© Zoological Institute, St.Petersburg, 2002

Checklist and nomenclatural notes on the Chinese (). II.

D.A. Rider, L.Y. Zheng & I.M. Kerzhner

Rider, D.A., Zheng, L.Y. & Kerzhner, I.M. 2002. Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. ZoosystematicaRossica, 11(1): 135-153. A checklist of all known Chinese Pentatominae is presented with synonymical, distributional, and nomenclatural notes. New name: Eurydemalingi forE. montana Ling, 1989 Gunior homonym of E. dominulus montana Kerzhner, 1972). New synonymies: Carbula humerigera Uhler, 1860 (non auctorum!) = C. obtusangula Reuter, 1881; Menida disjecta (Uhler, 1860) = M. scotti Puton, 1886; Lakhonia Yang,1936 = Oedocoris Zheng & Liu, 1987; Lakhonia nigripes Yang, 1936 = Oedocoris eburneus Zheng & Liu, 1987; chapana (Distant, 1921), sp. dist. (not syn. of P. unicolorel/a Kirkaldy, 1909) = P. haemorrhoidalis Lindberg, 1934. The name Carbulaabbreviata (Motschulsky, 1866) is restored for C. humerigera auct. Lectotypes are designated for Carbula humerigera, Menida disjecta, and Palomena chapana.

David A. Rider, Department ofEntomology, Box 5346, UniversityStation, North Dakota State University, Fargo, North Dakota 58102, USA.

Le-yi Zheng, Department of Biology, Nankai University, Tianjin 3000071, P.R. .

l.M. Kerzhner, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.

Introduction Checklist of Chinese Pentatominae

Part I of this series (Rider & Zheng, 2002) gave Acesines laticeps Zheng & Liu, 1987a. Jiangxi, Yunnan. a brief history of the taxonomic work on the A. tridentatus Xiong & Liu, 1995. Yunnan. A. trifasciatusXiong & Liu, 1995. Yunnan. Chinese pentatomid fauna, and a checklist and Acrocorisellus serraticollis (Jakovlev, 1876a) (= nomenclatural notes on the subfamily Asopinae. Pentatomapulchra Hsiao & Cheng, 1977). Shanxi. Also This second part of the series contains a check­ known from eastern Russia and Korea. list of all known pentatomid species belonging Aednus obscurus Dallas, I 851 (= A. similis Haglund, 1868). Hunan, Xizang. , , , Malay­ in the subfamilyPentatominae. Once again, there sia, and . are many nomenclatural problems in the current A. ventralis Dallas, 1852. Fujian, Guangdong, Hainan. literature; this paper attempts to clarify the sta­ Also known fromnorthern India. tus of all known species, and to give distribu­ acuminata (Linnaeus, 1758) (= Cimex rostratus tional records for all taxa. There are undoubt­ De Geer, 1773 (unnecessary new name); C. tessaroph­ edly errors and/or omissions; the authors request thalmus Schrank, 1781; Aelia neglecta Dallas, I 851; A. burmeisteri Kiister, 1852; A. pallida Kiister, 1852; A. that these mistakes be brought to their attention turanica Horvath, 1895; A. acuminata f. nigripes Wagner, so these lists can be updated. Please note that in 1938). Sichuan, Xinjiang. Widespread Palaearctic spe­ the checklist, several species are presented in cies. - Note: Distribution records of Jilin, Hebei, Shanxi, square brackets. Some of these species have been Shaanxi, Jiangxi, Jiangsu (Yang, 1962) probably belong reported from China in error, or their Chinese to other Aelia species. A. chinensis Rider & Rolston, 1995 (= A. bifidaHsiao records are questionable. These species probably & Cheng, 1977). Nei Monggol, Shanxi, Sichuan. Note: do not occur in China, or at least their existence A. bifidaHsiao & Cheng ( 1977) is a primary junior homo­ in China needs verification. Informationis given nym of A. bifida Costa ( 1847), which is now considered a in the followingsequence: valid name, synonyms valid species in the . Rider & Rolston (in parentheses), distribution in China (provinces (1995) proposed A. chinensis as a replacement name. listed in alphabetic order), distribution outside Derzhansky (personal communication) has indicated that this species may be a junior synonym of A. k/ugii. of China, notes. A. fieberi Scott, 1874 (= A. nasuta Wagner, 1960). 136 D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae • ZOOSYST. ROSSICA Vol. 11 ! Gansu, Hebei, Heilongjiang, Henan, Hubei, Hunan, Alcimocoris coronatus (Stal, 1876). Guangdong, Hai­ Jiangsu, Jiangxi, Jilin, Liaoning, Nei Monggol, Shaanxi, nan, Yunnan. Also recorded from India, Myanmar, and Shandong, Shanxi, Sichuan, Xizang, Yunnan, Zhejiang. Vietnam. Also recorded from eastern Russia, Korea, and Japan. - A. flavicornis(Distant, 1887). Yunnan.Originally de­ Note: Even though Kerzhner (1978) placed A. nasuta as scribed from northern India. a junior synonym of A. fieberi, several modem workers A. japonensis (Scott, 1880). Jiangsu, Shanxi, Taiwan, continue to use the name A. nasuta; these references Zhejiang. Also reported from Japan and Korea. should be transferredto A. fieberi. A. potanini Horvath, 1892. Gansu, Shaanxi. A. furcula Fieber, 1868 (= A. simillima Reuter, 1900). Ambiorix aenescens Stal, 1876. Xizang. Originally Xinjiang. Known from Afghanistan, Armenia, Austria, described from India. Azerbaijan, Greece, Iran, Kasakhstan, Kyrgyzstan, south­ Amblycara gladiatoria Stal, 1876. Also recorded from ern Russia, Tajikistan, Turkey, Ukraine, and Yugoslavia. India, , Japan, the , and Australia. A. granum Jakovlev, 1903a (= A. demissa Kiritshenko, Amyntor unicolor (Walker, 1867) (=' Halys obscura 1914b ). Nei Monggol. Also known from Kyrgyzstan and Dallas, 1849 (preoccupied); Oestopis terra Distant, Tadjikistan. - Note: Derzhansky (personal communica­ 1879a). Gansu, Guangxi, Guizhou, Jiangxi, Sichuan, tion) has indicated that this species may be a junior syno­ Yunnan. Also recorded fromBhutan, India, and Vietnam. nym of A. klugii. Note: Halys obscura Dallas, 1849, is a junior primary A. klugii Hahn, 1833. Nei Monggol. Widespread homonym of Ha/ys obscura Westwood, 1837 (which is Palaearctic species. now a species in the genus Sarju).Th e next available name A. sibirica Reuter, 1884 ('" A. henschi Montandon, is Bo/aca unico/or Walker, I 867 (see Rider & Rolston, 1886; A. aspersa Jakovlev, 1903a). Gansu, Nei Monggol, 1995). All known records of A. obscurus should be re­ Ningxia, Qinghai, Shanxi, Xinjiang. Austria, Iran, Italy, ferred to A. unicolor. Russia, Kazakhstan, Kyrgyzstan, Mongolia, Ukraine. Anaca fasciata (Distant, I 900a). Fujian, Guangdong, [Aeliomorpha lineatocollis (Westwood, 1837)]. Jiangxi, Yunnan.Also known from India and Sri Lanka. Hainan. Also recorded from Afghanistan, Israel, Syria, A. florens (Walker, 1867) (= Hyllus aeruginosus India, Sri Lanka, and Pakistan. -Note: The Hainan record Haglund, 1868). Fujian, Guangdong, Hainan. India and (Chen & Huang, 1996) is quite far from any other known Sri Lanka, southeast Asia, , Indonesia. record for this species; it needs verification. [A. punctiventris Breddin, 1904]. China?, Vietnam.­ A. yunnanensis Zheng & Liu, 1987a. Yunnan. Note: This species is only doubtfullyincluded in the Chi­ Aenaria lewisi (Scott, 1874). Fujian, Gansu, Guangxi, nese fauna.Breddin (1904) originally described this spe­ Hainan, Hunan, Jiangsu, Jiangxi, Sichuan, Taiwan, cies from Vietnam; Kirkaldy (1910) gave a record of Zhejiang. Also known from India, Korea, and Japan. China in double brackets. This was repeated by Hoffmann A. pinchii Yang, 1934. Anhui, Fujian, Guangdong, (1935) and Tang( 1935). There has been no verifiedrecord Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, froma specific locality in China. Jiangxi, Sichuan, Zhejiang. Antestiopsis anchora (Thunberg, 1783) (= Strachia A. zhangi Chen, 1989. Guangdong. pardalis Walker, 1867; S. platyspila Walker, 1867). Aeschrocoris ceylonicus Distant, 1899. Fujian, Guang­ Guangdong, Guangxi, Hainan, Xizang, Yunnan. Bangla­ dong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, desh and India eastward through China and southeast Asia Taiwan, Yunnan, Zhejiang. Also recorded from India and to Japan, Malaysia, and Indonesia. -Note: Leston (1952) Sri Lanka. described the genus Antestiopsis, and designated anchora A. obscurus (Dallas, 1851). Fujian, Hunan, Yunnan, as type species. Although several African species have Zhejiang. India, Sri Lanka, Myanmar, Indonesia (Java). been placed in this genus, most workers from the Indian [A. rugulosus (Distant, 1921)]. Vietnam. -Note: Dis­ and Oriental regions continue to refer to this species as tant (1921) described this species from Vietnam. Tang Antestia anchora. (1935) listed China (without specific location); he prob­ A. cruciata (Fabricius, 1775) (c pantherina ably was repeating Hoffmann's(1932a) listing ofrugu/osa Westwood, 1837; Strachia geomelrica Nietner, 1861; S. fromChina (but Hoffmann'slist of Chinese Pentatomidae velata Walker, 1867; S. subacta Walker, 1867; P/autia also included those fromKorea and lndo-China). No one picturata Distant, 19 l 8 ). Yunnan. Pakistan, India, Sri has reported a specific recordof this species fromChina. Lanka eastward through southeast Asia to the Philippines A. tuberculatus (Stal, 1865). Fujian, Hunan, Zhejiang. and Indonesia. - Note: Leston (1952) also placed this Also known fromIndia and Myanmar. species in his newly erected genus Antestiopsis. Several Agaeus longirostris Lin & Zhang, 1984. Yunnan. Africanworkers have referredto cruciata in Antestiopsis, A. mimus Distant, 1887. Jiangxi, Yunnan. Also re­ but most workers from India and from China continue to corded from India and Vietnam. refer to this species as Antestia cruciata. A. tessellatus Dallas, 1851. Hainan. Also recorded aliena (Reuter, 1891) (== from India, Myanmar, and Vietnam. laticol/is Jakovlev, 1905). Nei Monggol. Also reported Agathocles limbatus Stal, 1876. Fujian. Originally from Belarus, Estonia, Mongolia, Russia, and Korea. described fromIndia. A. eurynota eurynota Horvath, 1908 (= Carpocoris A. yunnanensis Zhang & Lin, 1984. Yunnan. oshanini