ISSN 0206-0477

PROCEEDINGS OF THE ZOOLOGICAL INSTITUTE OF THE RUSSIAN ACADEMY OF SCIENCES

ANNOTATED CATALOGUE OF THE OF RUSSIA

VOLUME I

SYMPHYTA AND : ACULEATA

SUPPLEMENT № 6, 2017 PROCEEDINGS OF THE ZOOLOGICAL INSTITUTE OF THE RUSSIAN ACADEMY OF SCIENCES

SUPPLEMENT № 6

ANNOTATED CATALOGUE OF THE HYMENOPTERA OF RUSSIA

VOLUME I

SYMPHYTA AND APOCRITA: ACULEATA

SAINT PETERSBURG 2017 69. Family Formicidae 197

Superfamily FORMICOIDEA Subfamily DOLICHODERINAE Number of taxa: World – 28 genera, 707 species, 69. Family FORMICIDAE – Palaearctic – 8/53, Russia – 4/16. D.A. Dubovikoff and Z.M. Yusupov BOTHRIOMYRMEX Emery, 1869. T y p e s p e c i e s Bothriomyrmex costae Emery, 1869. Temporary Ants are eusocial aculeate hymenopterous social parasites of Tapinoma species. Number of and currently include 13240 extant species belongs species: World – 24, Palaearctic – 22, Russia – 5. to 334 genera of 17 subfamilies and 738 fossil species Bothriomyrmex anastasiae from 151 genera of 14 subfamilies (3 subfamilies are Dubovikov, 2002. Russia: EP known only by fossils) (Bolton, 2016). There are 1777 (NC). Bothriomyrmex communista B meri- species in 97 genera of 11 subfamilies in the Palaearctic Santschi, 1919 ( . dionalis communista region (AntWeb, 2017). Ants, as eusocial insects, have a var. Santschi, 1919). Russia: EP clear cast differentiation, e.g. sexual (females (queens) (NC, CR). – Europe (WE, SE, EE). Bothriomyrmex corsicus B meridio- and males) and workers (or also soldiers). Colonies Santschi, 1923 ( . nalis corsica B gallicus include one (or more in polyginous colonies) queen and var. Santschi, 1923; . Emery, B gibbus B menozzii from dozens to millions of workers. Ants are one of the 1925; . Soudek, 1925; . Emery, EP most evolutionarily successful groups of invertebrates. 1925). Russia: (NC, CR). – Europe (WE, SE, EE). Bothriomyrmex modestus They are of great importance in various ecosystems, Radchenko, 1985. Russia: EP primarily as effective predators, but also as consumers (S). – Europe (EE). Bothriomyrmex turcomenicus B syrius and distributors of plant seeds, play a significant role Emery, 1925 ( . turcomenicus EP in soil formation and much more. Ants nesting is much Emery, 1925). Russia: (S, NC). – Turkmenistan, Kazakhstan. different: in soil, leaflitter, dead wood, in epiphytes, dif- ferent plants and with different special constructions DOLICHODERUS Lund, 1831 (Hypoclinea Mayr, like mounds. Synonymic and nomenclature data are 1855). Type species Formica attelaboides given according to «An online catalog of the ants of Fabricius, 1775. Number of species: World – 131, the world» (Bolton, 2016). The total number of taxa Palaearctic – 2, Russia – 2. is given excluding fossil and infraspecific taxa, unless Dolichoderus quadripunctatus (Linnaeus, 1771) explicitly indicated. [Formica] (Hypoclinea quadripunctata (Linnaeus, Sections for the genera Leptothorax, , 1771), auct.). Russia: EP (C, E, S, NC, CR), UR. – Temnothorax and most part of Tetramorium species Europe (WE, NE, SE, EE), Armenia, Iran. were prepared by Z.M. Yusupov, the rest of the text Dolichoderus sibiricus Emery, 1889 (D. quadripunc- was prepared by D.A. Dubovikoff. tatus sibiricus Emery, 1889). Russia: ES (KS, KR, Number of taxa: World – 485 genera, 13978 BR), FE (AM, KH, PR, KU). – Mongolia, China species (including fossils), Palaearctic – 94/1350, (NE, NC), Korean Peninsula, Japan (Hok, Hon, Russia – 44/264. Shi, Kyu). R e f e r e n c e s. Ruzsky, 1905, 1907; Karawajew, 1926; Kuznetsov-Ugamsky, 1929; Dlussky, Soyunov, LIOMETOPUM Mayr, 1861. T y p e s p e c i e s For- Zabelin, 1990; Kupianskaya, 1990, 1995, 2012; Rad- mica microcephala Panzer, 1798. Number of species: chenko, 1991, 1992, 1996a, 1996b, 1997a,b,c,d, 2004, World – 7, Palaearctic – 3, Russia – 2. 2005a,b, 2016; Seifert, 1992, 2000, 2003, 2007, 2012; Liometopum microcephalum (Panzer, 1798) [For- Arakelian, 1994; Czechowski et al., 2002; Dubovikoff, mica]. Russia: EP (S, NC). – Europe (WE, SE, EE). 2002, 2005; Csösz et al., 2007, 2010, 2015; Pfeiffer et al., Liometopum orientale Karawajew, 1927 (L. microce- 2007; Seifert, Schultz, 2009; Yusupov, 2009, 2014; Rad- phalum var. orientalis Karawajew, 1927). Russia: chenko, Elmes, 2010; Czechowski et al., 2012; Douwes FE (PR). – Korean Peninsula. et al., 2012; Guénard, Dunn, 2012; Radchenko, Yusupov, 2012, 2017; Borowiec, 2014; Radchenko, Scupola, 2015; TAPINOMA Förster, 1850. Ty p e s p e c i e s Tapi- Radchenko et al., 2015; Bolton, 2016; Radchenko et al., noma collina Förster, 1850 =T. erraticum (Latreille, 2016; Ward et al., 2016; AntWeb, 2017; Grebennikov, 1798). Number of species: World – 69, Palaearctic – Zryanin, 2017; Dubovikoff, Yusupov, 2017. 18, Russia – 7. 198 D.A. Dubovikoff and Z.M. Yusupov

Tapinoma christophi Emery, 1925. Russia: EP (S). Camponotus (Camponotus) obscuripes Mayr, 1879 Tapinoma emeryanum Kuznetsov-Ugamsky, 1927 (C. herculeanus ligniperdus var. obscuripes Mayr, (T. emeryanum dagestanicum Kuznetsov-Ugam- 1879). Russia: FE (SA, KU). – Korean Peninsula, sky, 1927). Russia: EP (NC). – Turkmenistan, Japan (Hok, Hon, Shi, Kyu). Kazakhstan. Camponotus (Camponotus) sachalinensis Forel, Tapinoma erraticum (Latreille, 1798) [Formica] 1904 (C. herculeanus var. sachalinensis Forel, 1904; (T. breve Emery, 1925; T. tauridis Emery, 1925). C. herculeanus var. wymperi Forel, 1904; C. hercu- Russia: EP (C, E, S, NC, CR), UR. – Europe (WE, leanus var. montana Ruzsky, 1905 (part); C. hercu- SE, EE), Caucasus, Turkmenistan, Kazakhstan. leanus var. altaica Ruzsky, 1915; C. herculeanus var. Tapinoma kinburni Karawajew, 1937. Russia: EP (C, eudokiae Ruzsky, 1926; C. herculeanus var. jacutica S). – Europe (EE). Karawajew, 1929). Russia: WS (AL), ES (KS, TU, Tapinoma melanocephalum (Fabricius, 1793) KR, IR, BR, YA, ZB), FE (KH, PR, SA, KU, KA, [Formica]. Tramp species. Russia: EP (Moscow, MG). – Mongolia, China (NE), Korean Peninsula, introduced). – Cosmopolitan. Japan (Hok, Hon, Shi). Tapinoma sinense Emery, 1925. Russia: ES (IR), FE Camponotus (Camponotus) saxatilis Ruzsky, 1895 (KH, PR). – Mongolia, China (NE, NC, NW, CC, (C. herculeanus saxatilis Ruzsky, 1895; C. penn- SW, WP, SE), Korean Peninsula. sylvanicus saxatilis Ruzsky, 1905). Russia: EP (E, UR WS ES Tapinoma subboreale Seifert, 2012 (T. erraticum ambi- S), , (TM, OM, TK, NS, KM, AL), (KS, FE guum Emery, 1925; T. ambiguum Emery, 1925, auct.). TU, KR, IR, BR, YA, ZB), (AM, KH, PR, SA, Russia: EP (C, S, NC). – Europe (WE, NE, SE, EE). MG). – Mongolia, China (NE), Korean Peninsula. Camponotus (Camponotus) vagus (Scopoli, 1763) [Formica] (C. vagus var. kodorica Forel, 1913). Subfamily FORMICINAE Russia: EP (C, E, S, NC, CR), UR, WS (TM, OM, TK, NS, KM, AL). – Europe (WE, NE, SE, EE), N Number of taxa: World – 51 genera, 3050 species, Africa, Turkey, Kazakhstan. Palaearctic – 23/382, Russia – 12/98. Camponotus (Myrmentoma) atricolor (Nylander, CAMPONOTUS Mayr, 1861. Ty p e s p e c i e s For- 1849) [Formica] (C. merula Losana, 1834; C. piceus mica ligniperda Latreille, 1802. Number of species: (Leach, 1825), part, auct.). Russia: EP (C, E, S, NC, World – 1021, Palaearctic – 113, Russia – 16. CR), UR. – Europe (WE, SE, EE), N Africa, Cau- casus, Turkey, Iran, Kazakhstan. Camponotus (Camponotus) herculeanus (Lin- Camponotus (Myrmentoma) fallax (Nylander, 1856) naeus, 1758) [Formica] (C. herculeanus var. shitkovi [Formica] (C. marginatus Latreille, 1798 (part, C herculeanus altaica Ruzsky, 1904; . var. Ruzsky, auct.); C. marginatus var. ruzskyi Emery, 1898; C herculeanus caucasicus 1915; . Arnoldi, 1967). C. marginatus var. kamensis Ruzsky, 1903). Rus- EP UR WS Russia: (N, NW, C, E, NC), , (TM, OM, sia: EP (C, E, S, NC, CR), UR. – Europe (WE, NE, NS). – Europe (WE, NE, SE, EE), Caucasus, Tur- SE, EE), N Africa, Caucasus, Turkey, Kazakhstan. key, Kyrgyzstan, Kazakhstan. Camponotus (Myrmentoma) quadrinotatus Forel, Camponotus (Camponotus) japonicus Mayr, 1866 1886 (C. marginatus var. quadrinotatus Forel, 1886; (C. herculeanus var. aterrima Emery, 1894; C. her- C. marginatus var. rubicolor Ruzsky, 1925; C. macu- culeanus japonicus var. manczhuricus Ruzsky, latus quadrinotatus var. nigricolor Ruzsky, 1925; 1915; C. japonicus var. sanguinea Karawajew, 1929; C. marginatus brunni Ruzsky, 1926). Russia: FE C. japonica var. miltotus Wheeler, 1929; C. japo- (KH, PR). – China (NC, NW, CC, SW, WP, SE), nica wui Wheeler, 1929; C. aterrimus Emery, 1895: Korean Peninsula, Japan (Hok, Hon, Shi, Kyu). Kupianskaya, 2012). Russia: ES (KS, KR, IR, BR, Camponotus (Myrmentoma) interjectus Mayr, 1877. ZB), FE (AM, KH, PR, SA, KU). – Mongolia, China Russia: EP (NC). – Israel, Iraq, Iran, Afghanistan, (NE, NC, NW), Korean Peninsula, Japan (Hok, Central Asia. Hon, Shi, Kyu). Camponotus (Myrmentoma) lateralis (Olivier, Camponotus (Camponotus) ligniperda (Latreille, 1792) [Formica]. Russia: EP (NC, CR). – Europe 1802) [Formica]. Russia: EP (N, NW, C, NC), UR. – (WE, SE, EE), N Africa, Caucasus, Turkey, Iran, Europe (WE, NE, SE, EE), Caucasus, Turkey. Turkmenistan. 69. Family Formicidae 199

