csiki, 1927; nom.pro jureceki Born "Mts.nr.Simbirsk"(=Ulyanovsk) ssp. jenissoni Dejean, 1831 Altays, Sayans, C-Siberia, N-Mongolia ~dubius Motschulsky, 1846, nec Herbst, 1779 Irkutsk(LT) =jacutus Mäklin, 1881 "Potapowski"(=Potapovo SE of Noril’sk) ~cyanescens Kraatz, 1882, nec Sturm, 1815 ?"Altay" ~nigritulus Kraatz, 1882, nec Kraatz, 1876 ?"Altay" =fraudator Lapouge, 1906 Irkutsk =hunnorum Lapouge, 1909 "Dzhungaria" 14 rothi Dejean, 1829 Romania, NE-Hungaria, W-Ukraine ssp. rothi Dejean, 1829 C-Romania:Sibiu env.; "Transsylvania" =aequistriatus Kraatz, 1878 =latestriatus Kraatz, 1878 =quadricatenatus Kraatz, 1878; nec ? =varistriatus Kraatz, 1878 =birthleri Reitter, 1885 ~rugulosus Birthler, 1885; un ~ignoscus Csiki, 1906; un; nom.pro quadricatenatus Kraatz ssp. comptus Dejean, 1831 CW-Romania:Retezat Mts.to Bihor Mts.; "Banat" ~hopffgarteni Kraatz, 1880, nec Kraatz, 1878 "S-Hungary"(inexact; ?Godeanu & Tarcu Mts.) ~merkli Kraatz, 1880, nec merklii Hopffgarten, 1878 ~merklianus Géhin, 1885; un; nom.pro merkli Kraatz =dieneri Merkl, 1900 "Szöreny Mts."(=Godeanu/Tarcu Mts.) =diffinis Csiki, 1906 "Komitat Kolosz"(=Cluj) =szoerenyensis Csiki, 1908; nom.pro hopffgarteni Kraatz "S-Hungary"(inexact; ?Godeanu/Tarcu Mts.) ~ulrichhofmanni Kleinfeld & Korell, 1986; un SW-Romania:30km SE of Lugoj:Cavaran ~cavarani Lie, 1989; un SW-Romania:30km SE of Lugoj:betw.Cavaran/Tincova =ulrichhoffmanni Lie, 1989 SW-Romania:ca.20km E of Lugoj:Nadrag env.:Gosta Hut nr.Nadrag ~borlovensis Lie, 1994; un SW-Romania:ca.50km SE of Lugoj:Mt.Muntele Mic near Borlova ~micotensis Lie, 1999; un SW-Romania:ca.40km SE of Lugoj:Poiana Rusca massif:Micota valley nr.Boutari ssp. hampei Küster, 1846 NW,C,?SW-Romania; NE-Hung.:Szabolcz-Szatmár; W-Ukraine:Mukachevo; "Siebenburgen" ~schaumi Birthler, 1886, nec Gaubil, 1849 "Serbia"(?inexact) =aurosericeus Kraatz, 1880 "Maros-Vasarhely"(=Tirgu Mures) =marusii Ormay, 1890 "Gyula-Fehervar & Nagy Enyed"(=Alba Iulia & Aiud) =ormayi Reitter, 1896 "Beregszasz"(=Beregovo) =liebli Dietl, 1897 "Komitat Kis-Küküllö"(=Tirnava Mic) ~marginatus Kraatz, 1900; un ?"Maros-Vasarhely"(=Tirgu Mures) =zoppai Kraatz, 1900 ?"Maros-Vasarhely"(=Tirgu Mures)

14 =dacicus Csiki, 1906 "Szamosujvar & Naszod"(=Gherla & Bistrita Nasaud) =eximius Csiki, 1906 "Nagyvarad"(=Oradea) =fraternus Csiki, 1906 "Maros-Vasarhely"(=Tirgu Mures) ~mendax Csiki, 1906, nec Rossi, 1790 "Komitat Marmaros"(=Maramureş) ~validus Csiki, 1906, nec Kraatz, 1884 "Komitat Hunyad:Nagayg"(=Hunedoara:ca.40km NE of Deva:Sacarimb) ~vitiosus Csiki, 1906; un ~zilahiensis Csiki, 1906; un "Komitat Szilagy:Zilah"(=Salaj:Zalau) ~concinnus Lapouge, 1916; un; nom.pro schaumi Birthler "Serbia"(?inexact) =hunyadensis Csiki, 1926; nom.pro validus Csiki "Komitat Hunyad:Nagayg"(=Hunedoara:ca.40km NE of Deva:Sacarimb) =bokori Csiki, 1927; nom.pro concinnus Lapouge "Serbia"(?inexact) =telekii Csiki, 1937 ?"Banat"(?inexact) =gutiiensis Takács & Lie, 1992 NW-Romania:NE of Baia Mare:Gutii Mts. =praedacicus Takács & Lie, 1992 C-Romania:WNW of Alba Iulia:Zlatna ssp. incompsus Kraatz, 1880 NC,C-Romania:Rodna Mts.to Harghita Mts.; "S-Hungary:Brasso env."(=Braşov) ~elegantissimus Csiki, 1906; un =mehelyanus Csiki, 1906 "Komitat Csik:Vigyazok-Tarko"(=ca.Harghita Mts.) =spectabilis Csiki, 1906 Rodna Mts. ssp. alutensis Savulescu, 1972 C-Romania:Olt River:Rimnicu Vilcea env.(Calimaneşti) 15 scheidleri Panzer, 1799 CE-Europe, W-Carpathians, Balcans ssp. scheidleri Panzer, 1799 N,E-Austria(Wien); SE-Germany:Passau, Regensburg; S-Bohemia; S-Moravia; C,W-Hungary ~caeruleus Panzer, 1812, nec coeruleus Steven, 1808 Wien =preyssleri Duftschmid, 1812 Niederösterrieich; Puchberg am Schneeberg(LT) =aeneipennis Sturm, 1815 =purpuratus Sturm, 1815 ~virens Sturm, 1815, nec Müller, 1776 ~atrocoeruleus Letzner, 1850; un ~aureus Letzner, 1850; un ~cyaneus Letzner, 1850; un ~niger Letzner, 1850; un ~purpureoaureus Letzner, 1850; un ~purpureomarginatus Letzner, 1850; un ~purpureus Letzner, 1850; un ~viridiaeneus Letzner, 1850; un ~viridimarginatus Letzner, 1850; un ~viridis Letzner, 1850; un =gibbicollis Motschulsky, 1865 =ambitiosus Schaufuss, 1872 =improbus Schaufuss, 1872 =principatus Schaufuss, 1872 =superbus Schaufuss, 1872 =viridicinctus Schaufuss, 1872 ~aerescens Dalla Torre, 1877; un ~aeruginescens Dalla Torre, 1877; un ~aurescens Dalla Torre, 1877; un ~coerulescens Dalla Torre, 1877; un ~cuprescens Dalla Torre, 1877; un ~elegans Dalla Torre, 1877; un ~fuscescens Dalla Torre, 1877; un ~metallescens Dalla Torre, 1877; un ~nigrescens Dalla Torre, 1877; un ~violescens Dalla Torre, 1877; un ~virescens Dalla Torre, 1877; un =dominus Reitter, 1885 ~graciosus Géhin, 1885; un =styriacus Kraatz, 1887 "Styria"(=Steiermark) =limbifer Reitter, 1889 =burghauseri Fleischer, 1899 =floriani Penecke, 1905 Steiermark:Gratwein ~distinguendus Csiki, 1906, nec Duftschmid, 1812 E of Budapest:Rakosfeld =jucundus Csiki, 1906 W of Budapest:Pilis Mts. =pannonicus Csiki, 1906 C-Hungary:Fejér:Székesfehérvár ~duchoni Lapouge, 1906; un "Krain"(error) ~prolifer Lapouge, 1906; un "Krain"(?error) =kollarisans Sokolar, 1907 "Moravia" ~coeruleocinctus Fleischer, 1925; un ~elegans Fleischer, 1925; un =gebhardti Bodemeyer, 1927 SW-Hungary:Tolna:Dombovar =maderianus Mandl, 1957 Niederösterrieich:Aspang ~pseudoscheidleri Mandl, 1964; un NE-Austria:Leitha Mts.:Winden env. ~punctatocostatus Mandl, 1964; un Leitha Mts.:Winden env. (reductecostatus Mandl, 1964; un Leitha Mts.:Winden env. ~baderlei Mandl, 1965; un NE-Austria:Burgenland:Parndorfer Platte(SE of Wien) ~praescheidleri Mandl, 1965; un SW-Hungary:Baranya:Mescek Mts. ~pseudojucundus Retezár, 1974; un NW-Hungary:Komárom:Gerecse Mts. ~vertesensis Retezár, 1974; un NW-Hungary:Fejér:Vértes Mts. ssp. helleri Ganglbauer, 1893; ?non priority Slovakia("Kom.Neutra"=Nitra); N,C,E-Moravia; N-Bohemia; ?W-Ukraine =repercussus Drapiez, 1819; ?priority "Karpathen" ~morawitzi Kraatz, 1887, nec Ganglbauer, 1887 ?"Bohemia" ~cupripennis Fleischer, 1925; un ~pseudoambitiosus Fleischer, 1925; un ~pseudoexcellens Fleischer, 1925; un ~pseudosuperbus Fleischer, 1925; un ~viridipennis Fleischer, 1925; un ~pseudopreyssleri Breuning, 1932; un; nom.pro morawitzi Kraatz ?"Bohemia" =giuseppemuelleri Hildebrand, 1955 S-Moravia:"Brünn"(=Brno)

15 =beheimi Niedl, 1957 W-Ukraine:Mukachevo ~bieneri Mandl, 1965; un C-Slovakia:Kralovany ~marmaroschensis Mandl, 1965; un "Marmaros" ~subparvulus Mandl, 1965; un NC-Hungary:Börzsöny Mts.:Dios Jenö ~subtilepunctatus Mandl, 1965; un ssp. kollari Palliardi, 1825 SW-Romania:Banat Mts.(Mehadia) =magnificus Kraatz, 1878 =semetricus Kraatz, 1878 "Banat" =negotinensis Reitter, 1896 Serbia:Negotin ~paroecus Csiki, 1906; un ~sejunctus Csiki, 1906; un =joergeri Born, 1927 "Walachei" ssp. praecellens Palliardi, 1825 Serbia:Novi Sad, Fruska Gora Mts.("Futtak"=Futog) ssp. illigeri Dejean, 1826, nec Panzer, 1806; lsu Bosnia; W-Croatia:Dinara planina =laticollis Sturm, 1826 =stentzi A.& J.B.Villa, 1833 "Hungary"(?error) ~parallelus Kraatz, 1878, nec Faldermann, 1835 Pljesevica Mts. =curtulus Ganglbauer, 1892 Mts.nr.Sarajevo =bjelasnicensis Apfelbeck, 1902 Bosnia:Bjelasnica Mts. =koschanini Csiki, 1904 Serbia:Kopaonik Mts. ~transfugus Apfelbeck, 1907, nec Semenov, 1898 Sarajevo env. =unanensis Born, 1909 Bosnia:Bihac ~kollarioides Mandl, 1966; un Serbia:Fruska Gora Mts. ~cerensis (Toševski, 1990); un NW-Serbia:Cer Planina(=SW of Sabac), Korenita nr.Loznica ~tarensis (Toševski, 1990); un W-Serbia:Tara Mts.:Mitrovac ssp. versicolor Frivaldsky, 1835 Bulgaria:Stara planina Mts.(Sliven) =buresi Hanuš, 1923 Stara planina Mts. ssp. zawadzkii Kraatz, 1854 E-Slovakia, NE-Hungary, W-Ukraine =dissimilis Csiki, 1906 "Kom.Ung"(=W-Ukraine:Uzhgorod env.) ssp. seriatissimus Reitter, 1896 W-Ukraine:Carpathians:Hoverla Massif; ?"Marmaros" ssp. ronayi Csiki, 1906 E-Slovakia("Kom.Zemplén & Ung"), NE-Hungary ssp. simulator Kraatz, 1876 NE-Serbia:Majdanpek env. =serbicus Hopffgarten, 1878 ?"Serbia" =polychloros Beuthin, 1885 ~tekijensis (Toševski, 1990); un NE-Serbia:Tekija:Djerdapska klisura, 50m(=Dunav right bank opposite to Orşova) 16 spasskianus Fischer, 1823 Altays Mts.to Transbaykalia; N-Mongolia ssp. spasskianus Breuning, 1932 SC-Siberia:Krasnoyarsk env. to Transbaykalia; N-Mongolia; "Dauria" =gryphus Motschulsky, 1846 "Baykal Lake"; "Sib.or.:Amaga"(LT) =teicherti Mandl, 1963 N-Mongolia:"Dsun Charaa" ssp. tungus Obydov, 1999 C-Siberia:Tunguska (Ilimpeya & Dovogpokan Riv.) & Podkam.Tunguska Riv.Basin ssp. putus Motschulsky, 1846 C,W-Altays Mts.(Zmeinogorsk, Kolyvan; LT) ssp. schestopalovi Plutenko, 1995 NE-Kazakhstan:Sarym-Sakty Mts.:Soldatovo env. ssp. geblerianus Obydov, 1999 N-Altays Mts.:Barnaul env.(Barnaul) ssp. saragaschensis Obydov, 1999 SC-Siberia:Hakassia:betw.Krasnoyarsk/Abakan:Saragash ssp. cracens Obydov, 1999 E-Kazakhstan:Kalbinsky Mt.R.:Ognevka village ssp. aeruginosiformis Breuning, 1932 West Sayans:Kazyr Suk River ssp. microputus O.Berlov & Shilenkov, 1996 West & East Tannu-Ola Mt.R.:S-macroslope(Tuva:Khandarayty env.) ssp. zinaidae Obydov, 1997 SE-Tuva:Sangilen Mts.:Kargy River valley:50:31N/97:01E, 1800m ssp. shoriensis Obydov, 1999 SC-Siberia:Gornaya Shoriya(=SE of Novokuznetsk) 17 tarbagataicus Kraatz, 1878 E-Kazakhstan:Tarbagatay Mt.R., Saur Mts. ssp. tarbagataicus Kraatz, 1878 E-Kazakhstan:Saur Mts.("Tarbagatai"-probably inexact; LT) ssp. dshungaricus Csiki, 1927; nom.pro motschulskyi Lapouge E-Kazakhstan:Tarbagatay Mt.R.:C, E-part; "Dzhungaria" ~motschulskyi Lapouge, 1909; un "Dzhungaria" ssp. pietrorattii Deuve, 1991 E-Kazakhstan:Tarbagatay Mt.R.:W-part(Urdzhar env.) =pseudogebleri Deuve, 1994 Tarbagatay Mts.:Alekseevka nr.Urdzhar ssp. fumidicolor Deuve, 1998 "Sibérie, Zlatoust"(=Zlatoust betw.Ufa/Chelyabinsk); 1f; ?regalis Fischer ssp. necincertus Deuve, 1998 "Sibérie orientale"; 1m 18 venustus Morawitz, 1862 Far East, N-Korea, NE-China ssp. venustus Morawitz, 1862 Russia:Far East(E-Siberia:Bureya Mts.) =carinulatus Chaudoir, 1869; nec ? ?"Amur" =acutesculptus Chaudoir, 1877; nom.pro carinulatus Chaudoir ?"Amur" =christophi Kraatz, 1878 "Amur" ssp. kaesongensis Imura, 1993 CW-Korea:Kaesong; 1f 19 wulffiusi Morawitz, 1862 Far East, N-Korea, NE-China ssp. wulffiusi Morawitz, 1862 Russia:Far East("Bai Possiet"=Pos’et SW of Vladivostok); NE-Korea:Chongjin, Hoeryong =dekraatzi Kraatz, 1881 ~distinctus Haury, 1886, nec Rossi, 1792 ssp. opacipennis Reitter, 1895 NE-Korea:"Gensan"(=Wonsan) ssp. odaesanus (Ishikawa & Kim, 1983) CE-Korea:Taebeaek-san Mt.R.:Mt.Odae-san ssp. taebeagsanus (Ishikawa & Kim, 1983) CE-Korea:Taebeaek-san Mt.R.:Mt.Taebaek-san ~alpinus (Kwon & Lee, 1984), nec Paykull, 1790 Mt.Taebaek-san ssp. ssangnamensis (Kwon & Lee, 1984) NC-Korea:Huchang:Ssangnamdong; 1f ssp. jilinensis Deuve & Imura, 1990 NE-China:Jilin:Hunjiang:Changbaishan Mts.:NW-slope:Bai-he =?yonghongi (Li, 1992) Heilongjiang:Shangzhi:Mao’er Shan =?yonghongianus Deuve, 1992; nom.pro yonghongi (Li) - ssp. liaoningensis Deuve & Imura, 1990 NE-China:E-Liaoning:Fengcheng:Fenghuang Shan; ?NW-Korea:Sinuiju:Chonma-san ssp. flavifemoratus Imura & Yamaya, 1994 NW-Korea:Sinuiu env.:Mt.Chonma-san ssp. onjinsanicus Rapuzzi, 1994 N-Korea:Hvanghe-pukto:Onjinsan Massif, 1000m(=ca.60km SE of Pyongyang); 2m ssp. myohyangicus Deuve & Li, 1999 NC-Korea:Hyangsan:Mt.Myohyang-san 20 zherichini Shilenkov, 1990 N-Siberia:Polar Urals to Kolyma River ssp. zherichini Shilenkov, 1990 N-Siberia:Polar Urals to Taymyr Peninsula(Gydan Peninsula:Nosok village) =pjasinensis Gottwald, 1993 NC-Siberia:Taymyr Peninsula:Pyasina River ~taimyrensis Shilenkov, in litt. - ssp. kuzminae Shilenkov, 1990 NE-Siberia:Kolyma basin(Kolyma Lowland:middle Alazeya River)

16 ****

PSEUDOCRANION Reitter, 1896; type species Carabus gansuensis Semenov =Cratocranion Semenov, 1898; type species Carabus benjamini Semenov species: 22

1 alliciens Deuve, 1990 N-Sichuan:Songpan Xian(exactly:nr.Huanglong, 3400m; humid forest) 2 benjamini Semenov, 1898 N-Sichuan & S-Gansu:Pingwu, Nanping, Wenxian env. ssp. benjamini Semenov, 1898 N-Sichuan:"Ta-tzao-pin nr.Lun-ngan-fu"(=pass betw.Pingwu/Wenxian; LT) ssp. meissonnieri Deuve & Mourzine, 1997 S-Gansu:N of Wenxian, 3000m ssp. roberti Deuve & Mourzine, 1998 N-Sichuan:Mts.N of Nanping, 2000m(=20km N of Nanping) 3 fumigatus Semenov, 1898 N-Sichuan:Songpan, Zhangla env. ssp. fumigatus Semenov, 1898 N-Sichuan:Songpan env.; "Sun-pan"(=Songpan; LT) ssp. zhanglaicola Deuve, 1996 N-Sichuan:Zhangla env., 3700m 4 gansuensis Semenov, 1887 S-Gansu, N-Sichuan; ?E-Qinghai ssp. gansuensis Semenov, 1887 S-Gansu:"Min-Tchjaou"(=Min Xian; LT); ?E-Qinghai:Qinghai Lake env. ~laetus Semenov, 1887, nec Fabricius, 1794 S-Gansu:"Ndami"(=Ponggartang) etc. ssp. venerandus Deuve & Mourzine, 1997 S-Gansu:betw.Wenxian/Wudu:Toqizi, 2300-2500m(=nr.Linjiang) ssp. tiro Semenov, 1898 N-Sichuan:Songpan env.; "Sun-pan"(=Songpan; LT) =subgraniferus Semenov, 1898 N-Sichuan:"Sun-pan"(=Songpan) ssp. dilacerans Deuve, 1992 S-Gansu:betw.Linxia/Xiahe; 1f ssp. jedompansis Deuve, 1989 N-Sichuan:Zoige env.:Jedompa; Gansu:Dogcanglhamo(=Langmusi) ssp. silviphilus Deuve, 1993 S-Gansu:Tewo ssp. achillecasalei Deuve, 1997 N-Sichuan:"betw.Huanglong/Jiuzhaigou, 3200m"(=?pass betw.Songpan/Nanping) ssp. luhuoensis Deuve, 1994 NW-Sichuan:betw.Luhuo/Sertar:pass 35km NNE of Luhuo 5 gonggaicus Deuve, 1989 W-Sichuan:Gongga Shan; 2m 6 jiudingensis Deuve, 1994 C-Sichuan:Maowen Xian:Jiuding Shan 7 kitawakiellus Imura, 1995 S-Shaanxi:Qinling Shan Mts.:Mt.Taibai Shan:Houzhenzi Zhen, 3700m(nr.summit) 8 labrangicus Deuve, 1992 S-Gansu:Xiahe(=Labrang), Ponggartang env. ssp. labrangicus Deuve, 1992 S-Gansu:Xiahe env.(=Labrang) ssp. macilentior Deuve, 1996 S-Gansu:valley E of Ponggartang 9 lazikouensis Deuve, 1997 S-Gansu:Lazikou env., 3100m; pass N of Lazikou 10 meditabundus Deuve, 1992 N-Sichuan:"Sunpanting"(=Songpan); 1f 11 pseudoremondianus Deuve, 1997 N-Sichuan:Pingwu:Maoniu Shan, 3100m; Nanping env. 12 pseudosackeni Deuve, 1989 W-Sichuan:Garze Xian; 1m 13 remondianus Deuve, 1996 N-Sichuan:Pingwu env.:Sanlukou, 2800m 14 sackeni Semenov, 1898 N-Sichuan:Pingwu, Songpan, Nanping env. ssp. sackeni Semenov, 1898 N-Sichuan:"Ta-tzao-pin nr.Lun-ngan-fu"(=pass betw.Pingwu/Wenxian) ssp. mephistopheles Deuve, 1989 N-Sichuan:Songpan env:Huanglong, 4000m(alpine meadows) ssp. gamisiensis Deuve, 1990 N-Sichuan:pass betw.Nanping/Songpan; Jiuzhaigou =jiuzhaiensis Deuve, 1992 Jiuzhaigou ssp. gerardchareti Deuve, 1993 N-Sichuan:Songpan Xian:Zhaga(=village 15km W of Songpan) 15 sackenioides Deuve, 1991 N-Sichuan:"Xindian"(=?NE of Ya’an, 30:11N/103:12E); 1m/1f 16 shuamaluko Deuve, 1991 C-Sichuan:Barkam env. ssp. shuamaluko Deuve, 1991 C-Sichuan:E of Barkam:Shuamaluko ssp. tatsienluioides Deuve, 1992 C-Sichuan:NW of Lixian:Zhegushan Pass ssp. barkamicola Deuve, 1996 C-Sichuan:valley SW of Barkam 17 sinicus Breuning, 1950 W-Sichuan:"Tongho River"(=?Dadu Ho valley:Luding env.); 1f 18 taibaishanicus Deuve, 1989 S-Shaanxi:Qinling Shan Mt.R. ssp. taibaishanicus Deuve, 1989 S-Shaanxi:Qinling Shan Mt.R.:Taibai Massif =fani Imura, 1993 S-Shaanxi:Qinling Shan Mt.R.:Chang’an Xian:2410-2760m:Maijieshi env. ssp. pressicollis Cavazzuti, 1999 S-Shaanxi:Huo Di Tang:Ping He Liang, 2050-2250m; 3f 19 tibetanophilus Deuve, 1991 N-Sichuan:Hongyuan, Waqen, Sanggarpar, Mewa env. ssp. tibetanophilus Deuve, 1991 N-Sichuan:Hongyuan, Waqen ssp. yak Deuve, 1991 N-Sichuan:Sanggarpar =prunicolor Deuve, 1996 N-Sichuan:Sanngarmai, 3650m ssp. confusipennis Deuve, 1996 N-Sichuan:Mewa env. 20 viatorum Deuve, 1992 S-Gansu:Lanzhou env., Xinglongshan Massif 21 wenxianicola Deuve, 1996 S-Gansu:Wenxian, Wudu env. ssp. wenxianicola Deuve, 1996 S-Gansu:Wenxian env. ssp. wuduensis Deuve, 1992 S-Gansu:70km W of Wudu, 1800-2000m 22 zhanglaensis Deuve, 1991 N-Sichuan:Zhangla env.

SHUNICHIOCARABUS Imura, 1995; type species Carabus uenoianus Imura species: 1

1 uenoianus Imura, 1995 NE-Sichuan, S-Shaanxi; ?W-Hubei:Shennongjia ssp. uenoianus Imura, 1995 NE-Sichuan:Dabashan Mts.:Chengkou Xian:Bashan env.:1600-1900m; ?W-Hubei:Shennongjia ssp. confectus Cavazzuti, 1999 S-Shaanxi: betw.Xi’an/Shiquan:65km S of Xi’an, 2150m; 1m

CUPREOCARABUS Deuve, 1997; type species Carabus huangi Deuve species: 10

1 alpherakii Semenov, 1898 W-Sichuan to NW-Yunnan:Kangding to Zhongdian ssp. alpherakii Semenov, 1898 Sichuan:"betw.Khotchukoi/Snino"(=betw.Kangding/Litang; LT); Litang env.; ?Jiulong env. ssp. budha Breuning, 1950 Sichuan:"Tongho River"(=?Dadu Ho); Kangding env.:Zheduo Pass, Mugecuo & Dagong Lake =tongensis Breuning, 1950 Sichuan:"Tongho River"(=?Dadu Ho) =mugeko Deuve, 1992 Sichuan:"Mugeko Lake"(=Mugecuo Lake N of Kangding) =tagongensis Deuve, 1992 Sichuan:"Red Lake nr.Tagong"(=Dagong N of Kangding, nr.Mugecuo Lake)

17 ssp. chareti Deuve, 1993 NW-Yunnan:Zhongdian env. ssp. pignataroi Cavazzuti, 1996 SW-Sichuan:pass ca.40km NE of Muli 2 brezinai Deuve, 1994 Sichuan:Barkam env. 3 huangi Deuve, 1992 Sichuan:NW of Lixian:Zhegushan Pass; Sanggarpar env. 4 laevithorax Breuning, 1935 C-Sichuan:Siguniang Shan Massif, Wolong env.; ?Kangding env. ssp. laevithorax Breuning, 1935 Sichuan:"Konker Shan"(=Gongga Shan; ?inexact); ?Yingxiuwan & Wolong env. ssp. pecoudianus Deuve, 1992 Sichuan:Gongga Shan,"Tatsienlu"(=Kangding; ?inexact) ssp. siguniangensis Deuve, 1997 Sichuan:Siguniang Shan Massif:Xiaojin Xian:Suangqiao(=Shuangqiao) 5 lixianensis Deuve, 1990 N,C-Sichuan:Barkam, Kangding, Garze env. ssp. lixianensis Deuve, 1990 Sichuan:NW of Lixian:Zhegushan Pass; Shuamaluko, Sanggarpar, Barkam env. =goergneri Kleinfeld, 1991 Sichuan:Barkam ssp. kangdingi Korell, Kleinfeld & Görgner, 1992 Sichuan:Kangding, 2800m, forest ssp. sculptior Deuve, 1992 Sichuan:S of Garze 6 miroslavi Deuve, 2000 Sichuan:Rilong:Qionglai Shan:Mt.Siguniang Shan:31:02.58N/102:16.85E/3800-4300m 7 morettoi Deuve, 1997 Sichuan:betw.Barkam/Xiaojin:Lianghekou env., 3400m; Xiaojin env.; 3250m 8 pseudohuangi Deuve, 1995 Sichuan:Mts.NNW of Barkam 9 saueri Deuve, 1993 C-Sichuan:Siguniang Shan Mt.R.:Rilong, Shuangqiao; "Zhilong"(=Rilong) 10 sinotibeticola Mandl, 1975 "Tibet:Jin Long Shien"(=?Jiulong S of Kangding); 1m; ?alpherakii ssp.

