Acarina 27 (2): 117–144 © Acarina 2019

IN MEMORIAM OF ANDREI VLADIMIROVICH BOCHKOV: SCIENTIFIC BIOGRAPHY Sergey V. Mironov1, Dmitry A. Apanaskevich1,2 and Sergey G. Medvedev1

1Zoological Institute of the Russian Academy of Sciences, Saint Petersburg, Russia 2United States National Collection, James H. Oliver, Jr. Institute for Coastal Plain Science, Georgia Southern University, Statesboro, USA

Slightly more than a year ago, the world aca- rological and parasitological community suffered from the loss of Andrei (André) Vladimirovich Bochkov (27 August 1968—9 May 2018)—an eminent acarologist, one of the greatest experts on systematics of parasitic acariform —as he died on the 50th year of his life, in the prime of his scientific career. Andrei’s death is an irretrievable loss for the national and world science. His col- leagues and friends dedicate this special issue of Acarina to the memory of this outstanding scientist and extraordinary man. The entire 32-year scientific career of Andrei Bochkov was associated with the Zoological Insti- tute of the Russian Academy of Sciences (Saint Petersburg). At this institute, he undertook his long lection and looking after in the vivarium. and challenging academic path from Lab Assistant His interest in acarology prompted Andrei to ask to Head Researcher, becoming one of the greatest his supervisor, Dr. H.V. Dubinina, an expert on experts in acarology. -associated acariforms and house dust From Laboratory Assistant to PhD (1986– mites, for an opportunity to work on a research 1999). Andrei Bochkov was born on 28 August project in this field. This request resulted in his 1968 in Novosibirsk (Russia). In the mid-1970s, first scientific works dealing with the systematics his family moved to a suburb of Leningrad (now of myobiid mites (Myobiidae) parasitizing on Saint Petersburg). Since his early years, Andrei murid rodents. expressed great interest in biology, which was in- Parallel with his work at the Institute, he was spired by his school teacher E.V. Kudryashov. In a part-time graduate student at the Department of 1986, after graduating from high school in a small Entomology of the Biological Faculty in St. Peters- town of Taitsy (Gatchina District, Leningrad Re- burg State University, from which he graduated in gion, Russia), he came to the Zoological Institute 1996. Later, Andrei declared that the Department with a naive desire “to study tigers”. However, at is where “the real systematists are formed”. By the that time, the Institute could only offer young An- time of his graduation from the university, his sci- drei a more prosaic position of a lab assistant at the entific interest was clearly focused on the system- Department of Parasitology. Nevertheless, he was atics of a particular category of acariform mites— delighted by the opportunity to work at the Institute permanent parasites of . He remained with parasites of vertebrates. Although the labora- faithful to this group of until the end of tory assistant duties in the facility were not his life. Upon graduation, while still working as a particularly exciting, later in his mature years, lab assistant, he independently and enthusiasti- Andrei spoke with considerable pride of the fact cally began conducting research on his favorite that he had passed all of the steps of the scientific group of mites—myobiids (Myobiidae). hierarchy, starting from the very bottom. Gradually, his research began to expand to From the first days of his work at the Depart- other groups of parasitic mites associated with ment, Andrei was not willing to limit himself with small mammals, both Astigmata and . simple tasks, like technical work in the col- In particular, under the supervision of H.V. Du-

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Fig. 1. Andrei Vladimirovich Bochkov and some parasitic mite specimens. A—searching through feathers of a live eared owl Asio otus infected by quill mites (Syringophilidae) (“Rybachy” banding station, Kaliningrad Region, September 2006). B—European robin Erythacus rubecula with web huts of Bakericheyla chanayi (Chetyletidae). C—working tent near Kyzylkol Lake (expedition to Kazakhstan, October 2007). D—Ornithocheyletia sp. (). E—Soricilichus scutisorex (Chirodiscidae), precopulatory guarding of tritonymph by male. F—in the office at the University of Michigan, Ann Arbor, USA (September 2003). binina, Andrei began studying fur mites Chirodis- scientific mindset was significantly broadened by cidae, and Myocoptidae, mange contacts and collaborations with senior researchers psoric mites , house dust mites Pyro- in the Department of Parasitology. Under the influ- glyphidae (Astigmata) and predatory mites Chey- ence of Dr. S.V. Mironov—a prominent expert on letidae (Prostigmata). The development of Andrei’s feather mites, who subsequently became his PhD