Kiritshenko, 1952). Xinjiang. Also recorded from Agonoscelis femoralis Walker, 1868. Guangxi, Gui­ Kyrgyzstan, Tajikistan, and Mongolia. zhou, Yunnan. Also known fromIndia, Sri Lanka, Myan­ A. lindbergi (Tamanini, 1962). Gansu, Nei Monggol, mar, and Vietnam. Qinghai, Shaanxi, Shanxi. Also known from Mongolia A. nubilis (Fabricius, 1775) (= Pentatoma grata Palisot and eastern Russia. de Beauvois, 1811; Agonoscelis indica Spinola, 1837; A. lunulata (Goeze, 1778) (� Cimexlynx Fabricius, 1794; Ae/ia crucifera Westwood, 1837; Nevroscia sulciventris Carpocoris lynx var. decolor Ferrari,1874; C. lunu/atusvar. Ellenrieder, 1862). Fujian, Guangdong, Guangxi, Gui­ pallidior Kirkaldy, 1909; C. lunulatus var. nigricornis zhou, Hainan, Hunan, Jiangxi, Sichuan, Xizang, Yunnan, Kormilev,1936). Xinjiang. Widespread Palaearctic species. - Zhejiang. Occurs from Pakistan, India, and Sri Lanka Note: The northern China records of Yang, 1962 (Beijing, eastward through China and southeast Asia to Japan, the Hebei, Heilongjiang, Henan, Jilin, Nei Monggol, Shanxi) Philippines, and Indonesia. probably refer to other Antheminia species. ZOOSYST. ROSSICA Vol.11 • D.A. Rideret al.: Chinese Pentatomidae. II. Pentatominae 137 A. mongolica (Kerzhner & Josifov, 1966). Nei Mong­ Qinghai, Shaanxi, Xinjiang, Xizang. Widespread gol. Originally described fromMongolia. Palaearctic species. A. pusio pusio (Kolenati, 1846) (= Mormidea helian­ B. signatum Jakovlev, 1879 (= B. turanicum Horvath, themi Garbiglietti, 1869; Carpocoris kirgisicus Jakovlev, 1889b; B. signatum var. immaculiscutum Kiritshenko, 1905; C. bergi Jakovlev, I 905; C. absinthii orientalis Ho­ 1922). Xinjiang. Also known fromArmenia, Azerbaijan, berlandt, 1956). Xinjiang. Widespread Palaearctic species. Iran, Kazakhstan, Turkey, and Turkmenistan. A. pusio longiceps (Reuter, 1884) (= pusio Cahara tibetana Zheng & Liu, 1986. Xizang. esakii Tamanini,1962). Hebei, Jilin, Liaoning, Nei Mong­ Cappaea taprobanensis (Dallas, 1851) (= C. multi­ gol, Shaanxi, Shanxi, Xinjiang. Also known from Mon­ linea Ellenrieder, 1862). Fujian, Guangdong, Guangxi, golia and eastern Russia. Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Tai­ A. varicornis (Jakovlev, 1874c) (= baical­ wan, Yunnan. Also reported from India, Sri Lanka, Bang­ ensis Jakovlev, 1894; D. varicornis montandoni Sien­ ladesh, Myanmar, and Indonesia. kiewicz, 1954). Hebei, Heilongjiang,Nei Monggol, Qing­ C. tibialis Hsiao & Cheng, 1977. Shandong. - Note: hai, Shaanxi, Shanxi, Xinjiang. Occurs fromeastern and This may actually be a species of Homalogonia. southern Europe eastward to Korea. - Note: Many work­ Carbula abbreviata (Motschulsky, 1866) (= Penta­ ers continue to include this species in the genus Codophila. toma humerigera: of authors (not Uhler, 1860); Arma Axiagastus mitescens Distant, 190 I b. Fujian, Guang­ japonica Walker, I868). Jilin. Described fromJapan; also dong, Hainan. Also known from southeast Asia and the known fromKorea and easternRussia. - Note: Wehave Philippines. examined the type material (a single femalespecimen) of A. rosmarus Dallas, 1851. Fujian, Guangdong, Jiangxi, Pentatoma humerigera Uhler, 1860, and have determined Taiwan,Zhejiang. Also recorded fromIndia, Japan, south­ that it is not conspecific with what most workers have eastAsia, Philippines, Indonesia, New Guinea, and north­ called C. humerigera. The humeral angles are not spinose. ernAustralia. They are rounded and quite prominent, but not as nar­ hilaris (Burmeister, 1835) (= Cimex pictus rowly rounded as in C. crassiventris (Dallas, 1849). Fabricius, 1775 (preoccupied); C. hebraicus Germar, Uhler's specimen most closely resembles either C. putoni 1838; C. jucundus Klug, 1845; Bagrada cruciferarum (Jakovlev, 1876) or C. obtusangula Reuter, 1881. Speci­ Kirkaldy, 1909). Sichuan, Xinjiang, Yunnan.Widespread mens of C. putoni have the head and anterior areas of the southern Palaearctic species, also occurring throughout pronotal disk shiny aeneous black, contrasting with the Africa, and in India, Sri Lanka, and Myanmar. - Note: dark brown of the remainder of the dorsum; specimens This species was originally described as Cimex pictus of C. obtusangula are more uniformly dark brown. The Fabricius, 1775, which is a junior primary homonym of coloration of the type specimen of C. humerigera is more C. pictus Drury, 1773, a species in the Coreidae. Kirkaldy uniformly dark brown. The descriptions of both C. ( 1909) proposed Bagrada cruciferarumas a replacement abbreviataand C. japonica indicate that the humeral an­ name, but there were several earlier names available. The gles are spinose as in this species. See also comments oldest available name is Cimex hi/aris Burmeister, 1835, under C. humerigera. which was originally synonymised with C. pictus by C. crassiventris (Dallas, 1849). Anhui, Fujian, Guang­ Horvath (1909); this synonymy has been recently con­ dong, Guangxi, Guizhou, Hainan, Heilongjiang, Hubei, firmed(Linnavuori, 1975). All records of Bagrada cruci­ Hunan, Jiangsu, Jiangxi, Shaanxi, Shanxi, Sichuan, Tai­ ferarum should be referredto this species. wan, Xizang, Yunnan, Zhejiang. Also known from Bhu­ B. kaufmanni (Oshanin, 1871) (= B. kaefmanni var. tan, India, Myanmar, and Japan (Okinawa, Ryukyu Is­ pallida Kiritshenko, 1914b). Xinjiang.Also known from lands, Tokara Islands). Afghanistan, Iran, Kazakhstan, Tajikistan, Turkmenistan, C. eoa Bergroth, 1891. Sichuan. and Uzbekistan. C. humerigera (Uhler, 1860) (= C. obtusangula B. qinlingensis Zheng, 1982. Shaanxi. Reuter, 1881b, syn. n.). Anhui, Fujian, Gansu, Guang­ B. stolata Horvath, 1936 (= B. stolata var. quadri­ dong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, maculata Horvath, 1936). Xinjiang. Also known from Jiangxi, Qinghai, Shaanxi, Shanxi, Sichuan, Yunnan, Bulgaria, Czech Republic, Greece, Hungary, Italy, Ka­ Zhejiang. Also known from Japan (the Ryukyu Islands). zakhstan, Kyrgyzstan, Romania, Russia (south of Euro­ - Note: Wehave examined the type material of C. humeri­ pean territory), Slovakia, Turkey, and Ukraine. - Note: gera (Uhler, 1860), and it appears to be conspecificwith The Bagrada sp. illustrated by Yang (1964: 101, pl. VI, C. obtusangula Reuter, 188 I. The humeral angles are fig.53) is this species; also there is a specimen of B. stolata prominent, but they are rounded, not spinose apically; from China, Xinjiang (Kiines River), in the Zoological the overall coloration is rather uniform dark brown, not Institute, Russian Academy of Sciences, St.Petersburg. distinctly darker on the head and anterior disk of scutellata Zheng & Liu, 1987a. Yunnan. pronotum. See also comments under C. abbreviata. It is B. ,sinica Zheng & Liu, 1987a. Guangdong. not possible to tell how many specimens Uhler originally Belopis unicolor Distant, I 879a. Guangdong, Guizhou, had when he made his description, so the one femalespeci­ Yunnan. Also recorded fromIndia and Myanmar. men located is here designated lectotype. It has the fol­ Brachymna bificeps Chen, 2000. Guangdong. lowing label data: a) "PR Uhler Collection"; b) [red square B. castanea Lin & Zhang, 1993. Fujian, Jiangxi. label with no writing]; c) "Pentatoma humerigera Uhler B. humerata Chen, 1989. Guandong. Tokanosima"; and d) Pentatoma humerigera Uhl. Toka. B. tenuis Stal, 1861 (= Balsa extenuata Walker,1867). Det UHLER". This specimen is conserved in the U.S. Anhui, Fujian, Guangdong, Guizhou, Henan, Hubei, National Museum, Washington, D.C. Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang. C. indica (Westwood, 1837) (= C.fuscaDistant, 1887). Brachynema germarii (Kolenati, 1846) (= Cimex Sichuan, Xizang. Also recorded from India and Nepal. virens Klug, I 845 (preoccupied); Pentatoma anabasis [C. insocia (Walker, 1868)]. Macao. Described from Becker, I 867; Rhaphigaster biplaga Walker, 1867; India, and also known fromPakistan. - Note: The Macao Oncoma germari var. flavomarginatus Jakovlev, 1871; record (Kirkaldy 1910) needs verification.Kirkaldy was 0. germari var. griseus Jakovlev, 1871; B,:achynema unsure of his determination as evidenced from his state­ melanota Jakovlev, 1874b; B. virens var. a/ternatum ment: "Ifl have rightly identifiedthis, it is not a Carbula." Horvath, 1899). Gansu, Hebei, Nei Monggol, Ningxia, C. maculata Hsiao & Cheng, 1977. Sichuan, Yunnan. 138 D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae • ZOOSYST. ROSSICA Vol. 11 C. producta Distant, 1901a. Yunnan. Also recorded C. varia (Fabricius, 1787) (= Cimex lunula Fabricius, from lndia and Myanmar. 1794; C. eryngiiGermar, 1814 (preoccupied); Carpocoris C. putoni (Jakovlev, 1876b) (= C. amurensis Reuter, bilunulata Kolenati, 1846; C. bilunulata var. persicae 1881b). Beijing, Hebei, Heilongjiang, Liaoning, Shan­ Kolenati, 1846; C. bilunulata var. nubica Kolenati, 1846; dong, Shanxi, Sichuan, Xinjiang. Also known fromeast­ Pentatoma laborans Costa, 1847; P. distinguenda Costa, ern Russia and Korea. 1847). Xinjiang. Widespread Palaearctic species. C. scutellata Distant, 1887. Guangdong, Guizhou, Cratonotus coloratus Distant, 1879a. Guangdong. Yunnan. Also known from India and Myanmar. Described from northeastern India, and reported from C. sinica Hsiao & Cheng, 1977. Gansu, Hubei, Hunan, Myanmar. Jiangxi, Shanxi, Sichuan, Zhejiang. Cresphontes monsoni (Westwood, 1837) (= C. nigro­ Carpocoris coreanus Distant, 1899 (= C. iranus Tama­ maculatus Haglund, 1868). Yunnan. Also known from nini, 1958). Gansu, Nei Monggol, Ningxia, Qinghai, India. Shaanxi, Xinjiang. Also known fromAfghanistan, Arme­ Critheus indicus (Distant, 1900c ). Yunnan. Also re­ nia, Azerbaijan, Georgia, Iran, Iraq, Israel, Kazakhstan, ported from India and Myanmar. - Note: Ghauri (1963) Kyrgyzstan, Mongolia, Russia (lower Volgaand Siberia), reviewed the genus Critheus and placed Fernelius as a Tajikistan, Turkey, Turkmenistan, Uzbekistan, and Paki­ junior synonym, thus creating the new combination stan. Critheus indicus. No one has followedthese changes; all C. fuscispinus (Boheman, 1850) (= Pentatoma hahni records of this species have still been under the combina­ Flor, 1856; var. rugicollis Seabra, tion Fernelius indicus. 