NEOPLESIUS Reitter, 1896; type species Carabus tanguticus Semenov = Carabus wagae tanguticus Semenov =Indocarabus Lapouge, 1930; type species Carabus wagae Fairmaire =Sigenacarabus Kleinfeld, 2000; type species Carabus sigena Kleinfeld species: 27

1 borodini Heinz, 1996 E-Tibet:Nyingtri:Mila-la Pass, 4650m(=pass betw.Lhasa/Nyingtri:ca.160km E of Lhasa) 2 bruggeianus Deuve, 1992 Sichuan:"Tatsienlu"(=Kangding); 1f 3 danae Kaláb, 1995 SC-Tibet:Kangtissu Shan:Nakaerh-Chushui(=Mts.ca.70km W of Lhasa:Nakar env.) 4 draco Březina, 1999 C-Sichuan:Wenchuan:Jiuding Shan:valley 5km N of Wenchuan, 31:28N/103:41E, 3300m 5 folwarcznyi Deuve, 1997 C-Tibet:50km of Jambajing:Suge La valley, 4600m; "Suge La Pass, 5100m"(=Shogula Pass) 6 glasunowi Heinz, 1997 Tibet:Karo-la Pass, W-slope, 4800-5000m(=pass ca.120km SW of Lhasa); 2m 7 glinkai Heinz, 1996 C-Tibet:Kampa-la Pass, 4650m(=pass ca.60km SW of Lhasa) 8 hengduanicola Deuve, 1996 Yunnan:Hengduan Shan:Yanmen env.:28:06N/98:52E 9 hummelioides Deuve, 1989 E-Tibet:"Qarchang"; 1f 10 kamensis Semenov, 1903 SE-Oinghai:Huashixia env.; W-Sichuan:Maniganggo, Garze env. ssp. kamensis Semenov, 1903 SE-Qinghai("Tibet:Dza-tschu R."=Za Qu), Huashi Gorge nr.Huashixia; Sichuan:Maniganggo =insidiosus Semenov, 1903 "Tibet:Dza-tschu River"(=SE-Qinghai:Za Qu) ssp. viridirugulosior Deuve, 1996 W-Sichuan:betw.Garze/Barong:pass 25km S of Garze ssp. garzensis Deuve, 1991 W-Sichuan:Garze Xian; 1m 11 kaschkarowi Semenov, 1903 W-Sichuan:Kangding, Dawu, Luhuo, Jiulong env. ssp. kaschkarowi Semenov, 1903 W-Sichuan:Kangding("A-njan-pu W of Tatsienlu"), Dawu, Qianning, Luhuo, Jiulong env. =dawuensis Deuve, 1992 W-Sichuan:SW of Dawu ssp. grahamianus Mandl, 1975 W-Sichuan:Zya-Ha Pass(=?W of Kangding); 1m 12 kocmani Kaláb, 1997 E-Tibet:ca.120km ENE of Lhasa:16km NW of Drigung til village(=Menpa), 4500m; 3m 13 lama Semenov, 1898 W-Sichuan ssp. lama Semenov, 1898 W-Sichuan:Litang, Maniganggo env.; "betw.Khotchukoi/Snino"(=Kangding/Litang; LT) ssp. batangensis Deuve, 1989 W-Sichuan:Batang Xian ssp. yajiangensis Deuve, 1994 W-Sichuan:betw.Yajiang/Litang:pass 20km W of Yajiang ssp. daochengensis Deuve, 1994 W-Sichuan:betw.Daocheng/Gaqag:pass 35km S of Daocheng, 60km NNE of Gaqag ssp. derongensis Deuve, 1994 W-Sichuan:betw.Xiangcheng/Derong:10km NW of Xiangcheng ssp. garzeicus Deuve, 1995 W-Sichuan:road Garze-Barong:30km SW of Garze ssp. pseudowagae Deuve, 1995 W-Sichuan:road Garze-Barong:25km S of Garze 14 ludmilae Deuve, 1992 E-Tibet:Nyingtri env. ssp. ludmilae Deuve, 1992 E-Tibet:NE of Nyingchi(=Nyingtri; LT) ssp. dongjugensis Deuve, 1997 E-Tibet:Nyingtri:20km of Dongjug:Lenang, 3000m 15 markamensis Deuve, 1992 E-Tibet:Markam, Wangda, Rawu env. ssp. markamensis Deuve, 1992 E-Tibet:Markam env. ssp. rawuensis Imura, Su & Osawa, 1997 E-Tibet:N of Rawu, 29:33N/96:47E ssp. wangdanus Imura, Su & Osawa, 1997 E-Tibet:Wangda env.:SE of Zogang, 29:38N/97:54E 16 morvanellus Deuve, 1998 W-Sichuan:Luding Xian:Jiajin Shan, 3300m(exactly: Mts. 5-10km E of Luding); 1m/1f 17 nanschanicus Semenov, 1898 E-Qinghai, NW-Sichuan ssp. nanschanicus Semenov, 1898 E-Qinghai:Qinghai Lake, Qilian env.("Nanschan:Babo-cho & Che-cho R."=Hei-he R.nr.Qilian) ssp. rugulosipennis Deuve & Kaláb, 1992 E-Qinghai:Gonghe Xian:pass NE of Heka; Ertala(error) ssp. noctirugulosus Deuve & Kaláb, 1996 E-Qinghai:Zhubgyugoin, Chumda env.; Qinghai/Sichuan:pass 15km E of Xiwu ssp. nocticolor Deuve & Kaláb, 1992 N-Sichuan(Huanglong); S-Gansu; ?Qinghai:N of Xining:Daban Shan ssp. katkae Deuve, 1994 N-Sichuan:10km S of Barkam; ?Dogcanglhamo/Langmusi ssp. piceophilus Deuve, 1994 N-Sichuan:betw.Luhuo/Sertar:35km NNE of Luhuo; 2f 18 panda Deuve, 1988 W-Sichuan:"Tatsienlu"(=Kangding); 1m 19 paulusi Kaláb, 1995 SC-Tibet:Lhasa env. ssp. paulusi Kaláb, 1995 SC-Tibet:Cak-la(=pass ca.70km NNE of Lhasa) ssp. rosaliae Deuve, 1997 SC-Tibet:70km of Lhasa:Taktse:"Shuga La" 20 pseudomarkamensis Deuve, 1992 W-Sichuan:Dege, Batang env. ssp. pseudomarkamensis Deuve, 1992 W-Sichuan:Dege:S-environs:4200-4300m ssp. aquilarum Cavazzuti, 1997 W-Sichuan:betw.Litang/Batang:ca.75km W of Litang, 4200m 21 puetzi Kleinfeld, 2000 W-Sichuan:Yajiang:32km WNW Yajiang:Shaluli Shan, 30:08:07N/100:42:36E, 4300m 22 sichuanicola Deuve, 1989 W-Sichuan:"Qiaoqi, Dong He"(=Qiaoqi N of Baoxing), Baoxing env. 23 sigena Kleinfeld, 2000 C-Sichuan:Ya’an:Tianquan:W of Erlang Shan Pass, 29:51:27N/102:15:47E, 2780m 24 tatsienlui Breuning, 1950 W-Sichuan:Kangding env. ssp. tatsienlui Breuning, 1934 W-Sichuan:"Tatsienlu"(=Kangding), ?Litang =persimilis Breuning, 1950 W-Sichuan:"Litan, 2800m"(=?Litang) ~wittmeri Mandl, 1975, nec Mandl, 1974 "Tibet:Jala-la Pass"(=?W of Kangding) =wittmerianus Mandl, 1975; nom.pro wittmeri Mandl, 1975 "Tibet:Jala-la Pass"(=?W of Kangding) ~similissimus Deuve, 1988; un; ios; nom.pro persimilis Breuning "Sichuan, Litan, 2800m"(=?Litang) =simillimus Deuve, 1988; emend.pro similissimus Deuve "Sichuan, Litan, 2800m"(=?Litang) ssp. krali Deuve, 1994 W-Sichuan:Kangding:"Pauma Shan Mts."(=pass betw.Kangding/Moxi)

18 25 trachynodes Bates, 1891 W-Sichuan:Kangding, Luding env. ssp. trachynodes Bates, 1891 W-Sichuan:"Tatsienlu"(=Kangding) ssp. moxiensis Deuve, 1992 W-Sichuan:Luding Xian:Moxi, Hailuogou valley 26 tsogoensis Deuve, 1997 E-Tibet:Nyingtri:"Tsogo, 3800m"(=ca.90km NNW of Nyingtri:N of Bayi:Basum Tso env.) =antonini Deuve, 1997 Nyingtri:50km of Bayi:Basum Tso, 3700m(=ca.90km NNW of Nyingtri:N of Bayi) 27 wagae Fairmaire, 1882 N-India:Uttarpradesh:Almora env.; N-Nepal; China:S-Tibet, E-Qinghai, N-Yunnan ssp. wagae Fairmaire, 1882 N-India:Uttarpradesh:Almora env.; N-Nepal; China:S-Tibet ssp. nyalamensis Deuve, 1996 C-Tibet:Nyalam, 3700m(=nr.Nepal borders; ca.28:06N/85:59E) ssp. yunglaensis Deuve, 1997 C-Tibet:50km NE of Gyanze:Yung La Pass, 5100m(=betw.Gyantse/Rinpung) ssp. tanguticus Semenov, 1887 SE-Qinghai:Zhubgyugoin, Chumda env.; "Mts.betw.Hoang-ho & Jan-tse-kiang"; "Amdo"(LT) ~rufipes Semenov, 1887, nec DeGeer, 1774 "Mts.betw.Hoang-ho & Jan-tse-kiang"; "Amdo"(LT) =semirufipes Semenov, 1898; nom.pro rufipes Semenov "Mts.betw.Hoang-ho & Jan-tse-kiang"; "Amdo"(LT) ssp. pseudotanguticus Deuve & Kaláb, 1992 SE-Qinghai:Yushu env. ssp. sanchari Andrewes, 1921 Sikkim:Gnatong ssp. thibeticola Breuning, 1964 "Tibet:Phari"(=Pagri betw.Sikkim/Bhutan); 1m ssp. schmidi Breuning, 1973 Sikkim:Chumtang ssp. alboequus Deuve & Imura, 1993 NW-Yunnan:Deqen env.:Hengduan Shan Mts.:Mt.Baimaxue Shan; Mts.SE of Deqen =pseudosanchari Deuve & Mourzine, 1997 NW-Yunnan:Deqen env., 4400-4600m(=ca.30km SE of Deqen:Meiliuxueshan Mts.) ssp. degeicola Deuve, 1993 Sichuan:Dege:S-environs:4200-4300m ssp. qamdoi Korell & Richter, 1996 E-Tibet:30km E of Qamdo, 4600-4800m ssp. cholashanus Imura, 1998 E-Tibet: betw.Yanjing/Markam:ca.50km S of Markam:Chola Shan Pass, 29:16N/98:38E, 4400m

EOCECHENUS Semenov & Znojko, 1932; type species Carabus kaznakovi Semenov & Znojko species: 10

1 ditomoides Deuve, 1991 E-Qinghai:Yushu, Chumda env. 2 kaznakovi Semenov & Znojko, 1932 SE-Qinghai:Nangqen env.; E-Tibet ssp. kaznakovi Semenov & Znojko, 1932 "E-Tibet:Tshok-tshu River, Mekong tributary"(=SE-Qinghai:ca.32:26N/97:00E; LT) ssp. nangqenicus Deuve, 1995 SE-Qinghai:betw.Doramarkog/Nangqen:20km N of Nangqen, valley 10km NNE of Nangqen ssp. graziosii Deuve, 1995 E-Tibet:road betw.Nangqu/Qamdo:Ka La ssp. angenensis Deuve, 1996 E-Tibet:Angen La 40km of Yanga, 4800m 3 kozloviellus Semenov & Znojko, 1932 "E-Tibet:Ba-tshu River, Jan-tse-kiang tributary"(=SE-Qinghai:ca.33:06N/96:48E); 1m 4 leptoplesioides Deuve, 1992 E-Tibet ssp. leptoplesioides Deuve, 1992 E-Tibet:Markam env.; 2f ssp. dongdaensis Deuve, 1996 E-Tibet:Taniantaweng Shan:Dongda La:4800m ssp. tarasovi Deuve, 1996 E-Tibet:"Dujela, 3900m"; 1m ssp. qilaensis Deuve, 1997 NE-Tibet:30km of Rongbo:Qi La, 5000-5200m 5 micromentum Deuve & Imura, 1993 Sichuan:Dege; 1m 6 mouthiezianoides Deuve & Kaláb, 1992 E-Qinghai:Zhubgyugoin 7 noctivagus Deuve, 1992 Sichuan:Dege env. 8 takashimai Deuve & Imura, 1993 E-Tibet:betw.Dege/Jomda; Qamdo env.:Kaqi La Pass 9 yanjinganus Imura & Mizusawa, 1998 E-Tibet: betw.Yanjing/Markam:ca.50km S of Markam:Chola Shan Pass, 29:16N/98:38E, 4400m 10 yushuensis Deuve, 1991 E-Tibet:Yushu:Baitang, Yushu env.

CALOCARABUS Semenov, 1887; type species Carabus gratus Semenov =Calocechenus Reitter, 1896; type species Carabus malachiticolor Reitter = Carabus przewalskii Morawitz species: 9

1 aristochroides Deuve, 1992 W-Sichuan:Maniganggo, Qagca env. ssp. aristochroides Deuve, 1992 W-Sichuan:Maniganggo ssp. qagcaensis Deuve, 1995 NW-Sichuan: pass 20km S Qagca; betw.Zogqen/Qagca:60km NW of Zogqen 2 gratus Semenov, 1887 E-Qinghai:Heka, Qingshuihe, Zhubgyugoin; NW-Sichuan:Serxü env. ssp. gratus Semenov, 1887 E-Qinghai:Ngola Pass SW of Heka; Qingshuihe; "Kon-chum-chu, By-dshu River"; "Amdo"(LT) ssp. nishidai Deuve & Imura, 1993 SE-Qinghai:Zhubgyugoin; betw.Qingshuihe/Zhenqin ssp. serxuensis Deuve, 1995 NW-Sichuan:Serxü env.; 1m 3 guinanensis Deuve, 1991 Qinghai:Guinan, "Ertala"(error; in fact:pass NE of Heka) 4 janatai Březina, 1996 S-Gansu:Dogcanglhamo/Langmusi:S-environs, 4100m 5 juengerianus Kleinfeld, 1995 SW-Gansu:Zhingshui env.:Wolong Pass, 35:33N/102:48E; valley E of Ponggartang 6 linxiaensis Deuve, 1992 S-Gansu, N-Sichuan ssp. linxiaensis Deuve, 1992 S-Gansu:betw.Linxia/Xiahe:Mts.W of Ponggartang ssp. pseudogratus Deuve, 1994 NW-Sichuan:betw.Luhuo/Sertar:pass 35km NNE of Luhuo ssp. darcangicus Deuve, 1995 NW-Sichuan:road Sertar-Darcang:20km SSE of Darcang ssp. sertarensis Deuve, 1995 NW-Sichuan:road Sertar-Darcang:pass 20km W of Sertar ssp. ponggartangensis Deuve, 1995 S-Gansu:Mts.E of Ponggartang; 1m ssp. n.ssp./n.sp. NC-Sichuan:betw.Sanggarmai/Luhua:pass ca.20km E of Sanggarmai; 3f 7 przewalskii Morawitz, 1886 Qinghai:Qinghai Lake env. ssp. przewalskii Morawitz, 1886 Qinghai:Qinghai Lake env.:Obo, Daotanghe, Xinyuan, Daban Shan; "Mts.of Kansu Province" =malachiticolor Reitter, 1896 "Kuku-noor"(=Qinghai Lake) ssp. sewertzowi Semenov, 1898 Qinghai:Qilian env.(Nanshan S of Kuku-nor:Babo-cho & Che-cho River"=Hei-he R.nr.Qilian) 8 trichothorax Březina & Imura, 1997 N-Sichuan:pass betw.Zhangla/Nanping, 4000m(=ca.40km N of Zhangla); Nanging; 1m/1f 9 turnaianus Deuve, 1995 NW-Sichuan:15km NE of Serxu; 1m/1f

CRYPTOCECHENUS Semenov, 1898; type species Carabus sifanicus Semenov species: 2

1 kalabi Deuve, 1990 S-Gansu:Dogcanglhamo env.(=Langmusi); N-Sichuan:Zoige env. ssp. kalabi Deuve, 1990 S-Gansu:Dogcanglhamo(=Langmusi) =cuprifuscatus Deuve, 1995 S-Gansu:Dogcanglhamo(=Langmusi) ssp. tanggorensis Deuve, 1995 N-Sichuan:Zoige:Tanggor env., 3600m(=betw.Zoige/Waqen) 2 sifanicus Semenov, 1898 Sichuan:Songpan env.; ca.40km NW of Songpan:Mts.betw. Zhangla/Zoige plateau, 3700m

19 ARISTOCARABUS Semenov, 1896; type species Carabus romanowi Semenov = Carabus viridifossulatus romanowi Semenov species: 1

1 viridifossulatus Fairmaire, 1887 N,W-Sichuan; S-Gansu; W-Hubei ssp. viridifossulatus Fairmaire, 1887 W-Sichuan:"Moupin"(=Baoxing); Baoxing env. ssp. phamianus Deuve, 1990 W-Sichuan:"Moupin env.:Yao Gi"(=Qiaoqi N of Baoxing) ssp. gonggashanensis Deuve, 1992 W-Sichuan:Luding Xian:Moxi; Kangding env. ssp. jiajiensis Deuve, 1998 W-Sichuan:Luding Xian:Jiajin Shan, 3300m(exactly: Mts. 5-10km E of Luding); 1f ssp. romanowi Semenov, 1896 N-Sichuan:"Lung-ngan-fu"(=Pingwu nr.Songpan), Jiuzhaigou ssp. lamaorum Deuve, 1990 N-Sichuan:pass betw.Songpan/Nanping; 1f ssp. ventrosior Deuve, 1990 N-Sichuan:Songpan Xian:Huanglong ssp. businskyorum Imura, 1995 W-Hubei:Shennongia env.:Mt.Dashennongjia:near summit, 2500-2900m ssp. andreii Imura, 1998 S-Gansu: ca.70km W of Wudu, 1800-2100m; 1m ssp. rizeanus Imura & Su, 1998 N-Sichuan:Jiuzhaigou Xian(=Nanping Xian):Rizegou:"Primitive Forest", 3100m ssp. seticollis Deuve & Mourzine, 2000 N-Sichuan: 53km W of Lixian, 2500m; 130km W of Wenchuan

PAGOCARABUS Morawitz, 1886; type species Carabus diruptus Morawitz = Carabus crassesculptus diruptus Morawitz species: 1

1 crassesculptus Kraatz, 1881 China:Hebei, Shanxi, Nei Menggu, Shaanxi, Sichuan, Gansu, Qinghai, Henan ssp. crassesculptus Kraatz, 1881 Hebei, Shanxi, Nei Menggu; "Mts.W of Peking" ssp. shimenicus Cavazzuti, 1999 C-Shaanxi:Tongchuan:Shimen Shan, S-slope, 1300-1600m ssp. diruptus Morawitz, 1886 Sichuan, Gansu, Qinghai ssp. garangensis Deuve, 1997 E-Qinghai:Laji Shan:Garang env., 3000-4200m ssp. paradiruptus Imura, 1999 W-Sichuan:Kangding Xian:ca.20km N of Sabde:Liqiu He River, 29:35N/101:23E ssp. rugosissimus Breuning, 1943 S-Gansu:"Hwei-sin, Peiling Shan, Tauping River"(=Hui Xian:Mts.betw.Min Shan/Qinling Shan) ssp. qunqingicolor Imura, 1993 Shaanxi:Qinling Shan:Hu Xian, Baoji Xian & Zhouzhi Xian ssp. jollyi Deuve & Mourzine, 1997 S-Gansu:betw.Wenxian/Wudu:Toqizi, 2300-2500m(=nr.Linjiang) ssp. eleganticolor Deuve, 1997 S-Gansu:70km W of Wudu, 1800-2000m ssp. articervix Cavazzuti, 1999 S-Shaanxi:Huo Di Tang:Ping He Liang, 2050-2250m ssp. crassethoracis Kleinfeld, 1999 Henan:Laojun Mts.:Luangchuan, 33:43N/111:37E; 1m

MEGODONTOIDES Deuve, 1991; type species Carabus erwini Mandl species: 4

1 erwini Mandl, 1975 C-Sichuan:Wenchuan, Lixian env. ssp. erwini Mandl, 1975 C-Sichuan:Wenchuan env.(="betw.Wenchuan/Meng Tseo") ssp. plesiolabrus Deuve, 1991 C-Sichuan:Lixian env. 2 poschingerianus Korell & Kleinfeld, 1989 C-Sichuan:Aba:Barkam ssp. danbaensis Kleinfeld, 1999 C-Sichuan:Danba env., 31:51N/101:54E, Maiduoshan 3 promachus Bates, 1891 W-Sichuan:Kangding, ?Litang env. ssp. promachus Bates, 1891 W-Sichuan:Kangding env.; 10km S of Litang; ?Rilong; Tatsienlu"(=Kangding); ssp. konkerianus Breuning, 1969 W-Sichuan:"Konker Shan, 3000m"(=Gongga Shan) 4 thibetanus Breuning, 1950 W-Sichuan:"Tongho River, 4000m"(=?Dadu Ho valley nr.Luding; altitude data incorrect)

PSEUDOCOPTOLABRUS Reitter, 1896; type species Carabus taliensis (Fairmaire) =Protodamaster Imura & Kezuka, 1989; type species Protodamaster aesculapius Imura & Kezuka = Carabus taliensis aesculapius (Imura & Kezuka) =Nigracoptolabrus Deuve, 1991; type species Carabus burmanensis Breuning species: 5

1 armiger Imura, 1997 SC-Sichuan ssp. armiger Imura, 1997 SC-Sichuan:Meigu Xian:Daliang Shan Mts.:Dafengding:Wahou env., 2600m; pass N of Meigu ssp. cataphractus Cavazzuti, 1998 SC-Sichuan:Xichang:Lunan Shan:Luoji, 3500m; 1f 2 branaungi Imura, 1999 N-Myanmar:130km E of Putao:Gaoligong Shan:ca.28km N of Nitadi, 27:39N/98:40E 3 burmanensis Breuning, 1932 N-Myanmar:N of Putao:Hkakabo Razi Massif("Seinghku, Admy vall.=Seingku, ?Adung valley) 4 nosei Imura, 1997 N-Myanmar:"NE of Putao:near the northern end of Myanmar"(=27:55N/97:23E) 5 taliensis (Fairmaire, 1886) W-Yunnan, SW-Sichuan, N-Myanmar ssp. taliensis (Fairmaire, 1886) Yunnan; ?Dali env. ssp. aesculapius (Imura & Kezuka, 1989) Yunnan:Dali env. ssp. lijiangensis Deuve, 1990 Yunnan:Lijiang env. =sazonovi Cavazzuti, 1999 NW-Yunnan:N of Lijian:"Jade Dragon Mt., 3600m"(=Mt.Yulongxue Shan) ssp. yulongxuensis Deuve, 1990 Yunnan:N of Lijiang:Yulongxue Shan Mts.; alpine form ssp. atentsensis Deuve, 1990 Yunnan:"Atentse"(=Deqen); 1m ssp. kezukai Deuve & Imura, 1991 Yunnan:S of Deqen:Baimaxue Shan Mts.; 1m ssp. xueshanicola Deuve, 1992 Yunnan:NNW of Lijiang:Habaxue Shan Mts., Zhongdian env. ssp. weibaoensis Deuve, 1992 Yunnan:Weibao, 3000m(=Mts.nr.Weicheng); 1m/1f ssp. wengshuiensis Deuve, 1994 Sichuan/Yunnan:betw.Xiangcheng/Wengshui:pass 15 km N of Wengshui ssp. mulianus Deuve, 1995 SW-Sichuan:30km NW of Muli ssp. stupaensis Cavazzuti, 1996 SW-Sichuan:pass betw.Yanyuan/Muli; 3f ssp. cangshanensis Deuve, 1996 Yunnan:Dali env.:Mt.Cangshan, 4000m:25:42N/100:06E; 1m/1f ssp. yanmenensis Deuve, 1996 Yunnan:Hengduan Shan:Yanmen env.:28:06N/98:52E ssp. firmatus Cavazzuti, 1997 N-Yunnan:Dayao Xian:Santai:"Shao Bei Tsaolin"(=Baicao Ling Shan); Dayao Xian:Tang Huan ssp. myanmarensis Imura, 1999 N-Myanmar:130km E of Putao:Gaoligong Shan:Mts. above Nitadi, 27:25N/98:41E ssp. liqiuensis Imura, 1999 W-Sichuan:Kangding Xian:ca.20km N of Sabde:Liqiu He River, 3200m; 2f ssp. tenuicervix Imura, 1999 NE-Yunnan:Qiaojia Xian:Yao Shan, 27:11N/103:01E, 3300m; 1m ssp. jingdongensis Deuve & Pan, 1999 C-Yunnan:Jingdong:Wuliang Shan, 2600m; 1f ssp. peterheisei Kleinfeld, 2000 W-Yunnan:Xuebang Shan:Lujing River, Lanping, 26:27N/ 99:23E, 4000m

IMAIBIODES Deuve, 1991; type species Carabus businskyi Deuve species: 1

20 1 businskyi Deuve, 1990 Yunnan:Lijiang:Yulongxue Shan:E-slope

ACATHAICUS Reitter, 1896; type species Carabus idolon Semenov = Carabus alexandrae idolon Semenov =Cathaicodes Reitter, 1896; type species Carabus alexandrae Semenov species: 1

1 alexandrae Semenov, 1887 S-Gansu:Wenxian, Tewo env.; N-Sichuan:Nanping env. ssp. alexandrae Semenov, 1887 S-Gansu:"Mts.betw.Quan-tin/Ljan-tscha-pa nr.Si-gu"(=Guanjin/Lianghekou nr.Zhugqu) ssp. subidolon Deuve, 1994 S-Gansu:Xiahe(?inexact); ?interruptocostatus Semenov ssp. cratocephaloides Semenov, 1887 S-Gansu:Wenxian env.("betw.Wen-sjan/Tse-Tscheu:Tao-tschi-tsa") ssp. idolon Semenov, 1898; nom.pro potanini Semenov, 1887; ?non priority S-Gansu:Wenxian env.; "Wen-sian"(LT) ~potanini Semenov, 1887, nec (Semenov, 1887) "Wen-sian"(LT) =interruptocostatus Semenov, 1887; ?un; ?priority "Tchoun-tcha-van village nr.Wen-sjan"(=Wenxian) ssp. amplior Deuve, 1994 S-Gansu:Minshan(?inexact) ssp. nanpingicus Deuve, 1995 N-Sichuan:Nanping ssp. tewoicus Deuve, 1996 S-Gansu:Tewo ssp. fantingi Imura & Su, 1998 N-Sichuan:Jiuzhaigou Xian(=Nanping Xian):N-bank of Bai He River:Jiuzhai Zhen, 2100m

CATHAICUS Bates, 1870; type species Carabus swinhoei Bates = Carabus brandti Faldermann species: 1

1 brandti Faldermann, 1835 Jilin, Shandong, Shanxi, Gansu, Shaanxi, Ningxia, Hebei; Beijing env. ssp. brandti Faldermann, 1835 Jilin, Shandong, Shanxi, Hebei; Beijing env.; "Nordchina" =swinhoei Bates, 1870 "Pekin"(LT) =tuerkheimi Harold, 1877 "Peking" =grossegranulatus Breuning, 1943 "S-Mandshuria" ssp. gansuanus Breuning, 1934 Gansu("Hoeishien"=Hui Xian), Shaanxi ssp. reginae Deuve, 1997 Ningxia:C-Hela Shan Mts.:Suyuko, 2800m ssp. aishanensis Kleinfeld, 1999 NE-Shandong:WSW of Yantai:Aishan, 37:25N/120:46E

EUPACHYS Chaudoir, 1857; type species Carabus glyptopterus Fischer species: 2

1 glyptopterus Fischer, 1827 N-Mongolia; SE-Siberia:Chita env.(Nerchinsk); China:Shanxi, Nei Menggu, Hebei, ?Jilin 2 acutithorax Deuve, 1989 Shanxi:Zhongtiao Shan; 1f