Camponotus (Tanaemyrmex) aethiops (Latreille, FORMICA Linnaeus, 1758. Ty p e s p e c i e s Formica 1798) [Formica] (F. nigrata Nylander, 1849; Campo- rufa Linnaeus, 1761. Number of species: World – notus aethiops var. glaber Emery, 1925). Russia: EP 176, Palaearctic – 53, Russia – 32. (C, E, S, NC, CR), UR. – Europe (WE, SE, EE), N Formica (Coptoformica) exsecta Nylander, 1846 Africa, Caucasus, Turkey, Israel, Iran, Afghanistan, (F. exsecta var. rubens Forel, 1874; F. nemoralis Central Asia, Kazakhstan. Dlussky, 1964). Russia: EP (N, NW, C, E, S, NC, Camponotus (Tanaemyrmex) fedtschenkoi Mayr, CR), UR, WS (TM, OM, TK, NS, KM, AL), ES 1877 (C. tichomirovi Ruzsky, 1905; C. fedtschenkoi (IR, BR, ZB), FE (AM, KH, SA, KA, MG). – Europe var. adi Stitz, 1930). Russia: EP (S, NC). – Armenia, (WE, NE, SE, EE), Mongolia, China (NE). Azerbaijan, Syria, Iran, Afghanistan, Turkmen- Formica (Coptoformica) fennica Seifert, 2000. Rus- istan, Uzbekistan, Kyrgyzstan, Kazakhstan. sia: EP (N, NC). – Europe (NE). Camponotus (Tanaemyrmex) turkestanicus Emery, Formica (Coptoformica) foreli Bondroit, 1918 (F. ex- C sylvaticus turkestanicus 1887 ( . st. Emery, 1887; secta pressilabris var. foreli Bondroit, 1918; F. tama- C attila EP . Stärcke, 1935). Russia: (S, NC). – Iran, rae Dlussky, 1964). Russia: EP (C, NC). – Europe Afghanistan, Turkmenistan, Uzbekistan, Kyrgyz- (WE, NE, SE, EE), Turkey, Kazakhstan. stan, Kazakhstan. Formica (Coptoformica) forsslundi Lohmander, Camponotus (Tanaemyrmex) turkestanus André, 1949 (F. brunneonitida Dlussky, 1964; F. fossilabris C sylvaticus turkestanus 1881 ( . var. André, 1882; Dlussky, 1965). Russia: EP (N, C, NC), ES (TU). – C ferganensis EP . Pisarski, 1967). Russia: (S, Europe (WE, NE, EE). NC). – Israel, Iran, Afghanistan, Turkmenistan, Formica (Coptoformica) manchu Wheeler, 1929 Uzbekistan, Kyrgyzstan, Kazakhstan. (F. exsecta var. manchu Wheeler, 1929; F. longi- CATAGLYPHIS Förster, 1850. Ty p e s p e c i e s ceps Dlussky, 1964, nom praeocc., nec Smith, 1863; Cataglyphis fairmairei Förster, 1850. Number of F. dlusskyi Bolton, 1995). Russia: ES (BR, YA, ZB), species: World – 91, Palaearctic – 65, Russia – 3. FE (AM, PR). – Mongolia, China (NE), Korean Peninsula. Cataglyphis aenescens (Nylander, 1849) [Formica] Formica (Coptoformica) pisarskii Dlussky, 1964. (Myrmecocystus cursor r. tancrei Forel, 1901; M. cur- Russia: ES (BR, YA, ZB), FE (AM). – Mongolia. sor var. caspius Ruzsky, 1902; M. altisquamis var. Formica (Coptoformica) pressilabris Nylander, 1846 jakobsoni Ruzsky, 1905; Cataglyphis aenescens ater- (F. exsecta pressilabris var. rufomaculata Ruzsky, rima Pisarski, 1967; C. aenescens georgica Arnoldi, 1895; F. exsecta pressilabris Kuznetsov-Ugamsky, 1968; C. aenescens chatkalenis Tarbinsky, 1976). 1928; F. rufomaculata: Kupianskaya, 1995). Russia: Russia: EP (C, E, S, NC, CR), UR, WS (OM, NS, EP (N, NW, C, E, S, NC), UR, WS (AL), ES (IR, BR, AL), ES (KS, TU, BR). – Europe (SE, EE), Caucasus, ZB), FE (AM, KH, PR). – Europe (WE, NE, SE, EE), Turkey, Iran, Afghanistan, Central Asia, Kazakh- Turkey, Mongolia, China (NE), Korean Peninsula. stan, Mongolia, China (NE, NC, NW). Formica (Coptoformica) suecica Adlerz, 1902. Rus- Cataglyphis nodus (Brullé, 1833) [Formica] (Cata- sia: EP (N), WS (OM). – Europe (NE). glyphis nodus caucasicola Arnoldi, 1964). Russia: Formica (Formica) aquilonia Yarrow, 1955. Russia: EP (NC). – Europe (SE, EE), Caucasus, Turkey, EP (N, NW, C), UR, WS (TM, OM, TK, NS, KM, Israel, Saudi Arabia, Iran, United Arab Emirates, AL), ES (KS, KR, IR, BR, YA, ZB), FE (KH, PR, Afghanistan, Central Asia. SA, MG). – Europe (WE, NE, SE, EE), Mongolia, Cataglyphis pallida Mayr, 1877. Russia: EP (S, NC). – China (NE), Korean Peninsula. Afghanistan, Central Asia, Kazakhstan. Formica (Formica) kupyanskayae Bolton, 1995 F opacа COLOBOPSIS Mayr, 1861. Ty p e s p e c i e s Formica ( . Kupianskaya, 1980, nom. praeocc., nec FE truncata Spinola, 1808. Number of species: World – Nylander, 1856). Russia: (AM, KH, PR, SA). – 94, Palaearctic – 1, Russia – 1. Korean Peninsula. Formica (Formica) lugubris Zetterstedt, 1838 (F. con- Colobopsis truncatus (Spinola, 1808) [Formica] gerens Nylander, 1846; F. rufa var. rufo-pratensis (Camponotus (Colobopsis) truncatus (Mayr, 1861), Forel, 1874; F. pratensis unicolor Ruzsky, 1926; auct.). Russia: EP (C, E, S, NC, CR). – Europe (WE, F. rufa var. nylanderi Bondroit, 1919). Russia: EP SE), N Africa, Caucasus, Turkey, Turkmenistan. (N, NW, C), UR, WS (TM, OM, TK, NS, KM, AL), 200 D.A. Dubovikoff and Z.M. Yusupov