1925; C. fuscispinus var. subrufus Seabra, 1925; C. C. lineatifrons Stal, 1869. Guangdong, Guangxi, fuscispinus var. flavescens Seabra, 1925; C. fuscispinus Hainan. Originally described from Myanmar, and also var. infuscatus Seabra, 1925). Nei Monggol, Xinjiang. known from India, , and Vietnam. Widespread Palaearctic species. Dabessus albovittatus Hsiao & Cheng, 1977. Guang­ [C. pudicus (Poda, 1761)] (= Cimex cinctus Schrank, dong, Hainan. 1776 (preoccupied); C. incarnatus Goeze, 1778 (preoc­ D. malayanus (Distant, 1900b). Guangdong, Hainan. cupied; part); C. carneusGmelin, 1790; Pentatoma wilkin­ Described fromMalaysia, and also known fromLaos and sonii Westwood,1837; P. pa/IidaDallas, 1851; Carpocoris the Philippines. pudicus var. atroscute//atus Halaszfy, 1955). Western Dalpada cinctipes Walker, 1867. Anhui, Fujian, Gan­ Palaearctic (see note below). - Note: This species has su, Guangdong, Guangxi, Guizhou, Hainan, Hebei, been recorded frommany Chinese provinces, but accord­ Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shanxi, Sichuan, ing to Derzhansky (1990), C. pudicus is distributed in W Taiwan, Yunnan, Zhejiang. Also recorded fromKorea and Palaearctic only. The Chinese records of this species are Japan. probably misidentificationsof other Carpocoris species, D. distincta Hsiao & Cheng, 1977. Anhui, Fujian, Gan­ especially C. purpureipennis. su, Guangdong, Guangxi, Guizhou, Hainan, Hebei, He­ C. purpureipennis (De Geer, 1773) (= Cimex nigri­ nan, Hunan, Jiangsu, Jiangxi, Shanxi, Sichuan, Zhejiang. cornis Fabricius, 1775; C. incarnatus Goeze, 1778 (pre­ D. maculata Hsiao & Cheng, 1977. Guangdong, occupied; part); C. porphyropterus Gmelin, 1790; Carpo­ Hainan, Hubei. coris tarsata Mulsant & Rey, 1866; C. nigricornis var. D. neoclavata Rider, 1998 ( = Cimex clavatus Fabricius, pyrosoma Westhoff, 1884; C. pudicus var. fumarius 1798 (preoccupied)). Xizang. Described from India, and Stichel, 1924). Gansu, Hebei, Heilongjiang, Jilin, Liao­ also known fromSri Lanka and southeast Asia. ning, Nei Monggol, Ningxia, Qinghai, Shaanxi, Shan­ D. nodifera Walker, 1867. Guangdong, Guangxi, dong, Shanxi, Xinjiang. Widespread Palaearctic species. Hainan, Jiangsu. Also reported from India, Japan, Myan­ C. seidenstueckeri Tamanini, 1959. Beijing, Jilin, mar, Malaysia, and Indonesia. -· Note: Distant (1899) Liaoning, Nei Monggol, Shandong, Shanxi. Also known considered this to be a junior synonym of D. oculata fromMongolia, eastern Russia, and Korea. (Fabricius, 1775); China (1928) determined it to be a good incarnatus (Drury, 1773) ( = Cimex species. nigripes Fabricius, 1775 (preoccupied); C. aurantius D. oculata (Fabricius, 1775) (= D. aspersa Ellenrieder, Sulzer, 1776; C. melanopus Gmelin, 1790). Guangdong, 1862). Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hainan, Jiangxi, Yunnan. Pakistan, India, and Sri Lanka Hunan, Jiangsu, Sichuan, Yunnan, Zhejiang. India, Ko­ eastward through China and southeast Asia to Korea, Ja­ rea, Japan, southeast Asia, Malaysia, Indonesia. pan, the Philippines, and Indonesia. D. perelegans Breddin, I 904. Guangdong, Guangxi, Caystrus obscurus (Distant, 1901a). Guizhou, Hainan, Sichuan, Xizang, Yunnan. Also recorded from Vietnam. Jiangxi, Xizang, Yunnan. Also known fromPakistan, In­ D. smaragdina (Walker, 1868) (� Amasenoides dia, and Myanmar. virescens Shiraki, 1913). Anhui, Fujian, Gansu, Guang­ C. orientalis Zhang & Lin, 1985. Fujian, Jiangxi. dong, Guangxi, Guizhou, Heilongjiang, Hcnan, Hubei, Cellobius abdominalis Jakovlev, 1885 (= C. abdo­ Hunan, Jiangsu, Jiangxi, Shanxi, Sichuan, Taiwan, Xi­ minalis var..ferrugatus Horvath, 1889b; Brachynema rufi­ zang, Yunnan, Zhejiang. corne Reuter, 1900; Cellobius genti/is Jakovlev, 1901). Dendrites kershawi Kirkaldy, 1909. Guangdong. Also Gansu, Nei Monggol, Xinjiang. Also recorded from Af­ reported from Vietnam. - Note: The Guangdong record ghanistan, Kazakhstan, Kyrgyzstan, Mongolia, Tajikistan, of Wu (1933) needs verification. Turkmenistan, and Uzbekistan. Desertomenida albula Kiritshenko, 1914a. Nei Mong­ Cervicoris omeiensis Hsiao & Cheng, 1977. Sichuan. gol. Also recorded from Iran, Kazakhstan, Kyrgyzstan, juniperina (Linnaeus, 1758) (= Cimex Turkmenistan, Uzbekistan, and Mongolia. jlavoviridis Goeze, 1778; C. trico/or Gmelin, 1790 (pre­ D. quadrimaculata (Horvath, 1892). Gansu, Nei occupied); C. unicolor Turton, 1802 (preoccupied); Penta­ Monggol, Xinjiang. Also known from Kazakhstan, Kyr­ toma confusa Westwood, 1837). Nei Monggol. Wide­ gyzstan, Mongolia, Tajikistan, Turkey, and Turkmenistan. spread Palaearctic species. Dolycoris baccarum (Linnaeus, 1758) (= Cimex sub­ Codophila maculicollis (Dallas, 1851) (= Pentatoma ater Harris, 1780; C. a/bidus Gmelin, 1790 (preoccupied); arabica ·st1H, 1854). Xinjiang. Northern Africa, Middle Aelia depressa Westwood, 1837; Pentatoma confusa East, India. Westwood, 1837; P. ineoncisa Walker, 1867; Dolycoris ZOO SY ST. ROSSICA Vol. 11 • D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae 139 baccarum var. brevipi/is Reuter, 1891). Fujian, Gansu, E. maracandica Oshanin, 1871. Gansu, Nei Monggol, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Qinghai, Xinjiang. Also known from Afghanistan, Iran, Heilongjiang, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Kazakhstan, Kyrgyzstan, Mongolia, southern Russia,Taji­ Jilin, Liaoning, Nei Monggol, Ningxia, Qinghai, Shaanxi, kistan, Turkmenistan, and Uzbekistan. - Note: Many of Shandong, Shanxi, Sichuan, Xinjiang, Xizang, Yunnan, the Chinese records under the combination E. festivaand Zhejiang. Widespread Palaearctic species. E. festiva var. chlorotica may actually be misidentifica­ D. formosanus Distant, 1887. Taiwan. tions of this species. D. indicus Stal, 1876. Sichuan, Xizang, Yunnan. Af­ E. oleracea (Linnaeus, 1758) (= Cimexdubius Scopoli, ghanistan, Pakistan, India, Sri Lanka, Myanmar. 1763; C. flavatus Schrank, 1776; C. albomarginatus D. penicillatus Horvath, 1904. Xinjiang,Xizang. Mid­ Schrank, 1776; C.jlavomaculatusGoeze, 1778; C. albo­ dle East to western Siberia. marginellus Fabricius, 1794; oleraceum var. Drinostia angulata Hsiao & Cheng, 1977. Sichuan. angularis Kolenati, 1846; Pentatoma oleraceavar. zetter­ D. fissipesStal, 1865. Fujian,Hunan, Jiangsu, Jiangxi, staedtii Gorski, 1852; Strachia lineola Baerensprung, Zhejiang. - Note: Stal (1865) originally spelled this spe­ 1858; S. oleracea var. insidiosa Mulsant & Rey, 1866; S. ciesflssipes, but everyone since has spelled itflssiceps. consobrina Puton, 1871; there are at least 27 more vari­ D. megacephala Zheng & Liu, 1987b. Guizhou. etal names proposed forvarious colour formsof this spe­ D. planiceps Stal, 1861. Guangdong. cies). Xinjiang. Widespread Palaearctic species. - Note: Dunnius fulvescens (Dallas, 1851). Guizhou, Hainan, The records of E. rugulosa (Yang 1962 and others) are Yunnan. Also known from India and Myanmar. misidentifications of this species. D. minor Zheng & Liu, 1987a. Yunnan. E. ornata (Linnaeus, 1758) (= Cime.xfestivusLinnaeus, Dymantiscus alticola (Cheng & Li, 1979). Xizang. 1767; Pentatoma pictum Herrich-Schiiffer, 1833; P. D. marginatus Hsiao, 1981. Xizang. decoratum Herrich-Schiiffer, 1833; Eurydema pustulata fullo(Thunberg, 1783) (= Cime.xmucoreus Fieber, 1837; E. pictum var. conjuncta Kolenati, 1846; Fabricius, 1794; Dalpadajaponica Walker, 1867). Anhui, Cimex fallax Scholtz, 1847; fal/eni Beijing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Gorski, 1852; Cime.xumbra/is Gistel, 1857; there are at Hainan, Hebei, Henan, Hubei, Hunan, Jiangsu, Jiangxi, least 13 more varietal names proposed forvarious colour Liaoning, Nei Monggol, Shaanxi, Shandong, Shanxi, forms of this species). Beijing, Gansu, Henan, Ningxia, Sichuan, Taiwan,Xinjiang, Yunnan,Zhejiang. Also from Qinghai, Xinjiang, Xizang, Yunnan. Widespread Afghanistan, Pakistan, India, and Sri Lanka eastward Palaearctic species. through China and southeast Asia to Japan and Indone­ E. pulchra (Westwood, 1837) (= E. sumatrana sia. Ellenrieder, 1862; Strachia designata Walker, 1867; E. ilia China, 1925. Fujian, Shandong, Yunnan, Zhe­ Eurydemaamoena Horvath, 1879). Guangdong, Hainan, jiang. -Note: This may possibly be a synonym of E.fullo. Hebei, Jiangsu, Jiangxi, Taiwan, Yunnan. Occurs from Eurydema alpina Ling, 1989. Sichuan. India through China to Korea and Japan, and through E. dominulus (Scopoli, 1763) (= Cimex cordiger southeast Asia, Indonesia to northernAustralia. - Note: Goeze, 1778; Pentatoma fimbriolatum Germar, 1834; Zheng (1983) indicated that this might only be a colour Eurydema daurica Motschulsky, 1860; Strachia minus­ variety of E. dominulus. cula Walker, 1867; S. festiva var. albiventris Jakovlev, E. rugosa Motschulsky, 1861 (= Strachia signatum 1876a; Eurydema dominulus var. schirmeri Schumacher, Walker, 1867; S. marginiferaWalker, 1867). Beijing. Also 19 I 1b; E. dominulus var. immaculata Stiebel, 1924 ). known fromeastern Russia(Sakhalin, Kunashir) and Japan. Fujian, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang, E. ventralis Kolenati, 1846 (= Pentatoma ornata hof­ Henan, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Nei Mong­ fmanseggi Gorski, 1852; Strachia ornata var. pectoralis gol, Shaanxi, Shandong, Shanxi, Sichuan, Xizang, Yun­ Fieber, 1861; S. ornata var. dissimilis Fieber, 1861; Tama­ nan, Zhejiang. Widespread Palaearctic species. - Note: nini ( 1957) proposed nine more varietal names for vari­ Many early authors usedfestiva forthe specificname for ous color forms of this species, three of which were pre­ this species;festiva Linnaeus (1767) is actually a junior occupied so he (Tamanini 1961) later proposed replace­ synonym of E. ornata. See also note under E. mara­ ment names). Nei Monggol, Sichuan, Xinjiang. Wide­ candica. spread Palaearctic species. -- Note: Early workers often E. fieberiFieber , 1837 (= Pentatoma ornataschaejferi referred to this species under the specificepithet ornata. Gorski, I 852; Strachia meyeri Fieber, 1861; S. dominula These were all misidentifications. var. caucasica Jakovlev, 1874a; S. scutellata Jakovlev, E. wilkinsi Distant, 1879a (= Strachia conspicua 1883; Eurydema fieberi var. melanogaster Horvath, Jakovlev, 1881). Nei Monggol, Ningxia, Xinjiang. Also 1917). Xinjiang. Widespread Palaearctic species. known from Kazakhstan, Kyrgyzstan, Mongolia, Tajiki­ E. gebleri Kolenati, 1846 (= Strachia picturata Stal, stan, Turkmenistan, and Uzbekistan. 