FULGENTICARABUS Deuve & Li, 1998; type species Carabus flutschi Deuve & Li species: 1

1 flutschi Deuve & Li, 1998 "Heilongjiang:Yichang, Hongxing, Linchang"(=?ca.60km of NE of Yichun:Hongxing)

TERATOCARABUS Semenov & Znojko, 1932; type species Carabus azrael Semenov & Znojko species: 1

1 azrael Semenov & Znojko, 1932 NE-China:Liaoning; ?Heilongjiang ssp. azrael Semenov & Znojko, 1932 "Tchoulu:San-dzjao-dze"(=?Heilongjiang:Sandaohezi 75km N of Mudanjiang) ssp. mizunumaianus Imura, 1991 E-Liaoning:Benxi:Xiaoshi:Shuidong ssp. gaizhouensis Imura, 1996 S-Liaoning:Chi Shan nr.Gaizhou(=Gai Xian); Shi-zhi-jie E of Gaizhou

CEPHALORNIS Semenov, 1889; nom.pro Ornitocephalus Semenov ~Ornitocephalus Semenov, 1887, nec Sömmering, ?; type species Ornitocephalus potanini Semenov = Carabus potanini (Semenov) =Cychrocarabus Morawitz, 1889; nom.pro Ornitocephalus Semenov species: 1

1 potanini (Semenov, 1887) S-Gansu & N-Sichuan:Wenxian, Nanping env. ssp. potanini (Semenov, 1887) S-Gansu: Wenxian env.; "Wen-sian"(LT) ssp. remondorum Deuve, 1995 N-Sichuan:Nanping

CYCHROSTOMUS Reitter, 1896; type species Carabus anchocephalus Reitter species: 2

1 anchocephalus Reitter, 1896 E-Qinghai, S-Gansu, N-Shanxi; ?Shaanxi ssp. anchocephalus Reitter, 1896 E-Qinghai:Qinghai Lake env.("Kuku-nor"):Huangyuan, Xinyuan; N-Shanxi:Hunyuan env. =facetus Semenov, 1903 "Burkhan-Budha Mts.SW of Kuku-kor:Chati"(=Burhan Budda Mts.) ssp. lanzhouensis Deuve, 1991 S-Gansu:Mts.S of Lanzhou 2 pseudoprosodes Semenov & Znojko, 1932 S-Gansu:nr.Lanzhou

ECCOPTOLABRUS Semenov, 1898; type species Carabus exiguus Semenov species: 1

1 exiguus Semenov, 1898 C,N-Sichuan; S-Gansu; W-Shaanxi ssp. exiguus Semenov, 1898 N-Sichuan:"Sun-pan"(=Songpan env.); Zhangla env.; 4f ssp. nivium Breuning, 1933 C-Sichuan:Mianning; W of Kangding:Zheduo Pass, pass W of Yajiang; pass SE Dawu =cordimeatus Cavazzuti, 1997 SW-Sichuan:betw.Sumdo/Cogsum:pass N of Sumdo, 4600m; 1m ssp. lanzhouicus Deuve, 1989 S-Gansu:Xinlongshan(=50km SSE of Lanzhou:Xinglong Shan Massif) ssp. fanianus Imura, 1993 W-Shaanxi:Qinling Shan:Zhouzhi Xian:pass betw.Bafangzi/Xindian:2000m ssp. wudumontanus Imura, 1998 S-Gansu: ca.70km W of Wudu, 2000-2400m

21 ssp. ochotonarum Březina, 1996 S-Gansu:valley E of Ponggartang; ?Zhingshui env.:Wolong Pass, 35:33N/102:48E

LASIOCOPTOLABRUS Imura, 1993; type species Carabus sunwukong Imura species: 1

1 sunwukong Imura, 1993 Shaanxi:Qinling Shan:Zhouzhi Xian:SW-part:1300m

COPTOLABRUS Solier, 1848; type species Carabus smaragdinus Fischer =Coptolabrinus Reitter, 1897; type species Carabus pustulifer Lucas =Eocarabus Semenov, 1898; type species Carabus jankowskii Oberthür =Eucoptolabrus Semenov, 1898; type species Carabus gemmifer Fairmaire =Nesocoptolabrus Lapouge, 1930; type species Carabus fruhstorferi Roeschke species: 14

1 augustus Bates, 1888 Jiangxi, Hunan, Guizhou, Fujian ssp. augustus Bates, 1888 NC,CW-Jiangxi:"Kiu-kiang"(=Jiujiang), De’an; Guling env.:Lu Shan Mts. =divus Roeschke, 1908 CW-Jiangxi:"Pinghsiang"(=Pingxiang) ~angusticollis Hauser, 1911, nec Fabricius, 1801 N-Jiangxi:"nr.Kuling"(=Guling W of Poyang Hu) =lueshanensis Born, 1918 N-Jiangxi:Lu Shan Mts. ~aereus Hauser, 1919, nec Dejean, 1826 Jiangxi:"Kiukiang, Nan-kang-fu, Luschan" ~spoliatus Hauser, 1919, nec Rossi, 1792 "Kiukiang, Nan-kang-fu, Luschan" ~viridicollis Hauser, 1919, nec Kraatz, 1822 "Kiukiang, Nan-kang-fu, Luschan" ~chlorotrachelus Hauser, 1921; un; nom.pro viridicollis Hauser, 1919 "Kiukiang, Nan-kang-fu, Luschan" ~ziegenhorni Hauser, 1921; un; nom.pro aereus Hauser, 1919 "Kiukiang, Nan-kang-fu, Luschan" ssp. ignigena Hauser, 1914 C,CE-Hunan:"Siantang, Tao Huaping, vallis Lokiang"(=Xiangtan, Tao-hua-ping, Zi Shiu River) ~pauperatus Hauser, 1914; un "Siantang, Tao Huaping, vallis Lokiang" ~purpureus Hauser, 1914; un "Siantang, Tao Huaping, vallis Lokiang" ~semiviridis Hauser, 1914; un "Siantang, Tao Huaping, vallis Lokiang" ~viridanus Hauser, 1914; un "Siantang, Tao Huaping, vallis Lokiang" ~costatus Vogt & Vogt, 1938, nec Germar, 1824 ~subpustulatus Breuning, 1961; un Hunan ~planipustulatus Mandl, 1975; un Jiangxi:Kuling ssp. yangminshanicus Font, 1997 S-Hunan:Yangming Shan(=E of Yongzhou/Lingling); NE-Guangxi:Guanyang:Qianjiadong ssp. cristianofonti Deuve & Font, 1998 CE-Guizhou:Wanshan; 1m =demissus Cavazzuti & Ratti, 1998 C-Guizhou:Tushan, "Fanzzhusum"; N-Guizhou:"Zuny"(=Zunyi) ssp. chalingxian Kleinfeld, 1998 E-Hunan:Batang Mts.:N of Chaling Xian, 1600m, 26:58N/113:31E ssp. pustuliferus Kleinfeld, 1998 S-Hunan: ca.20km S of Lanshan, 1600m, 25:23N/112:10E ssp. guglielmofonti Deuve & Font, 1999 Fujian:"Wuyi Shan, Guangzhi" 2 elysii Thomson, 1856 Anhui, Hubei, Hunan, Henan, Jiangxi, Jiangxu, Shaanxi; ?E-Sichuan ssp. elysii Thomson, 1856 Anhui:N of Anqing; Hubei, Hunan; ?Sichuan:Wanxian, Chia-ting-fu(=Leshan); "Chine boreale" =rothschildi Born, 1889 "Mongolei, Waschan"(?error) =rutishauseri Hauser, 1912 Hubei:"Hankow"(=Wuhan) =hunanensis Born, 1910 Hunan ~smaragdinus Hauser, 1921, nec Duftschmid, 1812 "Nordchina" =yuianus Deuve, 1991 "Kiangsu, Nanking"(=SW-Jiangsu:Nanjing); 1m ssp. connectens Hauser, 1912 Jiangxi:Jiujiang, De’an, Poyang Hu; S-Anhui:S of Chang Jiang; ?S-Jiangsu ~obscurior Hauser, 1912; un "Kiangsi"(=Jiangxi) ~viridipennis Hauser, 1912; un Jiangxi:"Jao-chow"(=Boyang) ~coeruleoviridis Hauser, 1913; un "Jao-chow" ~hilaris Hauser, 1913; un "Jao-chow" ~semicupreus Hauser, 1913; un "Jao-chow" ~thieffryi Hauser, 1913; un "Jao-chow" ~viridiobscurus Hauser, 1913; un "Jao-chow" ~viridissimus Hauser, 1913; un Jiangxi:"Kiukiang"(=Jiujiang) ~brevis Hauser, 1921; un "Jao-chow"(=Boyang) ~coelestiformis Hauser, 1921; un ?"Jao-chow"; ?"Ngan Hoei"(=Anhui) ~fossulatus Hauser, 1921; un "Jao-chow" ~kiangsianus Hauser, 1921; un Jiangxi:"ad lacum Pojang"(=Pojang Ho) ~lampros Hauser, 1921; un ?"Jao-chow" ~latilimbatus Hauser, 1921; un "Jao-chow" =maignaudi Hauser, 1921 NC-Jiangxi:"Te-ngan-hien"(=De’an) ~nigrescens Hauser, 1921; un "Te-ngan-hien" =tenganensis Hauser, 1921 "Te-ngan-hien" ~coactus Csiki, 1927; un; nom.pro brevis Hauser, 1921 "Jao-chow"(=Boyang) ~fossulatulus Csiki, 1927; un; nom.pro fossulatus Hauser "Jao-chow" ~superviridis Csiki, 1927; un; nom.pro viridissimus Hauser, 1913 "Kiukiang"(=Jiujiang) =anhweiensis Hauser, 1932 S-Anhui:S of Chang Jiang River:"Anking"(=Anqing) ssp. chikongschanensis Hauser, 1914 SE-Hunan:"Chikongschan"(=Zhuguang Shan E of Chenzhou/Chen Xian) ~atratus Hauser, 1914; un "Chikongschan" ~obscurior Hauser, 1914; un "Chikongschan" ~viridiobscurus Hauser, 1914; un "Chikongschan" ssp. lopinensis Hauser, 1914 NE-Jiangxi:"Minschan, Lopin"(=Leping) ~nigripennis Hauser, 1914; un NE-Jiangxi:"Minschan, Lopin" ~viridicollis Hauser, 1914; un "Minschan, Lopin" ~viridiobscurus Hauser, 1914; un "Minschan, Lopin" ~viridicoeruleus Hauser, 1921; un "Minschan, Lopin" ssp. lijingkei Font, 1995 N-Zhejiang:Anji:Mt.You-shi lu; 1m/1f; ?lafossei ssp. ssp. pulcher Kleinfeld, 1998 S-Henan:S of Xinyang:Jigong Shan, 31:49N/114:06E ssp. magnificens Kleinfeld, 1999 NW-Henan:Lushi:Xiong’er Mts., 34:03N/111:02E; Laojun Mts.:Luangchuan, 33:43N/111:37E ssp. idai Kleinfeld, 2000 Shaanxi:Hanyin:Langton Shan, 32:53N/108:31E 3 formosus Semenov, 1887 S-Gansu; N,C-Sichuan, Shaanxi; ?E-Qinghai ssp. formosus Semenov, 1887 S-Gansu:"Li-tcha-pou betw.Min-tchou/Si-gou"(=betw.Min Xian/Zhugqu); Wudu, Cheng Xian =berezowskii Semenov, 1898 S-Gansu:"Chuj-sjan env."(=Hui Xian)

22 ~spurius Semenov, 1898, nec Kraatz, 1882 S-Gansu:"Chuj-sjan" =margaritophorus Hauser, 1912 S-Gansu:"Hoei-shien"(=Hui Xian) =stoetzneri Hauser, 1921 "W of Peking"(error) =tener Csiki, 1927; nom.pro spurius Semenov, 1898 S-Gansu:"Chuj-sjan"(=Hui Xian) ssp. subformosus Semenov, 1887 S-Gansu:betw.Wudu/Wenxian("Nai-ti-kha village nr.Wen-sjan"=ca.16km NE of Wenxian) ssp. grumorum Semenov, 1898 E-Qinghai:"Sinin-Shan"(=?Mts.near Xining) ssp. sunpanensis Semenov, 1898 N-Sichuan:"Sunpan"(=Songpan) ssp. rostandi Deuve, 1991 C-Sichuan:Lixian env., Wenchuan ssp. bousqueti Deuve, 1993 S-Gansu:"Wenxian" ssp. latiformosus Deuve, 1993 S-Gansu:"Minchow"(=Min Xian); Shaanxi:"Han-chung"(=Hanzhong),"Hsing-ngan"(=Xi’an) ssp. akane Imura & Yamaya, 1994 C-Shaanxi:Xi’an env.:Tongchuan ssp. wanxianicus Imura, 1995 NE-Sichuan:Dabashan Mts.:Chengkou Xian:Bashan env. ssp. brockmanni Kleinfeld & Schütze, 1997 S-Shaanxi:Lintong:Li-shan Mts.:34:21N/109:13E ssp. brockmannioides Deuve & Mourzine, 1997 S-Gansu:Wudu, 900m; 1f ssp. purpureodiscoides Kleinfeld, 1999 NW-Hubei:Dalong Mts.:Yunxian, 1350m, 32:50N/110:44E ssp. flutschianus Deuve & Li, 1999 S-Shaanxi:Shangzhou:Shahezhizheng, Banshanpe(=Shang Xian ca.120 km ESE Xi’an) ssp. fimbriatus Cavazzuti, 1999 SE-Shaanxi:pass betw. Hujiayuan/Daping, 1300m; 2f 4 fruhstorferi Roeschke, 1900 Japan:Tsushima Island, SW-Honshu 5 gemmifer Fairmaire, 1887 Yunnan, SW-Sichuan ssp. gemmifer Fairmaire, 1887 ?S-Yunnan:S of Kunming; "Yunnan" =cyaneofemoratus Born, 1910 "Yunnan:southernmost part" ssp. mesites Fairmaire, 1897 SW-Sichuan:Xichang env.; ?Kangding env.("Tibet:Siao-Lou-Cha"=betw.Ya’an/Kangding) ssp. loczyi Csiki, 1927; nom.pro yunnanus Born W-Yunnan:Kunming, Lijiang, Dali,"Kouang-hsi"(=Luxi); "Yunnan" ~yunnanus Born, 1910, nec Fairmaire, 1886 "Yunnan" 6 ignimitella Bates, 1888; cos Fujian:Fuzhou; Guangdong, E-Guangxi, E-Hunan ssp. ignimitella Bates, 1888; cos CE-Fujian:Fuzhou, Min Jiang R.; "Fu-chau"(=Fuzhou); SC-Fujian:Yongding Xian(=Fengcheng) ~ignimetalla Bates, 1888; iss of ignimetalla Bates - =ertli Born, 1910 CE-Fujian:"Foutschou"(=Fuzhou) =marginithorax Donckier, 1915 "Foutscheou"(=Fuzhou) ~aureocinctus Hauser, 1916; un Fujian:"Minkiang"(=Minjiang River) ~ignicorona Hauser, 1916; un "Minkiang"(=Minjiang River) ~nigripennis Hauser, 1916; un "Minkiang"(=Minjiang River) ~sculpturatus Hauser, 1916; un "Minkiang"(=Minjiang River) ~amabilis Mandl, 1975; un Fujian:"Foochow"(=Fuzhou) =ngyukmingi Deuve, 1991 Fujian:"Ueshan" ssp. stillatus Cavazzuti & Ratti, 1998 C-Fujian:Youxi ssp. angulicollis Hauser, 1911 E-Guangdong:Meixian, Longchuan; SW-Guandong:Mt.Datian Ding ssp. antaeus Hauser, 1914 NC-Guangdong:Lianping env., Ruyuan Xian; Guangdong/Hunan borders region =mellianus Kolbe, 1914 Guangdong:"Tsha-pin-san Mts.nr.Siu-cao"(=massif ca.50km ENE of Shaoguan) ~costatus Hauser, 1921; un "Guandong/Hunan borders region:nr.Tschao-tschou"(=Shaoguan) ~nudipes Hauser, 1921; un "Guandong/Hunan borders region:nr.Tschao-tschou" ssp. guangxicola Deuve, 1998 CE-Guangxi:Dayao Shan ssp. guidongicus Cavazzuti & Ratti, 1998 E-Hunan:Guidong:Bamian Shan; 2f ssp. hainanensis Deuve & Tian, 1999 SW-Hainan:Ledong Xian; 1m 7 ishizukai Deuve & Ohshima, 1989 E-Sichuan:"Xiufeng"(=?village in Wushan Mt.R.) 8 jankowskii Oberthür, 1883 Russia:Far East; Korea; NE-China:Liaoning ssp. jankowskii Oberthür, 1883 Russia:Far East:Vladivostok; N,C-Korea =seoulensis Deuve, 1998 "Corée, environs de Séoul"(=Soul env.) =sorakensis Deuve, 1998 C-Korea:Taebeaek-san Mt.R.:N-part:Mt.Sorak-san, 700m ssp. fusanus Born, 1907 SE-Korea:Pusan env.; "Fusan"(=Pusan) ssp. kojensis (Kurosawa & Kudo, 1980) S-Korea:Koje-do Island ssp. obtusipennis (Ishikawa & Kim, 1983) SC-Korea:Chiri-san Massif("Mt.Jirisan") =elegans (Kwon & Lee, 1984) Chiri-san Massif(Mt.Chiri-san) ssp. taebeagsanensis (Ishikawa & Kim, 1983) SE-Korea:Taebaek-san Mt.R.(Mt.Odae-san:Dongsanri) =kangwonensis (Kwon & Lee, 1984) SE-Korea:Kangwon-do:Taebaek-san Mt.R.:Mt.Taebaek-san =sobaekensis Deuve & Remond, 1998 Taebeaek-san Mt.R.:N-part:Mt.Sobaek-san, 300m ssp. chindoensis Kwon & Park, 1989 SW-Korea:Chindo Island ssp. benxiensis Imura, 1991 NE-China:Liaoning(Benxi Shi:Guanmen Shan Mts.); S-Jilin ~gaixianensis (Li, 1992), nec (Li, 1992) E-Liaoning:Gai Xian:Wanhu:Haiba, 500m =lii Deuve, 1998; nom.pro gaixianensis (Li, 1992) E-Liaoning:Gai Xian:Wanhu:Haiba, 500m =pseudosobaekensis Deuve & Li, 1998 E-Liaoning:Kuandian Xian:Qigedingzhi, 950m; S-Jilin:Tonghua:Liuhe Xian ssp. fallettii Deuve, 1993 SE-Korea:Kyongsang-pukto:Mt.Chuwang =laurelinae Lassalle, 1999 SE-Korea:Kyongsang-pukto:Hayang ssp. quelpartianus Breuning, 1932 S-Korea:"Quelpart"(=Jeju-do Island) 9 kubani Deuve, 1990 Yunnan:Lijiang env.:Yulongxue Shan:Daju(=Jinsha Jiang right bank:Hutiao Gorge near Daju) 10 lafossei Feisthamel, 1845 Zhejiang, Fujian, Jiangxi, S-Anhui, E-Hubei ssp. lafossei Feisthamel, 1845 Zhejiang:Zhousan Archipelago; "Chusan"(=Zhousan) ~donckieri Born, 1898, nec Boileau, 1896 ~incertus Meyer-Darcis, 1902, nec Motschulsky, 1846 "China" ~chlorothyreus Hauser, 1921; un "Chusan"(=Zhousan) ~costatus Hauser, 1921; un "Tschusan"(=Zhousan) ~euprepes Hauser, 1921; un "Chusan"(=Zhousan) ~fuscoviolaceus Hauser, 1921; un "Chusan" =langurius Roeschke, 1921 Zhejiang:"Chusan:Ou-Hou"(=Zhousan Island:?) ~anceps Csiki, 1927, nec Lapouge, 1924; nom.pro incertus Meyer-Darcis "China" ~baptistatus Csiki, 1927; un; nom.pro costatus Hauser "Tschusan"(=Zhousan) =gyoerffyi Csiki, 1927; nom.pro anceps Csiki Zhejiang:Zhousan Island; "China" ssp. coelestis Stewart, 1845 N,C-Zhejiang:Shanghai env., Hangzhou, Ningbo, Jinhua =?dux Semenov, 1898 "prov.Tsche-kiang"(=Zhejiang) =semenowi Meyer-Darcis, 1902 Shanghai(?error) ~aurescens Hauser, 1913; un Zhejiang:"Yuyao"(=Yuao, 30:04N/121:09E) ~azurescens Hauser, 1913; un "Yuyao bei Ning-po"(=Yuao nr.Ningbo) ~coeruleipennis Hauser, 1913; un "Yuyao"(=Yuao, 30:04N/121:09E) ~nigrescens Hauser, 1913; un "Yuyao" ~smaragdulus Hauser, 1913; un "Yuyao"

23 =hangtschouensis Hauser, 1919 NC-Zhejiang:"Hangtschou"(=Hangzhou) =azurescens Hauser, 1921 ?"Hangtschou"(=Hangzhou) =melanelytrius Hauser, 1921 "Yuyao" ~mimetes Hauser, 1921, nec mimethes Kraatz, 1878 "Yuyao" =obscuratus Hauser, 1921 "Yuyao" ~obscuratus Hauser, 1921, nec Hauser, 1921 ?"Hangtschou" ~perviridis Hauser, 1921; un ?"Hangtschou" =plusios Hauser, 1921 "Yuyao" =pretiosissimus Hauser, 1921 "Yuyao" =pseudochlorothyreus Hauser, 1921 "Yuyao" =schenklingi Hauser, 1921 "Yuyao" =spengeli Hauser, 1921 "Yuyao" ~viridicollis Hauser, 1921, nec Kraatz, 1882 "Yuyao" =diabolicus Hauser, 1932 "Yuyao" ~olivaceus Hauser, 1932, nec Géhin, 1876 "Yuyao" ssp. buchi Hauser, 1913 SE-Zhejiang:"Haimen, Wenchow"(=Rui’an, Wenzhou) ~auratus Hauser, 1913; un "Haimen, Wenchow"(=Rui’an, Wenzhou) ~coerulescens Hauser, 1913; un "Haimen, Wenchow" ~cyanopterus Hauser, 1913; un "Haimen"(=Rui’an) ~holomelas Hauser, 1913; un "Haimen" ~hebes Hauser, 1913; un "Haimen, Wenchow" ~holochlorus Hauser, 1913; un "Haimen, Wenchow" ~nigripennis Hauser, 1913; un "Haimen, Wenchow" ~perviridis Hauser, 1913; un "Haimen, Wenchow" ~coerulescens Hauser, 1929; un SE-Zhejiang:Pingyang(=27:43N/120:31E) ~cyanopterus Hauser, 1929; un Pingyang ~hebes Hauser, 1929; un Pingyang ~melanelytrius Hauser, 1929, nec Hauser, 1921 Pingyang =pingyangensis Hauser, 1929 Pingyang =anjiensis Li, 1993 Zhejiang:Fang Shan:Anji Feng, Youzhihu =tyrannous Li, 1993 Zhejiang:Fang Shan:Anji Feng, Yu Shan, Huchun ssp. montigradus Hauser, 1920; nom.pro montanus Hauser NE-Jiangxi:"Heou-Leang, King-te-chen, Chang-tsin"(=Fuliang, Jingdezhen); C,S-Zhejiang ~lampros Hauser, 1913; un Zhejiang ~montanus Hauser, 1913, nec Géhin, 1882 NE-Jiangxi:"Heou-Leang, King-te-chen, Chang-tsin" =lungtschuanensis Hauser, 1919 S-Zhejiang:"Lungtschuan"(=Longquan) ~pseudolampros Hauser, 1919; un "Lungtschuan" ~coeruleatus Hauser, 1921; un ?"Jiangxi:Chang-tsin-tscheng"(=?Chengxiangzhen) ~coerulescens Hauser, 1921; un ?"Jiangxi:Chang-tsin-tscheng" ~coerulescens Hauser, 1921; un ?"Lungtschuan" ~hebes Hauser, 1921; un ?"Lungtschuan" ~lampros Hauser, 1921; un ?"Lungtschuan" ~magnificentissimus Hauser, 1921; un Jiangxi ~nigrescens Hauser, 1921; un ?Jiangxi ~perviridis Hauser, 1921, nec Reitter, 1896 ?"Lungtschuan" ~reticulatus Hauser, 1921; un Zhejiang:"Tschutschou"(=Lishui) ~roehmi Hauser, 1921; un "Tschutschou" ~schmidti Hauser, 1921; un "Tschutschou"(=Lishui) ~subcostatus Hauser, 1921; un ssp. tungchengensis Li, 1993 SC-Anhui:Tongcheng Xian:Longming:Shanling =dabieshanus Imura, 1996 SW-Anhui & E-Hubei:Dabie Shan:Yuexi(Mt.Miaodashan), Qianshan X., Luotian X., Yingshan ssp. tiantai Kleinfeld, 1997 NE-Hubei:Hong’an:Mt.Tiantai, 31:23N/114:37E; 2f ssp. pseudocoelestis Kleinfeld, 1999 NC-Hubei:Shuizhou:Dahong Mts., 1200m, 31:29N/112:58E ssp. saturatus Hauser, 1913; ?non priority N-Fujian:"Yungling-shan, Yenpin-fu"(=Shan Ling, Nanping); NE,CE-Jiangxi:Guixi, Nancheng ~giganteus Born, 1899, ?nec Heer, 1837; ?priority "Fokien:Gebirge"(=Mts.of Fujian Province) ~croesus Hauser, 1912, nec Fabricius, 1801 Jiangxi:"Kweiki"(=Guixi) ~atratus Hauser, 1913, nec Duftschmid, 1812 N-Fujian:"Yung-ling-shan, Yen-pin-fu"(=Shan Ling, Nanping) ~coeruleatus Hauser, 1913; un "Yung-ling-shan"(=Shan Ling) ~cyanescens Hauser, 1913; un "Yung-ling-shan" ~dauverchaini Hauser, 1913; un "Yung-ling-shan" ~ochraceus Hauser, 1913; un "Yung-ling-shan, Yen-pin-fu"(=Shan Ling, Nanping) =hephaestus Hauser, 1913 Jiangxi:"Chang-tsin-tscheng"(=?Chengxiangzhen) ~olivaceus Hauser, 1913; un "Yung-ling-shan, Yen-pin-fu"(=Shan Ling, Nanping) ~ornaticollis Hauser, 1913; un "Yung-ling-shan"(=Shan Ling) ~pervirens Hauser, 1913; un Jiangxi:"Kweiki"(=Guixi) ~prasinus Hauser, 1913; un "Yen-pin-fu"(=Nanping) ~pullus Hauser, 1913; un Jiangxi:"Chang-tsin-tscheng"(=?Chengxiangzhen) =scialdonei Hauser, 1913 Jiangxi:"Kweiki, Chang-tsin-tscheng"(=Guixi, ?Chengxiangzhen) ~valenciae Hauser, 1913; un "Yen-pin-fu"(=Nanping) ~coerulescens Hauser, 1921; un ?"Yung-ling-shan"(=Shan Ling) ~nigrescens Hauser, 1921; un ?"Yung-ling-shan" ~atratus Hauser, 1926; un CE-Jiangxi:"Mts.nr.Kienchang"(=Nancheng) =kienchanganus Hauser, 1926 CE-Jiangxi:"Kienchang"(=Nancheng) =sumptuosus Csiki, 1927; nom.pro croesus Hauser Jiangxi:"Kweiki"(=Guixi) 11 nankotaizanus Kano, 1932 N,E-Taiwan("N-Formosa:Nankotaizan Mts., 3000m") =taiwanicus Kryzhanovskij, 1956 Taiwan:"Tshibon-Shan" =poschingeri Korell, 1957 "Formosa:Taipe"(=?Taipei) =miwai Kurozawa, 1959 "C-Formosa:Mt.Sylvia, Mt.Pashen, Mokabubu, Malippa, Mt.Ali"(unreliable data) =shimonoyai (Kurosawa & Kudo, 1980) "Formosa:Mt.Lala-Shan"(=ca.24:44N/121:26E) =inopinatus Mandl, 1984 "Formosa:Peinanchushan, 1800m" 12 principalis Bates 1889 Hubei:Yichang:Changyang(LT), Schinan(=Lichuan); Hunan:Dongmen; ?Zhejiang:Hangzhou =hunanicus Deuve, 1995 Hunan; 1m/1f =gabrielischaeferi Font, 1996 N-Hubei 13 pustulifer Lucas, 1869 Sichuan, N-Yunnan, S-Gansu, S-Shaanxi, Guizhou, N-Guangxi, E-Hubei, S-Hunan ssp. pustulifer Lucas, 1869 C-Sichuan:Guan Xian, Baoxing, Kangding, Leshan env. =borni Meyer-Darcis, 1902 C-Sichuan:Emei Shan