ES (KR, IR, BR, YA, ZB), FE (KH, PR, KA, MG). – 1902, auct.). Russia: EP (N, NW, C, E, S, NC, CR), Europe (WE, NE, SE, EE), Mongolia, China (NE), UR, WS (TM, AL). – Europe (WE, NE, SE, EE), Korean Peninsula. Caucasus, Kazakhstan. Formica (Formica) polyctena Förster, 1850. Russia: Formica (Serviformica) cinereofusca Karawajew, EP (N, NW, C, E, S), UR, WS (TM, OM, TK, NS, 1929 (F. cinerea var. cinereofusca Karawajew, 1929). KM, AL), ES (KS, KR, IR, BR, YA, ZB). – Europe Russia: EP (NC). – Europe (SE, EE), Caucasus. (WE, NE, SE, EE), Kazakhstan. Formica (Serviformica) clarissima Emery, 1925 (F. ru­ Formica (Formica) pratensis Retzius, 1783 (F. pra- fibarbis subpilosa var. clarior Ruzsky, 1914, nom. tensis var. ciliata Ruzsky, 1926). Russia: EP (N, praeocc., nec Ruzsky, 1905; F. rufibarbis var. cla- NW, C, E, S, NC, CR), UR, WS (TM, OM, TK, rissima Emery, 1925; F. subpilosa ruzskyi Dlussky, NS, AL), ES (KR, IR, BR, ZB), FE (AM). – Europe 1965: Kupianskaya, 2012). Russia: ES (BR, ZB), FE (WE, NE, SE, EE), Caucasus, Turkey, Central Asia, (AM, KH). – Mongolia, China (NE). Kazakhstan, Mongolia, China (NE). Formica (Serviformica) cunicularia Latreille, 1798 Formica (Formica) rufa Linnaeus, 1761 (F. rufa var. (F. cunicularia fuscoides Dlussky, 1967). Russia: EP meridionalis Nasonov, 1889). Russia: EP (N, NW, (C, E, S, NC, CR), UR, WS (OM, TK, NS, AL). – C, E, S), UR, WS (TM, OM, NS, AL), ES (KS, Europe (SE, EE), Caucasus, Turkey, Iran, Central KR, IR). – Europe (WE, NE, SE, EE), Turkey, Asia, Kazakhstan. Kazakhstan. Formica (Serviformica) fusca Linnaeus, 1758. Rus- Formica (Formica) truncorum F trun- Fabricius, 1804 ( . sia: EP (N, NW, C, E, S, NC, CR), UR, WS (OM, cicola F rufa rufotruncicola Nylander, 1846; . var. TK, NS, KM, AL), ES (KS, TU, KR, IR, BR, ZB). – F truncorum menozzii Ruzsky, 1896; . var. Krausse, Europe (WE, NE, SE, EE), Caucasus, Turkey, Cen- 1926). Russia: EP (N, NW, C, E, S, NC), UR, WS tral Asia, Kazakhstan, Mongolia, China (NE), N (TM, NS, AL), ES (KS, TU, KR, IR, BR, ZB), FE America (introduced). (AM, KH, PR, SA, KU). – Europe (WE, NE, SE, Formica (Serviformica) gagates Latreille, 1798. Rus- EE), Caucasus, Turkey, Afghanistan, Central Asia, sia: EP (C, S, NC, CR). – Europe (WE, SE, EE), Kazakhstan, Mongolia, China (NE), Korean Pen- Georgia, Turkey. insula, Japan (Hok). Formica (Serviformica) gagatoides Ruzsky, 1904 Formica (Formica) yessensis Forel, 1901 (F. truncicola (F. fusca var. gagatoides Ruzsky, 1904). Russia: EP var. yessensis Forel, 1901). Russia: FE (PR). – China UR WS (NW, SE), Korean Peninsula, Japan (Hok, Hon). (N, NW), , (TM, OM, TK, NS, KM, AL), ES FE Formica (Raptiformica) sanguinea Latreille, 1798 (KS, TU, KR, IR, BR, YA, ZB), (KH, KA, (F. dominula Nylander, 1846; F. sanguinea var. MG). – Europe (NE), Japan (Hon). Formica (Serviformica) glauca F rufi- fuscipes Emery, 1895; F. sanguinea var. mollesonae Ruzsky, 1896 ( . Ruzsky, 1903; F. sanguinea var. clarior Ruzsky, 1905; barbis var. glauca Ruzsky, 1896; F. rufibarbis var. F. sanguinea arenicola Kuznetsov-Ugamskij, 1928). katuniensis Ruzsky, 1915). Russia: EP (C, E, S, NC, Russia: EP (NW, C, E, S, NC, CR), UR, WS (OM, CR), UR, WS (OM, TK, NS, KM, AL), ES (KS, KR, TK, NS, AL), ES (KS, TU, KR, IR, BR, YA, ZB), IR). – Europe (SE, EE), Caucasus, Turkey, Central FE (AM, KH, PR, SA, KU, MG). – Europe (WE, Asia, Kazakhstan, Mongolia. NE, SE, EE), Caucasus, Turkey, Iran, Afghanistan, Formica (Serviformica) japonica Motschulsky, 1866 Kyrgyzstan, Kazakhstan, Mongolia, China (NE, (F. fusca var. nipponensis Forel, 1901; F. fusca var. NW, CC), Korean Peninsula, Japan (Hok, Hon). japonica Wheeler, 1913). Russia: FE (AM, PR, SA, Formica (Serviformica) candida F. Smith, 1878 KU). – Mongolia, China (NE, NC, NW, CC, SW, WP, (F. fusca picea var. inplana Ruzsky, 1914; F. fusca SE), Korean Peninsula, Japan (Hok, Hon, Shi, Kyu). picea var. orientalis Ruzsky, 1914; F. picea auct.). Formica (Serviformica) lemani Bondroit, 1917 (F. fus­ Russia: WS (AL), ES (KS, TU, KR, IR, BR, YA, ca var. borealis Waschkewitsch, 1924). Russia: EP ZB), FE (AM, KH, PR, SA, KU, MG). – Kyrgyz- (N, NW, NC), UR, WS (TM, OM, TK, NS, KM, stan, Kazakhstan, Mongolia, China (NE, NW, CC), AL), ES (KR, IR, BR, YA, ZB) FE (AM, KH, PR, Korean Peninsula, Japan (Hok, Hon, Kyu). SA, KU, KA, MG, CH). – Europe (WE, NE, SE, Formica (Serviformica) cinerea Mayr, 1853 (F. cine- EE), Mongolia, China (NE), Korean Peninsula, rea var. armenica Ruzsky, 1905; F. imitans Ruzsky, Japan (Hok, Hon, Shi). 69. Family Formicidae 201

Formica (Serviformica) picea Nylander, 1846 Lasius (Chthonolasius) jensi Seifert, 1982 (L. umbra- (F. transcaucasica Nasonov, 1889). Russia: EP (N, tus: Кupianskaya, 1995). Russia: FE (AM, PR). – NW, C, E, NC, CR), UR, WS (TM, OM, TK, NS, Europe (WE, EE), Armenia, Korean Peninsula. KM, AL). – Europe (WE, NE, SE, EE), Georgia, Lasius (Chthonolasius) longiceps Seifert, 1988. Rus- Armenia, Turkey, Kazakhstan. sia: EP (C, S), UR. – Europe (EE), Kazakhstan. Formica (Serviformica) rufibarbis Fabricius, 1793. Lasius (Chthonolasius) meridionalis (Bondroit, Russia: EP (C, E, S, NC, CR), UR, WS (OM, TK, 1920) [Formicina]. Russia: EP (C, S, NC), FE NS, KM, AL). – Europe (WE, NE, SE, EE), Cau- (PR). – Europe (WE, NE, SE, EE), Georgia, Ar- casus, Turkey, Iran. menia, Kazakhstan, Korean Peninsula, Japan (Hok, Formica (Serviformica) subpilosa Ruzsky, 1902 Hon, Shi, Kyu). (F. rufibarbis subpilosa Ruzsky, 1902; F. cinerea var. Lasius (Chthonolasius) mixtus (Nylander, 1846) bipilosa Karawajew, 1926). Russia: EP (S, NC). – [Formica]. Russia: EP (NW, C, E, S, NC, CR), WS Caucasus, Central Asia. (TM, OM, TK, NS, KM, AL), ES (ZB), FE (AM, KH, Formica (Serviformica) uralensis Ruzsky, 1895. Rus- PR). – Europe (WE, SE, EE), Georgia, Armenia, sia: EP (C, NC), UR, WS (OM, TK, NS, KM, AL), Kazakhstan, Korean Peninsula. ES (KS, TU, KR, IR, BR, ZB), FE (AM, PR, SA, Lasius (Chthonolasius) sabularum (Bondroit, 1918) KU). – Europe (WE, NE, EE), Mongolia, China [Formicina]. Russia: EP (C, S, NC), ES (IR). – (NE), Korean Peninsula. Europe (WE, NE, SE, EE). Lasius (Chthonolasius) umbratus (Nylander, 1846) LASIUS Fabricius, 1804. Ty p e s p e c i e s Formica [Formica] (L. umbratus var. exacutus Ruzsky, 1902). nigra Linnaeus, 1758. Number of species: World – Russia: EP (NW, C, E, S, NC, CR), UR, WS (TM, 112, Palaearctic – 55, Russia – 31. OM, TK, NS, KM, AL), ES (ZB), FE (AM, PR, SA, KU). – Europe (WE, NE, SE, EE), Caucasus, Tur- Lasius (Austrolasius) carniolicus Mayr, 1861. Russia: key, Kazakhstan, Korean Peninsula, Japan (Hok, EP (C, S), UR, WS (Radchenko, 2016), ES (Rad- Hon, Shi, Kyu), N America. chenko, 2016), FE (PR, SA). – Europe (WE, NE, Lasius (Dendrolasius) capitatus (Kuznetsov-Ugam- SE, EE), Kazakhstan. sky, 1928) [Acanthomyops] (A. fuliginosus capita- Lasius (Cautolasius) flavus (Fabricius, 1782) [For- tus Kuznetsov-Ugamsky, 1928). Russia: FE (PR). – mica] (L. flavus var. fuscoides Ruzsky, 1902; L. fla- Japan (Hok, Hon, Shi). vus odoratus L flavus olivacea Ruzsky, 1905; . var. Lasius (Dendrolasius) fuji Radchenko, 2005 (L. fuli- EP Karawajew, 1926). Russia: (N, NW, C, E, S, NC, ginosus: Кupianskaya, 1995). Russia: FE (AM, KH, UR WS ES CR), , (TM, OM, TK, NS, KM, AL), PR, SA, KU). – China (NE), Korean Peninsula, (KS, TU, KR, IR, BR, YA, ZB), FE (AM, KH, PR, Japan (Hok, Hon, Shi). SA, KU). – Europe (WE, NE, SE, EE), Caucasus, Lasius (Dendrolasius) fuliginosus (Latreille, 1798) Iran, Kazakhstan, Korean Peninsula, Japan (Hok, [Formica]. Russia: EP (N, NW, C, E, S, NC, CR), Hon, Shi, Kyu), N America. UR, WS (OM, TK, NS, KM, AL). – Europe (WE, Lasius (Chthonolasius) bicornis (Förster, 1850) [For- NE, SE, EE), Caucasus, Kazakhstan. mica]. Russia: EP (C, E, S, NC). – Europe (WE, NE, Lasius (Dendrolasius) morisitai Yamauchi, 1978. SE, EE), Caucasus. Russia: FE (PR). – Korean Peninsula, Japan (Hon). Lasius (Chthonolasius) citrinus Emery, 1922 (L. affi- Lasius (Dendrolasius) nipponensis Forel, 1912 nis Schenck, 1852, nom praeocc., nec Leach, 1825; (L. fuliginosus var. nipponensis Forel, 1912; L. cris- L. bicornis var. citrine Emery, 1922; L. affinis auct.). pus Wilson, 1955). Russia: FE (PR). – Korean Pen- Russia: EP (C, E, S, NC, CR), ES (BR), FE (AM, insula, Japan (Hok, Hon, Shi). KH, PR, SA, KU). – Europe (WE, SE, EE), Cau- Lasius (Dendrolasius) orientalis Karawajew, 1912 casus, Kazakhstan, Korean Peninsula. (L. fuliginosus var. orientalis Karawajew, 1912; Lasius (Chthonolasius) distinguendus (Emery, 1916) L. teranishii Wheeler, 1928; L. ouchii Teranishi, [Formicina] (F. umbrata distinguenda Emery, 1916; 1940). Russia: FE (AM, PR, SA, KU). – Korean Lasius hybrida (Bondroit, 1918)). Russia: EP (C, Peninsula, Japan (Hok, Hon). E, S, NC), ES (ZB), FE (AM, PR). – Europe (WE, Lasius (Dendrolasius) spathepus Wheeler, 1910. SE, EE), Georgia, Armenia, Kazakhstan, Mongolia, Russia: FE (PR). – Korean Peninsula, Japan (Hok, Korean Peninsula. Hon, Shi, Kyu). 202 D.A. Dubovikoff and Z.M. Yusupov