1858; Eurydema gebleri var. tsherskii Reuter, 1910). Eurysaspis Oavescens Distant, 1911. Anhui, Fujian, Anhui, Gansu, Hebei, Heilongjiang,Henan, Hubei, Jiang­ Guizhou, Hebei, Hubei, Hunan, Jiangsu, Jiangxi, Zhe­ su, Jilin, Liaoning, Shaanxi, Shandong, Shanxi, Sichuan, jiang. Also recorded from the Philippines and Indonesia Xizang, Yunnan. Also known from Kazakhstan, Russia, (Borneo).- Note: The original spelling of the genus name Mongolia, and Korea. is EurysaspisSignore!, but many workers continue to use E. lingi nom. n. (= E. montana Ling, 1989 (preoccu­ the unjustified emended form Euryaspis. pied)). Sichuan. -Note: E. montana Ling, 1989, is a jun­ Exithemus assamensis Distant, 1902. Fujian, Guang­ ior primary homonym of E. dominulusmontana Kerzhner, dong, Guizhou, Hunan, Sichuan, Zhejiang. Originally 1972b, which is still considered to be a valid subspecies described from northern India. of E. dominulus. aenescens (Walker, 1867) (= Hoplistodera E. liturifera (Walker, 1867) (= E. vicaria Horvath, schwaneri Vollenhoven, 1868). Jiangxi. Described from 1889a; E. vicaria var. supplens Horvath, 1889a; E. Borneo, and also reported from Myanmar. lituriferavar. hypomelan Breddin, 1907; E. lituriferavar. E. aeneus (Scopoli, 1763) (= Cimex fucatus Rossi, hypopoecilaBreddin, 1907; E. lituriferavar. yunnanensis 1790; C. perlatus Fabricius, 1794; Eysarcoris per/atus China, 1925). Xizang, Yunnan.Also recorded fromPaki­ var. spinicollis Puton, 1881; E. perlatus var. ventralis stan, Sri Lanka, and Myanmar. Horvath, 1882 (preoccupied); E. parva Uhler, 1896; E. 140 D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae • ZOOSYST. ROSSI CA Vol. 11 aeneus f. nigriventris Stichel, 1962). Gansu, Heilongjiang, ( 1904) replacement name, Eysarcoris fabricii, but the Jilin, Nei Monggol, Shanxi. Widespread Palaearctic spe­ oldest correct name is Eysarcoris venustissimus (Schrank, cies. - Note: Josifov & Kerzhner (1978) placed parva as 1776). See Rider (1998) foran account of the taxonomic a junior synonym of this species, yet references to E. history of this species. parvus persist. Glaucias beryllus (Fabricius, 1787). Fujian, Guang­ E. annamita (Breddin, 1909). Anhui, Fujian, Gansu, dong, Macao, Taiwan. Also recorded from India. Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, G. crassus (Westwood, 1837). Fujian, Guangdong, Jiangsu, Jiangxi, Shanxi, Sichuan, Xizang, Yunnan, Guangxi, Henan, Yunnan. Also known from India and Zhejiang. Also recorded fromKorea, Japan, and Vietnam. Vietnam. - Note: At one time, this species was consid­ E. gibbosus Jakovlev, 1904b. Fujian, Heilongjiang, ered to be a variety of G beryl/us. Jiangxi, Jilin, Nei Monggol, Shaanxi, Sichuan. Described G dorsalis (Dohrn, I 860) (= Zangis virgin ea Stal, from Korea, and also recorded from eastern Russia, Ja­ 1876). Guangdong, Guangxi. India, Sri Lanka, Maldive pan, and Vietnam. - Note: This species has often been Islands, Malaysia, the Philippines. confused with E. venustissimus, a western Palaearctic G nigromarginatus (Stlll, 1871 ). Yunnan. Also known species. Chinese records of E. venustissimus, or its syno­ fromthe Philippines and Indonesia. nyms (melanocephalus, fabricii), should be referred to G subpunctatus (Walker, 1867) (= Rhaphigaster this species. melanosticticus Vollenhoven, 1868). Fujian, Guangdong, E. guttiger (Thunberg, 1783) (= Pentatoma nepalensis Guangxi, Hunan, Yunnan. Also recorded fromKorea, Ja­ Westwood, 1837; P punctipes Westwood, 1837 (preoc­ pan, and Indonesia. cupied); Eysarcoris potanini Jakovlev, 1890; E. brevi­ Halyabbas unicolor Distant, 1900c. Guangdong, Hai­ usculus Jakovlev, 1902a; Bainbriggeanus jletcheri Dis­ nan, Taiwan, Yunnan. Also known from India, southcast tant, 1918). Anhui, Fujian, Gansu, Guangdong, Guangxi, Asia, and Indonesia. Guizhou, Hainan, Hebei, Heilongjiang, Henan, Hubei, halys (Stal, 1855) (� Poecilometis Hunan, Jiangsu, Jiangxi, Liaoning, Nei Monggol, Ningxia, mistus Uhler, 1860; Dalpada brevis Walker, 1867; D. re­ Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Xizang, mota Walker,1867). Anhui, Fujian, Guangdong, Guangxi, Yunnan, Zhejiang. Occurs from Nepal, India, and Sri Guizhou, Hebei, Heilongjiang, Henan, Hubei, Hunan, Lanka eastward through China and southeast Asia to Jiangsu, Jiangxi, Jilin, Liaoning, Neu Monggol, Shaanxi. Korea, Japan, and the Philippines. - Note: Sto/lia Ellen­ Shanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang. Also rieder has for quite some time been considered a junior recorded fromKorea and Japan. --- Note: This species has synonym of Eysarcoris, but we still see an occasional had a long and confusing history. It has appeared in the reference to this species as Stollia guttiger. literature under halys and all three junior synonyms; in E. latus Walker, 1868. Taiwan. fact, it appears that many workers in Japan still referto it E. montivagus (Distant, 1902). Anhui, Fujian, Guang­ as H. mista. It has also frequently been confusedwith H. dong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan, picus, an Indian species. Josifov & Kerzhner (1978) de­ Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang. Also known termined that only one species of Halyomorpha is present from Afghanistan, Pakistan, India, Sri Lanka, and south­ in eastern China, Japan, and Korea, and it is different east Asia. - Note: Several recent records have been un­ from H. picus; the oldest available name is H. halys (Stal). der the combination Stollia montivagus. Halys rugosus Distant, 1921. Yunnan. Described from E. pustulatus Walker, 1868. Taiwan. Laos. - Note: Ghauri (1988) indicated that this species E. rosaceus Distant, 1901a. Guangdong, Guangxi, did not belong in the genus Halys, but he did not place it Sichuan, Taiwan, Xizang, Yunnan.Described fromMyan­ in any other genus. The Yunnan record of China ( 1925) mar, and also recorded from Vietnam. - Note: A couple needs verification. recent records have been under the name Stolliarosaceus. H. serriger (Westwood, 1837) (= Cimex dentatus Fab­ E. trigonus (Kiritshenko, 1931). Sichuan, Xizang. - ricius, 1775 (preoccupied)). Guangdong. Afghanistan, Note: Once again, recent records of this species have been India, Sri Lanka, Myanmar, and Japan. - Note: All re­ under Stollia trigonus. cent records of this species have been under Halys E. ventralis (Westwood, 1837) (= Pentatoma incon­ dentatus (Fabricius, 1775), but that name is a primary spicuum Herrich-Schaffer, 1844; Eysarcoris distactus homonym of Cimex dentatus De Geer, 1773, currently a Dallas, 1851; E. misellus Stal, 1853; E. he/Jeri Fieber, species in the Acanthosomatidae. The single Chinese 1861; Stollia rectipes Ellenrieder, 1862; E.ysarcoris epi­ record of Hsiao & Cheng ( 1977) needs confirmation. stomalis Mulsant & Rey, 1866; E. mayeti Mulsant & Rey, H. sulcatus (Thunberg, 1783) (= H. magnus Chopra, 1868; E. pseudoaeneus Jakovlev, 1869; E. inconspicuus 1974; H. qadrii Abbasi & Ahmad, 1976). Guangdong var. simplex Puton, 1881; E. scutellaris Jakovlev, 1885; (Hong Kong). Also known from Pakistan, India, Sri E. egenus Jakovlev, 1900; E. sindellus Distant, 1902; E. Lanka, and Japan. schmidti Jakovlev, 1902a; E. inconspicuus var. kwan­ Hippotiscus dorsalis (Stal, 1869). Anhui, Fujian, vonensis Yang, 1934; E. tangens Stichel, 1961). Fujian, Guangdong, Guangxi, Guizhou, Henan, Hunan, Jiangxi, Guangdong, Guangxi, Guizhou, Hainan, Hebci, Henan, Sichuan, Xizang, Zhejiang. Also reported from India. - Hubei, Jiangxi, Shanxi, Taiwan, Zhejiang. Widespread Note: This species was originally placed in the preoccu­ Palaearctic species, also occurring in Africa, the Indian pied genus Plexippus Stal. Bergroth (l 891) proposed region, southeast Asia, Indonesia, and the Philippines. llippota as a replacement name, but this name was also Also has been introduced into Hawaii. preoccupied. So, Bergroth (1906) proposed Hippotiscus [E. venustissimus (Schrank, 1776)] (= Cimex melano­ as a new replacement name. This species is still being cephalus Fabricius, 1775 (preoccupied); Eysarcoris fab­ referredto as Hippotadorsalis in the applied literature; it ricii Kirkaldy, 1904). Widespread western Palaearctic is a pest on bamboo (Xu et al., 1991a, 199\b; Yu et al., species. - Note: This species has been recorded often from 1999). China, but these are probably all misidentificationsof E. breviceps (Horvath, 1897a). Heilong­ gibbosus. All records of E. venustissimus, or its syno­ jiang. Also known from eastern Russia, Korea. nyms (melanocephalus,fabricii) fromeastern Palaearctic