24 ~amoenus Hauser, 1913; un C-Sichuan:"Kiating-fu"(=Leshan) =pratti Born, 1915 C-Sichuan:"Wa Shan, Omei Shan, Mou-pin"(=Mts.betw.Fulin/Ya’an, Emei Shan, Baoxing) ~szetschuanicus Csiki, 1927; un; nom.pro amoenus Hauser C-Sichuan:"Kiating-fu"(=Leshan) ~pulvillatus Breuning, 1934; un W-Sichuan:"Tatsienlu"(=Kangding) =grundmanni Mandl, 1955 "Westchina:Tschen-fu"(=?Chengdu) ssp. szechwanensis Hauser, 1931 SE-Sichuan:"Kinfu Shan"(=Jinfo Shan SE of Chongqing); ?NW-Guizhou:Lou Shan nr.Tongzi ~tenuis Breuning, 1934; un SE-Sichuan:"Kinfu Shan"(=Jinfo Shan SE of Chongqing) ssp. katsumii Deuve & Ohshima, 1992 S-Gansu:Wenxian; SW-Shaanxi:Lüeyang, Ankang env. ssp. mirandior Deuve, 1993 N-Yunnan:"Pe-yen-tsin"(=Yanfeng nr.Dayao, betw.Dali/Kunming) ssp. adriaenssensi Lassalle & Prunier, 1993 NE-Guangxi:Longsheng:Miao’er Shan; S-Hunan:Yangming Shan(=E of Yongzhou/Lingling) ssp. guizhouicus Deuve, 1994 SE-Guizhou:Leigong Shan Massif nr.Kaili ssp. wakoi Imura, 1995 NE-Sichuan:Dabashan Mts.:Chengkou Xian:Bashan env. ssp. ming Cavazzuti, 1995 NE-Guizhou:Jiangkou:Fanjing Shan; 1f =elisabettianus Font, 1997 NE-Guizhou:Huan-Jan Shan; 1m ssp. pseudoignigena Kleinfeld, 1996 NE-Guangxi:Yuanbao Shan, 1200m; 1m ssp. inusitatus Cavazzuti, 1997 NE-Guizhou:Jiangkou:Fanjing Shan ssp. wenshuicus Font, 1998 NW-Guizhou:Wenshui, Linjin(=ca.50km NW of Tongzi) ssp. qiannaicus Deuve & Tian, 2000 SE-Guizhou:Libo Xian:Maolan Nat.Park, Dong Shan Zan; 1f ssp. xiongi Imura & Su, 2000 NE-Sichuan:Nanjiang Xian:pass betw.Shanliangqu/Taoyuan:Mt.Guangwu Shan; 1f ssp. mirificus Kraatz, 1894 SW-Hubei("Changyang nr.Ichang"=Yichang) =variefucatus Font, 1997 N-Hubei:Wudang Shan ssp. guerryi Born, 1903 N-Yunnan:"Yu-Nan-Sen"(=Kunming),"Kouang-hsi"(=nr.Luxi); ?S-Sichuan =rectiperlatus Maindron, 1906 N-Yunnan:"Yunnan-fu"(=Kunming) ssp. stephanoforus Cavazzuti, 1998 N-Yunnan:Dongchuan env.; 3500m 14 smaragdinus Fischer, 1823, nec Duftschmid, 1812; lsu SE-Siberia:Chita region; Far East, Korea; NE,NC-China ssp. smaragdinus Fischer, 1823, nec Duftschmid, 1812; lsu SE-Siberia:Chita(Nerchinsk); Far East:Khabarovsk; China:NE-Heilongjiang; ?Sakhalin =paschkoviensis Hauser, 1921 "Middle Amur" ssp. mandschuricus Semenov, 1898; nom.pro dohrni Kraatz; ?non priority Vladivostok env.( Suyfun River), Sikhote-Alin Mt.R. ~major Kraatz, 1878; ?nud =pyroclopus Kraatz, 1878; ?un "Sidemi nr.Vladivostok" ~dohrni Kraatz, 1887, nec Gebler, 1847 Suyfun River(=W of Vladivostok) =costulatus Kraatz, 1889; ?un; ?priority Askold Island =pyrophorus Kraatz, 1889; ?iss of pyroclopus Kraatz - =ussuricus Born, 1914 Sikhote-Alin Mt.R. =steinwehri Hauser, 1932 "Mukden"(=Liaoning:Shenyang; error) ssp. monilifer Tatum, 1847 S-Korea:Jeju-do Island; "Eastern Seas, Korean Archipelago" =tyrannus Lapouge, 1911 "Quelpart"(=Jeju-do Island) ssp. longipennis Chaudoir, 1863 Heilongjiang, Jilin, Liaoning, Hebei; E-Nei Menggu:Honhot; N-Shanxi:Hunyuan; "N-China" =damasteroides Géhin, 1885 "Amur"(error) =cyanelytrius Kraatz, 1889 ?"Mukden"(=Liaoning:Shenyang) =chinganensis Semenov, 1898 "Chingan Majus, Dolon-nor, Wei-tschen"(=Hebei:Mts.E of Duolun) =mandarinus Born, 1898 "China" =meyerianus Born, 1898 "Amur"(error) =arachnopus Hauser, 1914 "Peking" ~cupreus Hauser, 1914; un "Mongolia:Inn-shan:Ta-tsing-shan nr.Kuku-khoto"(=Nei Menggu:Honhot env.) =innshanensis Hauser, 1914 "Mongolia:Inn-shan:Ta-tsing-shan nr.Kuku-khoto" =obbergeni Hauser, 1914 "Mongolia or.:Chinchow-fu"(=Liaoning:Jinzhou) ~obscurior Hauser, 1914; un "Mongolia:Inn-shan:Ta-tsing-shan nr.Kuku-khoto"(=Nei Menggu:Honhot env.) ~viridiobscurus Hauser, 1914; un "Mongolia or.:Chinchow-fu"(=Liaoning:Jinzhou) =antungensis Hauser, 1921 "Chikuan-shan, Antung"(=Liaoning:Zi Yang Guan, Dandong) ~coerulans Hauser, 1921, nec caerulans Morawitz, 1886 "Mongolia:Inn-shan:Ta-tsing-shan nr.Kuku-khoto"(=Nei Menggu:Honhot env.) =lanceolatus Hauser, 1921 "Mandschurei" =mucdenensis Hauser, 1921 "Mucden"(=Liaoning:Shenyang; error) =tschiliensis Hauser, 1921 "prov.Tschili"(=Jilin) ~viridicollis Hauser, 1921, nec Kraatz, 1882 "Mongolia:Inn-shan:Ta-tsing-shan nr.Kuku-khoto"(=Nei Menggu:Honhot env.) ~viridis Hauser, 1921, nec Dejean, 1826 "Mongolia:Inn-shan:Ta-tsing-shan nr.Kuku-khoto" ~maoershanensis (Li, 1992), nec Li, 1992 S-Heilongjiang:Shangzhi:Mao’er Shan ssp. honanensis Hauser, 1921 NC-Henan:"Honan-fu"(=Luoyang); Dengfeng env.:Mt.Song Shan ssp. liaodongensis (Li, 1992) SE-Liaoning:Gai Xian, Gaodun:Minyang Ding, 1000m =laopingdingensis Deuve & Li, 1998 SE-Liaoning:Xiuyan:Dayingzhi:Shajinchun:Laoping Ding; Kuandian:Shihugouxiang ~qianshanensis Deuve & Li, 1998, nec (Li, 1992) E-Liaoning:Anshan:Mt.Qian Shan ssp. furumiellus Deuve, 1994 E-Liaoning:Benxi Xian:Xiaoshi ssp. zhongtiaoshanus Imura & Yamaya, 1994 S-Shanxi:Yuanqu env.:Zhongtiao Shan Mts., Li Shan Mts. ssp. branickii Taczanowski, 1887 C-Korea(Soul env.) =oudoti Hauser, 1913 "Chemulpo" =fulminifer Hauser, 1921 CE-Korea:"Gensan"(=Wonsan) ~planus Born, 1922, nec Fabricius, 1792 "Korea" =horvathi Csiki, 1927; nom.pro planus Born "Korea" ssp. pinganensis Hauser, 1920 N-Korea:"Pingan"(=?Pyongyang); Pu Ryong(=Puryong N of Chongjin) =coreicus Hauser, 1921 "Pu Ryong, Seishin"(=Puryong, Chongjin) ~obscurior Hauser, 1921; un "Pu Ryong, Seishin" ssp. euviridis (Ishikawa & Kim, 1983) SC-Korea:Chiri-san Massif("Mt.Jirisan") =sangjaei (Kwon & Lee, 1984) Chiri-san Massif(Mt.Chiri-san) ssp. hoenggandoensis (Kwon & Lee, 1984) S-Korea:betw.Jeju Island/Korean continent:Chujagundo Isles(Hoenggando Island) ssp. hongdoensis (Kwon & Lee, 1984) S-Korea:Hong-do Island ssp. dolichognathus Deuve, 1995 N-Korea:Tonggon(=?); 1m/1f ssp. yichunensis Deuve & Li, 1999 "Heilongjiang:Yichun, Hongxing, Linchang"(=?ca.60km of NE of Yichun:Hongxing) ssp. yanggangensis Deuve & Li, 2000 N-Korea:Yanggang Do:Samjiyon San ssp. shantungensis Born, 1910 E-Shandong:Shandong Peninsula("Tsingtau"=Qingdao); SE-Liaoning:Dalian env. =tsingtauensis Hauser, 1913 "Tsingtau"(=Shandong:Qingdao) ~tristis Hauser, 1921; un "Tsingtau"

COPTOLABRODES Březina & Imura, 1997; type species Carabus haeckeli Březina & Imura species: 1

25 1 haeckeli Březina & Imura, 1997 SW-Shaanxi:Qinling Mts.:NW of Houzhenzi, 33:35N/107:43E

SHENOCOPTOLABRUS Imura, Zhou & Su, 1999; type species Carabus osawai Imura, Zhou & Su species: 1

1 osawai Imura, Zhou & Su, 1999 W-Hubei, NE-Sichuan ssp. osawai Imura, Zhou & Su, 1999 W-Hubei:Shennongjia:Dashennongjia Massif:E-slope: Guanmenshan, 1460m ssp. micangshanus Imura & Su, 2000 NE-Sichuan:Nanjiang Xian(Micang Shan Mts.:S-bank of Jiaojia Riv.:below Daba)

ACOPTOLABRUS Morawitz, 1886; type species Carabus schrencki Motschulsky =Koreacoptolabrus Kwon & Lee, 1984; type species Damaster constricticollis (Kraatz) = Carabus constricticollis Kraatz species: 7

1 changeonleei (Ishikawa & Kim, 1983) SC-Korea:Chiri-san Massif("Mt.Jirisan") 2 constricticollis Kraatz, 1886 E-Russia:Vladivostok env.; NE-China; N-Korea ssp. constricticollis Kraatz, 1886 E-Russia:Vladivostok env.(Suyfun River); NE-China:E-Jilin; NE-Korea =grallatorius Roeschke, 1921 NE-Korea:"Port Seishin"(=Chongjin) =latioricollis (Mandl, 1954) ~jilinensis (Li, 1992), nec Deuve & Imura, 1990 SC-Jilin:Ji’an =jilinicus Deuve, 1992; nom.pro jilinensis (Li) SC-Jilin:Ji’an(=China/Korea borders S of Tonghua) ssp. furumielloides Deuve, 1997 NE-China:Liaoning:Benxi:Liu’erhe Shan ssp. heilongjiangensis Deuve & Li, 1998 "Heilongjiang:Yichung, Hongxing, Linchang"(=?ca.60km of NE of Yichun:Hongxing); 2f ssp. pihyonensis Deuve & Li, 1998 N-Korea:Pyongan-pukto:Pihyon(=ESE of Sinuiju); 1m/1f 3 gehinii Fairmaire, 1886 Japan:Hokkaido ssp. gehinii Fairmaire, 1886 CW-Hokkaido:Shiribeshi:E-part to Sorachi; "Japan" =grandis Bates, 1883 Sapporo ~cyaneoviolaceus Hauser, 1921, nec Motschulsky, 1844 ?Rumoi:Teshio =katomelas Hauser, 1921 ?Rumoi:Teshio ~viridis Hauser, 1921, nec Dejean, 1826 ?Rumoi:Teshio =shinnai (Nakajima, 1993) SW-Hokkaido:Shimanaki-gun:Ohira River right bank:Shimamaki-mura area ~tagawai (Nakajima, 1993), nec (Nakajima, 1993) SW-Hokkaido:Yamakoshi-gun:Mt.Shamanbé-yama ssp. aereicollis Hauser, 1921 N,C-Hokkaido; ?LNF ~gustavhauseri Nakane, 1961; un TNF ssp. sapporensis Uchida & Tamanuki, 1927 SC-Hokkaido:Hidaka:W-part; "Sapporo"(error) ssp. konsenensis (Ishikawa, 1968) CE-Hokkaido:Nemuro(Yoroushi) & Kushiro:Konsen Plateau ssp. radiatocostatus (Ishikawa, 1968) SC-Hokkaido:Tokachi(Osharibetsu) & Hidaka:Hidaka Mt.R. ssp. manoianus (Imura, 1989) C-Hokkaido:Sorachi-gun(Minami-furano-cho:Ochiai:Sorachi River:left bank:Ecchu-dantai) ssp. shimizui Imura, 1994 W-Hokkaido:Shakotan Peninsula:Tomaru-togé Pass, Furubira-cho, Furu-u-gun, Shakotan-cho ssp. munakatai (Ishikawa, 1968) SW-Hokkaido:Hiyama, Oshima, S-Shiribeshi; Hiyama/Oshima:Mt.Daisengendake =furumii Mandl, 1982 Hiyama/Shiribeshi:Mt.Karibayama =hashinaukamui (Nakajima, 1993) SW-Hokkaido:Kamiiso-gun/Hiyama-gun:Umezuke-togé Pass =icalari (Nakajima, 1993) SW-Hokkaido:Yamakoshi-gun/Nisi-gun:Unseki-togé Pass =kimunkamui (Nakajima, 1993) SW-Hokkaido:Nishi-gun/Kudoo-gun/Setana-gun:Mt.Yurappu-daké =kunikanei (Nakajima, 1993) SW-Hokkaido:Matsumae-gun:Mt.Shirakami-daké =okikulumi (Nakajima, 1993) SW-Hokkaido:Nisi-gun/Yamakoshi-gun:Mt.Otobe-daké =pilicasius (Nakajima, 1993) SW-Hokkaido:Shimamaki-gun:Chiwasé =ponlui (Nakajima, 1993) SW-Hokkaido:Shimamaki-gun:Ohira-gawa River drainage middle reaches =shiuni (Nakajima, 1993) SW-Hokkaido:Yamakoshi-gun:Seiyobetsu =suganoe (Nakajima, 1993) SW-Hokkaido:Kameda-gun/Hiyama-gun:Nakayama-togé Pass =tagawai (Nakajima, 1993) SW-Hokkaido:Yamakoshi-gun/Hiyama-gun:nr.Atsu-un Tunnel =teunni (Nakajima, 1993) SW-Hokkaido:Kudoo-gun:Mt.Matsukura-yama =australofinis (Nakajima, 1994) SW-Hokkaido:Matsumae-gun:Fukushima-cho:Mt.Shirakami-daké ssp. nishijimai (Imura, 1991) SW-Hokkaido:Shiribeshi:Shimamaki-gun:Mt.Obira-yama =ninetopa (Nakajima, 1993) SW-Hokkaido:Shimamaki-gun:Ohira =origin (Nakajima, 1993) SW-Hokkaido:Shimamaki:low altitudinal zone =pulepilica (Nakajima, 1993) SW-Hokkaido:Shimamaki-gun:Mt.Obira-yama 4 leechi Bates, 1888 Korea ssp. leechi Bates, 1888 CE-Korea:"Gensan"(=Wonsan); 1f ssp. colasellus Deuve, 1990 NE-Korea:"Seishin"(=Chongjin) ssp. yooni Deuve, 1990 CE-Korea:Taebaek-san Mt.R.(Mt.Sorak-san) ssp. auvrayorum Deuve & Mourzine, 1993 NE-Korea:Hamgyong-namdo:50km ENE of Hungnam:Sinp’o ssp. chonmasanensis Deuve, 1993 NW-Korea:NE of Sinuiju:Mt.Chonma-san ssp. myohyangensis Deuve & Li, 1998 NC-Korea:Hyangsan:Mt.Myohyang-san; 1m/1f ssp. huisaensis Deuve & Li, 1999 NC-Korea:Hamgyong-namdo:Mt.Huchi/Huisa ssp. samjiyonensis Deuve & Li, 2000 N-Korea:Yanggang Do:Samjiyon San ssp. chiricola (Kwon & Lee, 1984) SC-Korea:Chiri-san Massif(Mt.Chiri-san) 5 lopatini Morawitz, 1886 S-Sakhalin(W-coast); Moneron Island; ?C-Sakhalin:Onor:Khondasa River 6 mirabilissimus (Ishikawa & Deuve, 1982) Korea ssp. mirabilissimus (Ishikawa & Deuve, 1982) CE-Korea:Taebaek-san Mt.R.:Mt.Taebaek-san, Mt.Sorak, Mt.Sobaek, Mt.Odae-san ~furumianus Mandl, in litt. - ssp. kana Imura, 1989 SC-Korea:Chollabuk-to:E of Chongju:Togyu-san Massif ssp. igniferescens Deuve, 1992 CE-Korea:N-Taebaek-san Mt.R.:Kangwon-do:Kumgang-san Massif:Mammul-san ssp. rigoutianus Deuve & Li, 1998 N-Korea:Pyongan-pukto:"Kuyang:Mt.Ryongmo San"(=Kujang env.); 1f 7 schrencki Motschulsky, 1860 Russia:Far East; NE-China:Heilongjiang, Jilin, Liaoning; N-Korea ssp. schrencki Motschulsky, 1860 Russia:Vladivostok, Khabarovsk; E-Heilongjiang, E-Jilin; NE-Korea; "Amur nr.Djaré" =hauryi Géhin, 1885 ?Rimsky-Korsakov Archipelago(=SW of Vladivostok) ~pyrrhophorus Hauser, 1921, nec pyrophorus Kraatz, 1889 ~reductus Hauser, 1921, nec Roeschke, 1896 =fastuosior Deuve, 1990 "Transbaikalia" ssp. seonhyeongae (Kwon & Lee, 1984) NC-Korea:Pyonganbukto:Huch’ang env.; ?Nangnim Mts. ssp. kimieae (Kezuka & Imura, 1989) E-Liaoning:Benxi env.(Mt.Maokui-shan) =qianshanensis (Li, 1992) E-Liaoning:Qian Shan Mt.R.(=Anshan env.)

26 =huairenensis Deuve & Li, 1998 E-Liaoning:"Huai-Ren Xian:Pulepuzhen:Niumaogou, 800m"(=Huanren Xian) ssp. hisakai (Imura, 1990) E-Liaoning:Fengcheng env.:Mt.Feng-Huang

DAMASTER Kollar, 1836; type species Carabus blaptoides Kollar =Adamaster Reitter, 1896; type species Carabus rugipennis Motschulsky species: 1

1 blaptoides Kollar, 1836 Japan, Kuriles; ?Taiwan ssp. blaptoides Kollar, 1836 S,C-Honshu; Kyushu, Shikoku, Iki Island; ?LNF =lewisi Rye, 1872 Kyushu(?Simabara), S-Honshu(?Hyogo) =goliath Morawitz, 1886 ~lineatipennis Hauser, 1921; un =multiseriatus Hauser, 1921 =pseudooxuroides Hauser, 1921 =hanae Chu, 1967 "Taiwan:Taichung pref.:Chia-pao-tai, 1150m"(?error); 1m; =brevicaudus Imura & Mizusawa, 1995 SW-Honshu:Shimane:Oki Islands:Dogo Island:Goka-mura:Fuku-ura ssp. rugipennis Motschulsky, 1861 Hokkaido(Hakodate); S,C-Kuriles =auricollis Waterhouse, 1867 Hokkaido:Hakodate =anurus Semenov, 1898 Japan ssp. fortunei Adams, 1861 N-Honshu:Tabu-shima Island(=Awa-shima) ssp. viridipennis Lewis, 1880 N-Honshu:Aomori ~hauserianus Born, 1922, nec Reitter, 1896 ?N-Honshu:Iwate:Karumai =niponicus Csiki, 1927; nom.pro hauserianus Born ?N-Honshu:Iwate:Karumai ssp. capito Lewis, 1881 Sado Island =formosanus Uchida & Tamanuki, 1927 "Taiwan:Mt.Funkiko"(?error) ssp. babaianus Ishikawa, 1983; nom.pro montanus Baba C-Honshu; ?LNF ~montanus Baba, 1939, nec Géhin, 1882 ?LNF ssp. oxuroides Schaum, 1862 C-Honshu; ?LNF =pandurus Bates, 1873 =cyanostola Lewis, 1882 C-Honshu:Tochigi:Mts.nr.Chiuzenji N of Nikko ~swinhoei Oberthür, 1883, nec Bates, 1870 "Formosa"(?error) ~angulatus Hauser, 1921, nec Fabricius, 1781 ~multiseriatus Hauser, 1921, nec Hauser, 1921 =subreticulatus Hauser, 1921 =viridipennis Hauser, 1921 =matsumurai Csiki, 1927; nom.pro swinhoei Oberthür "Formosa"(?error) ~multiordinatus Csiki, 1927; un; nom.pro multiseriatus Hauser =shiranensis Kanda, 1928 C-Honshu:Gunma/Tochigi:Mt.Shirane-san =paraoxuroides Baba, 1938 =pseudofortunei Baba, 1938

27 HYBRIDS

MESOCARABUS urculoensis Nicolas, 1919; hyb Mesocarabus problematicus/lusitanicus probletanicus Meyer, 1992; hyb Mesocarabus problematicus/lusitanicus AULONOCARABUS ~kusnetzovi Shilenkov, 1996, nec Semenov, 1903; =careniger victorianus Obydov, 1997; ?hyb careniger/canaliculatus jankowskiellus ?victorianus Obydov, 1997; nom.pro kusnetzovi Shilenkov, 1996; ?Aulonocarabus careniger ssp.; ?hyb careniger/canaliculatus jankowskiellus PLATYCARABUS gspani Kobmann, 1925; hyb Platycarabus creutzeri/irregularis konscheggi Born, 1912; hyb Platycarabus creutzeri/irregularis pseudonothus Kraatz, 1878; hyb Platycarabus creutzeri/irregularis ?meschniggi Born, 1920; ?hyb fabricii/depressus sustenensis Born, 1920; hyb Platycarabus fabricii/depressus EOTRIBAX ?celsus Semenov, 1898; ?hyb Eotribax eous/Alipaster pupulus; =?eous eous Morawitz dissimulatus Deuve, 1991; hyb Eotribax valikhanovi/Alipaster pupulus ?karkarensis Kabak & Ovchinnikov, 1994; Eotribax sp.; ?hyb Eotribax valikhanovi/Alipaster pupulus ?manap Březina & Kabak, 1993; Eotribax sp.; ?hyb Eotribax eous/Alipaster pupulus siniaevi Deuve, 1993; hyb Eotribax eous/Alipaster pupulus ?eugeniellus Obydov, 2000; ?hyb Eotribax eous/Alipaster pupulus EOTRIBAX/CRATOCECHENUS wrzecionkoi Deuve, 1991; hyb Eotribax valikhanovi/Cratocechenus akinini puellus CRATOCECHENUS niedli Gottwald, 1987; hyb Cratocechenus akinini loudai/Cratocephalus corrugis PANTOPHYRTUS/CRATOCEPHALUS ?kleinfeldorum Kabak & Putchkov, 1995; Leptoplesius sp.; ?hyb Pantophyrtus turcomanorum/Cratocephalus corrugis evstigneevi Obydov, 1997; ?hyb Pantophyrtus brachypedilus/Cratocephalus cicatricosus PANTOPHYRTUS/DEROPLECTES ?alajensis Semenov, 1896; nom.pro punctatostriatus Heyden; Pantophyrtus sp.; ?hyb turcomanorum/staudingeri TRIBAX ?balkaricus Belousov & Abdurachmanov, 1991; Tribax sp.; ?hyb Tribax kasbekianus/fossiger MACROTHORAX olcesei Putzeys, 1872; hyb Macrothorax rugosus rugosus/morbillosus aumonti INIOPACHUS/CHRYSOCARABUS bigorrensis (Camard, 1995); hyb Iniopachus pyrenaeus/Chrysocarabus auronitens CHRYSOCARABUS andronnensis Nicolas, 1923; hyb Chrysocarabus splendens/auronitens barthei Le Moult, 1912; hyb Chrysocarabus splendens/auronitens boreli Le Moult, 1912; hyb Chrysocarabus splendens/auronitens breiti Barthe, 1913; nom.pro barthei Le Moult; hyb Chrysocarabus splendens/auronitens clarofemoratus Nicolas, 1924; hyb Chrysocarabus splendens/auronitens esquieryensis Nicolas, 1923; hyb Chrysocarabus splendens/auronitens eugenieae Raynaud, 1936; hyb Chrysocarabus splendens/auronitens faustulus Lapouge, 1910; hyb Chrysocarabus splendens/auronitens ignipennis Le Moult, 1912; hyb Chrysocarabus splendens/auronitens kricheldorffi Le Moult, 1913; hyb Chrysocarabus splendens/auronitens lanieli Le Moult, 1912; hyb Chrysocarabus splendens/auronitens lemoulti Lapouge, 1910; hyb Chrysocarabus splendens/auronitens lucaniusae Fradois, 1940; hyb Chrysocarabus splendens/auronitens nigricornis Lebis, 1926; hyb Chrysocarabus splendens/auronitens pseudofaustulus Le Moult, 1912; hyb Chrysocarabus splendens/auronitens pseudoholochrysus Le Moult, 1912; hyb Chrysocarabus splendens/auronitens pseudopumicatus Le Moult, 1912; hyb Chrysocarabus splendens/auronitens ~rufofemoratus Barthe, 1924; hyb Chrysocarabus splendens/auronitens ~rugosus Le Moult, 1912, nec Fabricius, 1792; hyb Chrysocarabus splendens/auronitens salvanerensis Raynaud, 1970; hyb Chrysocarabus splendens/auronitens vacheri Basilewsky, 1931; nom.pro rufofemoratus Barthe; hyb Chrysocarabus splendens/auronitens vulcanus Le Moult, 1912; hyb Chrysocarabus splendens/auronitens annae Raynaud, 1936; hyb Chrysocarabus lineatus/auronitens lorenzoi Raynaud, 1968; hyb Chrysocarabus lineatus/auronitens arraizensis Raynaud, 1938; hyb Chrysocarabus lineatus/splendens bartheensis Nicolas, 1923; hyb Chrysocarabus lineatus/splendens jeannae Raynaud, 1936; hyb Chrysocarabus lineatus/splendens portoi Raynaud, 1968; hyb Chrysocarabus lineatus/splendens CHRYSOCARABUS/CHRYSOTRIBAX bugareti Jeannel, 1941; hyb Chrysocarabus auronitens/Chrysotribax hispanus helenae Mollard, 1992; ?emend.pro helenais Mollard; hyb Chrysocarabus auronitens/Chrysotribax rutilans ~helenais Mollard, 1992; ios; hyb Chrysocarabus auronitens/Chrysotribax rutilans olieri Raynaud, 1943; hyb Chrysocarabus splendens/Chrysotribax hispanus ~croesus Oberthür, 1883, nec Fabricius, 1801; hyb Chrysotribax rutilans/hispanus MEGODONTUS deubeli Reitter, 1896; hyb Megodontus violaceus/planicollis ~altitudinorum Breuning, 1934; ios; ?hyb exaratus/septemcarinatus ?altitudinum Breuning, 1934; emend.pro altitudinorum Breuning; ?hyb exaratus/septemcarinatus ?lebedewi Lutshnik, 1927; nom.pro seriatus Motschulsky; ?exaratus Quensel; ?hyb Megodontus exaratus/violaceus aurolimbatus ~seriatus Motschulsky, 1850, nec Wiedemann, 1821; ?exaratus Quensel; ?hyb Megodontus exaratus/violaceus aurolimbatus PROCRUSTES/OXYCARABUS ~incertus Haury, 1885, nec Motschulsky, 1846; hyb Procrustes coriaceus/Oxycarabus saphyrinus DEROPLECTES ?darvazensis Deuve, 1990; ?dokhtouroffi dokhtouroffi Ganglbauer; ?hyb Deroplectes dokhtouroffi/sphinx

28 ACOPTOLABRUS/DAMASTER kamuiterrestris (Nakajima, 1994); hyb Acoptolabrus gehinii munakatai/Damaster blaptoides rugipennis lamtui (Nakajima, 1993); hyb Acoptolabrus gehinii munakatai/Damaster blaptoides rugipennis

FOSSIL FORMS andrzejowskii Lomnicki, 1914; fos; violaceus Linnaeus arietinus Lapouge, 1902; fos; ?violaceus Linnaeus dzieduszyckii Lomnicki, 1894; fos; glabratus Paykull ~heeri Scudd., 1895, nec Germar, 1824; fos; ?Carabus (Oreocarabus) sp.; ?Calosoma sp. humerosus Lapouge, 1902; fos; nitens Linnaeus maeandroides Lomnicki, 1894; fos; cancellatus Illiger malacopterus Lapouge, 1902; fos; nemoralis Müller neli Deuve, 1998; fos; ?Morphocarabus sp. orcinus Lapouge, 1902; fos; violaceus violaceus Linnaeus praearvensis Lomnicki, 1894; fos; arcensis arcensis Herbst praeviolaceus Lomnicki, 1894; fos; violaceus Linnaeus sangamon Wickh., 1917; fos; maeander Fischer thuerachi Flach, 1884; fos; menetriesi Hummel winkleri Weyenberg, 1869; fos; ?Carabus sp.