Lasius (Lasius) alienus (Förster, 1850) [Formica]. Lepisiota semenovi Ruzsky, 1905 [Acantholepis] Russia: EP (NW, C, E, S, NC, CR), UR, WS (OM, (Acantholepis frauenfeldi var. semenovi Ruzsky, TK, NS, KM, AL), ES (BR, ZB), FE (AM, KH, PR, 1905). Russia: EP (NC). – Europe (SE, EE), Cau- SA). – Europe, Caucasus, Turkey, Israel, Iran, Kaz- casus, Turkey, Iran, Central Asia, Kazakhstan. akhstan, Mongolia, Japan (Hok, Hon, Shi, Kyu), N NYLANDERIA Emery, 1906. Ty p e s p e c i e s For- America. mica vividula Nylander, 1846. Number of species: Lasius (Lasius) brunneus (Latreille, 1798) [Formica]. World – 110, Palaearctic – 9, Russia – 3. Russia: EP (C, E, S, NC, CR). – Europe (WE, NE, SE, EE), Caucasus, Turkey, Israel, Iran. Nylanderia colchica (Pisarski, 1960) [Paratrechina]. Lasius (Lasius) emarginatus (Olivier, 1792) [Formica] Russia: EP (NC). – Abkhazia, Georgia. (L. alienus var. pontica Stärcke, 1944). Russia: EP Nylanderia flavipes (Smith, 1874) [Tapinoma]. Rus- (C, S, NC, CR). – Europe (WE, NE, SE, EE), Cau- sia: EP (NC, introduced), FE (KU). – Abkhazia casus, Turkey, Israel, Iran. (introduced), China (NE, NC, CC, SW, WP, SE), Lasius (Lasius) hayashi Yamauchi et Hayashida, Korean Peninsula, Japan (Hok, Hon, Shi, Kyu, Ryu). 1970. Russia: FE (KU). – Korean Peninsula, Japan Nylanderia vividula (Nylander, 1846) [Formica]. (Hok, Hon, Shi, Kyu). Russia: EP (C, S, NC, CR) (introduced). – Tramp Lasius (Lasius) japonicus Santschi, 1941 (L. emar- species. Copsmopolitan. ginatus japonicus L niger var. Santschi, 1941; . : PLAGIOLEPIS Mayr, 1861. Ty p e s p e c i e s For- FE Kupianskaya, 1995). Russia: (AM, KH, PR, mica pygmaea Latreille, 1798. Number of species: SA, KU). – Korean Peninsula, Japan (Hok, Hon, World – 62, Palaearctic – 14, Russia – 3. Shi, Kyu). Lasius (Lasius) neglectus Van Loon, Boomsma et Plagiolepis karawajewi Radchenko, 1989. Russia: EP Andrasfalvy, 1990. Russia: EP (S, NC, CR). – Europe (CR). (WE, SE, EE), Turkey, Israel, Iran, Uzbekistan, Plagiolepis pallescens Forel, 1889. Russia: EP (NC). – Kyrgyzstan. Europe (SE, EE), N Africa, Caucasus, Turkey, Cen- Lasius (Lasius) niger (Linnaeus, 1758) [Formica] tral Asia. (L. niger var. alienonigra Forel, 1874). Russia: EP Plagiolepis taurica Santschi, 1920 (P. maura var. (N, NW, C, E, S, NC, CR), UR, WS (TM, OM, TK, taurica Santschi, 1920; P. barbara var. pyrenaica NS, KM, AL), ES (BR, ZB). – Europe (WE, NE, SE, Emery, 1921; P. satunini Karawajew, 1931). EP (C, EE), Kazakhstan, Mongolia, N America. E, S, NC, CR). – Europe (WE, SE, EE), Caucasus, Lasius (Lasius) koreanus Seifert, 1982. Russia: FE Iran, Central Asia, Kazakhstan. (KU). – Korean Peninsula. POLYERGUS Latreille, 1804. Ty p e s p e c i e s For- Lasius (Lasius) obscuratus Stitz, 1930 (L. brun- mica rufescens Latreille, 1798. Number of species: neus var. obscurata Stitz, 1930). Russia: EP World – 14, Palaearctic – 3, Russia – 3. (NC). – Tajikistan. Lasius (Lasius) paralienus Seifert, 1992. Russia: EP Polyergus nigerrimus Marikovsky, 1963. Russia: ES (C, S, NC, CR). – Europe (WE, NE, SE, EE), Turkey. (TU, BR), FE (KH). – Mongolia. Lasius (Lasius) platythorax Seifert, 1991. Russia: EP Polyergus rufescens (Latreille, 1798) [Formica]. Rus- (N, NW, C, E, S, NC, CR), UR, WS (TM, OM, TK, sia: EP (C, E, S, NC, CR), UR, WS (OM, AL). – NS, KM, AL). – Europe (WE, NE, SE, EE), Iran. Europe (WE, NE, SE, EE), Caucasus, Turkey, Kyr- Lasius (Lasius) psammophilus Seifert, 1992. Russia: gyzstan, Kazakhstan. EP (C, S, NC, CR). – Europe (WE, NE, SE, EE). Polyergus samurai Yano, 1911. Russia: FE (AM, PR). – Lasius (Lasius) turcicus Santschi, 1921. Russia: EP China (NE, NC, CC, SW, WP, SE), Korean Penin- (NC). – Europe (WE, SE, EE), Georgia, Turkey, sula, Japan (Hok, Hon, Shi, Kyu). Iran. PRENOLEPIS Mayr, 1861. Ty p e s p e c i e s Tapi­ noma nitens LEPISIOTA Santschi, 1926 (Acantholepis Mayr, Mayr, 1853. Number of species: World – 1861). T y p e s p e c i e s Plagiolepis rothneyi Forel, 13, Palaearctic – 1, Russia – 1. 1894. Number of species: World – 83, Palaearctic – Prenolepis nitens (Mayr, 1853) [Tapinoma]. Russia: EP 25, Russia – 1. (NC, CR). – Europe (WE, SE, EE), Georgia, Turkey. 69. Family Formicidae 203

PROFORMICA Ruzsky, 1902. Ty p e s p e c i e s For- 1928; A. sinensis: Kupianskaya, 1995). Russia: FE mica nasuta Nylander, 1856. Number of species: (KU). – China (CC, SW, WP, SE), Korean Penin- World – 25, Palaearctic – 25, Russia – 3. sula, Japan (Hok, Hon, Shi, Kyu). Aphaenogaster kurdica Ruzsky, 1905 (A. subterra- Proformica epinotalis Kuznetsov-Ugamsky, 1927. nea var. kurdica Ruzsky, 1905). Russia: EP (NC). – Russia: EP (C, E, S, NC, CR), UR. – Europe (EE), Georgia, Turkey. Iran, Turkmenistan, Kyrgyzstan, Kazakhstan. Aphaenogaster obsidiana (Mayr, 1861) [Atta]. Russia: Proformica mongolica (Emery, 1901) [Formica]. Rus- EP (NC). – Europe (SE), Georgia, Turkey. sia: ES (KS, TU, KR, IR, BR, ZB). – Kyrgyzstan, Aphaenogaster splendida (Roger, 1859) [Atta] (Apha- Kazakhstan, Mongolia, China (NE, NC, NW), enogaster splendida transcaucasica Karawajew, Korean Peninsula. 1926). Russia: EP (NC, CR). – Europe (SE), N Proformica ossetica Dubovikoff, 2005. Russia: EP Africa, Armenia, Azerbaijan, Turkey, Israel, Iran. (NC). Aphaenogaster subterranea (Latreille, 1798) [For- ROSSOMYRMEX Arnoldi, 1928. Ty p e s p e c i e s mica]. Russia: EP (NC, CR). – Europe (WE, SE, Rossomyrmex proformicarum Arnoldi, 1928. Num- EE), N Africa, Caucasus, Turkey. ber of species: World – 4, Palaearctic – 4, Russia – 1. Aphaenogaster subterraneoides Emery, 1881 (A. pal- lida var. subterraneoides Emery, 1881). Russia: EP Rossomyrmex proformicarum Arnoldi, 1928. Russia: (NC, CR). – Europe (WE, SE, EE), N Africa, Cau- EP (S, NC). – Kazakhstan. casus, Turkey.