H. capitatus (Jakovlev, 1889b) (= peltatus should be referred to E. gibbosus. The majority of mod­ Jakovlev, 1894; P. vernalisvar. stramineus Horvath, 1895; ern records for this species have been under Kirkaldy's P. pa/lescens Jakovlev, 1902b ). Gansu, Xinjiang. Afghani- ZOOSYST. ROSSI CA Vol. 11 • D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae 141 stan, Iran, Kazakhstan, Kyrgyzstan, Tajikistan, Turk­ have ever been recorded. As the type species of Oedocoris menistan, Uzbekistan, and Mongolia. Zheng & Liu (0. eburneus) is now proved to be a junior H. ovatus (Jakovlev, 1889b) (= Peribalus pallidicornis synonym of L. nigripes, the type species Lakhonia Yang, Reuter, 1913 ). Nei Monggol, Shanxi. Also recorded from the genus Oedocoris is consequently synonymised with Mongolia, eastern Russia, and Korea. Lakhonia (new synonymy). Lakhonia is a genus with H. strictus vernalis (Wolff, 1804 ). N ei Monggol, extraordinary and isolated features(thick, compact body; Shanxi, Xinjiang. Widespread Palaearctic species. - Note: short dense dorsal pubescence; and perpendicular head). H. strictus and H. vernalis have until recently been con­ Yang (1936) tentatively suggested that it was related to sidered to be two separate species. Josifov (1986) treated Pentatoma Olivier, but this is certainly not so. them as subspecies; he has been followedin doing so by [Laprius gastricus (Thunberg, 1822)]. "China." Also Lis ( 1990). Many recent workers have continued to rec­ known fromKorea and Japan. - Note: Kirkaldy (1909) ognize vernalisas a separate species, but all of these have listed "China" in the distribution, with no specificlocal­ been in lists or simple records of distribution; no one has ity mentioned. Its occurrence in China needs verification. as of yet refuted Josifov's treatment. L. varicornis (Dallas, 1851). Anhui, Fujian, Guang­ Homalogonia chinensis Lin, Zhang & Xiong, 1992. dong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan, Hunan. Jiangsu, Jiangxi, Shaanxi, Shandong, Sichuan, Yunnan, H. confusa Kerzhner, 1972a. Heilongjang. Eastern Zhejiang. Also known fromPakistan, India, Japan, Myan­ Russia and Korea. mar, Vietnam, and the Philippines. H. crocemaculata Chen, 1983. Guizhou, Hebei, Sichu­ Lelia decempunctata (Motschulsky, 1860) (= L. por­ an, Yunnan. - Note: Chen (1983) originally spelled this rigens Walker, 1867). Anhui, Guizhou, Heilongjiang, species as listed above; all other workers have listed this Hunan, Jiangxi, Jilin, Lioaning, Nei Monggol, Shaanxi, species as H. croceomacu/ata. Sichuan, Xizang, Yunnan,Zhejiang. Also occurs in east­ H. grisea Josifov& Kerzhner, 1978. Guangxi, Hebei, ern Russia, Korea, and Japan. Henan, Hubei, Jiangxi, Shandong, Sichuan, Zhejiang. L. octopunctata (Dallas, 1849). Guizhou, Yunnan. Described from Korea. Also known from India and Bhutan. H. obtusa obtusa (Walker, 1868) (= H. macu/ata Liicoris tibetanus Zheng & Liu, 1987b. Xizang. Jakovlev, 1876a). Fujian, Gansu, Guangdong, Guangxi, Madates limbata (Fabricius, 1803). Guangdong, Gui­ Guizhou, Hebei, Heilongjiang, Henan, Hubei, Jiangsu, zhou, Yunnan.Also occurs from Pakistan, India, and Sri Jiangxi, Jilin, Liaoning, Nei Monggol, Shaanxi, Shan­ Lanka eastward through China and southeast Asia to In­ dong, Sichuan, Xizang, Yunnan, Zhejiang.Also recorded donesia. fromeastern Russia, Korea, Japan, India, and Vietnam. Meeidea indica Dallas, 1851. Sichuan, Yunnan. Also H. obtusa yunnana Zheng & Liu, 1987a. Yunnan. recorded fromPakistan and India. H. pinicola Lin & Zhang, 1992. Hunan, Jiangxi, Menida disjecta (Uhler, 1860) (= Stromatocoris amoe­ Zhejiang. nus Jakovlev, 1876a (preoccupied), syn. n.; Menidascotti H. sordida Zheng, 1994. Gansu, Shaanxi. Puton, 1886, syn. n.; M scotti var. sinensis Reuter, 1888a, Hoplistodera fergussoniDistant, I 9 II. Anhui, Fujian, syn. n.). Gansu, Guangdong, Guangxi, Hebei, Heilong­ Guangdong, Guangxi, Guizhou, Hainan, Hubei, Jiangxi, jiang, Hubei, Hunan, Jiangxi, Liaoning, Nei Monggol, Shaanxi, Sichuan, Xizang, Yunnan,Zhejiang. Qinghai, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, H. incisa Distant, 1887. Guangxi, Guizhou, Hainan, Xizang, Yunnan. Also known from eastern Russia, Ko­ Yunnan.Also recorded from India, Myanmar, and Viet­ rea, and Japan. - Note: We have examined the type ma­ nam. terial for Menida disjecta (Uhler), which consists of a H. longispina Hsiao & Cheng, 1977. Yunnan. single female specimen. It is a typical specimen of what H. pulchra Yang, 1934. Anhui, Fujian, Gansu, Guang­ most workers have called Menida scotti Puton. It is not dong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, possible to tell how many specimens Uhler had when he Shanxi, Sichuan, Xizang, Yunnan, Zhejiang. made his original description, so a lectotype designation [H. scutellomaculata Distant, 1921]. Laos. - Note: is in order. The femalespecimen (lacking the head) with Tang (I 935) listed "China: Yunnan (Mekong)" in his dis­ the labels a) "PR Uhler Collection"; b) "Rhaphigaster tribution, but Mekong is located in Laos. Otherwise, this disjectus Uhler_ Hong Kong." is designated lectotype. species has not been recorded fromChina. The lectotype is conserved in the U.S. National Museum, H. virescens Dallas, 1851. Guizhou, Xizang, Yunnan. Washington, DC. Also known from India and Myanmar. M. formosa(Westwood, 1837) (= Rhaphigaster spec­ Iphiarusa compacta (Distant, 1887) (= /. aratrix tandus Stal, 1860; R. a/bidens Ellenrieder, 1862; R. nitens Breddin, 1904). Fujian, Guangdong, Henan, Hunan, Vollenhoven, 1868). Hainan, Guangdong, Guangxi, Gui­ Jiangxi, Sichuan. Also reported fromIndia, Myanmar, and zhou, Jiangsu, Jiangxi, Taiwan, Xizang, Yunnan, Zhejiang. Vietnam. Occurs from India and Sri Lanka eastward to Indonesia. I. longicauda Hsiao & Cheng, 1977. Sichuan, Yunnan. M. lata Yang, 1934. Anhui, Fujian, Guangdong, Guang­ Jugalpada varia (Dallas, 1851) (= Dalpada apicifera xi, Guizhou, Hainan, Henan, Hubei, Hunan,Jiangsu, Jiangxi, Walker, 1867). Hong Kong, Jiangxi. Also known from Sichuan, Zhejiang. India, Bhutan, and Myanmar. - Note: Most records of M. maculiscutellata Hsiao & Cheng, 1977. Fujian, this species have been under the combination Dalpada Hubei, Sichuan. varia. M. megaspila (Walker, 1867) (= Antestia megaspila Lakhonia nigripes Yang,1936 (= Oedocoris eburneus Walker, 1868). Guangdong, Taiwan. Zheng & Liu, 1987a, syn. n.). Yunnan. Originally de­ M. metallica Hsiao & Cheng, 1977. Guizhou, Sichuan, scribed from Thailand. - Note: Yang (I 936) described Yunnan. Lakhonia nigripes based on a single female specimen M. mosaica Zheng & Liu, 1987c. Yunnan. collected from Lakhon, Thailand. A femaleparatype of M. musiva (Jakovlev, I 876a) (= Mjaponica Distant, 0edocoris eburneus collected from southwestern China 1883). Heilongjiang,Jilin, Sichuan, Taiwan,Yunnan. Also was compared with the L. nigripes type material in Paris recorded from eastern Russia, Korea, and Japan. Museum by Dr. Dominique Pluot-Sigwalt and doubtlessly M. ornata Kirkaldy, 1910. Guizhou, Sichuan, Yunnan. proved as conspecific. No males of this rare pentatomid M. pinicola Zheng & Liu, 1987c. Hubei, Yunnan. 142 D.A. Rider et al.: Chinese Pentatomidae.11. Pentatominae • ZOOSYST. ROSSICA Vol. II M. speciosa Zheng & Xiong, 2001. Yunnan. sericea Westwood, 1837; P leii Westwood, 1837; P tri­ M. szechuensis Hsiao & Cheng, 1977. Fujian, Gui­ punctigera Westwood, 1837; P chinensis Westwood, zhou, Jiangxi, Ningxia, Sichuan, Xizang, Yunnan. 1837; P. proxima Westwood, 1837; P propinqua West­ M. varipennis (Westwood, 1837). Fujian, Guangdong, wood, 1837; P chlorocephala Westwood, 1837; Cimex Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiang­ hemichloris Gerrnar, 1838; approximata Reiche xi, Sichuan, Yunnan,Zhejiang. Afghanistan, Pakistan, and & Fairmaire, 1848; Pentatoma plicaticollis Lucas, 1849; India eastward to the Philippines and Indonesia. Rhaphigaster orbus Stal, 1853; Pentatoma vicaria Walker, M. versicolor (Gmelin, 1790) (= Cimexhistrio Fabrici­ 1867; var. aurantiaca Costa, 1884; N. us, 1787 (preoccupied); Pentatoma bengalensis West­ viridula var. hepatica Horvath, 1903). Anhui, Fujian, wood, 1837; Rhaphigaster concinnus Dallas, 1851; R. Guangdong, Guangxi, Guizhou, Hainan, Hebei, Henan, strachioides Walker, 1867; Menida cey/anica Breddin, Hubei, Hunan, Jiangsu, Jiangxi, Ningxia, Shaanxi, . 