INCERTAE SEDIS adil Bogachev, 1936; inc amauryi Haury, 1898; inc; ?artifact ~collaris Motschulsky, 1850, nec Herbst, 1784; inc densepunctatus Mandl, 1955; inc; ?Semnocarabus sp. microchondrus Fischer, 1842; inc; odoratus ?dohrni Gebler simardianus Deuve, 1990; inc; ?artifact; Tomocarabus sp.

Note: All available names, including synonyms (except permanently invalid names, such as junior homonyms) are listed in bold italics. Unavailable names and permanently invalid names are listed in italics. GENUS GROUP TAXA

Acarabus Semenov, 1890; type species grombczewskii Semenov; =Goniocarabus Géhin Acathaicus Reitter, 1896; type species idolon Semenov; Carabus subgenus; p.81 Acoptolabrus Morawitz, 1886; type species schrencki Motschulsky; Carabus subgenus; p.91 Acoptopterus Lapouge, 1927; type species vigilax Bates; =Archaeocarabus Semenov Acrocarabus Lapouge, 1930; type species guerini Fischer; Carabus subgenus; p.23 Adamaster Reitter, 1896; type species rugipennis Motschulsky; =Damaster Kollar Adelocarabus Reitter, 1896; type species semiopacus Reitter; =Leptocarabus Géhin Ainocarabus Mandl, 1973; type species aino Rost; =Megodontus Solier Alagocarabus Morawitz, 1886; type species caerulans Morawitz; =Goniocarabus Géhin Alipaster Reitter, 1896; type species pupulus Morawitz; Carabus subgenus; p.52 Allocarabus Lapouge, 1921; type species aurocinctus Motschulsky; =Diocarabus Reitter Amorphocarabus Lapouge, 1930; type species henningi Fischer; =Morphocarabus Reitter Ancylocarabus Reitter, 1896; type species tarbagataicus Kraatz; =Morphocarabus Géhin Anthracocarabus Lapouge, 1930; type species erosus Motschulsky; =Semnocarabus Reitter Apatocarabus Semenov, 1898; type species favieri Fairmaire; =Eurycarabus Géhin Apocarabus Lapouge, 1930; type species stscheglowi Mannerheim; =Carabus Géhin Apoplesius Deuve, 1990; type species omphreodes Reitter; Carabus subgenus; p.80 Apostocarabus Reitter, 1896; type species odoratus Motschulsky; =Morphocarabus Géhin Apotomopterus Hope, 1838; type species prodigus Erichson; Carabus subgenus; p.23 Aptocarabus Reitter, 1896; type species rossii Dejean; =Archicarabus Seidlitz Araeocarabus Reitter, 1896; type species roborowskii Semenov; =Rhigocarabus Reitter Archaeocarabus Semenov, 1898; type species relictus Semenov; Carabus subgenus; p.3 Archicarabus Seidlitz, 1887; type species nemoralis Müller; Carabus subgenus; p.20 Archiplectes Gottwald, 1982; type species komarowi Reitter; Carabus subgenus; p.57 Aristocarabus Semenov, 1896; type species romanowi Semenov; Carabus subgenus; p.86 Asthenocarabus Lapouge, 1930; type species opaculus Putzeys; =Diocarabus Reitter Aulacocarabus Géhin, 1876; type species septemcarinatus Motschulsky; =Megodontus Solier Aulonocarabus Reitter, 1896; type species canaliculatus Adams; Carabus subgenus; p.46 Autocarabus Seidlitz, 1887; type species auratus Linnaeus; =Tachypus Weber Axinocarabus Morawitz, 1886; type species melanochrus Morawitz; Carabus subgenus; p.81

Bactrocarabus Semenov, 1927; type species bokharensis Csiki (=theanus Reitter); =Ulocarabus Reitter Basilicocarabus Lapouge, 1930; type species regalis Fischer; =Morphocarabus Reitter Baudiicarabus Ganglbauer, 1909; type species cychroides Baudi; =Platycarabus Morawitz

Callistocarabus Reitter, 1896; type species marginalis Fabricius; =Tomocarabus Reitter Calocarabus Semenov, 1887; type species gratus Semenov; Carabus subgenus; p.82 Calocechenus Reitter, 1896; type species malachiticolor Reitter (=przewalskii Morawitz); =Calocarabus Semenov Cancellocarabus Lutshnik, 1924; nom.pro Goniocarabus Reitter; =Tachypus Weber Carabulus Lutshnik, 1924; type species ermaki Lutshnik; =Megodontus Solier Carabus Linnaeus, 1758; type species granulatus Linnaeus; Carabini genus; p.3; Carabus subgenus; p.4 Carpathophilus Reitter, 1896; type species linnei Panzer; =Oreocarabus Géhin Cathaicodes Reitter, 1896; type species alexandrae Semenov; =Acathaicus Reitter Cathaicus Bates, 1870; type species swinhoei Bates (=brandti Faldermann); Carabus subgenus; p.81 Cathoplius Thomson, 1875; type species asperatus (Dejean); Carabus subgenus; p.61 Caucasocarabus Lapouge, 1930; type species koenigi Ganglbauer; =Pachycarabus Géhin Cavazzutiocarabus Imura, 1998; type species latreilleanus Csiki; Carabus subgenus; p.42 Cechenochilus Motschulsky, 1850; nom.pro Cechenus Fischer; Carabus subgenus; p.54 Cechenotribax Semenov & Znojko, 1932; type species petri Semenov & Znojko; Carabus subgenus; p.52 Cechenus Fischer, 1822, nec Illiger, ?; type species boeberi Adams; =Cechenochilus Motschulsky, 1850 Cephalornis Semenov, 1889; nom.pro Ornitocephalus Semenov; Carabus subgenus; p.82 Chaetocarabus Thomson, 1875; type species intricatus Linnaeus; Carabus subgenus; p.50 Chaetogaster Lapouge, 1930; type species saulcyi Piochard; =Lamprostus Motschulsky Chaetomelas Thomson, 1875; type species ehrenbergi Klug (=praestigiator Morawitz); =Procrustes Bonelli Chaetoprostus Reitter, 1896; type species hemprichi Dejean; =Lamprostus Motschulsky Chrysocarabus Thomson, 1875; type species auronitens Fabricius; Carabus subgenus; p.68 Chrysotribax Reitter, 1896; type species hispanus Fabricius; Carabus subgenus; p.68 Coptolabrinus Reitter, 1897; type species pustulifer Lucas; =Coptolabrus Solier Coptolabrodes Březina & Imura, 1997; type species haeckeli Březina & Imura; Carabus subgenus; p.90 Coptolabrus Solier, 1848; type species smaragdinus Fischer; Carabus subgenus; p.86 Cratocarabus Reitter, 1896; type species puer Morawitz; Carabus subgenus; p.52 Cratocechenus Reitter, 1896; type species akinini Morawitz; Carabus subgenus; p.53 Cratocephalus Kirsch, 1859; type species songoricus Kirsch; =cicatricosus Fischer; Carabus subgenus; p.53 Cratocranion Semenov, 1898; type species benjamini Semenov; =Pseudocranion Reitter Cratophyrtus Reitter, 1896; type species segregatus Morawitz; Carabus subgenus; p.54 Cryocarabus Lapouge, 1930; type species chamissonis Fischer; =Diocarabus Reitter Cryptocarabus Reitter, 1896; type species lindemanni Ballion; Carabus subgenus; p.19 Cryptocechenus Semenov, 1898; type species sifanicus Semenov; Carabus subgenus; p.82 Ctenocarabus Thomson, 1875; type species galicianus Gory; Carabus subgenus; p.61 Cupreocarabus Deuve, 1997; type species huangi Deuve; Carabus subgenus; p.84 Cyclocarabus Reitter, 1896; type species namanganensis Heyden; Carabus subgenus; p.20 Cychrocarabus Morawitz, 1889; nom.pro Ornitocephalus Semenov; =Cephalornis Semenov

30 Cychrocephalus Géhin, 1875; type species asperatus (Dejean); =Cathoplius Thomson Cychrostomus Reitter, 1896; type species anchocephalus Reitter; Carabus subgenus; p.82 Cytilocarabus Reitter, 1896; type species cribratus Quensel; Carabus subgenus; p.44

Damaster Kollar, 1836; type species blaptoides Kollar; Carabus subgenus; p.92 Deroplectes Reitter, 1895; type species sphinx Reitter; Carabus subgenus; p.80 Deuterocarabus Reitter, 1896; type species montivagus Palliardi; =Archicarabus Seidlitz Dichocarabus Reitter, 1896; type species rugosus Fabricius; =Macrothorax Desmarest Diocarabus Reitter, 1896; type species loschnikovi Fischer; Carabus subgenus; p.28 Dolichocarabus Lapouge, 1927; type species delavayi Fairmaire; =Apotomopterus Hope Dorcarabus Reitter, 1896; cos; type species morbillosus Fabricius; =Macrothorax Desmarest Dorcocarabus Reitter, 1896; iss of Dorcarabus Reitter Dysmictocarabus Puisségur, 1964; type species solieri Dejean; =Chrysocarabus Thomson

Eccoptolabrus Semenov, 1898; type species exiguus Semenov; Carabus subgenus; p.86 Entomopterus Lapouge, 1927; type species mecynodes: Lapouge non Bates (=achilles Hauser); =Apotomopterus Hope Eocarabus Semenov, 1898; type species jankowskii Oberthür; =Coptolabrus Solier Eocechenus Semenov & Znojko, 1932; type species kaznakovi Semenov & Znojko; Carabus subgenus; p.82 Eotribax Semenov, 1898; type species eous Morawitz; Carabus subgenus; p.52 Eremocarabus Lapouge, 1930; type species sculptipennis Chaudoir; =Scambocarabus Reitter Eucarabus Géhin, 1885; type species ullrichi Germar; Carabus subgenus; p.5 Eucoptolabrus Semenov, 1898; type species gemmifer Fairmaire; =Coptolabrus Solier Euleptocarabus Nakane, 1956; type species porrecticollis Bates; Carabus subgenus; p.27 Eupachycechenus Semenov, 1898; type species riffensis Fairmaire; =Mesocarabus Thomson Eupachys Chaudoir, 1857; type species glyptopterus Fischer; Carabus subgenus; p.82 Euporocarabus Reitter, 1896; type species hortensis Linnaeus; =Oreocarabus Géhin Eurycarabus Géhin, 1885; type species numidus Castelnau; Carabus subgenus; p.33 Eutelocarabus Reitter, 1896; type species arcensis Herbst; =Carabus Linnaeus

Fulgenticarabus Deuve & Li, 1998; type species flutschi Deuve & Li; Carabus subgenus; p.81

Gigantocarabus Semenov, 1898; type species gebleri Fischer; =Morphocarabus Géhin Goniocarabus Géhin, 1885; nom.pro Goniognathus Kraatz; Carabus subgenus; p.81 Goniocarabus Reitter, 1896, nec Géhin, 1855; type species cancellatus Illiger; =Tachypus Weber Goniognathus Kraatz, 1883, nec Agassiz, ?; type species gracilis Kraatz (=sogdianus Semenov); =Goniocarabus Géhin, 1885 Gnathocarabus Deuve, 1991; type species kusnetzovi Semenov; Carabus subgenus; p.23

Hadrocarabus Thomson, 1875; type species latus Fabricius; =Mesocarabus Thomson Haplocarabus Semenov, 1898; type species confinis Semenov (=variabilis Ballion); =Ophiocarabus Reitter Hemicarabus Géhin, 1885; type species nitens Linnaeus; Carabus subgenus; p.27 Heptacarabus Imura, 1998; type species ohshimaianus Deuve; =Qinlingocarabus Imura Heterocarabus Morawitz, 1886; type species marietti Cristoforis & Jan; Carabus subgenus; p.51 Homoeocarabus Reitter, 1896; type species maeander Fischer; Carabus subgenus; p.27 Hygrocarabus Thomson, 1875; type species nodulosus Fabricius; Carabus subgenus; p.51 Hypsocarabus Semenov, 1898; type species latro Semenov; Carabus subgenus; p.44

Imaibiodes Deuve, 1991; type species businskyi Deuve; Carabus subgenus; p.85 Imaibius Bates, 1889; type species barysomus Bates; Carabus subgenus; p.79 Indocarabus Lapouge, 1930; type species wagae Fairmaire; =Neoplesius Reitter Iniopachus Solier, 1848; cos; type species pyrenaeus Audinet-Serville; Carabus subgenus; p.67 Iniopachys Solier, 1848; iss of Iniopachus Solier Ischnocarabus Kraatz, 1877; type species tenuitarsis Kraatz; Carabus subgenus; p.23 Isiocarabus Reitter, 1896; type species fiduciarius Thomson; Carabus subgenus; p.9

Koreacoptolabrus Kwon & Lee, 1984; type species constricticollis Kraatz; =Acoptolabrus Morawitz

Lamprocarabus Thomson, 1875; type species humboldti Faldermann; =Lipaster Motschulsky Lamprostus Motschulsky, 1865; type species spinolae Cristoforis & Jan; Carabus subgenus; p.65 Laocarabus Lapouge, 1916; type species vitalisi Lapouge; =Apotomopterus Hope Lasiocoptolabrus Imura, 1993; type species sunwukong Imura; Carabus subgenus; p.86 Leptinocarabus Reitter, 1896; type species acutesculptus Chaudoir (=venustus Morawitz); =Morphocarabus Géhin Leptocarabus Géhin, 1885; type species procerulus Chaudoir; Carabus subgenus; p.47 Leptoplesius Reitter, 1896; type species marquardti Reitter; Carabus subgenus; p.53 Lichnocarabus Reitter, 1896; type species vinctus Weber; Carabus subgenus; p.5 Limnocarabus Géhin, 1876; type species clathratus Linnaeus; Carabus subgenus; p.26 Liocarabus Reitter, 1896; type species coarctatus Brullé; =Nesaeocarabus Bedel Lipaster Motschulsky, 1865; type species stjernvalli Mannerheim; Carabus subgenus; p.20 Lithariophorus Mandl, 1985; type species caschmirensis Redtenbacher; =Imaibius Bates Loxocarabus Reitter, 1896; type species obsoletus Sturm; =Eucarabus Géhin Lyperocarabus Lapouge, 1930; type species estreicheri Fischer; =Trachycarabus Géhin

Macrogenus Motschulsky, 1846; type species clypeatus Adams; =Procrustes Bonelli Macrothorax Desmarest, 1850; type species aumonti Lucas; Carabus subgenus; p.67 Makalubius Mandl, 1985; type species arunensis Heinertz; =Meganebrius Kraatz Meganebrius Kraatz, 1895; type species indicus Fairmaire; Carabus subgenus; p.48 Megodontoides Deuve, 1991; type species erwini Mandl; Carabus subgenus; p.86 Megodontus Solier, 1848; type species caelatus Fabricius; Carabus subgenus; p.75 Melancarabus Thomson, 1875; type species hungaricus Fabricius; =Pachystus Motschulsky Mesocarabus Thomson, 1875; type species catenulatus: Duftschmid, non Scopoli (=problematicus Herbst); Carabus subgenus; p.38 Microcarabus Ishikawa, 1979; type species subparallelus Ballion; =Cryptocarabus Reitter Microplectes Reitter, 1896; type species riedeli: Reitter, non Ménétriés, =argonautarum Semenov; Carabus subgenus; p.59 Microtribax Gottwald, 1982; type species fausti Dohrn; Carabus subgenus; p.55 Mimocarabus Géhin, 1885; type species maurus (Adams); Carabus subgenus; p.18 Morphocarabus Géhin, 1885; type species monilis Fabricius; Carabus subgenus; p.11

31 Nabicarabus Kwon & Lee, 1984; type species vietinghoffi Adams; =Megodontus Solier Nemoralus Schuler, 1976; type species nemoralis Müller; =Archicarabus Seidlitz Neocarabus Lapouge, 1930; type species taedatus Fabricius; Carabus subgenus; p.28 Neocarabus Hatch, 1949, nec Lapouge, 1930; type species granulatus Linnaeus; =Carabus Linnaeus Neodamaster Nakajima, 1993; type species lamtui (Nakajima); hyb Acoptolabrus gehinii/Damaster blaptoides Neoplectes Reitter, 1885; nom.pro Plectes: Fischer, 1823, nec Fischer, 1821; type species ibericus Fischer; Carabus subgenus; p.57 Neoplesius Reitter, 1896; type species tanguticus Semenov; Carabus subgenus; p.83 Nesaeocarabus Bedel, 1895; type species interruptus Dejean (=abbreviatus Brullé); Carabus subgenus; p.33 Nesocoptolabrus Lapouge, 1930; type species fruhstorferi Roeschke; =Coptolabrus Solier Nigracoptolabrus Deuve, 1991; type species burmanensis Breuning; =Pseudocoptolabrus Reitter Nippocarabus Lapouge, 1930; type species vanvolxemi Putzeys; =Carabus Linnaeus

Ohomopterus Reitter, 1896; type species dehaanii Chaudoir; Carabus subgenus; p.9 Ophiocarabus Reitter, 1896; type species striatulus Géhin; Carabus subgenus; p.19 Oreocarabus Géhin, 1876; type species errans Gory (=amplipennis Lapouge) ; Carabus subgenus; p.43 Orientocarabus Kwon & Lee, 1984; type species billbergi Mannerheim; =Parhomopterus Lapouge Orinocarabus Kraatz, 1878; type species sylvestris Panzer; Carabus subgenus; p.41 Ornitocephalus Semenov, 1887, nec Sömmering, ?; type species potanini (Semenov); =Cephalornis Semenov, 1889 Oxycarabus Semenov, 1898; type species saphyrinus Cristoforis & Jan; Carabus subgenus; p.62

Pachycarabus Géhin, 1876; type species staehlini Adams; Carabus subgenus; p.41 Pachycechenus Semenov, 1898; type species solskyi Ballion; =Cratocephalus Kirsch Pachycranion Solier, 1848; type species schoenherri Fischer; =Megodontus Solier Pachycranius Solier, 1848; iss of Pachycranion Solier Pachystus Motschulsky, 1865; type species hungaricus Fabricius; Carabus subgenus; p.32 Pagocarabus Morawitz, 1886; type species diruptus Morawitz; Carabus subgenus; p.85 Pancarabus Reitter, 1896; type species aeruginosus Fischer; =Morphocarabus Géhin Pantophyrtus Thieme, 1881; type species turcomanorum Thieme; Carabus subgenus; p.54 Paracarabus Reitter, 1896; type species planatus Chaudoir; =Macrothorax Desmarest Paracarabus Lapouge, 1930, nec Reitter, 1896; type species granulatus Linnaeus; =Carabus Linnaeus Parameganebrius Mandl, 1970; type species eberti Mandl (=lebretae Colas); =Meganebrius Kraatz Paraplesius Morawitz, 1886, nec ?, ?; type species staudingeri Morawitz; =Deroplectes Reitter, 1895 Parhomopterus Lapouge, 1930; type species sternbergi Reitter; Carabus subgenus; p.8 Parimaibius Breuning, 1958; type species gridellii Breuning; =Imaibius Bates Pentacarabus Ishikawa, 1972; type species harmandi Lapouge; =Diocarabus Reitter Phricocarabus Reitter, 1896; type species glabratus Paykull; =Oreocarabus Géhin Piocarabus Reitter, 1896; type species vladimirskyi Dejean; Carabus subgenus; p.44 Platycarabus Morawitz, 1886; type species depressus Bonelli; Carabus subgenus; p.49 Platychrus Kolenati, 1845; type species puschkini Adams; =Tribax Fischer Plectes: Fischer, 1823, non Fischer, 1821; =Neoplectes Reitter, 1885 Plectocarabus Deuve, 1991; nom.pro Neoplectes: Gottwald; =Neoplectes Reitter Plesius Semenov, 1905; nom.pro Paraplesius Morawitz; =Deroplectes Reitter Procerus Dejean, 1828; type species scabrosus: Fabricius, 1801, non Olivier, 1795 (=gigas Creutzer); Carabus subgenus; p.73 Procrustes Bonelli, 1809; type species coriaceus Linnaeus; Carabus subgenus; p.62 Procrusticus White, 1845; type species payafa White; =Procrustes Bonelli Procrustides Semenov, 1905; type species bessarabicus Fischer; =Tomocarabus Reitter Procrustocarabus Géhin, 1876; type species impressus Klug; =Procrustes Bonelli Promorphocarabus Reitter, 1896; type species gebleri Fischer; =Morphocarabus Géhin Proteocarabus Géhin, 1876; type species violaceus Linnaeus; =Megodontus Solier Protodamaster Imura & Kezuka, 1989; type species aesculapius (Imura & Kezuka); =Pseudocoptolabrus Reitter Pseudocechenus Morawitz, 1886; type species irregularis Fabricius; =Platycarabus Morawitz Pseudocoptolabrus Reitter, 1896; type species taliensis (Fairmaire); Carabus subgenus; p.85 Pseudocranion Reitter, 1896; type species gansuensis Semenov; Carabus subgenus; p.84 Pseudoprocrustes Morawitz, 1886; type species durvillei Dupont (=punctatus Castelnau); =Lamprostus Motschulsky Pseudotribax Kraatz, 1884; type species validus Kraatz; Carabus subgenus; p.54

Qinlingocarabus Imura, 1998; type species kitawakianus Imura; Carabus subgenus; p.45

Relictocarabus Ledoux, 1984; type species meurguesae (Ledoux, 1984) (=meurguesianus Ledoux); Carabus subgenus; p.81 Rhabdotocarabus Seidlitz, 1887; type species melancholicus Fabricius; =Ctenocarabus Thomson Rhigocarabus Reitter, 1896; type species morawitzianus Semenov; Carabus subgenus; p.45 Rhigoidocarabus Deuve, 1997; type species zhubajie Imura; Carabus subgenus; p.18 Rhipocarabus Reitter, 1896; type species alysidotus Illiger; Carabus subgenus; p.33

Sajanocarabus Lutshnik, 1924; type species kozhantschikowi Lutshnik; =Morphocarabus Géhin Scambocarabus Reitter, 1896; type species kruberi Fischer; Carabus subgenus; p.31 Sellaecarabus Sturani, 1947; type species olympiae Sella; =Chrysocarabus Thomson Semnocarabus Reitter, 1896; type species regulus Dohrn; Carabus subgenus; p.31 Shenocoptolabrus Imura, Zhou & Su, 1999; type species osawai Imura, Zhou & Su; Carabus subgenus; p.59 Shunichiocarabus Imura, 1995; type species uenoianus Imura; Carabus subgenus; p.85 Siamocarabus Imura, 1989; type species masumotoi (Imura); =Apotomopterus Hope Sigenacarabus Kleinfeld, 2000; type species: sigena Kleinfeld; =Neoplesius Reitter Sphodristocarabus Géhin, 1885; type species adamsi Adams; Carabus subgenus; p.59 Sphodristus Motschulsky, 1865; type species acuticollis Motschulsky; =Procrustes Bonelli Sphodristus Thomson, 1875, nec Motschulsky, 1865; type species varians Fischer; =Sphodristocarabus Géhin, 1885 Stephanocarabus Imura, 1995; type species fraterculoides Breuning; Carabus subgenus; p.30 Syzygocarabus Semenov, 1898; type species cateniger Morawitz; =Rhigocarabus Reitter

Tachycarabus Semenov, 1898; type species poeta Semenov; =Rhigocarabus Reitter Tachypus Weber, 1801; type species auratus Linnaeus; Carabus subgenus; p.34 Taiwanocarabus Imura & M.Satô, 1989; type species masuzoi (Imura & M.Satô); =Apotomopterus Hope Tanaocarabus Reitter, 1896; type species sylvosus Say; Carabus subgenus; p.40 Teratocarabus Semenov & Znojko, 1932; type species azrael Semenov & Znojko; Carabus subgenus; p.82 Titanocarabus Breuning, 1933; type species titanus Breuning; Carabus subgenus; p.82 Titanocechenus Breuning & Ruspoli, 1970; type species osellai Breuning & Ruspoli; =Lipaster Motschulsky

32 Tmesicarabus Reitter, 1896; type species cristoforii Spence; =Tachypus Weber Tomocarabus Reitter, 1896; type species convexus Fabricius; Carabus subgenus; p.29 Trachycarabus Géhin, 1885; type species scabriusculus Olivier; Carabus subgenus; p.17 Tribax Fischer, 1817; type species puschkini Adams; Carabus subgenus; p.55 Tropidocarabus Kraatz, 1895; type species opacus Kraatz (=dardiellus Bates); =Imaibius Bates Tylocarabus Reitter, 1896; type species cumanus Fischer; =Eucarabus Géhin

Ulocarabus Reitter, 1896; type species stschurovskii Solsky; Carabus subgenus; p.32

Weolseocarabus Kwon & Lee, 1984; type species koreanus Reitter; =Leptocarabus Géhin

Xystrocarabus Reitter, 1896; type species catenatus Panzer (=catenulatus Scopoli); =Eucarabus Géhin