Subfamily CARDIOCONDYLA Emery, 1869. Ty pe species Cardiocondyla elegans Emery, 1869. Number of Number of taxa: World – 142 genera, 6605 species, species: World – 70, Palaearctic – 31, Russia – 5. Palaearctic – 34/1000, Russia – 23/142. Cardiocondyla koshewnikovi Ruzsky, 1902. Russia: ANERGATES Forel, 1874. Ty p e s p e c i e s Myr- EP (S, NC). – Kyrgyzstan, Kazakhstan, Mongolia, mica atratula Schenck, 1852. The social parasites of China (NW). Tetramorium species. Number of species: World – 1, Cardiocondyla mauritanica Forel, 1890 (C. nuda Palaearctic – 1, Russia – 1. var. mauritanica Forel, 1890). Russia: EP (CR). – Europe (SE), N Africa, Afghanistan, Turkey, intro- Anergates atratulus (Schenck, 1852) [Myrmica]. duced into many countries. The social parasite of Tetramorium caespitum. Cardiocondyla sahlbergi Forel, 1913 (C. elegans var. Russia: EP (C, S, NC, CR). – Europe (WE, NE, sahlbergi Forel, 1913). Russia: EP (NC). – Europe SE, EE), Armenia, Turkey, Kazakhstan, N America (EE), N Africa, Caucasus, Turkey, Israel, Iran, (introduced). Uzbekistan. APHAENOGASTER Mayr, 1853. Type species Cardiocondyla stambuloffii Forel, 1892 (C. bogdanovi Aphaenogaster sardoa Mayr, 1853. Number of spe- Ruzsky, 1905; C. stambuloffii taurica Karawajew, cies: World – 187, Palaearctic – 78, Russia – 8. 1927). Russia: EP (C, E, S, NC, CR), UR. – Europe (SE, EE), Armenia, Turkey, Iran. Aphaenogaster famelica (Smith, 1874) [Ischno- Cardiocondyla ulianini Emery, 1889 (C. elegans myrmex] (A. famelica frontosa Wheeler, 1928; Emery, 1869, auct.; C. elegans var. ulianini Emery, A. famelica ruida Wheeler, 1928; A. vapida Wheel- 1889; C. elegans schkaffi Arnoldi, 1928). Russia: EP er, 1928). Russia: FE (KU). – China (CC, SW, WP, (E, S, NC, CR). – Europe (EE), Caucasus, Saudi SE), Korean Peninsula, Japan (Hok, Hon, Shi, Kyu). Arabia, Iran, Afghanistan, Kyrgyzstan, Kazakh- Aphaenogaster gibbosa (Latreille, 1798) [Formica] stan, Mongolia. (A. striola (Roger, 1859); A. gibbosa muschtaidica Emery, 1908). Russia: EP (NC). – Europe (WE, CHALEPOXENUS Menozzi, 1923 (Leonomyrma SE, EE), N Africa, Caucasus, Turkey, Israel, Iraq, Arnoldi, 1968). Ty p e s p e c i e s Chalepoxenus Iran, Turkmenistan. gribodoi Menozzi, 1923. The social parasites of Tem- Aphaenogaster japonica Forel, 1911 (A. schmitzi japo- nothorax species. Number of species: World – 9, nica Forel, 1911; A. smythiesii sinensis Wheeler, Palaearctic – 9, Russia – 1. 204 D.A. Dubovikoff and Z.M. Yusupov

Chalepoxenus tauricus Radchenko, 1989. The social EP (N, NW, C, E, NC), UR, WS (TM, OM, TK, parasite of Temnothorax tauricus. Russia: EP (CR). NS, KM, AL), ES (KS, TU, KR, IR, BR, YA, ZB), FE (AM, KH, PR, SA, KU, KA, MG, CH). – Europe CREMATOGASTER Lund, 1831. T y p e s p e c i e s (WE, NE, SE, EE), Caucasus, Turkey, Central Asia, Formica scutellaris Olivier, 1792. Number of species: Kazakhstan, Mongolia, China (NE, NC), Korean World – 506, Palaearctic – 36, Russia – 3. Peninsula, Japan (Hok, Hon, Shi, Kyu). Crematogaster schmidti (Mayr, 1853) [Acrocoelia]. Leptothorax gredleri Mayr, 1855. Russia: EP (NC). – Russia: EP (CR, NC). – Europe (WE, SE, EE), Cau- Europe (WE, NE, SE, EE). casus, Turkey, Iran, Turkmenistan. Leptothorax muscorum (Nylander, 1846) [Myrmica] Crematogaster sordidula (Nylander, 1849) [Myrmica] (L. muscorum var. fagi Ruzsky, 1905; L. muscorum (C. sordidula var. caspica Emery, 1912). Russia: EP var. betulae Ruzsky, 1916). Russia: EP (N, NW, C, (NC). – Europe (WE, SE, EE), Caucasus, Turkey, E, NC, CR), UR, WS (TM, OM, TK, NS, KM, AL), Israel, Iran, Turkmenistan. ES (KS, TU, KR, IR, BR, YA, ZB), FE (AM, KH, Crematogaster subdentata Mayr, 1877. Russia: EP PR). – Europe (WE, NE, SE, EE), Caucasus, Turkey, (S introduced, NC, CR introduced). – Caucasus, Mongolia, China (NE, NC). Turkey, Iran, Afghanistan, Central Asia, Kazakh- Leptothorax oceanicus (Kuznetsov-Ugamsky, stan, Mongolia. 1928) [Mychothorax] (L. muscorum oceanicum: Kupianskaya, 1990). Russia: FE (AM, KH, PR). – FORMICOXENUS Mayr, 1855. Ty p e s p e c i e s China (NE), Korean Peninsula. Myrmica nitidula Nylander, 1846. The social parasite Leptothorax pacis (Kutter, 1945) [Doronomyrmex]. species, in nest of Formica s. str. (xenobiotic). Num- Russia: EP (N), WS (KM). – Europe (WE, NE, ber of species: World – 7, Palaearctic – 2, Russia – 2. SE). Formicoxenus nitidulus (Nylander, 1846) [Myrmica] Leptothorax scamni Ruzsky, 1905 (L. muscorum var. (F. laeviuscula Förster, 1850). Russia: EP (NW, C, scamni Ruzsky, 1905). Russia: EP (NC). – Caucasus, E, NC), UR, WS (TM, OM, TK, NS, KM, AL), ES Turkey. (IR, YA, ZB), FE (KH, PR). – Europe (WE, NE, MANICA Jurine, 1807. Ty p e s p e c i e s Formica SE, EE), Turkey. rubida Latreille, 1802. Number of species: World – Formicoxenus sibiricus (Forel, 1899) [Leptothorax] 6, Palaearctic – 2, Russia – 1. (F. orientalis Dlussky, 1963). Russia: ES (BR, ZB), FE (PR). Manica rubida (Latreille, 1802) [Formica]. Russia: EP (NC, CR). – Europe (WE, SE, EE), Georgia, HARPAGOXENUS Forel, 1893 (Tomognathus Armenia, Turkey. Mayr, 1861). T y p e s p e c i e s Myrmica sublaevis Nylander, 1849. The social parasites of Leptothorax MESSOR Forel, 1890. Ty p e s p e c i e s Formica bar- species (dulosis). Number of species: World – 3, bara Linnaeus, 1767. Number of species: World – Palaearctic – 1, Russia – 1. 106, Palaearctic – 69, Russia – 4. Harpagoxenus sublaevis (Nylander, 1852) [Myrmica] Messor aciculatus (Smith, 1874) [Aphaenogaster]. (M. hirtula Nylander, 1849). The social parasite of Russia: FE (PR). – Mongolia, China (NE, NC, Leptothorax acervorum. Russia: EP (N, NW, C, E, NW), Korean Peninsula, Japan (Hok, Hon, Shi, NC), UR, ES (BR, ZB), FE (PR). – Europe (WE, Kyu). NE, SE, EE). Messor denticulatus Kuznetzov-Ugamsky, 1927 (M. barbarus meridionalis var. denticulatus Kuz- LEPTOTHORAX Mayr, 1855. T y p e s p e c i e s For- netzov-Ugamsky, 1927). Russia: EP (S, NC). – Cau- mica acervorum Fabricius, 1793. Number of species: casus, Iran, Central Asia, Kazakhstan. World – 19, Palaearctic – 9, Russia – 6. Messor kasakorum Arnoldi, 1970 (M. kasakorum nadez- Leptothorax acervorum (Fabricius, 1793) [Formica] hdae Arnoldi, 1970). Russia: EP (S). – Kazakhstan. (L. acervorum var. superus Ruzsky, 1905; L. acer- Messor structor (Latreille, 1798) [Formica] (Myr- vorum var. nigrescens Ruzsky, 1905; L. acervorum mica mutica Nylander, 1849; Messor structor var. kamtschaticus Ruzsky, 1920; Mychothorax acer- clivorum Ruzsky, 1905; M. tataricus Ruzsky, 1905; vorum orientalis Kuznetzov-Ugamsky, 1928). Russia: M. structor turanicus Kuznetzov-Ugamsky, 1927; 69. Family Formicidae 205