1909). Fujian, Guangdong, Guangxi, Guizhou, Hainan, Shandong, Shanxi, Sichuan, Xizang, Yunnan, Zhejiang. Jiangxi, Macao, Sichuan, Xizang, Taiwan, Yunnan, Zhe­ Widespread cosmopolitan species. jiang. Occurs from Pakistan, India, and Sri Lanka east­ N. yunnana Zheng, 1982. Yunnan. ward through China and southeastAsia to Japan, the Phil­ Niphe eiongata (Dallas, 1851 ). Anhui, Fujian, Guang­ ippines, and Indonesia. - Note: Nearly all recent records dong, Guangxi, Guizhou, Hainan, Henao, Hubei, Hunan, of this species have been under the name Menida histrio, Jiangsu, Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, but this is a junior primary homonym; the next available Yunnan,Zhejiang. Also known fromIndia, Japan, Myan­ name is versicolor.See Rider (1998) for further details. mar, and the Philippines. M. violacea Motschulsky, 1861.Anhui, Fujian, Gansu, N. subferruginea (Westwood, 1837) (= Pentatoma Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, cephalus Dallas, 185 I; P lateral is Walker, 1867 (preoc­ Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Nei Monggol, cupied)). Guangdong, Guangxi, Guizhou, Yunnan. Also Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Yunnan, recorded from India, southeast Asia, and Indonesia. Zhejiang. Also known from eastern Russia, Korea, Ja­ Ochrophara chinensis Zheng & Liu, 1987b. Yunnan. pan, and India. Ochyrotylus helvinus Jakovlev, 1885 (= 0. signoreti [M. vitalisana Distant, 1921]. Yunnan.Described from Jakovlev, 1885; 0. helvinus var. rubripennis Kiritshenko, Vietnam. - Note: The Yunnan record (Hoffmann, 1935) 1914b; 0. helvinus var. rubens Kiritshenko, 1914b). Nei needs verification. Monggol. Also recorded fromAzerbaijan, Iran, Kazakh­ Mimula dungana (Kiritshenko, 1912). Xinjiang. Also stan, Mongolia, Turkmenistan, and Uzbekistan. known fromKazakhstan and Kyrgyzstan. Okeanos quelpartensis Distant, 1911. Gansu, Hebei, Myrochea gyirongnensis (Lin & Zhang, 1985), Hubei, Hunan, Jiangxi, Jilin, Shaanxi, Sichuan, Yunnan. comb. n. Xizang. - Note: Lin & Zhang (1985) described Also recorded from Korea and eastern Russia. this species in the genus DorpiusDistant, but Linnavuori Otantestia heterospila (Walker, 1867) (= Pentatoma (1982) recently placed Dorpius as a junior synonym of pulchra Dallas, I 851 (preoccupied)). Guangdong, Hainan, Myrochea. Qinghai, Yunnan. Also known from Bhutan, Pakistan, M. kershawi (Kirkaldy, 1910), comb. n. Macao. - India, southeastAsia, and Indonesia. - Note: Even though Note: Originally described in the genus Dorpius and all Kirkaldy ( 1909) catalogued this species correctly, and subsequent workers have treated it in Dorpius, but Hoffmann (1932a, 1932b, 1935, 1954) referred to it as Linnavuori (1982) placed Dorpius as a junior synonym Otantestia heterospila, all other workers from India and of Myrochea. the Oriental region have referredto this species asAntestia Neojurtina typica Distant, I921. Fujian, Guangdong, pulchra. Pentatoma pulchra Dallas, 1851 is a primary Guangxi, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang.Also junior homonym of P. pulchra Westwood, 1837, a spe• known from Vietnam and Malaysia. cies now in the genus Eurydema. The next available name Neottiglossa leporina (Herrich-Schaffer, 1830) (= N. is Strachia heterospila Walker, 1867. Kirkaldy (1909) calva Jakovlev, 1903c). Heilongjiang, Nei Monggol, designated pulchra (= heterospila) type species of the Xinjiang. Widespread Palaearctic species. genus Otantestia. All records of Antestia pulchra should N. pusilla (Gmelin, 1790) (= Cimex griseusnigro-pzmc­ be referred to Otantestia heterospila. tatus De Geer, 1773 (unavailable trinomen); Cydnus injlexus 0. modificata(Distant, 1887). Yunnan. Also recorded Wolff, 1811; Platyso/en obscurus Sahlberg, 1870; Aelia fromIndia. - Note: Breddin (1900) placed this species in panzeri Krynicky, 1871; Neottiglossa seorsa Jakovlev, his newly erected genus Otantestia. This is where Kirkal­ 1903c; N. nigella Jakovlev, 1903c). Nei Monggol, Shaanxi, dy (1909) and Hoffmann (1935) also treated this species. Shanxi, Xinjiang. Widespread Palaearctic species. Hsiao & Cheng's (1977) record of Antestia modificata Neostrachia bisignata (Walker, 1867) (= N. hellenica from Yunnanshould be referredto Otantestia modificata. Saunders, 1877). Jiangxi. Recorded from Greece, Saudi Palomena angulosa (Motschulsky, 1861) (= Mormidea Arabia, Tunisia, and India. - Note: See Rider (1998) for basicornis Motschulsky, 1866; Palomena amurensis an explanation on this generic placement. Reuter, 1908). Guizhou, Heilongjiang, Henao, Jiangxi, Nevisanus nagaensis Distant, 1893. Xizang. Described Jilin, Liaoning, Nei Monggol, Shaanxi, Shanxi, Sichuan, from northeastern India. Xizang, Yunnan,Zhejiang. Also known from easternRus­ Nezara antennata Scott, 1874 (= N. antenna/a var. sia, Korea, and Japan. balteata Horvath, 1889a). Fujian, Gansu, Guangdong, P. cha pan a (Distant, 1921), sp. dist. (= P haemorrho­ Guangxi, Guizhou, Hainan, Hebei, Henan, Hubei, Hunan, idalis Lindberg, 1934, syn. n.). Gansu, Hebei, Hubei, Jiangsu, Jiangxi, Shanxi, Sichuan, Taiwan, Xinjiang, Hunan, Shaanxi, Sichuan, Xizang, Yunnan. Also known Xizang, Yunnan. India,Sri Lanka, Korea, Japan, the Phil­ from Nepal, Myanmar, and Vietnam. - Note: Distant ippines. ( 1921) described Epagathus chapana based on material N. viridula (Linnaeus, 1758) (= Cimex torquatusFabri­ fromChapa, Vietnam ("Tonkin"). China ( 1925) synonym­ cius, 1775; C. smaragdulus Fabricius, 1775; C. transver­ ised E. chapana with Pa/omena unicolorella Kirkaldy, sus Thunberg, 1783; C. variabilis Villers, 1789; C. spirans 1909. Zheng & Ling ( 1989) followedthis synonymy but Fabricius, 1798; C. viridissimus Wolff, 180 I (preoccu­ noted that this was rather tenuous as only femalesyntype pied); Pentatoma oblonga Westwood, 1837; P unico/or material was examined in that study. On the other hand, Westwood, 1837; P berylina Westwood, 1837; P sub- the Yunnan record of P unico/ore/la (China, 1925) was ZOOSYST. ROSSICA Vol. 11 • D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae 143 determined to be a misidentificationof P.haemorrhoidalis P. hingstoni Kiritshenko, 1931. Xizang. - Note: Al­ Lindberg (Zheng& Ling, 1989, studied the material upon though Kiritshenko ( 1931) originally spelled this species which Dr. China's work was based). Recently, one male hingstoni, many workers referto this species as hinstoni. syntype of Epagathus chapana was examined and is here P. hsiaoi Zheng & Wang, 1995. Gansu. designated lectotype (conserved in the BMNH). The de­ P. illuminata (Distant, 1890). Hubei, Shanxi. tails of the male genitalia indicate that chapana is P. japonica (Distant, 1882). Fujian, Gansu, Guizhou; conspecificwith haemorrhoidalis, but different fromthe Heilongjiang, Hubei, Hunan, Jilin, Liaoning, Nei Mong­ Indian unicolorella. Therefore,chapana is removed from gol, Qinghai, Shaanxi, Yunnan, Zhejiang. Also known the synonymy of P. unico/orella, and is resurrected as from eastern Russia, Korea, and Japan. senior synonym of haemorrhoidalis. P. kunmi.ngensisXiong, 1981. Guizhou,Sichuan, Y\mnan. P. hsiaoi Zheng & Ling, 1989. Guizhou, Hubei, Jiangxi, P. leliiformis Kirkaldy, 1909. Yunnan. Sichuan, Yunnait, Zhejiang. P. longirostrata Hsiao& Cheng, 1977. Sichuan, Xizang. P. huoaoeosis Lin & Zhang, 1992. Hunan. P. major Zheng & Jin, 1990. Xizang. P. limbata Jakovlev, 1904a. Gansu, Sichuan, Xizang, P. metallifera (Motschulsky, 1860) (= Tropicorisbas­ Yuonan. nini Oshanin, 1870; T. davidi Signoret, 1880). Beijing, P. prasioa (Linnaeus, 1761) (= Cimex viridis lotus De Gansu, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mong­ Geer, 1773 (unavailable trinomen); C. roseusMuller, 1776; gol, Ningxia, Qinghai, Shanxi. Also known from Mon­ C. viridis Harris, 1780; C. dissimilis Fabricius, 178 I; C. golia, easternRus sia, Korea, and Japan. -Note: Ling and disco/or Wolff, 1811 (preoccupied); Pentatoma prasina Zheng (1987) only tentatively placed davidi as a syno­ subrubescens Gorski, 1852; P. prasina var. rhododactyla nym of metalliferaas they were not able to examine type Horvath, 1883). Xinjiang. Widespread Palaearctic species. material of davidi. - Note: Several authors have reported this species from P. montana mootana Hsiao & Cheng, 1977. Sichuan, Yunnanand from Xizang (China, 1925; Hsiao& Cheng 1977; Yunnan. Zhang & Lin, 1981, 1987), but all have proven to be P. montana yulonglca Zheng & Liu, 1987c. Yunnan. misidentifications of P. tibetana. The Kiritshenko (1964) P. mosaica Hsiao & Cheng, 1977. Anhui, Fujian, Hu­ record fromXinjiang appears to be accurate. nan, Jiangsu, Jiangxi, Qinghai, Zhejiang. [P. rubricornis Scott, 1874). Hebei. Described fromJa­ P. nlgra Hsiao & Cheng, 1977. Sichuan, Yunnan. pan.- Note: No one in recent times hasreferred to this spe­ P. parametallifera Zheng& Li, 1991. Heilongjiang,Jilin. cies (other than in listsor catalogs).The only species gener­ Also known from eastern Russia and southern Korea. ally collected nearth e typelocality in Japanis P. angulosa; P. parataibaiensis Liu & Zheng, 1995. Ningxia,Shaanxi. the record fromChina is probably a misidentificationof P. P. puoctipes (Stdl, 1876) (= Compastes truncatus Dis­ viridissima. An examination of the type material will be tant, 1887). Xizang. Also known fromIndia. neededto clarify th e status of thisspecies. P. roselcornuta Zheng& Ling, 1983. Guizhou, Hunan, P. similis Zheng & Ling, 1989. Sichuan. Sichuan. [P. spinosa Distant, 1880]. Xizang. Also known from P. rufipes (Linnaeus, 1758) (= Cimex notatus Poda, Pakistan and India. - Note: The Xizang record (Horvath, 1761; Pentatoma viridiaenea Pali sot de Beauvois, 1811; 1897b) needs verification. Tropicoris nigricornis Reuter, 1879a; T. rufipes var. P. tibetana Zheng & Ling, 1989. Xizang. moestaReuter, 188 la). Gansu, Hebei, Heilongjiang,Jilin, P. viridissima (Poda, 1761) ( = Pentatoma rotundicollis Liaoning, Nei Monggol, Ningxia, Qinghai, Shaanxi, Westwood, 1837; Palomena amplificata Distant, 1880; Shanxi, Sichuan, Xizang, Xinjiang. Widespread Palae­ P. viridissima var. simulans Puton, I 881; P. viridissima arctic species. var. rubicundaWesthoff, 1884). Gansu, Hebei, Heilong­ P. semiannulata (Motschulsky, 1860) (= Tropicoris jiang, Jilin, Nei Monggol, Ningxia, Qinghai, Shaanxi, armandi Fallou, 1881; Gudeaichikawana Distant, 1911). Shanxi, Shandong, Yunnan. Widespread Palaearctic spe­ Gansu, Guizhou, Hebei, Heilongjiang, Henan, Hubei, cies. Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Nei Monggol, Paracritheus trimaculatus (Lepeletier & Serville, Ningxia, Qinghai, Shaanxi, Shanxi, Sichuan, Zhejiang. 1828) (= Hoplistodera gibba Vollenhoven, 1868). Guang­ Also known fromMongolia, eastern Russia, Korea, and xi. Also recorded from India, Myanmar, the Philippines, Japan. - Note: Ling & Zheng (1987) only tentatively ac­ and Indonesia. cepted Hsiao & Cheng's (1977) placement of armandi Paralcimocoris chinensis Zhang & Lin, 1988. Xizang. and semiannulataas synonyms, as they were not able to Paterculus affinis (Distant, 1900a). Hunan, Sichuan, examine type material of armandi. Xizang, Yunnan.Also known from India and Bhutan. P. sordlda Zheng& Liu, 1987b. Gansu, Hubei, Hunan, P. bidentatus Xiong & Liu, 1996. Yuonan. Zhejiang. P. elatus (Yang, 1934). Anhui, Fujian, Guangdong, P. tabalensls Zheng & Ling, 1983. Shaanxi. Guangxi, Guizhou, Hunan, Jiangsu, Jiangxi, Sichuan, P.zhengiRider, 1998 (=P.venosa Zheng& Ling, 1983 Yunnan, Zhejiang. (preoccupied)). Yunnan. -Note: P. venosa Zheng & Ling, P. ovatus Hsiao& Cheng, 1977. Hubei,Sichuan, Yunnan. 