Zoocarabus Reitter, 1896; type species bogdanowi Ballion; =Semnocarabus Reitter

SPECIES GROUP TAXA A aabadensis Lassalle, 1991; Sphodristocarabus rotundicollis ssp. abadzechus Zamotajlov, 1988; Tribax titan ssp. abagonensis Starck, 1894; Tribax circassicus ssp. abantayaginiensis Schweiger, 1962; =convexus erzeliki Schweiger abashicus Belousov, in litt.; ibericus ssp. abasinus Rost, 1893; =circassicus circassicus Ganglbauer abbreviatus Brullé, 1835; nom.pro interruptus Dejean; Neseaocarabus sp. abchasicus Motschulsky, 1850; =coriaceus excavatus Charpentier abchasicus Reitter, 1896, nec Motschulsky, 1850; =circassicus circassicus Ganglbauer abchasicus: auct., non Motschulsky; =clypeatus kurnakovi Kryzhanovskij, 1968 abdurachmanovi Belousov, in litt.; Microtribax nothus ssp. abruptus Lapouge, 1903; un; =koenigi Ganglbauer absentivus Belousov & Kabak, 1993; Pantophyrtus turcomanorum ssp. accuratus Chaudoir, 1848; =estreicheri Fischer acheicus Zamotajlov, 1991; =reitteri pshuensis Gottwald achillecasalei Deuve, 1997; Pseudocranion gansuensis ssp.. achilles Hauser, 1924; Apotomopterus grossefoveatus ssp. achunensis Gottwald, 1985; =reitteri reitteri Retowski acicularis Lapouge, 1902; un; =cancellatus carinatus Charpentier acorep Lassalle & Prunier, 1993; Apotomopterus longeantennatus ssp. acosmos Cavazzuti, 1997; Apotomopterus longeantennatus ssp. acuminatus Ménétriés, 1836, nec Olivier, 1790; =graecus morio Mannerheim acutangulatus Csiki, 1927; un; nom.pro acutangulus Eichler; =sibiricus errans Fischer acutangulus Chaudoir, 1879; Tomocarabus convexus ssp. acutangulus Eichler, 1924, nec Chaudoir, 1879; =sibiricus errans Fischer acutesculptus Chaudoir, 1877; nom.pro carinulatus Chaudoir; =venustus venustus Morawitz acuticollis Motschulsky, 1858; Procrustes payafa ssp. acutithorax Deuve, 1989; Eupachys sp. acutus Lapouge, 1904; =gilnickii gilnickii Deyrolle adamellicola Ganglbauer, 1904; Orinocarabus sp. adamsi Adams, 1817; Sphodristocarabus sp. adamsii (Fischer, 1817), nec adamsi Adams, 1817; =puschkini puschkini Adams adamsianus Mandl, 1975; =adamsi adamsi Adams adangensis Gottwald, 1983; Cechenochilus sp. adatarasanus Ishikawa,1966; Diocarabus harmandi ssp. adcharicus Gottwald, in litt.; Neoplectes ibericus ssp. adductus Cavazzuti & Ratti, 1998; Apotomopterus cantonensis ssp. adelaidae Starck, 1889; Archiplectes starckianus ssp. adelphus Rost, 1892; Tribax sp. adelphus Kolbe, 1912, nec Rost, 1892; =cancellatus cancellatus Illiger adelus Tarrier, 1974; un; =violaceus fiorii Born adeptus Kolbe, 1913; =cancellatus tibiscinus Csiki adil Bogachev, 1936; inc adjunctus Csiki, 1927; nom.pro conjunctus Born; =cantonensis Hauser adolescens Hauser, 1925; Apotomopterus sp. adonis Hampe, 1853; =arcadicus arcadicus Gistl adoxus Fischer, 1823; =estreicheri Fischer adriaenssensi Lassalle & Prunier, 1993; Coptolabrus pustulifer ssp. adspersarius Lapouge, 1917; un; =mulsantianus mulsantianus Morawitz adspersus Gmelin, 1788; =clathratus clathratus Linnaeus adsypschi Gottwald, 1985; Archiplectes reitteri ssp. adyg Zamotajlov, 1997; Cechenochilus boeberi ssp. aegyptiacus Motschulsky, 1844; =scabrosus ?scabrosus Olivier aemulus Fischer, 1823; =convexus convexus Fabricius aeneipennis Sturm, 1815; =scheidleri scheidleri Panzer aeneocupreus Hauser, 1932; Apotomopterus sp. aeneolus Morawitz, 1886; Ophiocarabus sp. aeneopunctatus DeGeer, 1774; =clathratus clathratus Linnaeus aenescens Depoli, 1929; un; =catenulatus catenulatus Scopoli aeneus Motschulsky, 1850; =scabrosus tauricus Bonnelli aeneus Letzner, 1850; un; =sylvestris sylvestris Panzer aequaliceps Reitter, 1896; Cechenochilus boeberi ssp. aequatus Motschulsky, 1845; =billbergi billbergi Mannerheim

33 aequatus Lapouge, 1902, nec Motschulsky, 1845; =rutilans rutilans Dejean aequicostatus Mandl, 1974; =franzi franzi Mandl aequicostalis Staven & Heinz, 1995; Sphodristocarabus rotundicollis ssp. aequilimes Semenov, 1898; =cateniger Morawitz aequistriatus Kraatz, 1878; =rothi rothi Dejean aeratus Géhin, 1885, nec Quensel in Schönherr, 1806; nom.pro ziegleri Géhin; =arcensis arcensis Herbst aeratus Culot, 1986; un; =clathratus clathratus Linnaeus aereicollis Hauser, 1921; Acoptolabrus gehinii ssp. aerescens Dalla Torre, 1877; un; =scheidleri scheidleri Panzer aereus Dejean, 1826; =aeruginosus aeruginosus Fischer aereus Dejean, 1826, nec Dejean, 1826; =arcensis arcensis Herbst aereus Hauser, 1919, nec Dejean, 1826; =augustus augustus Bates aeruginescens Dalla Torre, 1877; un; =scheidleri scheidleri Panzer aeruginosiformis Breuning, 1932; Morphocarabus spasskianus ssp. aeruginosus Fischer, 1822; Morphocarabus sp. aeruginosus Letzner, 1850; un; =sylvestris sylvestris Panzer aesculapius (Imura & Kezuka, 1989); Pseudocoptolabrus taliensis ssp. aethiops Cristoforis & Jan, 1837, nec Panzer, 1797; =graecus morio Mannerheim aetolicus Schaum, 1857; Carabus granulatus ssp. affictus Codina, 1913; un;=rutilans rutilans Dejean affinis Panzer, 1798, nec Schrank, 1781; =monilis Fabricius affinis Duftschmid, 1812, nec Schrank, 1781; =cancellatus emarginatus Duftschmid affinis Gistl, 1857, nec Schrank, 1781; =cancellatus excisus Dejean affinis (Kwon & Lee, 1984), nec Schrank, 1781; =fraterculus odaesaniculus Deuve, 1991 afflictus Csiki, 1927; un; nom.pro tristiculus Semenov, 1887; =imperfectus imperfectus Semenov agamemnon Breuning, 1943; Archaeocarabus sp. agassi Le Conte, 1850; cos; Neocarabus taedatus ssp. agassizi Le Conte, 1850; iss of agassi Le Conte agnatus Ganglbauer, 1889; Tribax sp. agrestis Creutzer, 1799; =scabriusculus scabriusculus Olivier ahascoanus Lapouge, 1924; =lusitanicus macrocephalus Dejean ahmetustaensis Cavazzuti, 1987; Lamprostus torosus ssp. aibgensis Starck, 1890; =starcki Heyden, 1885 aino Rost, 1908; Megodontus kolbei ssp. aishanenesis Kleinfeld, 1999; Cathaicus brandti ssp. aisrrensis Belousov, 1992; Archiplectes koltzei ssp. ajax Breuning, 1933; Apotomopterus sp. akaishiensis (Ishikawa, 1986); Diocarabus harmandi ssp. akane Imura & Yamaya, 1994; Coptolabrus formosus ssp. akbesianus Lapouge, 1914; un; =mulsantianus mulsantianus Morawitz akbesianus Breuning, 1975; Procerus scabrosus ssp. akbesiensis Toulgoet, 1987; nom.pro akbesianus Breuning; =scabrosus akbesianus Breuning akcakocaensis Blumenthal & Breuning, 1967; un; =wiedemanni wiedemanni Ménétriés akdagicola Mandl, 1973; un; =saphyrinus notabilis Roeschke akensis Haury, 1889; Heterocarabus marietti ssp. akibensis Belousov, in litt.; Archiplectes daphnis ssp. akinini Morawitz, 1886; Cratocechenus sp. akitanus (Ishikawa, 1992); Leptocarabus arboreus ssp. akkusanus Breuning, 1964; un; =wiedemanni congruus Motschulsky akkusensis Breuning, 1964; un; =akkusensis Breuning & Ruspoli, 1970 akkusensis Breuning & Ruspoli, 1970; Sphodristocarabus macrogonus ssp. aksuanus Gottwald, 1987; Cratocechenus akinini ssp. alagoensis Kraatz, 1896; =maurus maurus (Adams) alagoensis Reitter, in litt.; =varians alagoensoides Mandl alagoensoides Mandl, 1955; un; =alagoensoides Mandl, 1975 alagoensoides Mandl, 1975; Sphodristocarabus varians ssp. alainsis Meyer, 1996; un;=violaceus aurichalceus Kraatz. alajensis Semenov, 1896; nom.pro punctatostriatus Heyden; Pantophyrtus sp.; =?hyb turcomanorum/staudingeri alamannicus Csiki, 1927; nom.pro germanicus Sokolar; =ullrichi rhilensis Kraatz alanstivelli (Morvan, 1981); Meganebrius sp. alarici (Darnaud, 1983); un; =hispanus hispanus Fabricius alaricus (Meyer, 1990); un; =coriaceus coriaceus Linnaeus alaskanus Obydov, 1996; =vietinghoffi vietinghoffi Adams alaskensis Basilewsky, 1937; =truncaticollis truncaticollis Eschscholtz albanensis Géhin, 1885; =catenulatus catenulatus Scopoli albanicola Breuning, 1964; un; =hortensis neumeyeri Schaum albanicus Breuning, 1943; =coriaceus joniensis Breuning albarracinus Ganglbauer, 1886; =lusitanicus helluo Dejean alberensis (Darnaud, 1978); un; =rutilans rutilans Dejean alberginius (Remond, 1986); un; =splendens Olivier albiensis Depoli, 1938; ?nom.pro carnicus Csiki; =croaticus croaticus Dejean albinae Haury in Géhin, 1885; =violaceus ?violaceus Linnaeus alboequus Deuve & Imura, 1993; Neoplesius wagae ssp. albowi Reitter, 1896; =reitteri laevisternis Starck albrechti Morawitz, 1862; Ohomopterus sp. albrechti: Maindron, 1906, non Morawitz, 1862; =bornianus Hauser, 1922 alcaracinus Kraatz, 1886; =lusitanisus latus Dejean aldarkose Kabak, 1992; Acrocarabus guerini ssp. aleppensis Lapouge, 1914; =mulsantianus mulsantianus Morawitz alessiensis Apfelbeck, 1901; Tachypus cancellatus ssp. alexandrae Semenov, 1887; Acathaicus sp. algiricus Géhin, 1885; =famini maillei Solier aliai Escalera, 1944; Cathoplius asperatus ssp. alicantinus Ganglbauer, 1886; =lusitanicus helluo Dejean allevardensis Nicolas, 1919; =auronitens auronitens Fabricius

34 alliciens Deuve, 1990; Pseudocranion sp. alioni Lie, 1990; un; ?nom.pro alionidiernae Lie; =ullrichi fastuosus Palliardi alionidiernae Lie, 1989; ?nud; =ullrichi fastuosus Palliardi alluaudi Colas, 1936; =problematicus harcyniae Sturm almushensis Mandl, 1975; Sphodristocarabus tokatensis ssp. alotrus Lassalle & Remond, 1998; =latro latro Semenov alpestris Sturm, 1815; Orinocarabus sp. alpherakii Semenov, 1898; Cupreocarabus sp. alpicola Heer, 1837; =arcensis sylvaticus Dejean alpigradus Sermet, 1985; =monilis Fabricius alpinus Dejean, 1826, nec Paykull, 1790; =concolor concolor Fabricius alpinus (Kwon & Lee, 1984), nec Paykull, 1790; =wulffiusi taebeagsanus Ishikawa & Kim altaicus Gebler, 1847; =massagetus Motschulsky altamontanus Ochs, 1965; =solieri solieri Dejean altercoriaceus Kolbe, 1925; =coriaceus coriaceus Linnaeus alternans Palliardi, 1825, nec Fabricius, 1792; lsu; Macrothorax morbillosus ssp. alternans Kraatz, 1887, nec Fabricius, 1792; =hortensis hortensis Linnaeus alternatus Haury, 1855; =catenulatus catenulatus Scopoli alterviolaceus Kolbe, 1925; =violaceus andrzejuscii Fischer alticola Bellier, 1880; =monilis Fabricius altitudinorum Breuning, 1934; un; ios of altitudinum Breuning altitudinum Breuning, 1934; emend. pro altitudinorum Breuning; ?hyb exaratus/septemcarinatus altuvensis Shilenkov, in litt.; Carabus arcensis ssp. altyntuanus Shilenkov, 1996; Morphocarabus odoratus ssp. alutensis Savulescu, 1972; Morphocarabus rothi ssp. alveolatus Semenov, 1898; un; =balassogloi Dohrn alysidotus Illiger, 1798; Rhipocarabus sp. alzonai Deuve, 1990; Isiocarabus kiukiangensis ssp. amabilis Mandl, 1975; un; =ignimitella ignimitella Bates amasicus Csiki, 1927; nom.pro modestus Kraatz; Procerus scabrosus ssp. amauryi Haury, 1898; inc; ?artifact amayensis Lassalle, 1984; un; =lusitanicus macrocephalus Lapouge ambicornis Sokolar, 1907; nom.pro nigricornis Dejean; =cancellatus mimus Kolbe ambiguus Lapouge, 1914, nec Paykull, 1790; =calleyi chalconatus Mannerheim ambitiosus Schaufuss, 1872; =scheidleri scheidleri Panzer amitinus Kolbe, 1912; =cancellatus cancellatus Illiger ammonius Lapouge, 1909; =splendens Olivier amoenus Baudet-Lafarge, 1836; =monilis Fabricius amoenus Chaudoir, 1852, nec Baudet-Lafarge, 1836; =ermaki ermaki Lutshnik, 1924 amoenus Hauser, 1913; un; =pustulifer pustulifer Lucas amplicollis A.& J.B.Villa, 1844; =concolor concolor Fabricius amplicollis Bleuse, 1913, nec A.& J.B.Villa, 1844; =problematicus planiusculus Géhin amplior Deuve, 1994; Acathaicus alexandrae ssp. amplipennis Lapouge, 1924; Oreocarabus sp. amurensis Géhin, 1885; nom.pro femoralis Motschulsky; =arcensis conciliator Fischer amurlandicus Shilenkov, 1994; nom.pro viridimarginatus Kraatz; =hummeli smaragdulus Kraatz anami (Ledoux, 1977); Imaibius sp. ananowi Semenov, 1896; =kasakorum tschetschenicus Semenov anatolicus Chaudoir, 1857; Procrustes sp. anceps Lapouge, 1924; =wiedemanni congruus Motschulsky anceps Csiki, 1927, nec Lapouge, 1924; nom.pro incertus Meyer-Darcis; =lafossei lafossei Feisthamel anchocephalus Reitter, 1896; Cychrostomus sp. andalusiacus Géhin, 1885; un; =rugosus baeticus Deyrolle anderseni Beuthin, 1896; un; =cancellatus cancellatus Illiger andiaensis Forel & Leplat, 1998; =guadarramus La Ferté-Sénectere andorranus Lapouge, 1902; =problematicus planiusculus Géhin andreii Imura, 1998; Aristocarabus viridifossulatus ssp. andronnensis Nicolas, 1923; hyb Chrysocarabus splendens/auronitens andrzejowskii Lomnicki, 1914; fos; =violaceus Linnaeus andrzejuscii Fischer, 1823; Megodontus violaceus ssp. angelinus Reitter, 1899; =slovtzovi Mannerheim angenensis Deuve, 1996; Eocechenus kaznakovi ssp. angladensis (Jeanne, 1969); =auronitens farinesi Dejean anglicus Motschulsky, 1865; =arcensis sylvaticus Dejean anglicus Lapouge, 1906; un; =monilis Fabricius angulatus Hauser, 1921, nec Fabricius, 1781; =blaptoides oxuroides Schaum angulicollis Hauser, 1911; Coptolabrus ignimitella ssp. angulihabitus Kleinfeld, 1998; Apotomopterus sp. angustatus Panzer, 1812, nec Fabricius, 1787; =linnei linnei Panzer angustatus Redtenbacher, 1849, nec Fabricius, 1787; =sylvestris redtenbacheri Géhin, 1876 angustatus Ballion, 1878, nec Fabricius, 1787; =ballionis Kraatz, 1879 angustatus Petri, 1912, nec Fabricius, 1787; =glabratus glabratus Paykull angusticollis Motschulsky, 1859, nec Fabricius, 1801; =coriaceus coriaceus Linnaeus angusticollis Haller, 1885, nec Fabricius, 1801; =scabrosus scabrosus Olivier angusticollis Hauser, 1911, nec Fabricius, 1801; =augustus augustus Bates angustior Born, 1895; =problematicus problematicus Herbst angustipennis Mandl, 1970; Meganebrius koganae ssp. angustulus Haury, 1876; =intricatus intricatus Linnaeus angustus Roeschke, 1898; Parhomopterus sp. anhuinus Imura, 1996; Archaeocarabus sp. anhweiensis Hauser, 1932; =elysii connectens Hauser anjiensis Li, 1993; =lafossei buchi Hauser ankarensis Breuning, 1961; Ischnocarabus tenuitarsis ssp. annae Raynaud, 1936; hyb Chrysocarabus lineatus/auronitens annettae Korell, 1982; Meganebrius quinlani ssp.

35 annisus Kolbe, 1913; =cancellatus intermedius Dejean annikae (Darnaud, 1979); un; =lineatus lateralis Chevrolat annikae Kleinfeld, 1996; Apotomopterus grossefoveatus ssp. anomalus Géhin, 1880; =monilis Fabricius antaeus Hauser, 1914; Coptolabrus ignimitella ssp. antaiensis Deuve, 1994; Apotomopterus longeantennatus ssp. antakyae Korell, 1985; =mulsantianus mulsantianus Morawitz anthracinus Everts, 1918, nec Illiger, 1798; =problematicus harcyniae Sturm anthracinus Barthe, 1921; un; =auratus auratus Linnaeus antipai Panin, 1942; =convexus convexus Fabricius antiquus Dejean, 1826; Mesocarabus lusitanicus ssp. antonellii Luigioni, 1921; =clathratus auraniensis Müller antonini Deuve, 1997; =tsogoensis Deuve antonrichteri Heinz, 1983; Imaibius pachtoun ssp. antropovi Shilenkov, 1996; Morphocarabus odoratus ssp. antungensis Hauser, 1921; =smaragdinus longipennis Chaudoir anurus Semenov, 1898; =blaptoides rugipennis Motschulsky anxiensis Kleinfeld, 1998; Apotomopterus sp. anxius Lapouge, 1914, nec Duftschmid, 1812; =impressus impressus Klug apfelbecki Born, 1904; =cancellatus intermedius Dejean apollo Zolotarev, 1913; Archiplectes sp. appenninus Depoli, 1927; =convexus convexus Fabricius apricalis Cavazzuti, 1997; Apotomopterus patroclus ssp. apschuanus Rost, 1893; Tribax sp. arabikensis Gottwald, 1985; =reitteri gagrinus Starck aquatilis Bates, 1883; =clathratus maacki Morawitz aquilarum Cavazzuti, 1997; Neoplesius pseudomarkamensis ssp. aquitaniensis Mollard, 1996; =nitens Linnaeus arachnopus Hauser, 1914; =smaragdinus longipennis Chaudoir aragonensis Lapouge, 1913; =rutilans rutilans Dejean aragonicus Ganglbauer, 1886; =lusitanicus brevis Dejean araneipes Lapouge, 1903; =creutzeri creutzeri Fabricius araratensis Kraatz, 1896; =pumilio Küster araxenus Jablokov-Khnzorjan, 1976; =calleyi pseudoprasinus Lapouge arborensis Kraatz, 1908; =morbillosus constantinus Lapouge arboreus Lewis, 1882; Leptocarabus sp. arcadicus Gistl, 1850; Chaetocarabus sp. arcanus Semenov, 1898; nom.pro concinnus Semenov; Deroplectes sp. arcensis Herbst, 1784; cos; Carabus sp. arcticus Sparre-Schneider, 1888, nec Olivier, 1795; =violaceus violaceus Linnaeus arelatensis Lapouge, 1903; =clathratus auraniensis Müller argonautarum Semenov, 1898; nom.pro cupreus Chaudoir; Microplectes convallium ssp. arietinus Lapouge, 1902; fos; =?violaceus Linnaeus aristochroides Deuve, 1992; Calocarabus sp. arlequinus Deuve, 1994; Apotomopterus yuae ssp. armeniacus Mannerheim, 1830; Sphodristocarabus varians ssp. armeniensis Korge, ?in litt.; =calleyi ?pseudoprasinus Lapouge armenius Zaitzev, 1930; Procerus scabrosus ssp. armiger Imura. 1997; Pseudocoptolabrus sp. armoricanus Oberthür, 1935; =auronitens auronitens Fabricius arpaalaniensis Machard, 1985; =cribratus tazieffi Breuning & Ruspoli arradoyensis (Lequet, 1975); un; =splendens Olivier arraizensis Raynaud, 1938; hyb Chrysocarabus lineatus/splendens arrogans Schaum, 1858; Eucarabus ullrichi ssp. arrogantior Deuve, 1991; Apotomopterus arrowi ssp. arrowi Hauser, 1913; Apotomopterus sp. arrowianus Breuning, 1934; Ohomopterus insulicola ssp. arsiyandagensis Machard, 1985; =varians armeniacus Mannerheim artemis Kleinfeld, 2000; Apotomopterus sp. articervix Cavazzuti, 1999; Pagocarabus crassesculptus ssp. artusi Schweiger, 1964; un; =saphyrinus saphyrinus Cristoforis & Jan artusi (Le Mao, 1984); un; =auronitens costellatus Géhin arunensis Heinertz, 1980; Meganebrius sp. arvensis Paykull, 1790; iss of arcensis Herbst arvernus Barthe, 1908; un; =monilis Fabricius arvernus Lapouge, 1902; un; =problematicus harcyniae Sturm ascendens Semenov, 1898; Apotomopterus sp. asidoides Lapouge, 1902; un; =cancellatus carinatus Charpentier askhicus Belousov, 1992; Archiplectes daphnis ssp. asperatus (Dejean, 1826); Cathoplius sp. asperatus Mulsant, 1853, nec (Dejean, 1826); =mulsantianus mulsantianus Morawitz, 1886 asperipennis Lapouge, 1921; =violaceus purpurascens Fabricius asperulus Kraatz, 1878; =violaceus purpurascens Fabricius asperus Beuthin, 1892; =monticola monticola Dejean aspromontanus Born, 1918; =intricatus lefebvrei Dejean assamensis Mandl, 1965; =indicus indicus Fairmaire assiduus Kolbe, 1913; =cancellatus scythicus Schaum assimilis Duftschmid, 1812, nec Paykull, 1790; =cancellatus graniger Palliardi, 1825 assimilis Cristoforis & Jan, 1837, nec Paykull, 1790; =chevrolati chevrolati Cristoforis & Jan assimilis (Kwon & Lee, 1984), nec Paykull, 1790; =fraterculus taebaeksaniculus Deuve, 1991 astrabadensis Semenov, 1903; =elegantulus Motschulsky astridae Fiévet, 1991; un; =auratus honnoratii Dejean astur Lapouge, 1898; =cancellatus carinatus Charpentier asturicus Born, 1925; =lusitanicus cantabricus Chevrolat asturiensis Born, 1925; =violaceus aurichalceus Kraatz

36 atavus Lapouge, 1908; =?nemoralis prasinotinctus Heyden atbaschensis Obydov, 1998; =eous eous Morawitz atchibachi Zamotajlov, 1992; Archiplectes reitteri ssp. atentsensis Deuve, 1990; Pseudocoptolabrus taliensis ssp. ater Letzner, 1850; un; =glabratus glabratus Paykull ater Fleischer, 1898, nec Villers, 1789; =cancellatus graniger Palliardi ater (Raynaud, 1970); un; =solieri solieri Dejean athesinus Depoli, 1927; =convexus dilatatus Dejean athleta Semenov, 1896; Lamprostus nordmanni ssp. athletus Semenov, 1896; iss of athleta Semenov ativsengueni Schweiger, 1962; =marietti marietti Cristoforis & Jan atlanticus Lapouge, 1924; =maeander lapilayi Castelnau atlantis Alluaud, 1923; Eurycarabus famini ssp. atratus Heer, 1841, nec Duftschmid, 1812; =auronitens auronitens Fabricius atratus Hauser, 1913, nec Duftschmid, 1812; =lafossei saturatus Hauser atratus Hauser, 1914; un; =elysii chikongschanensis Hauser atratus Hauser, 1926; un; =lafossei kienchanganus Hauser atripes Géhin, 1885; un; =auratus honnoratii Dejean atrocoeruleus Letzner, 1850; un; =scheidleri scheidleri Panzer atrocyanescens Lamy, 1973; un; =granulatus granulatus Linnaeus atronitens Henderickx, 1998; un; =auronitens auronitens Fabricius atropurpureus Dalla Torre, 1877; un; =catenulatus catenulatus Scopoli atrovirescens Dalla Torre, 1877; un; =catenulatus catenulatus Scopoli atroviridulus Müller, 1898; =cancellatus corpulentus Kraatz attrepidans Deuve, 2000; Archaeocarabus fontellus ssp. aubersoni (Raynaud, 1973); un; =nemoralis nemoralis Müller aubryi (Colas, 1969); =auronitens farinesi Dejean auctus Semenov & Znojko, 1932; =careniger Chaudoir audouini Brullé, 1835; Procerus scabrosus ssp. aufrerei Colas, 1964; =auratus honnoratii Dejean augustus Bates, 1888; Coptolabrus sp. aulacocnemus Semenov, 1896; Cyclocarabus sp. auliensis Lapouge, 1916; Semnocarabus erosus ssp. auliensoides Mandl, 1955; =carbonicolor korolkowi Semenov aumonti Lucas, 1849; Macrothorax morbillosus ssp. auraniensis Müller, 1902; Limnocarabus clathratus ssp. auratoides Reitter, 1896; =auratus auratus Linnaeus auratonitens Illiger, 1798; un; error for auronitens Fabricius auratopumicatus Sirguey, 1924; un; =auronitens festivus Dejean auratus Linnaeus, 1761; Tachypus sp. auratus Hauser, 1913; un; =lafossei buchi Hauser auratus Heuer, 1926, nec Linnaeus, 1761; =nemoralis nemoralis Müller aureocinctus Hauser, 1916; un; =ignimitella ignimitella Bates aureocupreus Reitter, 1880; =obsoletus obsoletus Sturm aureomicans Letzner, 1850; un; =nitens Linnaeus aurescens Dalla Torre, 1877; un; =scheidleri scheidleri Panzer aurescens Hauser, 1913; un; =lafossei coelestis Stewart aureus DeGeer, 1774; =nitens Linnaeus aureus Letzner, 1850; un; =scheidleri scheidleri Panzer aureus Sirguey, 1931, nec DeGeer, 1774; =auronitens auronitens Fabricius aurichalceus Kraatz, 1879; Megodontus violaceus ssp. auricollis Waterhouse, 1867; =blaptoides rugipennis Motschulsky auricollis Born, 1922, nec Waterhouse, 1867; =fiduciarius kirinicus Csiki, 1927 auriculatus Putzeys, 1872; Iniopachus sp. auritus Cavazzuti, 2000; Scambocarabus sp. aurocinctus Motschulsky, 1846; Diocarabus sp. aurocostatus Lapouge, 1902; =auronitens montanus Géhin aurolimbatus Dejean, 1829; Megodontus violaceus ssp. auronitens Fabricius, 1792; Chrysocarabus sp. auropurpureus Lapouge, 1898; =auronitens auronitens Fabricius aurosericeus Kraatz, 1880; =rothi hampei Küster aurosplendens Born, 1910; =cancellatus emarginatus Duftschmid aurulentus Lapouge, 1902; un; =intricatus intricatus Linnaeus austerus Cavazzuti, 1997; Apotomopterus yuae ssp. australis Bernau, 1914; collective name; =cancellatus Illiger, s.lat. australofinis (Nakajima, 1994); =gehinii munakatai Ishikawa austriacus Sturm, 1815, nec Fabricius, 1801; =problematicus ?problematicus Herbst austriae Sokolar, 1907; =arcensis arcensis Herbst authionensis Ochs, 1953; =violaceus ?picenus A.& J.B.Villa auverloti (Thibaudeau, 1983); un; =auronitens auronitens Fabricius auvrayi Charet, 1992; =tenuitarsis ankarensis Breuning auvrayorum Deuve & Mourzine, 1993; Acoptolabrus leechi ssp. auzati Le Moult, 1912; =hispanus Fabricius avadcharensis Kurnakov, 1970; Archiplectes satyrus ssp. avenionensis (Schaefer, 1973); =auratus lotharingus Dejean aveyronensis Beuthin, 1895; =cancellatus carinatus Charpentier aveyronnensis (Raynaud, 1970); un; =auronitens festivus Dejean avilensis Breuning, 1966; un; =lineatus lineatus Dejean avinovi Semenov & Znojko, 1932; Megodontus sp. avunculus Kolbe, 1912; =cancellatus cancellatus Illiger awajiensis Imura, Dejima & Mizusawa, 1993; Ohomopterus japonicus ssp. awakazusanus Ishikawa, 1981; Ohomopterus lewisianus ssp. awashimae Ishikawa & Takami, 1996; Ohomopterus albrechti ssp. axiomorphus Tarrier, 1971; un; =solieri clairi Géhin ayderensis Blumenthal & Breuning, 1967; un; =ayrdensis Breuning & Ruspoli, 1976