M. rufitarsis jakowlevi Arnoldi, 1977; M. clivorum Myrmica bergi Ruzsky, 1902 (M. rugosa var. kirgi- auct., M. rufitarsis auct.). Russia: EP (C, E, S, NC, sica Ruzsky, 1903; M. bergi var. barchanica Ruzsky, CR), UR. – Europe (WE, SE, EE), Caucasus, Israel, 1905; M. bergi var. fortior Crawley, 1920; M. kamy- Iran, Afghanistan, Central Asia, Kazakhstan. schiensis Arnoldi, 1934; M. bergi persiana Weber, EP WS MONOMORIUM Mayr, 1855. Ty p e s p e c i e s 1947). Russia: (S, NC, CR), (AL). – Europe Monomorium monomorium Bolton, 1987, replace- (EE), Azerbaijan, Iran, Central Asia. Myrmica commarginata ES ment name for Monomorium minutum Mayr, 1855. Ruzsky, 1905. Russia: Number of species: World – 358, Palaearctic – 76, (KR, BR, ZB). – Mongolia. Russia – 2. Myrmica deplanata Emery, 1921 (M. lobicornis var. deplanata Emery, 1921; M. moravica Soudek, 1922; Monomorium barbatulum Mayr, 1877. Russia: EP M. lobicornis var. plana Karawajew, 1927; M. mora- (S). – Israel, Saudi Arabia, Iran, Oman, United Arab vica var. dagmarae Sadil, 1939). Russia: EP (C, S, Emirates, Afghanistan, Turkmenistan, Kyrgyzstan, NC, CR), WS (AL). – Europe (WE, SE, EE), Cau- Kazakhstan. casus, Iran, Central Asia. Monomorium pharaonis (Linnaeus, 1758) [Formica]. Myrmica displicentia Bolton, 1995 (M. bicolor Kupian­ Synanthropic species with cosmopolitan distri- skaya, 1986). Russia: FE (MG, KA). bution, in Russia only in heated rooms. Russia, Myrmica divergens Karawajew, 1931 (M. bergi var. introduced. – Cosmopolitan. divergens Karawajew, 1931). Russia: WS (AL), ES MYRMECINA Curtis, 1829. Ty p e s p e c i e s Myr- (KR, IR, BR, YA, ZB). – Mongolia. mecina latreillii Curtis, 1829. Number of species: Myrmica eidmanni Menozzi, 1930 (M. lobicornis litto- World – 53, Palaearctic – 7, Russia – 2. ralis Weber, 1948). Russia: WS (AL), ES (KS, TU, KR, IR, BR, ZB), FE (AM, KH, PR). – Mongolia, Myrmecina graminicola (Latreille, 1802) [Formica] China (NE), Korean Peninsula. (Myrmica striatula Nylander, 1849). Russia: EP Myrmica elbrusi Radchenko et Yusupov, 2012. Russia: (C, E, S, NC, CR). – Europe (WE, NE, SE, EE), N EP (NC). Africa, Caucasus, Turkey, Israel. Myrmica excelsa Kupianskaya, 1990 (M. sinica Wu Myrmecina nipponica Wheeler, 1906 (M. grami- et Wang, 1995; M. cadusa B. Kim, Park et J. Kim, nicola nipponica Wheeler, 1906; M. graminicola: 1997). Russia: FE (PR). – China (NE), Korean Pen- Kupianskaya, 1995). Russia: FE (AM, KH, PR, KU). – China (SE), Korean Peninsula, Japan (Hok, insula, Japan (Hok, Hon). Myrmica forcipata ES Hon, Shi, Kyu). Karawajew, 1931. Russia: (KR, IR, BR, YA, ZB). – Mongolia. MYRMICA Latreille, 1804. Ty p e s p e c i e s Formica Myrmica gallienii Bondroit, 1920 (M. rugulosa limanica rubra Linnaeus, 1758. Number of species: World – Arnoldi, 1934; M. jacobsoni Kutter, 1963; M. lima- 188, Palaearctic – 75, Russia – 44. nica obensis Arnoldi, 1970). Russia: EP (NW, C, E, Myrmica ademonia Bolton, 1995 (replacement name S, NC, CR), UR, WS (Radchenko, Elmes, 2010). – for M. aspersa Kupianskaya, 1990). Russia: FE (KH, Europe (WE, NE, EE). PR, KU). – Korean Peninsula. Myrmica hellenica Finzi, 1926 (M. rugulosa var. helle- Myrmica angulinodis Ruzsky, 1905 (M. scabrinodis nica Finzi, 1926; M. rugulosa var. rugulososcabrinodis angulinodis Ruzsky, 1905; M. incurvata Colling- Karawajew, 1929; M. rugulosa caucasica Arnoldi, wood, 1976; M. baikalensis Dmitrienko, 1979; 1934). Russia: EP (NC, CR). – Europe (SE, EE), M. kasczenkoi: Kupianskaya, 1979; M. forcipata: Caucasus, Turkey. Kupianskaya, 1990). Russia: WS (NS, KM, AL), ES Myrmica hirsuta Elmes, 1978. Russia: EP (C). – (KS, TU, KR, IR, BR, YA, ZB), FE (AM, KH, PR, Europe (WE, NE, SE, EE). MG). – Mongolia, China (NE), Korean Peninsula. Myrmica jennyae Elmes, Radchenko et Aktac, 2002. Myrmica arnoldii Dlussky, 1963. Russia: ES (TU, KR, Russia: EP (NC). – Turkey. IR, BR, ZB). – Mongolia. Myrmica kamtschatica Kupianskaya, 1986 (M. abori- Myrmica bakurianica Arnoldi, 1970 (M. pilosi- genica Zhigulskaya, 1991). Russia: WS (KM, AL), scapus bakurianica Arnoldi, 1970). Russia: EP ES (Radchenko, Elmes, 2010), FE (AM, KH, PR, (NC). – Georgia. MG, KA). – Mongolia, Korean Peninsula. 206 D.A. Dubovikoff and Z.M. Yusupov

Myrmica karavajevi (Arnoldi, 1930) [Symbiomyrma] Myrmica rugulosa Nylander, 1849. Russia: EP (N W, (Sifolinia karavajevi Arnoldi, 1930). Russia: EP C, E, NC, CR), UR. – Europe (WE, NE, SE, EE), (NW, C). – Europe (WE, NE, SE, EE). Caucasus. Myrmica kasczenkoi Ruzsky, 1905 (M. scabrinodis Myrmica sabuleti Meinert, 1861. Russia: EP (C, E, S, kasczenkoi Ruzsky, 1905). Russia: WS (KM, AL), NC, CR), UR, WS (Radchenko, 2016). – Europe ES (KS, TU, KR, IR, BR, ZB). – Mongolia. (WE, NE, SE, EE), Caucasus, Turkey. Myrmica koreana Elmes, Radchenko et Kim, 2001. Myrmica salina Ruzsky, 1905 (M. scabrinodis var. Russia: WS (KM, AL), ES (KS, TU, KR, IR, BR, salina Ruzsky, 1905). Russia: UR, WS (Radchenko, ZB), FE (AM, KH, PR). – Kazakhstan, Mongolia, Elmes, 2010). – Kazakhstan. China (NE), Korean Peninsula. Myrmica scabrinodis Nylander, 1846 (M. scabrinodis Myrmica kotokui Forel, 1911 (M. rubra kotokui var. rugulosoides Forel, 1915; M. scabrinodis var. Forel, 1911; M. ruginodis orientalis Karawajew, ahngeri Karawajew, 1926). Russia: EP (N, NW, C, E, 1926; M. orientalis: Kupianskaya, 1990). Russia: S, NC, CR), UR, WS (Radchenko, 2016), ES (KR, FE (PR). – China (NE), Korean Peninsula, Japan BR, ZB). – Europe (WE, NE, SE, EE), Caucasus, (Hok, Hon). Turkey, Kazakhstan. Myrmica kozakorum Radchenko et Elmes, 2010. Rus- Myrmica schencki Viereck, 1903 (M. rubra var. sia: EP (S, NC, CR). – Europe (EE). schencki Viereck, 1903; M. schenckioides Boer, EP Myrmica kurokii Forel, 1907 (M. rubra kurokii Forel, Noordijk, 2005). Russia: (NW, C, E, S, NC, CR), UR WS ES 1907). Russia: FE (PR). – China (NE, NC), Korean , (Radchenko, 2016), (KR). – Europe Peninsula, Japan (Hok, Hon). (WE, NE, SE, EE), Caucasus, Turkey, Central Asia. Myrmica slovaca M curvithorax Myrmica lobicornis Nylander, 1846 (M. lobicornis Sadil, 1952 ( . Bondroit, 1920). Russia: EP (C, S, NC, CR), UR, WS (Rad- alpestris Arnoldi, 1934). Russia: EP (N, NW, C, E, chenko, 2016). – Europe (WE, SE, EE), Caucasus, NC), UR, WS (Radchenko, 2016), ES (KS, TU, Turkey, Kazakhstan. KR, IR, BR, YA, ZB). – Europe (WE, NE, SE, EE), Myrmica specioides Bondroit, 1918 (M. scabrinodis Caucasus, Turkey, Kazakhstan, Mongolia. var. sancta Karawajew, 1926; M. sancta tshuliensis Myrmica lonae Finzi, 1926 (M. scabrinodis lonae Finzi, Arnoldi, 1976). Russia: EP (C, S, NC, CR), UR, WS 1926). Russia: EP (N, NW, C, E), UR, WS (Rad- (Radchenko, Elmes, 2010). – Europe (WE, NE, ES chenko, 2016), (BR). – Europe (WE, NE, SE, SE, EE), Caucasus, Turkey, Iran, Turkmenistan, EE), Turkey, Kazakhstan. Kazakhstan. Myrmica luteola FE Kupianskaya, 1990. Russia: (PR). – Myrmica stangeana Ruzsky, 1902 (M. bergi stangeana China (NE), Korean Peninsula, Japan (Hok, Hon). Ruzsky, 1902; M. stangeana medvedevi Arnoldi, Myrmica pisarskii Radchenko, 1994. Russia: WS (KM, 1970). Russia: EP (E, S), UR, WS (AL). – Europe AL), ES (KS, TU, KR, IR, BR, ZB). – Mongolia. (EE), Kazakhstan. Myrmica ravasinii Finzi, 1923. Russia: EP (NC). – Myrmica sulcinodis Nylander, 1846 (M. sulcinodis Europe (SE), Georgia, Armenia, Turkey. var. nigripes Ruzsky, 1895; M. myrmecophila Was- Myrmica rubra (Linnaeus, 1758) [Formica] (M. lae- mann, 1910; M. sulcinodis var. sulcinodoscabrinodis vinodis Nylander, 1846; M. longiscapus Curtis, 1854; Forel, 1915). Russia: EP (N, NW, C, NC, CR), UR, M. laevinodis var. europea Finzi, 1926; M. laevinodis WS (Radchenko, 2016), ES (Radchenko, 2016), var. bruesi Weber, 1947; M. microrubra Seifert, FE (AM, KH, PR, SA, MG). – Europe (WE, NE, 1993). Russia: EP (N, NW, C, E, S, NC, CR), UR, SE, EE), Caucasus, Turkey, Kazakhstan, Mongolia, WS (TM, OM, TK, NS, KM, AL), ES (KS, TU, KR, Korean Peninsula. IR, BR, YA, ZB). – Europe (WE, NE, SE, EE), Cau- Myrmica transsibirica Radchenko, 1994 (M. taediosa casus, Turkey, Central Asia. Bolton, 1995). Russia: WS (KM, AL), ES (Rad- Nylander, 1846 (M. diluta Ny- chenko, Elmes, 2010), FE (AM, KH, PR, SA). – lander, 1849). Russia: EP (N, NW, C, E, S, NC, China (NE), Korean Peninsula, Japan (Hok). CR) UR, WS (TM, OM, TK, NS, KM, AL), ES Myrmica tschekanovskii Radchenko, 1994. Russia: (KS, TU, KR, IR, BR, YA, ZB), FE (AM, KH, PR, ES (KR, YA). SA, KU). – Europe (WE, NE, SE, EE), Caucasus, Myrmica vandeli Bondroit, 1920 (M. scabrinodis var. Turkey, Kazakhstan, Mongolia, China (NE, NC), vandeli Bondroit, 1920). Russia: EP (NC). – Europe Korean Peninsula, Japan (Hok, Hon, Shi, Kyu). (WE, NE, SE, EE). 69. Family Formicidae 207