1983 is a junior primary homonym of P. venosa Forster, P. parvus Hsiao & Cheng, 1977. Guangxi, Hunan, Yun­ 1891, which is still considered a valid species of fossil nan, Zhejiang. Pentatomidae. P. vittatus (Distant, I 90 I a). Guizhou. Originally de­ Phaeocorisellipticus ellipticus (Herrich-Schiiffer,1840) scribed fromMyanmar. (= P. semenoviJakovlev, 1887; Timuria melanoceraHorvath, Pentatoma acuticornuta Zheng& Ling, 1983. Sichuan. 1903; Tancreisca breddiniJensen-Haarup, 1937). Qinghai, P. angulata Hsiao & Cheng, 1977. Gansu, Hunan, Jilin, Xinjiang, Xizang. Also recorded from Mongolia, Sichuan, Xizang, Yunnan. Kazakhstan,and Russia. -Note: Thisspecies has had a con­ P. bruonea Zheng & Ling, 1983. Sichuan. fusedhistory. Some believe it belongs in the ; oth­ P. carlnata Yang, 1934. Gansu, Guizhou, Hunan, Jiang­ ers in theMyrocheini. Zhang et al. ( 1995) had it listed as a xi, Yunnan. junior synonymof Dymantisplana (Fabricius), although they P. distincta Hsiao & Cheng, 1977. Guizhou, Sichuan, listed it as Dymantis elliptica in the figure legend. At this Xizang, Zhejiang. time, it seems best to leave it asPhaeocoris ellipticusuntil P. emeiensis Ling, 1988. Sichuan. its actual taxonomic placement can be studied. 144 D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae • ZOOSYST. ROSSICA Vol. 11 hybneri (Gmelin, 1790) (= Cimex rubro­ P. stali Scott, 1874 (= Nezara amurensis Reuter, fasciatus Fabricius, 1787 (preoccupied); C. flavescens 1888b). Fujian, Gansu, Guangdong, Guangxi, Hebei, He­ Fabricius, 1798 (preoccupied); Rhaphigasterflavolineatus nan, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Westwood, 1837; R. virescens Amyot & Serville, 1843; Shaanxi, Shandong, Shanxi, Sichuan, Zhejiang. Also Nezarapellucida Ellenrieder, 1862). Anhui, Henan, Jiang­ known fromeastern Russia, Korea, Japan, and Hawaii. - su, Shaanxi, Shandong, Shanxi. Widespread Palaearctic Note: Several workers in Japan have treated this species species that also occurs in Africa, southeast Asia, and In­ as a subspecies of crossota. donesia. - Note: Cimex rubrofasciatus Fabricius, 1787, Praetextatus chinensis Hsiao & Cheng, 1977. Hainan, is a junior primary homonym of C. rubrofasciatus De Hunan, Sichuan, Zhejiang. Geer, 1783, which is a species ofReduviidae. Yet, many P. typicus Distant, 1901 b. Taiwan. Also recorded from workers continue to use P. rubrofasciatus.Gmelin (1790) Myanmar. proposed hybneri as a replacement name. Priassus excoffieri Martin, I 902. Yunnan. P. Iituratus (Fabricius, 1794) (= Cimex bilobus Gmelin, P. exemptus (Walker, 1868) (= P. carinatus Horvath, 1790 (preoccupied); Pentatoma incarnatum Germar, 1889a). Fujian?, Taiwan, Xizang, Zhejiang? Also known 1821; P. alliaceum Germar, 1824; P.purpuripennis Hahn, from Afghanistan, Tajikistan, India, and. Myanmar. - 1834; Rhaphigaster punctulatus Westwood,1837; Penta­ Note: The Fujian (Lin & Zhang, 1993) and Zhejiang toma purpuripennis griseoviridis Gorski, 1852; P. purpu­ (Chen & Xu, 1995) records need verification. ripennis rosea Gorski, 1852; Piezodorus degeeri Fieber, P. spiniger Haglund, 1868. F�jian, Guangxi, Guizhou, 1861; Rhaphigaster parilis Walker, 1867). Xinjiang. Hubei, Hunan, Jiangxi, Sichuan, Xizang,Yunnan, Zhejiang. Widespread Palaearctic species. Also recorded from India, Myanmar, and Indonesia. Placosternum alces StAI, 1876. Shanxi, Yunnan.Also P. testaceus Hsiao & Cheng, 1977. Gansu, Guizhou, recorded from Pakistan, India, and Sri Lanka. Hubei, Sichuan, Xizang, Yunnan. [P. dama (Fabricius, 1794)]. India. - Note: Distant Prionaca hubeiensis Zhang, Lin & Zhao, 1990. Hubei. ( 1902) recorded this species fromCochin, India; this was P. hunanensis Lin & Zhang, 1989. Hunan. misinterpreted as Cochin China by Kirkaldy (1909), which P. jiangxiensis Lin & Zhang, 1989. Hunan, Jiangxi. was furthermisinterpreted as Cochin, China by Hoffmann P. tlbetana Zheng & Jin, 1990. Xizang. (1935). Hoffmann(1954) corrected his mistake and indi­ P. tonkinensis Distant, 1921. Guizhou, Yunnan.Origi­ cated that P. dama does not occur in China. nally described from Vietnam. P. jiangleense Lin & Zhang, 1993. Fujian, Hunan. P. yunnanensis Zhang & Lin, 1992. Guizhou, Yunnan. P. taurus (Fabricius, 1781). Guizhou, Hainan, Yunnan. Rhaphigaster brevispina Horvath, 1889b (= R. mon­ Also known from Pakistan, India, Sri Lanka, and south­ golica Puton, 1898). Gansu, Nei Monggol, Ningxia, Xin­ east Asia. jiang. Occurs from Israel and Syria through Kazakhstan P. urus StAI, 1876. Fujian, Guizhou, Henan, Hubei, and Central Asia eastward to Mongolia. - Note: This spe­ Hunan, Jiangxi, Shandong, Sichuan, Xizang, Yunnan, cies has often been confused with R. nebulosa. Records Zhejiang. Also reported from India and Sri Lanka. of R. nebulosa fromChina should be referredto this spe­ crossota (Dallas, 1851) (= Cimex fimbriatus cies. Fabricius, 1787 (preoccupied); Pentatoma fimbriolatum R. genitalia Yang, 1934. Henan, Jiangxi, Zhejiang. Herrich-Schiiffer, 1844 (new name forfimbriatus Fabri­ [R. nebulosa (Poda, 1761)] (= Cimexatomarius Fabri­ cius, 1787, but also preoccupied); Rhaphigaster rufoviri­ cius, 1775; C. sagittifer Goeze, 1778; C. lentiginosus dis Walker, 1867; Nezara rubripennis Jakovlev, 1890; Gmelin, 1790; C. punctipennis Illiger, 1807; Pentatoma Plautia elegans Linnavuori, 1975). Fujian, Guangdong, stigmatica Palisot de Beauvois, 1811; Menida juscaMo­ Guangxi, Guizhou, Hainan, Hubei, Hunan, Sichuan, Xi­ tschulsky, 1861; Rhaphigaster griseus var. impunctata zang, Yunnan. Widespread, occurring in parts of Africa, Garbiglietti, 1869). Widespread westernPalaearctic spe­ Afghanistan, India, Sri Lanka, southeast Asia, to the Phil­ cies. - Note: This species has often been confusedwith ippines and Indonesia. - Note: It appears that about half R. brevispina. Records of R. nebulosa fromChin a should of recent citations have been made using each of the two be referred to R. brevispina. namesfimbriata and crossota. Roche (1979) confirmed Rhynchocoris humeralis (Thunberg, 1783) (= Cimex that crossota andfimbriataare indeed synonyms. Cimex hamatus Fabricius, 1787). Fujian, Guangdong, Guangxi, fimbriatus Fabricius, 1787, is a primary junior homonym Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Macao, of C. fimbriatus Thunberg, 1783, a species now in the Shandong, Sichuan, Taiwan, Yunnan, Zhejiang. Occurs South African genus Dismegistus. Herrich-Schiiffer from Pakistan, India, and Sri Lanka eastward through (1844) recognised this case of homonymy and proposed China and southeast Asia to Indonesia. Pentatomafimbriolatum as a replacement name, but this R. nigridens StAI, 1871. Fujian, Guangdong, Hainan, also is a junior primary homonym, in this case of P.fimbri­ Yunnan. Originally described fromthe Philippines. olatum Germar, 1838, which is now a synonym of Euryde­ R. plagiatus (Walker, 1867) (= R. taprobanensis Berg­ ma dominulum. The next available name is Pentatoma roth, 1891). Yunnan. Also known from India, Sri Lanka, crossota Dallas, 1851. and Vietnam. P. flavifuscaLiu & Zheng, 1994. Guangdong, Hainan, Risibia xanthochila Horvath, 1888 (= Parastaria mi­ Xizang. - Note: This species was proposed to represent mula Kiritshenko, 1911). Xinjiang. Iran and Afghanistan the misidentifications of P. viridicollis (nee Westwood) east to Mongolia. by Hsiao & Cheng (1977) and Zheng & Jin (1990). Rolstoniellus boutanicus (Dallas, 1849). Sichuan, Xi­ P. Iushanica lushanica Yang, 1934. Fujian, Guizhou, zang, Yunnan. Originally described fromBhutan, and also Henan, Hubei, Jiangxi, Shaanxi, Shanxi, Sichuan, Yunnan, known from India and Myanmar. - Note: This species Zhejiang. has until recently been placed in the genus Compastes P. Iushanica yunnanensis Liu & Zheng, 1994. Yunnan. Stal, 1867, which is a junior homonym of Compastes P. propinqua Liu & Zheng, 1994. Gansu, Hubei, Si­ Gistel, I 848, a genus in the Orthoptera. As such, Rider chuan, Yunnan. (1997) proposed Rolstoniellus as a replacement name. P. sordida Xiong & Liu, 1996. Yunnan. R. exstimulatus (Yang, 1935). Yunnan. Originally P. spiendens Distant, 1900c. Fujian. Also recorded described from Bhutan. - Note: See comments under R. from Japan. boutanicus. ZOOSYST. ROSSI CA Vol. 11 • D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae 145 Rubiconla intermedla (Wolff, 1811) (= Pentatoma America. ·-Note: It appears that griseus Eversmann, 183 7 neglectum Herrich-Schaffer, I 853). Gansu, Guangdong, could compete forpriority forthe valid name of this tax.on. Guangxi, Guizhou, Hebei, Heilongjiang, Henan, Hubei, It was presented in a list, without description, but with an Hunan, Jilin, Liaoning, Ningxia, Qinghai, Shanxi, indication that this was a new name forthe misidentified Sichuan. Widespread Palaearctic species. umbrinus of Hahn. S. griseus has not, however, been used R. peltata Jakovlev, 1890. Anhui, Gansu, Hebei, Hei­ by anyone else since, and should at best be considered a longjiang,Henan, Hubei, Hunan, Jiangxi, Jilin, Liaoning, nomen oblitum. Nei Monggol, Shanxi, Sichuan, Zhejiang. Also known S. (A.) umbrlnus (Wolff,1804) (= S. brevicollisFieber, from eastern Russia, Korea, and Japan. 