37 ayensis (Darnaud, 1980); un; =auronitens farinesi Dejean ayrdensis Breuning & Ruspoli, 1976; ?iss of ayderensis Blumenthal & Breuning; Tribax puschkini ssp. azaleae Cavazzuti, 1986; Sphodristocarabus macrogonus ssp. azari Deuve, 1998; Procrustes piochardi ssp. azrael Semenov & Znojko, 1932; Teratocarabus sp. azureoauratus Dalla Torre, 1877; un; =auronitens auronitens Fabricius azurescens Dejean, 1826; Megodontus violaceus ssp. azurescens Dalla Torre, 1877; un; =monilis Fabricius azurescens Dalla Torre, 1877; un; =problematicus problematicus Herbst azurescens Hauser, 1913; un; =lafossei coelestis Stewart azurescens Hauser, 1921; =lafossei coelestis Stewart azurescens (Raynaud, 1970); un; =solieri clairi Géhin

B babai Ishikawa, 1968; Leptocarabus arboreus ssp. babaianus Ishikawa, 1983; nom.pro montanus Baba; Damaster blaptoides ssp. babaulti Le Moult, 1912; un; =splendens Olivier babinjensis Apfelbeck, 1919; =croaticus durmitorensis Apfelbeck babukensis Zamotajlov, 1988; Archiplectes starckianus ssp. baccivorus Fischer, 1823; =taedatus taedatus Fabricius baderlei Mandl, 1965; un; =scheidleri scheidleri Panzer baeri Ménétriés, 1851; Morphocarabus odoratus ssp. baeri Bleuse, 1914, nec Ménétriés, 1832; =auratus honnoratii Dejean baeterrensis Lapouge, 1901; Megodontus violaceus ssp. baeticus Deyrolle, 1852; Macrothorax rugosus ssp. baguenai Breuning, 1926; Mesocarabus lusitanicus ssp. baidulyensis Belousov & Kabak, 1994; =eous Morawitz baimanorum Deuve, 1999; Archaeocarabus sp. balachowskyi (Morvan, 1970); =kokujewi Semenov balanensis Csiki, 1942; =violaceus wolffii Dejean balassogloi Dohrn, 1882; Cratocephalus sp. balcanicus Born, 1899; Tachypus cancellatus ssp. balcanicus Lapouge, 1901, nec Born, 1899; =violaceus rilvensis Kolbe baldensis Schaum, 1857; Platycarabus creutzeri ssp. baldiai Mandl, 1963; un; =arcensis arcensis Herbst baldianus Mandl, 1955; un; =cancellatus excisus Dejean balearicus Lapouge, 1913; =morbillosus macilentus Lapouge balkaricus Belousov & Abdurachmanov, 1991; Tribax sp.; ?hyb Tribax kasbekianus/fossiger ballionis Kraatz, 1879; nom.pro angustatus Ballion; Ophiocarabus sp. baltalensis Ledoux, 1980; =dardiellus subpunctulus Roeschke balticus Bernau, 1915; un; =cancellatus cancellatus Illiger bamianicola Deuve & Tian, 2000; Apotomopterus arrowi ssp. bambousicola Deuve & Tian, 1999; Archaeocarabus fuscomarginatus ssp. bampsi Henderickx, 1998; un; =auronitens auronitens Fabricius banati Faldermann, 1835; iss of brandti Faldermann, 1835 (error by Li, 1992) banaticus Redtenbacher, 1849; Procrustes coriaceus ssp. banghaasi Reitter, 1905; =caerulans Morawitz banoni Dejean, 1829; Procrustes sp. baoshanensis Deuve, 1993; Apotomopterus solidior ssp. baptistatus Csiki, 1927; un; nom.pro costatus Hauser; =lafossei lafossei Feisthamel barakaicus Zamotajlov, 1991; Tribax biebersteini ssp. baranyensis Sokolar, 1908; =ullrichi rhilensis Kraatz bararii Deuve, 2000; Sphodristocarabus adamsi ssp. barbarus Dejean, 1826; =rugosus rugosus Fabricius barbulatus Cavazzuti & Ratti, 1999; Archaeocarabus nestor ssp. barcelecoanus Lapouge, 1924; Mesocarabus lusitanicus ssp. bargusinicus Shilenkov, 1996; Morphocarabus odoratus ssp. barkamensis Deuve, 1994; Rhigocarabus xiei ssp. barkamianus Korell & Kleinfeld, 1988; Hypsocarabus laotse ssp. barkamicola Deuve, 1996; Pseudocranion shuamaluko ssp. barnaulanus Born, 1922; =henningi henningi Fischer barnaulensis Born, 1922; =convexus convexus Fabricius barnolai Xaxars, 1921; =rugosus levantinus Lauffer baronii Heinertz, 1977; Imaibius sp. barovskii Semenov & Znojko, 1931; Alipaster sp. barrosi Lapouge, 1908; =steuartii Deyrolle bartheensis Nicolas, 1923; hyb Chrysocarabus lineatus/splendens barthei Lapouge, 1912; =auronitens farinesi Dejean barthei Le Moult, 1912; hyb Chrysocarabus splendens/auronitens barthei Lebis, 1924; un; =auratus auratus Linnaeus bartholomaei Motschulsky, 1852; =stjernvalli humboldti Faldermann bartoni Mařan, 1930; =violaceus dryas Gistl barysomus Bates, 1889; Imaibius sp. basalis Beuthin, 1896; =cancellatus graniger Palliardi baschkiricus Breuning, 1936; Carabus arcensis ssp. bashanensis Imura, 1995; Archaeocarabus pseudolatipennis ssp. basilewskianus Breuning & Ruspoli, 1970; =rotundicollis rotundicollis Mandl basilewskyi Breuning, 1966; =?theanus theanus Reitter basilianus Starck, 1890; Archiplectes prometheus ssp. basilicatus Born, 1902; =coriaceus coriaceus Linnaeus basilicus Chevrolat, 1837; =lineatus lineatus Dejean basirufus Marcu, 1937; =estreicheri Fischer basoglui Schweiger, 1964; =scabrosus sommeri Mannerheim basquini Darge, 1990; =kurdicus kurdicus Heinz

38 basquinianus Darge, 1992; =torosus cankirianus Blumenthal & Breuning bastiani Lassalle, 1982; =steuartii Deyrolle batangensis Deuve, 1989; Neoplesius lama ssp. batangicus Deuve, 1989; Rhigocarabus sp. batesi Lapouge, 1916; =maeander paludis Géhin batesianus Deuve, 1995; =odoratus odoratus Motschulsky batschkowskii Bodemeyer, 1928; un; =reitteri reitteri Retowski battalensis Heinz & Staven, 1998; Imaibus baronii ssp. battonianus Mandl, 1966; un; =italicus italicus Dejean battoniellus Deuve, 1991; =mellyi mellyi Chaudoir battoniensis Deuve, 1991; Archaeocarabus sp. battonii Mandl, 1966; un; =problematicus inflatus Kraatz baudii Kraatz, 1878; =fairmairei fairmairei Thomson bavaricus Beuthin, 1896; =monilis Fabricius bavaricus Kolbe, 1912, nec Beuthin, 1896; =cancellatus excisus Dejean bavariensis Lapouge, 1909; iss of bavaricus Beuthin bayanbulak Deuve, 1993; Alipaster barovskii ssp. bayardi Solier, 1835; Chaetocarabus intricatus ssp. bayeri Lapouge, 1910; =problematicus harcyniae Sturm beauvoisi Dejean, 1826; =problematicus ?harcyniae Sturm bebiai Gaskö, in litt.; ?Archiplectes sp. bechteli Charet, 1996; =enigmaticus citamgineus Lassalle & Charet beharrius Nicolas, 1916; =lineatus troberti Kraatz beheimi Niedl, 1957; =scheidleri ?helleri Ganglbauer beidensis Codina, 1927; =rugosus rugosus Fabricius beirensis Born, 1924; =galicianus Gory beizhenensis Deuve & Li, 2000; Aulonocarabus canaliculatus ssp. belgicus Lapouge, 1913; =problematicus harcyniae Sturm belloni (Tarrier, 1966); =solieri bonnetianus Colas bellus Kraatz, 1886; =arcensis faldermanni Dejean belousovi Kabak, 1992; Cyclocarabus sp. benabarrensis Meyer & Mollard, 1996; =lusitanicus trabuccarius Fairmaire benardi Breuning, 1931; Apotomopterus sp. benedictae (Lacroix, 1986); un; =auronitens festivus Dejean benjamini Semenov, 1898; Pseudocranion sp. benoiti (Devecis, 1975); =monilis Fabricius benxiensis Imura, 1991; Coptolabrus jankowskii ssp. bepmalei Lapouge, 1904; =problematicus planiusculus Géhin berardi (Raynaud, 1967); =hispanus Fabricius berberus Géhin, 1885; =famini numidus Castelnau berezowskii Semenov, 1898; =formosus formosus Semenov bernensis Born, 1895; =concolor concolor Fabricius bernhardinus Kraatz, 1878; =concolor concolor Fabricius bernhauerorum Heinz, 1977; Procrustes mulsantianus ssp. bertolinii Kraatz, 1878; Orinocarabus sp. besanconi Breuning & Ruspoli, 1970; =saphyrinus saphyrinus Cristofori & Jan bescidensis Reitter, 1896; =linnei linnei Panzer beskydicus Smetana, 1949; =auronitens auronitens Fabricius besleticus Kurnakov, 1972; Archiplectes satyrus ssp. bessarabicus Fischer, 1823; Tomocarabus sp. besseri Fischer, 1822; Trachycarabus sp. besucheti Schweiger, 1965; =wiedemanni wiedemanni Ménétriés beszedesi Depoli, 1915; un; =cancellatus corpulentus Kraatz bettingeri Barthe, 1921; =auratus auratus Linnaeus betuliae Csiki, 1940; =violaceus wolffii Dejean beuthini Schulz, 1899; =cristoforii Spence beybienkoi Kryzhanovskij, 1973; Diocarabus sp. bhadarwahensis Deuve, 1982; Imaibius boysi ssp. bhaktiai Deuve, 1993; Meganebrius dilatotarsalis ssp. bibanensis Lie, 1994; =cancellatus graniger Palliardi bicanaliceps Casey, 1920; Neocarabus taedatus ssp. bicinctus Born, 1905; =violaceus baeterrensis Lapouge bicolor Letzner, 1850; un; =violaceus violaceus Linnaeus bicolor Letzner, 1850; un; =intricatus intricatus Linnaeus bicolor Beuthin, 1896, nec (Walker, 1866); =cancellatus cancellatus Illiger; bicolor (Walker, 1866); =Calosoma moniliatum LeConte bicolor (Keith, 1989); un; =arcensis sylvaticus Dejean bicolor: auct., non Walker, 1866; =taedatus agassi LeConte bicoloratus Fleischer, 1930; un; =obsoletus obsoletus Sturm biebersteini Ménétriés, 1832; Tribax sp. biellensis Born, 1923; =monticola fontanae Born bielzi Birthler, 1886; =obsoletus carpathicus Palliardi bieneri Mandl, 1965; un; =scheidleri helleri Ganglbauer bifoveicollis Reitter, 1896; =modestulus modestulus Semenov bigerrio Lapouge, 1924; =auronitens montanus Géhin bigorrensis (Camard, 1995); hyb Iniopachus pyrenaeus/Chrysocarabus auronitens biharicus Reitter, 1896; =cancellatus graniger Palliardi bilineatus Géhin, 1885; un; =lineatus lateralis Chevrolat bilmenecus Lassalle, 1996; =cribratus cribratus Quensel billbergi Mannerheim, 1827; Parhomopterus sp. billieriensis (Schaefer, 1969); =auratus lotharingus Dejean birmanus Andrewes, 1929; Apotomopterus sp. biroi Csiki, 1927; nom.pro kindermanni Chaudoir; Sphodristocarabus sp. birthleri Reitter, 1885; =rothi rothi Dejean bischoffi Chaudoir, 1848; Heterocarabus marietti ssp. biseriatus Chaudoir, 1846; =decolor Fischer

39 bisioi Cavazzuti & Meli, 1999; Orinocarabus putzeysianus ssp. bithynicus Heinz & Korge, 1967; Lamprostus erenleriensis ssp. bitschnaui Born, 1926; =irregularis irregularis Fabricius bjelasnicensis Apfelbeck, 1902; =scheidleri illigeri Dejean blairi Breuning, 1934; Ohomopterus yaconinus ssp. blakistoni Newman, 1857; =gyllenhali Fischer blanci (Darnaud, 1978); un; =rutilans rutilans Dejean blandus Frivaldsky, 1865; Archicarabus montivagus ssp. blaptoides Kollar, 1836; Damaster sp. blayeulensis Machard, 1977; un; =auratus honnoratii Dejean bleonensis Raynaud, 1973; un; =monticola monticola Dejean bleusei Oberthür, 1884; =auronitens costellatus Géhin blumenthali Blumenthal, 1961; un; =violaceus violaceus Linnaeus blumenthalianus Mandl, 1963; un; =sylvestris sylvestris Panzer blumenthaliellus Deuve, 1988; Qinlingocarabus sp. blumenthaliensis Heinz & Korge, 1967; Sphodristocarabus rotundicollis ssp. bochmanni Ménétriés, 1832; iss of bohemani Ménétriés bodemeyeri Lapouge, 1914; un; =torosus jani Géhin bodemeyeri Lapouge, 1924; =coriaceus cerisyi Dejean bodemeyeri Kolbe, 1928, nec Lapouge, 1924; =morbillosus morbillosus Fabricius bodemeyerianus (Mařan, 1952); un; nom.pro bodemeyeri Lapouge, 1924; =coriaceus cerisyi Dejean boeberi Adams, 1817; Cechenochilus sp. bogatshevi Zamotajlov, 1990; =stjernvalli tabackuriensis Novotný & Voříšek bogdanowi Ballion, 1878; Semnocarabus sp. bogdoinus Breuning, 1933; Ophiocarabus imperfectus ssp. bohatschi Reitter, 1881; =cancellatus emarginatus Duftschmid bohemani Ménétriés, 1832; Sphodristocarabus sp. bohemicus Haury, 1881; =intricatus intricatus Linnaeus bohemicus Tanzer, 1934; un; =menetriesi pacholei Sokolar bokharensis Csiki, 1927; nom.pro hauseri Lapouge; =theanus theanus Reitter bokori Csiki, 1927; nom.pro concinnus Lapouge; =rothi hampei Küster bolivari Breuning, 1926; =lusitanicus castilianus Dejean boluensis Schweiger, 1964; un; =saphyrinus pseudosaphyrinus Schweiger bonadonai Colas, 1947; =solieri bonnetianus Colas bondonensis Mandl, 1967; un; =creutzeri baldensis Schaum bonellii Dejean, 1826, nec Adams, 1817; =depressus depressus Bonelli bonneti Le Moult, 1912; =auronitens festivus Dejean bonneti Colas, 1936, nec Le Moult 1912; =solieri bonnetianus Colas, 1937 bonnetianus Colas, 1937; nom.pro bonneti Colas; Chrysocarabus solieri ssp. bonplandi Ménétriés, 1837; =torosus spinolae Cristoforis & Jan bontempsi Lassalle, 1992; un; =stjernvalli gordius Reitter bonvouloiri Chaudoir, 1863; Megodontus sp. boomensis Kabak, 1994; Ophiocarabus aeneolus ssp. borchmanni Mandl, 1959; un; =coriaceus hopffgarteni Kraatz borealensis (Raynaud, 1973); =nemoralis nemoralis Müller borealis Géhin, 1885, nec Paykull, 1790; =convexus convexus Fabricius borealis Poppius, 1906, nec Paykull, 1790; =arcensis conciliator Fischer borealis Born, 1908, nec Paykull, 1790; =clathratus clathratus Linnaeus borealis (Kwon & Lee, 1984), nec Paykull, 1790; =vietinghoffi bowringi Chaudoir boreli Le Moult, 1912; hyb Chrysocarabus splendens/auronitens borensis Lapouge, 1916; =bogdanowi kuldshensis Ballion boreus Csiki, 1927; nom.pro borealis Born; =clathratus clathratus Linnaeus borlovensis Lie, 1994; un; =rothi comptus Dejean borni Meyer-Darcis, 1902; Coptolabrus pustulifer ssp. borni Csiki, 1905, nec Meyer-Darcis, 1902; =carinthiacus Sturm borni Krausse, 1908, nec Meyer-Darcis, 1902; =morbillosus alternans Palliardi borni Kolbe, 1917, nec Meyer-Darcis, 1902; =arcadicus merlini Schaum borni Barthe, 1921; un; =vagans Olivier borni Lapouge, 1924, nec Meyer-Darcis, 1902; =auronitens auronitens Fabricius bornianus Hauser, 1922; Archaeocarabus sp. borniellus Csiki, 1927; nom.pro borni Kolbe; =arcadicus merlini Schaum borodini Heinz, 1996; Neoplesius sp. boroensis Lapouge, 1916; iss of borensis Lapouge borschmanni Mandl, 1959; iss of borchmanni Mandl borussicus Csiki, 1927; nom.pro detritus Lapouge, 1908; =arcensis arcensis Herbst boscensis Nicolas, 1923; =auronitens farinesi Dejean boschniaki Faldermann, 1835; =stjernvalli stjernvalli Mannerheim bosiljevici Drovenik & Pavičevič, 1985; =croaticus zepcensis Reitter bosnicus Apfelbeck, 1890; Megodontus croaticus ssp. bosniensis Born, 1918; =intricatus intricatus Linnaeus bosogensis Obydov, 1996; =pupulus pupulus Morawitz bosphoranus Fischer, 1823; Trachycarabus sibiricus ssp. bosphoranus Motschulsky, 1844, nec Fischer, 1823; =scabrosus sommeri Mannerheim, 1844 boszdaghensis Breuning, 1966; =montivagus vellepiticus Hampe botchan Imura & Mizusawa, 1995; Ohomopterus kawanoi ssp. botezati Marcu, 1934; un; =excellens Fabricius boudeti Beuthin, 1892; =hispanus Fabricius bouilloni Breuning & Ruspoli, 1976; Sphodristocarabus scovitzi ssp. bouisseti Barthe, 1936; =auronitens farinesi Dejean boulbenei (Darnaud, 1979); un; =lineatus lateralis Chevrolat boulbeni (Blanc, 1980); un; =rutilans rutilans Dejean boulbenianus Deuve, 1996, Apotomopterus sp. bourdeaui (Darnaud, 1979); un; =lineatus lateralis Chevrolat bourgini Balazuc, 1946; un; =auronitens auronitens Fabricius bourgoini Deuve, 1998; Procrustes impressus ssp.

40 bousqueti Deuve, 1993; Coptolabrus formosus ssp. bousquetellus Deuve, 1998; Apotomopterus sp. bousquetianus Deuve, 1994; Apotomopterus tuxeni ssp. bowringi Chaudoir, 1863; Megodontus vietinghoffi ssp. boysi Tatum, 1851; Imaibius sp. bozdaghensis Breuning, 1966; iss of boszdaghensis Breuning brabeus Schaufuss, 1876; =lusitanicus macrocephalus Dejean brachyderus Wiedemann, 1824; =chamissonis Fischer brachypedilus Morawitz, 1886; Pantophyrtus sp. branaungi Imura, 1999; Pseudocoptolabrus sp. brandti Faldermann, 1835; Cathaicus sp. branickii Taczanowski, 1887; Coptolabrus smaragdinus ssp. brannani Schaufuss, 1871; =rugosus celtibericus Germar brauni Heinz, 1993; Imaibius stoliczkanus ssp. brdensis Sokolar, 1911; =cancellatus excisus Dejean breiti Barthe, 1913; nom.pro barthei Le Moult; hyb Chrysocarabus splendens/auronitens bremii Stierlin, 1881; ?priority; =latreilleanus Csiki, 1927 breuili Alluaud, 1930; =rugosus baeticus Deyrolle breuningi Csiki, 1927; nom.pro kricheldorffi Breuning; Mesocarabus lusitanicus ssp. breuningianus Le Moult, 1930; Apotomopterus sp. breuningiellus Deuve, 1988; nom.pro cicatricosoides Breuning; Scambocarabus sculptipennis ssp. brevicaudus Imura & Mizusawa, 1995; =blaptoides blaptoides Kollar brevicollis Lapouge, 1910, nec Fabricius, 1792; =rutilans rutilans Dejean brevicornis Kraatz, 1878; =alpestris hoppei Germar breviculus Cavazzuti & Ratti, 1998; Rhigocarabus morawitzianus ssp. breviformis Chaudoir, 1863; =kruberi ?bungii Gebler brevior Poppius, 1907; =canaliculatus canaliculatus Adams brevipennis Lapouge, 1913; =auronitens auronitens Fabricius brevis Dejean, 1826, nec Duftschmid, 1812; lsu; Mesocarabus lusitanicus ssp. brevis Hauser, 1921; un; =elysii connectens Hauser brevituberculatus Roubal, 1909; =cancellatus excisus Dejean breviusculus Kraatz, 1876; =scabrosus sommeri Mannerheim breviusculus Kraatz, 1885, nec Kraatz, 1876; =glabratus gibbosus Heyden brezinai Deuve, 1994; Cupreocarabus sp. brianskyi Lutshnik, 1909; =prometheus prometheus Reitter brignolii Breuning & Ruspoli, 1970; un; =gilnickii gilnickii Deyrolle brisouti Fauvel, 1863; un; =problematicus inflatus Kraatz, 1878 britannicus Born, 1908; =violaceus violaceus Linnaeus brittoni Mandl, 1965; =quinlani quinlani Mandl brockmanni Kleinfeld & Schütze, 1997; Coptolabrus formosus ssp. brockmannioides Deuve & Mourzine, 1997; Coptolabrus formosus ssp. brosciformis Semenov, 1896; Cyclocarabus sp. brostenensis Jacquet, 1886; =irregularis montandoni Buysson browni Deuve, 1999; =violaceus violaceus Linnaeus bruggei Gottwald, 1990; =pupulus pupulus Morawitz bruggeianus Deuve, 1992; Neoplesius sp. brullei Géhin, 1885; =auratus auratus Linnaeus brunieri Barthe, 1921; =auratus auratus Linnaeus brunneicupreus Deuve, 1993; Sphodristocarabus gilnickii ssp. brunneofemoratus Letzner, 1850; un; =linnei linnei Panzer brunneus Letzner, 1850; un; =sylvestris sylvestris Panzer brunnipes Beuthin, 1885; =irregularis montandoni Buysson brunnipes Lapouge, 1908; un; =nemoralis nemoralis Müller bruschii Tarrier, 1974; =violaceus picenus A.& J.B.Villa brussensis Lapouge, 1909; un; =coriaceus cerisyi Dejean bruttianus Born, 1906; =morbillosus alternans Palliardi bucciarellii Mandl, 1966; =convexus convexux Fabricius bucephalus Kraatz, 1879; un; =irregularis ramamus Sokolar bucharensis Csiki, 1927; iss of bokharensis Csiki bucheti (Ochs, 1949); =alysidotus Illiger buchi Hauser, 1913; Coptolabrus lafossei ssp. buchtarmensis Kryzhanovskij, 1953; Trachycarabus mandibularis ssp. bucovinensis Marcu, 1934; un; =intricatus intricatus Linnaeus bucovinianus Kolbe, 1913; =cancellatus scythicus Schaum bucsecsianus Born, 1902; =cancellatus scythicus Schaum buddaicus Semenov, 1887; Rhigocarabus sp. buddhaicus Semenov, 1887; iss of buddaicus Semenov budensis Csiki, 1946; =cancellatus tibiscinus Csiki budha Breuning, 1950; Cupreocarabus alpherakii ssp. buendneri Breuning, 1934; un; =violaceus neesi Hope & Hornschuch buffi Deuve & Kaláb, 1992; Cratocechenus akinini ssp. bugareti Jeannel, 1941; hyb Chrysocarabus auronitens/Chrysotribax hispanus bugnionii Chaudoir, 1837; =fabricii fabricii Panzer bulgaricus Csiki, 1927; nom.pro ponticus Lapouge; =montivagus montivagus Palliardi bulgarus Lapouge, 1908; Trachycarabus scabriusculus ssp. bulghardaghensis Breuning, 1926; Archicarabus wiedemanni ssp. bulgharmaadensis Bodemeyer, 1915; Procerus scabrosus ssp. bungii Gebler, 1830; Scambocarabus kruberi ssp. bureianus Shilenkov, 1996; =vietinghoffi fulgidus Fischer bureschianus Breuning, 1928; Procerus scabrosus ssp. buresi Hanuš, 1923; =scheidleri versicolor Frivaldsky burgassiensis Apfelbeck, 1904; =wiedemanni wiedemanni Ménétriés burghauseri Fleischer, 1899; =scheidleri scheidleri Panzer buriatus Lapouge, 1924; =hummeli smaragdulus Kraatz burmanensis Breuning, 1932; Pseudocoptolabrus sp.

41 burnaschewi Dejean, 1826; =hummeli hummeli Fischer buschi Semenov & Znojko, 1931; Cechenochilus boeberi ssp. businskyi Deuve, 1990; Imaibiodes sp. businskyorum Imura, 1996; Aristocarabus viridifossulatus ssp. buycki Hauser, 1924; Apotomopterus sp. buyssoni Barthe, 1929; =auronitens costellatus Géhin bythinus Lapouge, 1908; =wiedemanni wiedemanni Ménétriés bytinskii Schweiger, 1970; =piochardi labruleriei Géhin, 1883 bzybiensis Gottwald, 1982; Tribax agnatus ssp.