Myrmica zojae Radchenko, 1994. Russia: UR, WS (AL). STENAMMA Westwood, 1839. Ty p e s p e c i e s Stenamma westwoodi Westwood, 1839. Number of MYRMOXENUS Ruzsky, 1902 (Epimyrma Emery, species: World – 84, Palaearctic – 14, Russia – 6. 1915). Ty p e s p e c i e s Myrmoxenus gordiagini Ruzsky, 1902. The social parasites of Temnothorax Stenamma debile (Förster, 1850) [Myrmica] (S. west­ species. Number of species: World – 12, Palaearc- woodii Westwood, 1839, auct.; S. golosejevi Kara- tic – 12, Russia – 2. wajew, 1926; S. ucrainicum Arnoldi, 1928). Russia: EP (C, E, S, NC, CR). – Europe (WE, NE, SE, EE), Myrmoxenus gordiagini Ruzsky, 1902 (Myrmetaerus Armenia, Turkey. microcellatus Soudek, 1925). Russia: WS (TK, NS, Stenamma georgii Arnoldi, 1975. Russia: EP (NC). – AL). – Europe (SE, EE), Kazakhstan. Abkhazia, Georgia. Myrmoxenus ravouxi (André, 1896) [Formicoxenus] Stenamma kurilense FE (Epimyrma goesswaldi Menozzi, 1930). Russia: EP Arnoldi, 1975. Russia: (KU). Stenamma lippulum (NC, CR). – Europe (WE, SE, EE). (Nylander, 1849) [Myrmica] (S. westwoodi var. hirtula Emery, 1898; S. caucasi- PHEIDOLE Westwood, 1839. Ty p e s p e c i e s Atta cum Arnoldi, 1975). Russia: EP (NC). – Abkhazia, providens Sykes, 1835. Number of species: World – Georgia, Azerbaijan. 1004, Palaearctic – 19, Russia – 2. Stenamma striatulum Emery, 1895 (S. westwoodi var. striatulum S westwoodi tscherkes- Pheidole fervida Smith, 1874. Russia: FE (PR, KU). – Emery, 1895; . var. sicum EP China (CC, SW, WP, SE), Korean Peninsula, Japan Arnoldi, 1928). Russia: (NC). – Europe (Hok, Hon, Shi, Kyu). (SE, EE), Turkey.

Pheidole pallidula (Nylander, 1849) [Myrmica] (Ph. Stenamma ussuriense Arnoldi, 1975. Russia: FE pallidula var. arenarum Ruzsky, 1905; Ph. pallidula (PR). – Korean Peninsula. koshewnikovi Ruzsky, 1905). Russia: EP (S, NC). – STRONGYLOGNATHUS Mayr, 1853. Ty p e Europe (WE, SE, EE), N Africa, Caucasus, Turkey, species Eciton testaceum Schenck, 1852. The Israel, Iran, Afghanistan, Central Asia, Kazakhstan. social parasites of Tetramorium species (dulosis). SOLENOPSIS Westwood, 1840 (Diplorhoptrum Number of species: World – 24, Palaearctic – 24, Mayr, 1855). T y p e s p e c i e s Solenopsis mandibu- Russia – 5. laris Westwood, 1840. Number of species: World – Strongylognathus arnoldii Radchenko, 1985. Russia: 195, Palaearctic – 33, Russia – 5. EP (CR). Solenopsis deserticola Ruzsky, 1905 [Diplorhoptrum] Strongylognathus christophi Emery, 1889 (S. huberi (D. deserticola auct.). Russia: EP (S, NC). – Cau- ruzskyi Emery, 1909). Russia: EP (E, S, NC, CR), casus, Central Asia, Kazakhstan. UR. – Caucasus, Kazakhstan. Solenopsis fugax (Latreille, 1798) [Formica] (Myr- Strongylognathus karawajewi Pisarski, 1966. Russia: mica flavidula Nylander, 1849; Solenopsis fugax EP (S, NC, CR). – Caucasus, Turkmenistan. orientalis Ruzsky, 1905; S. orbula var. latroides Strongylognathus rehbinderi Forel, 1904. Russia: EP Ruzsky, 1905; S. (Diplorhoptrum) fugax var. pon- (S, NC). – Abkhazia. tica Santschi, 1934; Diplorhoptrum fugax auct.). Strongylognathus testaceus (Schenck, 1852) [Eci- Russia: EP (C, E, S, NC, CR). – Europe (WE, NE, ton] (Myrmus emarginatus Schenck, 1853; Strongy- SE, EE), N Africa, Caucasus, Turkey, Israel, Iran, lognathus diveri Donisthorpe, 1936). Russia: EP (C, Afghanistan, Central Asia, Kazakhstan. E, S, NC, CR), UR, WS (NS, AL). – Europe (WE, Solenopsis ilinei Santschi, 1936 (S. orbula var. oculata NE, SE, EE), Caucasus, Turkey, Kazakhstan. Karawajew, 1926; S. (Diplorhoptrum) orbula var. STRUMIGENYS Smith, 1860 (Pyramica Roger, 1862; ilinei Santschi, 1936). Russia: EP (CR). – Azerbaijan. Epitritus Emery, 1869).T y p e s p e c i e s Strumige- Solenopsis japonica S fugax Wheeler, 1928 ( . var. nys mandibularis Smith, 1860. Number of species: japonica Diplorhoptrum japonica Wheeler, 1928; : World – 837, Palaearctic – 19, Russia – 1. Kupianskaya, 1995). Russia: FE (KU). – Korean Peninsula, Japan (Hok, Hon, Shi, Kyu). Strumigenys argiola (Emery, 1869) [Epitritus] (Pyra- Solenopsis juliae (Arakelian, 1991) [Diplorhoptrum]. mica argiola auct.). Russia: EP (NC). – Europe Russia: EP (NC). – Armenia. (WE, SE), N Africa, Caucasus, Turkey, Israel. 208 D.A. Dubovikoff and Z.M. Yusupov