1851; S. fleberii Flor, I 860; S. rotermundi Schumacher, [Sabaeus humeralis (Dallas, I 851)] (= Cuspicona 191 la). Nei Monggol. Widespread Palaearctic species. smaragdina Walker,1867). China. Originally described Note: Most recent workers have considered S. umbrinus from India, and also known from Pakistan, Bangladesh, to be a member of the nominate subgenus, but Derzhansky and Myanmar. - Note: Distant (1902) in passing stated (1994) transferredit to the subgenus Aposciocoris. that he had a specimen of this species in his collection Sepontiella aenea (Distant, 1883). Henan, Zhejiang. fromChina. Catalogers (Kirkaldy, 1909; Hoffinann,1935; Also recorded fromKorea and Japan. - Note: Nearly all Tang, I 935) have repeated this record, but no one else references to this species have been under the combina­ has reported new records from China. Its existence in tion Sepontia aenea. China needs verification. Sinometis emeiensis Zheng & Liu, 1987b. Sichuan. Saceseurus insignis (Distant, 1900b) (= S. bicolor S. lingxianensis Lin & Zhang, 1992. Hunan. Breddin, 1900; Brachycoris thoracicus Distant, 190 I b; Spermatodes variolosus (Walker, 1867). Fujian, B. insignisvar.jlavinotum Kirkaldy, 1909; B. insignis var. Guangdong, Guangxi, Guizhou, Hainan, Jiangxi, Sichuan, nigrescensKirkaldy, 1909). Guangdong, Guangxi. India, Zhejiang. Also known from Pakistan, India, Sri Lanka, Sri Lanka, Myanmar, Indonesia. - Note: Leston (1954) southeast Asia, Japan (Ryukyu Islands), the Philippines, indicated that Saceseurusis the correctgeneric placement and Australia. - Note: Most recent records of this spe­ for this species. cies have been under Sepontia variolosa. Saontarana burmanica Distant, 1918. Yunnan. Type Stacbyomia rubra Chen, 1992. Guangdong. locality is in Myanmar. speciosum (Dallas, 1851) (= Strachia Sarju burmana burmana Ghauri, 1977. Guizhou, inornata Walker, 1867). Guangxi, Sichuan, Yunnan.Also Xizang. Originally described fromMyanmar. occurs fromIran, Pakistan, and India eastward into south­ S. obscura (Westwood,1837) (= Halysnigricollis West­ east Asia. wood, 1837). Guangx.i. Also known fromJapan, India, and Strachia crucigera Hahn, I 833 (= S. jlammulaEllen­ Nepal. -Note: Past records of thisspecies may be under the rieder, 1862; S. strangulata Walker, 1867). Guangdong, combination Dalpada obscura or D. nigricollis. Hainan, Yunnan.Also known fromPakistan, India, south­ S. taungylana Ghauri, 1977. Guangdong, Guangxi, east Asia, Malaysia, and Indonesia. Guizhou, Hainan, Jiangxi, Sichuan, Xizang, Yunnan. Also Tachengia ascra China, 1925. Guizhou, Yunnan. recorded fromVietnam. -Note: Zheng & Liu (1987) in­ T. viridula Hsiao & Cheng, 1977. Guizhou, Sichuan, dicated that Chinese records of Dalpada jugatoria are Xizang, Yunnan. actually misidentificationsand should be referred to this T. yunnana Hsiao & Cheng, 1977. Xizang, Yunnan. species. Tmetopis chin ensis Kiritchenko, 194 7. Guangxi, (Masthletinus) abbreviatus (Reuter, 1879b) Guizhou, Sichuan. -Note: No specificlocality was given (= Masthletinusnigriventris Jakovlev, 1889a). Hebei, Nei in the original description (Kiritshenko, I 947), nor has Monggol, Shandong. Also reported from Kazakhstan, any other specificrecord been published. The Zoological Mongolia, and Russia. - Note: Many records of this spe­ Institute, Russian Academy of Sciences in St. Petersburg cies are under the combination Masthletinus abbreviatus, contains a specimen from Guizhou and the Nankai Uni­ but Wagner (1965) reduced Masthletinus to a subgenus versity, specimens from Guangxi and Sichuan. of Sciocoris. basalis (Dallas, 1851). Fujian, Guangdong, S. (Sciocoris) deltocephalus Fieber, I 861. Xinjiang. Guangxi, Guizhou, Hainan, Jiangxi, Shaanxi, Yunnan, Occurring fromeastern Europe to western China. Zhejiang. Also recorded from Vietnam and Indonesia. S. (S,) dilutus Jakovlev, 1903b (= S. agnatus Jakovlev, T. gutta (Dallas, 1851) (= Pentatoma inobtrusa 1903b). Nei Monggol, Xinjiang. Also known from Af­ Walker, 1867). Fujian, Guangxi, Hainan, Shaanxi, ghanistan, Azerbaijan, Kazakhstan, Kyrgyzstan, Iran, Sichuan, Yunnan, Zhejiang. - Note: At one time this Mongolia, Tajikistan, Turkmenistanand Uzbekistan. species was considered to be a junior synonym of T. S. (S.} distinctus Fieber, 1851 (= S. ovatus Wagner, basalis. 1965). Nei Monggol. Widespread Palaearctic species. T. latipes (Dallas, 185 I) (= Dalpada obtusico/lis S. (S,) indlcus Dallas, 1851. Guangdong, Hainan, Ellenrieder, 1862; Pentatoma trispila Walker, 1867; P. Sichuan, Xizang, Yunnan. Originally described fromIn­ contingens Walker, 1867; Tolumnia ferruginescens dia, and also known from Pakistan. Breddin, 1904). Anhui, Fujian, Guangdong, Guangxi, S. (S.) lateralis Fieber, 1851. Guangdong, Hainan, Guizhou, Hainan, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Yunnan. Alsoknown fromPakistan, India, and Sri Lanka. Shanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang. S. Iugubrls (Walker,1867). Taiwan. -Note: The iden­ Occurs from India through southeast Asia and China to tity of this species needs verification.No one has reported Malaysia and Indonesia. - Note: Many Oriental workers to which subgenus it belongs. still referto T. latipes formcontingens and T. latipes form S. (Aposciocoris) macrocephalus Fieber, 1851 (= S. typica (which should actually be referred to as form basalis Fieber, I 851 ). Gansu, Xinj iang. Widespread latipes). Also, even though Distant (1921) placed T. Palaearctic species. ferruginescens as a junior synonym of this species, sev­ S. (A.) microphthalmus Flor, 1860 (= S. griseus eral workers still occasionally refer to T.ferruginescens Eversmann, 1837 (see note); S. curtipennis Moisant & as a valid species. Rey, 1866; S. microphthalmus poppiusi W.agner, 1965). T. trinotata (Westwood, 183 7) (= Pentatoma pallidi­ Hebei, Heilongjiang, Nei Monggol, Shanxi, Xizang. ventris Vollenhoven, 1868). Yunnan. Also known from Widespread Palaearctic species; also occurs in North Cambodia, Malaysia, and Indonesia. 146 D.A. Rider et al.: Chinese Pentatomidae. II. Pentatominae • ZOOSYST. ROSSI CA Vol. 11 Udo9:ga s�inidens Distant, 1921. Fujian, Guangdong, Guangdong Province. Entomotaxonomia, 22(2): 116- Guangx1, Gmzhou, Hainan, Hubei, Hunan, Jiangxi Ma­ 118. cao, Shaanxi, Shanxi, Xizang, Yunnan, Zhejiang. De;crib­ Chen, z.-Y. & Huang, W.-Z. 1996. A new record spe• ed fromLaos. cies ofAeliomorpha Stal fromChina. Acta Sci.naM Valescus jianhenensis Chen, 1983. Guizhou. Univ. Sunyatseni, 35(5): 126. V. omeiensis Hsiao & Cheng, 1977. Sichuan. Cheng, L. & LI, J. 1979. In: Li, J. Bugs of Ali, Tibet . Vit�llus orientalis Dis!ant, 1900c. Guangdong, Guang­ (Pentatomidae, Coreidae and Reduviidae� - Plants and x1, Gu1zhou, YUIIIlan. Onginally described from India. of Ali region, Xizang: 8-12. Science Press, Vitruvius insignis Distant, 1901 a. Guangxi, Hainan, Peking. Yunnan. Also known from Myanmar and Indonesia. China, W.E. 1925. The collected by Prof. Zhengius spiniferus (Zheng & Liu, 1987b). Xizang, J.W. Gregory 's expedition to Yunnan,with synonymic Yunnan. - Note: This species was originally described in notes on allied species. Ann. Mag. natur. Hist. (9), !he. genus Tibetocoris Zheng & Liu, 1987b, which is a 16: 449-485. Jumor homonym of Tibetocoris Hutchinson, 1934. Rider China, W.E. 1928. Results ofan expedition to �orinchi (1998) proposed Zhengius as a new name. Peak, Sumatra. XIII. Hemiptera: Pentatomo1dea col­ Z. zhangmuensis (Zhang & Lin, 1988). Xizang. - lected in Korind, West Sumatra by Messrs. H.C. Ro­ Note: See comments under z. spiniferus. binson and C. Boden Kloss. J. Fed. Malay States Zheng, 1986. Yunnan. Mus., ZOuicoris elegans 8: 185-196. . Chopra, N.P. 1974. Studies on the genus Halys (Hem1• References ptera: Heteroptera: Pentatomidae). Oriental , 8(4): 473-479. Costa, A. 1847. Cimicum regni Neapolitani. Centuria Abbasi, Q.A. & Ahmad, I. 1976. A new species ofthe ge­ secunda. Decas sexta, septima, octava. nona et Halys nus Fabr.(Pentatomidea: Haly �i) fro1!1East Ben­ decima. 41 pp. Napoli. gal withnotes on its genitalia andtheir beanng on clas­ Costa, A. 1884. Notizie ed osservazioni sulla ge -fauna sification. .J. Sci. Univ. Karachi, 4(1-2): 26-31. � Amyot, C.J.B. Serville, A. Histoire Nature/le Sarda. Memoria terza. Risultamento delle ncerche & 1843. fattein Sardegna nella estate de! 1883. Atti r. Accad. des Jnsectes. Hemipteres. lxxvi and 681 pp., 12 pls. Sci. fis. mat. Napoli (2), I: 1-64. . Roret, Paris. Dallas, W.S. 1849. Notice of some hem1pterous insects Baerensprung, F. von. 1858. Neue und seltene Rhyn­ Berlin. entomol. Z., from Boutan (East Indies), with descriptions of the choten der europiiischenFauna. new species. Trans. entomol. Soc. Lond., 5: 186-194. 2: 188-208, pl. II. Dallas, W.S. _ Becker, A. 1851. List ofthe specimens ofhem1pterous 1867. Noch einige Mittheilungen Uber Astra­ insects in the collection of the British Museum. 2 pts., chaner und Sareptaer Pflanzen und Insekten. Bull. Soc. imp. Natur. Moscou, 40: 592 pp., 15 pls. Brit. Mus., London. I 04-115. Dallas, W.S. Bergroth, E. 1852. Descriptions of some new species of 1891. Contributions ii l'etude des penta­ hemipterous insects belonging to the tribe Scutata. tomides. Rev. Entomol., 10: 200-235. Trans. entomol. Soc. Land. Bergroth, E. (2), 2: 6-17. 1906. Systematische und synonymische De Geer, C. 1773. Memoirespour servir a l'histoire des Bemerkungen Uber Hemipteren. Wien.Entomol. Ztg, Insectes, 25(1): 3: i-viii, 1-696. Stockholm. 1-12. Derzhansky, V.V. 1990. Shield bugs ofthe genus Carpo· Boheman, C.H. 1850. Bidrag till Gottlands insekt-fauna. coris Kong/. Vet.-Akad. Hand!., 70, Kol. (Heteroptera, Pentatomidae) in the USSR (1849): 195-267. Entomol. Obozr., 69(1): Breddin, 61-70. G 1900. Hemiptera Sumatrana collecta a dom. Derzhansky, V.V. Stettin. entomol. 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