V vacheri Basilewsky, 1931; nom.pro rufofemoratus Barthe; hyb Chrysocarabus splendens/auronitens vadoni Colas, 1962; =auratus honnoratii Dejean vafer Csiki, 1927, nec Duftschmid, 1812; nom.pro fallax Sokolar; =sylvestris haberfelneri Ganglbauer vagans Olivier, 1795; Tachypus sp. vagasensis Casey, 1913; iss of vegasensis Casey vagepunctatus Lapouge, 1907; un; =punctatus Castelnau vaitoiani Strasser, 1900; =wiedemanni wiedemanni Ménétriés valenciae Hauser, 1913; un; =lafossei saturatus Hauser valens Cavazzuti & Ratti, 1999; Apotomopterus cyanipennis ssp. validus Kraatz, 1884; Pseudotribax sp. validus Csiki, 1906, nec Kraatz, 1884; =rothi hampei Küster valikhanovi Kabak, 1990; Eotribax sp. vallierensis (Colas, 1965); Iniopachus pyrenaeus ssp. vallierensis (Jeanne, 1969), nec (Colas, 1965); =auronitens farinesi Dejean vancouvericus (Csiki, 1927); nom.pro bicolor Walker; =Calosoma moniliatum LeConte vancouvericus: auct., non Csiki, 1927; =taedatus bicanaliceps Casey vanensis Machard, 1985; Procrustes chavrolati ssp. vanvolxemi Putzeys, 1875; Carabus sp. variabilis Ballion, 1878; Ophiocarabus sp. varians Fischer, 1827; Sphodristocarabus sp. varicolor Joerin-Gerber, 1888; un; =monilis Fabricius variefucatus Font, 1997; =pustulifer mirificus Kraatz variolaris Ménétriés, 1849; =hungaricus cribellatus Adams variolatus Costa, 1839; Pachystus cavernosus ssp. variolosus Fabricius, 1787; Hygrocarabus sp. varistriatus Kraatz, 1878; =rothi rothi Dejean varius Born 1923, nec Olivier, 1795; =cantonensis Hauser varvasi Solier, 1835; =famini numidus Castelnau vasjurini Lafer, 1989; Aulonocarabus gossarei ssp. vaucheri Semenov, 1927; =erosus auliensis Lapouge vediensis Zamotajlov, 1992; Archiplectes komarowi ssp. vegasensis Casey, 1914; =serratus Say velebiticus Hampe, 1850; iss of montivagus vellepiticus Hampe vellepiticus Hampe, 1850; cos; Archicarabus montivagus ssp. velleri Marceron, 1937; =depressus depressus Bonnelli velox Cavazzuti & Ratti, 1999; Apotomopterus melli ssp. venerandus Deuve & Mourzine, 1997; Pseudocranion gansuensis ssp. veneti Auzat, 1915; un; =problematicus harcyniae Sturm venetianus Bernau, 1914; Carabus arcensis ssp. ventouxensis Nicolas, 1904; =auratus honnoratii Dejean ventricosus Bernau, 1915; =cancellatus corpulentus Kraatz ventrosior Deuve, 1990; Aristocarabus viridifossulatus ssp. ventusicus (Bellon & Tarrier, 1972); =monilis Fabricius venustoides Deuve, 1989; =gossarei imanensis Lafer venustus Morawitz, 1862; Morphocarabus sp. venustus (Raynaud, 1973), nec Morawitz, 1862; =nemoralis prasinotinctus Heyden verae Csiki, 1905; Megodontus planicollis ssp. verdonensis Puisségur, 1961; =nemoralis nemoralis Müller veris Gistl, 1857; =monilis Fabricius vermiculatus (Kocher, 1965); un; =famini fezzanus Lapouge vermiculosus Fons, Leplat & Ferrero, 1993; =rugosus levantinus Lauffer vernus Semenov & Znojko, 1932; Cyclocarabus namanganensis ssp. verrucosus Heer, 1837; =cancellatus excisus Dejean verrucosus Lapouge, 1903, nec Heer, 1837; =famini maillei Solier versicolor Frivaldsky, 1835; Morphocarabus scheidleri ssp. versicolor Letzner, 1850; un; =arcensis arcensis Herbst versicolor Letzner, 1850; un; =auronitens auronitens Fabricius versicolor Letzner, 1850; un; =intricatus intricatus Linnaeus versicolor Letzner, 1850; un; =irregularis irregularis Fabricius versicolor Letzner, 1850; un; =linnei linnei Panzer vertesensis Retezár, 1974; un; =scheidleri scheidleri Panzer verzhutzkii O.Berlov & Shilenkov, 1996; Morphocarabus aeruginosus ssp. veselyi Gottwald, 1980; Tribax kasbekianus ssp. vestigialis Lapouge, 1919; =carbonicolor carbonicolor Morawitz vesubiensis (Poncetton & Reymond, 1935); =solieri solieri Dejean vexator Schweiger, 1964; Lamprostus torosus ssp. viatorum Deuve, 1992; Pseudocranion sp. vicarius Cavazzuti, 1996; Apotomopterus patroclus ssp. vicinus Waltl, 1838; =coriaceus ?cerisyi Dejean vicinus Semenov, 1887, nec Waltl, 1838; =variabilis Ballion

42 victor Fischer, 1836; Archicarabus sp. victorianus Obydov, 1997; nom.pro kusnetzovi Shilenkov, 1996; ?Aulonocarabus careniger ssp.; ?hyb careniger/canaliculatus vieirae Oliveira, 1875; =lusitanicus lusitanicus Fabricius viennensis Kraatz, 1877; =hungaricus hungaricus Fabricius vietinghoffi Adams, 1812; Megodontus sp. vietinghoffianus Semenov, 1907; =vietinghoffi bowringi Chaudoir vigil Semenov, 1898; nom.pro striatus Semenov; Archaeocarabus sp. vigilax Bates, 1890; Archaeocarabus sp. vignatagliantii Breuning & Ruspoli, 1977; =torosus pseudescherichi Breuning villiersi (Morvan, 1972); =quinlani quinlani Mandl vincens Reitter, 1896; =koenigi Ganglbauer vincentianus Devecis, 1991; un; =monilis Fabricius vinctus Weber, 1801; Lichnocarabus sp. vinculatus Gebler, 1830; =arcensis conciliator Fischer vindobonensis Kubik, 1901; =auronitens kraussi Lapouge vinokurovi Deuve & Dolin; =mouthiezianus Deuve vinosus Le Moult, 1912; un; =splendens Olivier violaceocyaneus (Jeanne, 1972); un; =splendens Olivier violaceomarginatus Letzner, 1850; un; =convexus convexus Fabricius violaceomicans Letzner, 1850; un; =violaceus violaceus Linnaeus violaceopurpureus Barthe, 1908; un; =auronitens festivus Dejean violaceoviridis (Rouselle, 1978); un; =splendens Olivier violaceus Linnaeus, 1758; Megodontus sp. violaceus Letzner, 1850; un; =intricatus intricatus Linnaeus violaceus Breuning, 1972, nec Linnaeus, 1758; =guerini guerini Fischer violescens Dalla Torre, 1877; un; =scheidleri scheidleri Panzer violescens Dalla Torre, 1877; un; =scheidleri scheidleri Panzer virenquensis (Schaefer, 1969); =auratus lotharingus Dejean virens Sturm, 1815, nec Müller, 1776; =scheidleri scheidleri Panzer virens Schilsky, 1888, nec Müller, 1776; =fabricii fabricii Panzer virescens Letzner, 1850; un; =nemoralis nemoralis Müller virescens Letzner, 1850; un; =violaceus violaceus Linnaeus virescens Letzner, 1850; un; =glabratus glabratus Paykull virescens Letzner, 1850; un; =granulatus granulatus Linnaeus virescens Letzner, 1850; un; =intricatus intricatus Linnaeus virescens Letzner, 1850; un; =irregularis irregularis Fabricius virescens Letzner, 1850; un; =nitens Linnaeus virescens Letzner, 1850; un; =sylvestris sylvestris Panzer virescens Dalla Torre, 1877; un; =arcensis arcensis Herbst virescens Dalla Torre, 1877; un; =catenulatus catenulatus Scopoli virescens Dalla Torre, 1877; un; =scheidleri scheidleri Panzer viridanus Hauser, 1914; un; =augustus ignigena Hauser viridescens Dalla Torre, 1877; un; =monilis Fabricius viridiaeneus Letzner, 1850; un; =scheidleri scheidleri Panzer viridiaeneus Letzner, 1850; un; =arcensis arcensis Herbst viridiaeneus Dalla Torre, 1877; un; =cancellatus excisus Dejean viridiaeneus Dalla Torre, 1877; un; =ullrichi rhilensis Kraatz viridiaeneus Dalla Torre, 1877; un; =granulatus granulatus Linnaeus viridiaeneus Dalla Torre, 1877; un; =sylvestris haberfelneri Ganglbauer viridiaeneus Dalla Torre, 1877; un; =creutzeri creutzeri Fabricius viridiater (Raynaud, 1970); un; =solieri clairi Géhin viridiauratus Dalla Torre, 1877; un; =auratus auratus Linnaeus viridiauratus Dalla Torre, 1877; un; =auratus auratus Linnaeus viridiauratus Dalla Torre, 1877; un; =auronitens auronitens Fabricius viridiaureus Dalla Torre, 1877; un; =hortensis hortensis Linnaeus viridicans Obydov, 1993; =danilevskii Obydov viridicervix Csiki, 1917; un; nom.pro viridicollis Le Moult, 1912; =splendens Olivier viridicinctus Schaufuss,1872; =scheidleri scheidleri Panzer viridicinctus Kraatz, 1882, nec Schaufuss, 1872; =regalis regalis Fischer viridicoeruleus Letzner, 1850; un; =violaceus violaceus Linnaeus viridicoeruleus Kraatz, 1886; =arcensis faldermanni Dejean viridicoeruleus Hauser, 1921; un; =elysii lopinensis Hauser viridicollis Kraatz, 1882; =regalis regalis Fischer viridicollis Kraatz, 1886, nec Kraatz, 1882; =billbergi billbergi Mannerheim viridicollis Le Moult, 1912, nec Kraatz, 1882; =splendens Olivier viridicollis Hauser, 1913; un; =polemistes Hauser viridicollis Hauser, 1914; un; =elysii lopinensis Hauser viridicollis Hauser, 1919, nec Kraatz, 1882; =augustus augustus Bates viridicollis Hauser, 1921, nec Kraatz, 1882; =smaragdinus longipennis Chaudoir viridicollis Hauser, 1921, nec Kraatz, 1882; =lafossei coelestis Stewart viridicollis Sirguey, 1931, nec Kraatz, 1882; =auronitens auronitens Fabricius viridicolor Csiki, 1927; un; nom.pro viridis Ferrer y Vert; =rutilans rutilans Dejean viridicupreior Deuve & Mourzine, 1993; Aulonocarabus kolymensis ssp. viridicupreus (Raynaud, 1970); un; =solieri solieri Dejean viridifossulatipennis Breuning, 1964; un; =gilnickii gilnickii Deyrolle viridifossulatus Fairmaire, 1887; Aristocarabus sp. viridilimbatus Motschulsky, 1859; Morphocarabus odoratus ssp. viridilimbatus Kraatz, 1878, nec Motschulsky, 1859; =ullrichi fastuosus Palliardi viridilucens Barthe, 1909; =violaceus baeterrensis Lapouge viridimarginatus Letzner, 1850; un; =scheidleri scheidleri Panzer viridimarginatus Letzner, 1850; un; =convexus convexus Fabricius viridimarginatus Dalla Torre, 1877; un; =convexus convexus Fabricius viridimarginatus Kraatz, 1886; =hummeli smaragdulus Kraatz viridimicans Kraatz, 1878; =creutzeri creutzeri Fabricius viridimicans Enderlein, 1929, nec Kraatz, 1878; =abbreviatus Brullé

43 viridiniger Kraatz, 1886; =arcensis conciliator Fischer viridiobscurus Hauser, 1913; un; =elysii connectens Hauser viridiobscurus Hauser, 1914; un; =elysii lopinensis Hauser viridiobscurus Hauser, 1914; un; =elysii chikongschanensis Hauser viridiobscurus Hauser, 1914; un; =smaragdinus longipennis Chaudoir viridipennis Lewis, 1880; Damaster blaptoides ssp. viridipennis Kraatz, 1886, nec Lewis, 1880; =hummeli smaragdulus Kraatz viridipennis Bleuse, 1896, nec Lewis, 1880; =auronitens costellatus Géhin viridipennis Hauser, 1912; un; =elysii connectens Hauser viridipennis Hauser, 1921, nec Lewis, 1880; =blaptoides oxuroides Schaum viridipennis Fleischer, 1925; un; =scheidleri helleri Ganglbauer viridipunctus Kraatz, 1886; =clathratus maacki Morawitz viridirugulosior Deuve, 1996; Neoplesius kamensis ssp. viridis Dejean, 1826; =splendens Fabricius viridis Letzner, 1849; un; =cancellatus excisus Dejean viridis Letzner, 1850; un; =arcensis arcensis Herbst viridis Letzner, 1850; un; =auronitens auronitens Fabricius viridis Letzner, 1850; un; =scheidleri scheidleri Panzer viridis Schilsky, 1888, nec Dejean, 1826; =alpestris hoppei Germar viridis Ferrer y Vert, 1910; un; =rutilans rutilans Dejean viridis Everts, 1918; un; =cancellatus carinatus Charpentier viridis Hauser, 1921, nec Dejean, 1826; =smaragdinus longipennis Chaudoir viridis Hauser, 1921, nec Dejean, 1826; =gehinii gehinii Fairmaire viridis Fleischer, 1930; un; =obsoletus nagyagensis Birthler viridis Vicol, 1987; un; =auronitens escheri Palliardi viridissimus Kraatz, 1876; =scabrosus tauricus Bonnelli viridissimus Hauser, 1913; un; =elysii connectens Hauser viridissimus Pater, 1938; un; =pyrenaeus jeanneli Deuve, 1994 viridulus Kraatz, 1878; =ullrichi rhilensis Kraatz viridulus Ragusa, 1905, nec Kraatz, 1878; =morbillosus alternans Palliardi vitalisi Lapouge, 1916; Apotomopterus sp. vitiosus Csiki, 1906; un; =rothi hampei Küster vittatus Lapouge, 1902; un; =splendens Olivier vivesi Jeanne, 1973; =lusitanicus ?lusitanicus Fabricius vladimirskyi Dejean & Boisduval, 1830; Piocarabus sp. vlascanus Born, 1909; =violaceus wolffii Dejean vlasuljensis Apfelbeck, 1894; Megodontus violaceus ssp. vocontianus Keith, 1995; =violaceus mixtus Géhin vogeli Breuning, 1933; un; =cancellatus tibiscinus Csiki vogtae Beheim & Breuning, 1943; Apotomopterus sp. vogti Beheim & Breuning, 1943; Aulonocarabus pseudokoreanus ssp. vogtianus Beheim & Breuning, 1943; Leptocarabus sp. vojnitsi Mandl, 1980; Aulonocarabus canaliculatus ssp. volgensis Csiki, 1927; nom.pro jureceki Born, 1922; =regalis regalis Fischer volhynicus Lutshnik, 1927; =ullrichi ullrichi Germar volujakianus Apfelbeck, 1894; =caelatus caelatus Fabricius vomax Dejean, 1826; =hungaricus mingens Quensel vorax Bernau, 1913; un; =cancellatus scythicus Schaum voriseki Novotný, 1975; un; =varians armeniacus Mannerheim vulcani (Lebis, 1955); un; =auronitens auronitens Fabricius vulcanus Le Moult, 1912; hyb Chrysocarabus splendens/auronitens vulneratus Lapouge, 1915; un; =odoratus odoratus Motschulsky

W wagae Fairmaire, 1882; Neoplesius sp. wagneri Küster, 1846; =saphyrinus saphyrinus Cristoforis & Jan wakamatsuensis Imura & Mizusawa, 1994; Ohomopterus japonicus ssp. wakoi Imura, 1995; Coptolabrus pustulifer ssp. wallichi Hope, 1831; Meganebrius sp. wallichi Kraatz, 1895; =boysi boysi Tatum walteri Reitter, 1896; =hortensis neumeyeri Schaum waltersachi Mandl, 1984; =nemoralis nemoralis Müller wangdanus Imura, Su & Osawa, 1997; Neoplesius markamensis ssp. wangxingensis Deuve & Li, 1998; =hummeli pusongensis Imura wankae Sokolar, 1919; =cancellatus cancellatus Illiger wanxianicus Imura, 1995; Coptolabrus formosus ssp. washingtoni Casey, 1920; =chamissonis Fischer weibaoensis Deuve, 1992; Pseudocoptolabrus taliensis ssp. weidneri (Lassalle, 1990); Procerus scabrosus ssp. weigeli Panzer, 1801; =variolosus variolosus Fabricius weiratheri Breuning, 1934; un; =intricatus intricatus Linnaeus weiratherianus Breuning, 1936; un; =coriaceus banaticus Redtenbacher weisei Reitter, 1883; =convexus dilatatus Dejean weishanicola Deuve, 1999; Apotomopterus bousquetellus sp. wengshuiensis Deuve, 1994; Pseudocoptolabrus taliensis ssp. wenshuicus Font, 1998; Coptolabrus pustulifer ssp. wenxianensis Deuve, 1992; Archaeocarabus pseudolatipennis ssp. wenxianicola Deuve, 1996; Pseudocranion sp. weolseae (Kwon & Lee, 1984); =koreanus koreanus Reitter werneri Reitter, 1899; =ullrichi rhilensis Kraatz; Bosnia,Serbia werzhutzkii Berlov & Shilenkov, 1996; iss of verzhutzkii Berlov & Shilenkov, 1996 whitei Deyrolle, 1852; =lineatus lineatus Dejean wiedemanni Ménétriés, 1836; Archicarabus sp. wiedmanni Cristoforis & Jan, 1837, nec wiedemanni Ménétriés, 1836; =chevrolati thirki Chaudoir

44 wilkinsi Semenov, 1903; =pupulus pupulus Morawitz willemsi (Culot, 1986); un; =nitens Linnaeus wimmeli Schulz, 1900; =granulatus granulatus Linnaeus winkleri Weyenberg, 1869; fos; ?Carabus sp. winkleri Breuning, 1933, nec Weyenberg, 1869; =graecus morio Mannerheim winklerianus Breuning, 1933; =wiedemanni wiedemanni Ménétriés wirthumeri Mandl, 1964; un; =cancellatus excisus Dejean wittmeri Mandl, 1974; Meganebrius wallichi ssp. wittmeri Mandl, 1975, nec Mandl, 1975; =tatsienlui Breuning wittmerianus Mandl, 1975; nom.pro wittmeri Mandl, 1975, nec Mandl, 1974; =tatsienlui Breuning wittmerorum Heinertz, 1978; Imaibius sp. witzgalli Reiser in Freude, 1976; ?nud; =menetriesi pacholei Sokolar wockei Born, 1898; =problematicus harcyniae Sturm wolffii Dejean, 1826; Megodontus violaceus ssp. wolfianus Reitter, 1888; =prometheus prometheus Reitter wolongensis Deuve, 1993; Apotomopterus ascendens ssp. worleei Beuthin, 1894; =auratus auratus Linnaeus wrzecionkoi Deuve, 1991; ?hyb Cratocechenus akinini puellus/Eotribax valikhanovi wuangxianus Kleinfeld, 1998; Archaeocarabus sp. wuduensis Deuve, 1997; Pseudocranion wenxianicola ssp. wudumontanus Imura, 1998; Eccoptolabrus exiguus ssp. wufengensis Kleinfeld, 1998; Archaeocarabus nanosomus ssp. wulffiusi Morawitz, 1862; Morphocarabus sp. wuliangensis Cavazzuti, 1997; Apotomopterus ludivinae ssp. wumingensis Deuve, 1992; Apotomopterus sp. wushanling Kleinfeld, 1998; Archaeocarabus sp.

X xanthicus Darge, 1992; =torosus cankirianus Blumenthal & Breuning xanthiensis (Mařan, 1952); un; =coriaceus hopffgarteni Kraatz xaxarsi Born, 1917; =problematicus planiusculus Haury in Géhin xiangchengicus Deuve, 1993; Rhigocarabus sp. xianfengicus Deuve & Li, 2000; Archaeocarabus nestor ssp. xiaoxiangensis Deuve, 1995; Apotomopterus sp. xiaoxianglingensis Cavazzuti, 1996; Apotomopterus patroclus ssp. xiei Deuve, 1992; Rhigocarabus sp. xingshanensis Imura, 1996; Archaeocarabus sp. xingziensis Kleinfeld, 1996; =dargei Deuve xinlongicus Deuve, 1995; =buddaicus gracilicollis Semenov xinyingensis Kleinfeld, 1998; =dreuxi jinxiuensis Imura xiongi Imura & Su, 2000; Coptolabrus pustulifer ssp. xiuyanensis Deuve & Li, 1998; Parhomopterus sp. xueshanicola Deuve, 1992; Pseudocoptolabrus taliensis ssp. xui Deuve, 1996; Rhigocarabus roborowskii ssp. xunyangbanus Imura, 1998; Leptocarabus marcilhaci ssp. xupuensis Kleinfeld, 1998; Archaeocarabus sp.

Y yaanensis Deuve & Tian, 1999; Apotomopterus grossefoveatus ssp. yaconinus Bates, 1873; Ohomopterus sp. yajiangensis Deuve, 1994; Neoplesius lama ssp. yak Deuve, 1991; Pseudocranion tibetanophilus ssp. yakalamakus Lassalle, 1993; Sphodristocarabus gilnickii ssp. yamaokai Ishikawa & Kubota, 1994; Ohomopterus yaconinus ssp. yamato (Nakane, 1953); Ohomopterus sp. yanggangensis Deuve & Li, 2000; Coptolabrus smaragdinus ssp. yangmingensis Deuve, 1995; Isiocarabus dreuxi ssp. yangminshanicus Font, 1997; Coptolabrus augustus ssp. yangxianensis Deuve, 1999; Archaeocarabus pseudolatipennis ssp. yanjianganus Imura & Mizusawa, 1998; Eocechenus sp. yanmenensis Deuve, 1996; Pseudocoptolabrus taliensis ssp. yanyuanicus Cavazzuti, 1996; Archaeocarabus yunnanus ssp. yao Imura, 1999; Archaeocarabus morphocaraboides ssp. yaophilus Deuve, 1990; Apotomopterus sp. yaraligozi Battoni, 1982; Cytilocarabus cribratus ssp. yasudai (Ishikawa, 1971); Megodontus kolbei ssp. yenicensis Lassalle, 1998; =saphyrinus pseudosaphyrinus Schweiger yeongsiki (Kwon & Lee, 1984); =fraterculus jirisanensis Ishikawa & Kim yezoensis Bates, 1883; Carabus granulatus ssp. yichunensis Deuve & Li, 1999; Coptolabrus smaragdinus ssp. yingpanjienus Cavazzuti, 1999; Archaeocarabus pseudolatipennis ssp. yokoae Deuve, 1988; Leptocarabus sp. yonghongi (Li, 1992); =?wulffiusi jilinensis Deuve & Imura yonghongi (Li, 1992), nec (Li, 1992); =fraterculus fraterculus Reitter yonghongianus Deuve, 1992; nom.pro yonghongi Li; =?wulffiusi jilinensis Deuve & Imura yongshan Kleinfeld, 1999; Apotomopterus longeantennatus ssp. yooni Deuve, 1990; Acoptolabrus leechi ssp. yoroensis Ishikawa & Kubota, 1995; Ohomopterus insulicola ssp. yoshiyukii Imura & Mizusawa, 1999; Ohomopterus japonicus ssp. yuae Deuve, 1989; Apotomopterus sp. yuanshanensis Kleinfeld, 1996; Apotomopterus sp. yuanbaoensis Deuve, 1994; Apotomopterus sp. yubariensis (Ishikawa, 1966); =kolbei aino Rost

45 yubeicus Deuve & Tian, 1999; Apotomopterus arrowi ssp. yudanus Nakane, 1977; nom.pro nakamurai (Ishikawa, 1968), nec (Ishikawa,1966); Diocarabus harmandi ssp. yuellus Deuve, 1992; Apotomopterus sp. yuianus Deuve, 1991; =elysii elysii Thomson yulongxuensis Deuve, 1990; Pseudocoptolabrus taliensis ssp. yunanensis Born, 1905; Apotomopterus sp. yungeri Breuning & Ruspoli, 1970; un; ios of juengeri Breuning & Ruspoli yunglaensis Deuve, 1997; Neoplesius wagae ssp. yunkaicus Deuve, 1992; Apotomopterus sauteri ssp. yunlingensis Deuve, 1991; Apotomopterus sp. yunlingicus Deuve, 1994; Archaeocarabus oblongior ssp. yunlongensis Kleinfeld, 2000; =Apotomopterus delavayi ssp. yunnanicola Deuve, 1989; Apotomopterus sp. yunnanicus Deuve, 1998; Apotomopterus szetschuanus ssp. yunnanus Fairmaire, 1886; Archaeocarabus sp. yunnanus Born, 1910, nec Fairmaire, 1886; =gemmifer loczyi Csiki, 1927 yuwuensis Kleinfeld, 1996; Apotomopterus guizhouensis ssp. yushuensis Deuve, 1991; Eocechenus sp.

Z zabljakensis Eidam, 1927; =violaceus azurescens Dejean zaianus Antoine, 1941; =famini piraticus Fairmaire zaikai Obydov, 1999; Morphocarabus chaudoiri ssp. zakharschewskii Motschulsky, 1845; =stscheglowi Mannerheim zanderi Battoni & Blumenthal, 1973; =macrogonus resplendens Breuning zanteanus Breuning, 1964; =coriaceus joniensis Breuning zantunensis Cavazzuti & Ratti, 1998; Apotomopterus szetschuanus ssp. zaraensis Breuning & Ruspoli, 1970; =rumelicus rumelicus Chaudoir zarudnyi Semenov & Znojko, 1932; Cyclocarabus sp. zawadzkii Kraatz, 1854; Morphocarabus scheidleri ssp. zepcensis Reitter, 1902; Megodontus croaticus ssp. zeytinlikensis Charet, 1996; =puschkini ishikawaianus Breuning & Ruspoli zhanglaensis Deuve, 1991; Pseudocranion sp. zhanglaicola Deuve, 1996; Pseudocranion fumigatus ssp. zhaojueanus Imura, 1999; Archaeocarabus morphocaraboides ssp. zheduoshanensis Cavazzuti, 1997; Rhigocarabus jiulongensis ssp. zherichini Shilenkov, 1990; Morphocarabus sp. zhongnanshan Kleinfeld & Schütze, 1997; Apotomopterus grossefoveatus ssp. zhongtiaoshanus Imura & Yamaya, 1994; Coptolabrus smaragdinus ssp. zhubajie Imura, 1993; Rhigoidocarabus sp. ziegenhorni Hauser, 1921; un; nom.pro aereus Hauser, 1919; =augustus augustus Bates ziegleri Géhin, 1876, nec Duftschmid, 1812; nom.pro aereus Dejean; =arcensis arcensis Herbst ziegleri Kraatz, 1883, nec Duftschmid, 1812; =cancellatus emarginatus Duftschmid zilahiensis Csiki, 1906; =rothi hampei Küster zinaidae Obydov, 1997; Morphocarabus spasskianus ssp. znojkoi Semenov & Breuning, 1931; Cratocechenus sp. znojkoiellus Deuve, 1988; nom.pro pullus Semenov & Znojko; =pullus Semenov & Znojko zoigensis Deuve, 1989; Rhigocarabus roborowskii ssp. zolotarevi Zamotajlov, 1988; Archiplectes prometheus ssp. zoppai Kraatz, 1900; =rothi hampei Küster zorkae Novotný & Voříšek, 1988; =apschuanus pseudoplatessa Gottwald zoufali Fleischer, 1922; un; =cancellatus intermedius Dejean zugmayeriae Reitter, 1888; =prometheus prometheus Reitter zwickii Heer, 1837; =auronitens auronitens Fabricius zyrjanovskianus Shilenkov & O.Berlov, 1996; Morphocarabus aeruginosus ssp. zyzzyus Lassalle, 1992; Tribax puschkini ssp.

46 LEGEND: species major subspecies (well geographically and morphologically limited) minor subspecies (in global faunistic view, numerous taxa can be treated as synonyms) =synonym, local form etc. (in local faunistic view, some taxa can be treated as subspecies) ~permanently invalid name (mostly junior homonym or unavailable infrasubspecific name)

un unavailable (mostly infrasubspecific) name spn suppressed name nud nomen nudum cos correct original spelling ios incorrect original spelling iss incorrect subsequent spelling lsu junior homonym treated as valid name due to long standing use hyb hybrid form fos fossil form inc taxon incertae sedis HT holotype LT lectotype NT neotype TNF type not fixed LNF type locality not fixed underlined type locality 1m/1f only 1 male/1 female specimen known