TELEUTOMYRMEX Kutter, 1950. Ty pe species Temnothorax knipovitschi (Karawajew, 1916) [Lepto- Teleutomyrmex schneideri Kutter, 1950. Number thorax] (L. tuberum knipovitshi Karawajew, 1916; of species: World – 2, Palaearctic – 2, Russia – 1. L. nikitae Arnoldi, 1977; L. junipereti Arnoldi, 1977). Russia: EP (E, S, NC, CR), UR. – Europe (EE), Teleutomyrmex schneideri Kutter, 1950. Russia: EP Caucasus, Kazakhstan. (S). – Europe (WE), Turkmenistan. Temnothorax kurilensis (Radchenko, 1994) [Lepto- TEMNOTHORAX Mayr, 1861. Ty p e s p e c i e s thorax]. Russia: FE (KU). Myrmica recedens Nylander, 1856. Number of spe- Temnothorax mongolicus (Pisarski, 1969) [Lepto- cies: World – 390, Palaearctic – 200, Russia – 29. thorax] (L. serviculus mongolicus Pisarski, 1969). Russia: ES (ZB), FE (AM). – Mongolia, Korean Temnothorax affinis (Mayr, 1855) [Leptothorax]. Peninsula. Russia: EP (C, E, S, NC, CR). – Europe (WE, NE, Temnothorax nassonovi (Ruzsky, 1895) [Lepto- SE, EE), Caucasus, Turkey. thorax] (L. nassonovi var. subnudus Ruzsky, 1905; Temnothorax alinae (Radchenko, 1994) [Lepto- L. galeatus Wheeler, 1927; L. nassonovi firssovi FE thorax]. Russia: (PR). Kuznetzov-Ugamsky, 1928). Russia: EP (E, S), UR, Temnothorax alpinus (Ruzsky, 1902) [Leptothorax] WS (OM, TK, NS, AL), ES (BR, ZB), FE (AM, KH, L werneri EP ( . (Radchenko, 1994)). Russia: PR, SA). – Europe (EE), Kazakhstan, Mongolia, (NC). – Georgia. China (NE, NC), Korean Peninsula. Temnothorax arnoldii Radchenko et Fedoseeva, 2015. Temnothorax parvulus (Schenck, 1852) [Myrmica]. EP Russia: (NC). Russia: EP (NC). – Europe (WE, SE, EE), N Africa, Temnothorax brauneri (Ruzsky, 1905) [Leptothorax] Caucasus, Turkey, Iran, Turkmenistan. L serviculus brauneri ( . var. Ruzsky, 1905). Russia: Temnothorax recedens (Nylander, 1856) [Myrmica]. EP (NC). – Georgia. Russia: EP (S, NC, CR). – Europe (SE), N Africa, Temnothorax clypeatus (Mayr, 1853) [Myrmica]. Caucasus, Turkey, Iran, Turkmenistan. Russia: EP (CR). – Europe (WE, SE, EE). Temnothorax satunini (Ruzsky, 1902) [Leptothorax]. Temnothorax congruus (Smith, 1874) [Leptothorax]. Russia: EP (NC). – Caucasus, Turkey. Russia: FE (PR, KU). – China (NE), Korean Pen- Temnothorax semenovi (Ruzsky, 1903) [Leptothorax]. insula, Japan (Hok, Hon, Shi, Kyu). Russia: EP (S, NC, CR). – Central Asia. Temnothorax corticalis (Schenck, 1852) [Myrmica]. Temnothorax serviculus (Ruzsky, 1902) [Lepto- Russia: EP (S, NC, CR). – Europe (WE, NE, SE, thorax]. Russia: EP (E, S), UR, WS (Radchenko, EE), N Africa, Caucasus. 1996b), ES (Radchenko, 1996b). – Kazakhstan, Temnothorax crasecundus Seifert et Csősz, 2015. Mongolia. Russia: EP (NC, CR). – Europe (SE, EE), Caucasus, Temnothorax tauricus (Ruzsky, 1902) [Lepto- Turkey. thorax] (L. unifasciatus var. tauricus Ruzsky, 1902; Temnothorax crassispinus (Karawajew, 1926) [Lepto- L. pithyusae Arnoldi, 1977; L. pithyusae babuganicus thorax] (L. nylanderi var. crassispina Karawajew, Arnoldi, 1977). Russia: EP (NC, CR). – Europe (EE). 1926; L. nylanderi slavonicus Seifert, 1995). Russia: Temnothorax tembotovi Radchenko et Yusupov, 2015. EP (C, E, S). – Europe (WE, SE, EE). Russia: EP (NC). Temnothorax discoloratus (Arnoldi, 1977) [Lepto- Temnothorax tuberum (Fabricius, 1775) [Formica] thorax]. Russia: EP (NC). (F. tuberosa Latreille, 1802; Leptothorax tuberum Temnothorax dlusskyi Radchenko, Yusupov et Fedo- stipaceus Ruzsky, 1905; L. tuberum var. nigricephala seeva, 2015. Russia: EP (NC). Karawajew, 1930; L. tuberum alaicus Tarbinsky, Temnothorax interruptus (Schenck, 1852) [Myrmica]. 1976; L. pratostepposus Arnoldi, 1977; L. tuberum Russia: EP (NC). – Europe (WE, NE, SE, EE). acutinodis Arnoldi, 1977; L. tuberum ciscaucasicus Temnothorax jailensis (Arnoldi, 1977) [Leptothorax]. Arnoldi, 1977). Russia: EP (N, NW, C, E, S, NC, Russia: EP (CR). CR), UR, WS (NS, AL). – Europe (WE, NE, SE, Temnothorax kaszabi (Pisarski, 1969) [Leptothorax] EE), Caucasus, Turkey, Central Asia. (L. tuberum sachalinensis Kupianskaya, 1990). Rus- Temnothorax unifasciatus (Latreille, 1798) [Formica] sia: WS (AL), ES (TU, YA, ZB), FE (SA). – Mon- (Leptothorax tuberum kirillovi Ruzsky, 1905; L. tube- golia, Korean Peninsula. rum var. salina Karawajew, 1937; L. unifasciatus 69. Family Formicidae 209

ucrainicus Arnoldi, 1977). Russia: EP (C, E, S, NC, TRICHOMYRMEX Mayr, 1865. Ty p e s p e c i e s CR), UR. – Europe (WE, NE, SE, EE), N Africa, Trichomyrmex rogeri Mayr, 1865. Number of spe- Caucasus, Turkey, Iran, Turkmenistan. cies: World – 24, Palaearctic – 10, Russia – 1. Temnothorax volgensis (Ruzsky, 1905) [Leptothorax] Trichomyrmex perplexus (Radchenko, 1997) [Mono- (L. nassonovi volgensis Ruzsky, 1905). Russia: EP morium]. Russia: EP (NC). – Europe (SE), Cau- (C, E, S), UR. – Europe (EE), Kazakhstan. casus, Turkey, United Arab Emirates, Iran. TETRAMORIUM Mayr, 1855. Ty p e s p e c i e s Formica caespitum Linnaeus, 1758. Number of spe- Subfamily cies: World – 564, Palaearctic – 120, Russia – 10. Tetramorium caespitum (Linnaeus, 1758) [Formica]. Number of taxa: World – 142 genera, 6605 species, Russia: EP (N, NW, C, E, S, NC, CR), UR, WS Palaearctic – 12/33, Russia – 4/7. (OM, TK, NS, AL), ES (KS, TU, KR, IR, BR, ZB). – BRACHYPONERA Emery, 1900. T y p e s p e c i e s Europe (WE, NE, SE, EE), N Africa, Caucasus, Euponera (Brachyponera) croceicornis Emery, 1900. Turkey, Syria, Lebanon, Central Asia. Number of species: World – 19, Palaearctic – 2, Tetramorium chefketi Forel, 1911 (T. caespitum var. Russia – 1. chefketi Forel, 1911; T. caespitum var. sarkissiani Forel, 1911; T. caespitum st. turcomanicum Santschi, Brachyponera chinensis (Emery, 1895) [Ponera] 1921; T. taurocaucasicum Arnoldi, 1968). Russia: (P. nigrita chinensis Emery, 1895). Russia: EP (NC EP (S, NC), UR, WS (AL). – Europe (SE, EE), introduced), ?FE (PR). – Abkhazia (introduced), Caucasus, Turkey, Iran, Central Asia, Kazakhstan. Georgia (introduced), China (CC, SW, WP, SE), Tetramorium densopilosum Radchenko et Arakel- Korean Peninsula, Japan (Hon, Shi, Kyu, Ryu). ian, 1990. Russia: EP (S, NC). – Georgia, Armenia, Emery, 1893. Ty p e s p e c i e s Turkey. Cryptopone testacea Emery, 1893. Number of spe- Tetramorium ferox Ruzsky, 1903 (T. caespitum var. cies: World – 24, Palaearctic – 1, Russia – 1. ferox Ruzsky, 1903; T. caespitum st. ferox var. per- spicax Santschi, 1921; T. confinis Radchenko et Cryptopone ochracea (Mayr, 1855) [Ponera]. Rus- Arakelian, 1990; T. perspicax Radchenko, 1992). sia: EP (NC, CR). – Europe (WE, SE, EE), Saudi Russia: EP (S, NC, CR). – Europe (WE, SE, EE), Arabia. Caucasus, Turkey. HYPOPONERA Santschi, 1938. Ty p e s p e c i e s Tetramorium feroxoides Dlussky et Zabelin, 1985 Ponera abeillei André, 1881. Number of species: (T. bursakovi Radchenko, 1992). Russia: EP (S, World – 152, Palaearctic – 9, Russia – 2. NC). – Caucasus, Turkey, Iran, Central Asia, Kazakhstan. Hypoponera eduardi (Forel, 1894) [Ponera]. Russia: Tetramorium hungaricum Röszler, 1935 (T. caespitum EP (NC). – Europe (WE, SE, EE), N Africa, Turkey, hungarica Röszler, 1935). Russia: EP (S, CR). – Israel, Saudi Arabia, Iran, United Arab Emirates. Europe (WE, EE). Hypoponera punctatissima (Roger, 1859) [Po- Tetramorium impurum (Förster, 1850) [Myrmica] nera]. Tramp species. Russia, in heated (T. caespitum var. penninum Santschi, 1927; T. staer­ buildings. – Cosmopolitan. ckei EP Kratochvíl, 1944). Russia: (S, NC). – Europe PONERA Latreille, 1804. Ty p e s p e c i e s For- (WE, NE, SE, EE). mica coarctata Latreille, 1802. Number of species: Tetramorium inerme Mayr, 1877 (T. caespitum var. World – 57, Palaearctic – 5, Russia – 3. inermis Mayr, 1877). Russia: EP (S, NC). – Iran, Central Asia, Kazakhstan. Ponera coarctata (Latreille, 1802) [Formica] (P. coarc- Tetramorium moravicum Novák et Sadil, 1941 tata var. lucida Emery, 1898; P. coarctata coarctata (T. rhenanum Schulz, 1996). Russia: EP (S, NC, colchica Arnoldi, 1932; P. taurica Arnoldi, 1932). CR). – Europe (WE, SE, EE), Caucasus, Turkey. Russia: EP (C, E, S, NC, CR), UR. – Europe (WE, Tetramorium tsushimae Emery, 1925 (T. caespitum NE, SE, EE), N Africa, Caucasus, Turkey, Israel, jacoti Wheeler, 1927). Russia: FE (AM, KH, PR, Iran, Kyrgyzstan, Kazakhstan. KU). – China (NE), Korean Peninsula, Japan (Hok, Ponera japonica Wheeler, 1906 (P. japonica var. cro- Hon, Shi, Tsushima, Kyu, Yakushima). cea Santschi, 1941). Russia: FE (PR, KU). – China 210 D.A. Dubovikoff and Z.M. Yusupov

(SW, WP, SE), Korean Peninsula, Japan (Hok, Hon, PROCERATIUM Roger, 1863. Ty p e s p e c i e s Pro- Shi, Kyu). ceratium silaceum Roger, 1863. Number of species: Ponera testacea Emery, 1895 (P. coarctata var. testa- World – 83, Palaearctic – 7, Russia – 1. cea Emery, 1895). Russia: EP (S, NC, CR). – Europe Proceratium melinum (Roger, 1860) [Ponera] (Sys- (WE, SE, EE). phingta europaea Forel, 1886; S. fialai Kratochvíl, Novák et Snoflák, 1944, S. rossica Arnoldi, 1930). Subfamily PROCERATIINAE Russia: EP (S, NC, CR). – Europe (WE, SE, EE), Turkey. Number of taxa: World – 3 genera, 144 species, Palaearctic – 1/7, Russia – 1/1.