118 In memoriam of Andrei V. Bochkov advisor—scientific interests of Andrei have ex- and communicative person, Andrei has established panded to several groups of mites parasitic on ; scientific contact with many researchers world- namely, the families Syringophilidae and Harpi- wide. His interests have expanded to include such rhynchidae (Prostigmata). Based on his long-term mites as bee-associated cheyletids (Cheletophyes), study of myobiids, in 1999, Dr. Bochkov success- skin-dwelling feather mites of the families Derma- fully defended his Candidate of Sciences thesis tionidae and (Astigmata) and even (Russian equivalent of PhD) “Mites of the family gamasid mites (: ). The Myobiidae (: Prostigmata) and their position period of postdoc studies was crucial for Andrei’s in the system”. His thesis work resulted in a pub- development, when in less than a decade he became lication titled “New classification of myobiid mites a world expert in the systematics and phylogeny (Acari, )” (Bochkov, 1997c), which is of parasitic acariform mites. Among his most im- considered a significant contribution to this sub- portant generalization works are those on the clas- field of acarology. sification and phylogeny of mites of the superfam- Foreign postdoc research (2000–2006). Post- ily (Bochkov 2002); on the phylog- doc fellowships at foreign museums and universi- eny and host associations of the fur mite subgenus ties played a huge role in Andrei’s subsequent Listrophoroides (Atopomelidae) (Bochkov and development as a world-class scientist. They al- OConnor 2005b); and, in collaboration with S.V. lowed him to work under the mentorship of world Mironov and A. Fain, on the phylogeny and evolu- leading acarologists and to gain access to rich tion of in feather mites of the families materials deposited in some of the largest world Epidermoptidae and Dermationidae () collections. It is necessary to give Andrei credit for (Mironov et al. 2005). his determination and perseverance: his self-admit- Scientific peaks (2007–2012). After the post- ted less-than-mediocre knowledge of English did doc period of his life—being full of energy, enthu- not deter him from undertaking his first postdoc. siasm and creative plans—Andrei decisively fo- In 2000, Andrei participated in a study under cused on generalization studies and the preparation the supervision of Professor Alex Fain (the Institut of his Doctor of Sciences thesis (equivalent of a royal des Sciences naturelles de Belgique, Brussels, Habilitation Degree in many European countries). Belgium)—the greatest taxonomist-acarologist of During six subsequent years, he published 64 works the 20th century. The project was titled “Phylogeny on astigmatan and prostigmatan mites associated and systematics of the family Cheyletidae (Acari: with mammals, including three monographs: on the Cheyletoidea),—Grant from the Belgian Office for phylogeny and systematics of the endoparasitic Scientific, Technical and Cultural Affairs”. For the astigmatids of the families , Lemur- first time, Andrei had access to the richest collec- nyssidae and Pneumocoptidae (Bochkov et al. tions of parasitic mites stored in Brussels and 2008); a review of mites of the parvorder Eleuther- Tervuren (at the Musee royale de l’Afrique central, engona, permanent parasites of mammals (Bochkov Belgium). As a result of this and other subsequent 2009a); and a monograph on host-parasite relation- collaborative works with Professor Fain, 49 joint ships between the mite subgenus Microtimyobia publications were produced, among which was the (Myobiidae) and cricetid rodents (Bochkov 2011a). reconstruction of the phylogenetic system of Among the most important works of this period are predatory and parasitic mites of the family Chey- generalization papers on the origin and evolution letidae (Bochkov and Fain 2001). of parasitism in mites of the infraorder Eleutheren- From 2003 to 2006, Dr. Bochkov held a three- gona (Bochkov 2008c, 2009a); works on the phy- year postdoc position at the Museum of Zoology logeny and evolution of parasitism in mammal- of the University Michigan, Ann Arbor, USA, in associated psoroptidian mites (Astigmata) (Boch- the lab of Professor Barry M. OConnor—the great- kov 2011b; Bochkov and Mironov 2011); and works est expert on systematics of astigmatan mites. on the use of phylogeny and coevolutionary rela- There, Andrei participated in the project titled tionships of parasites in the study of the evolution “Phylogeny and Historical Ecology of Mite Para- of hosts, e.g., the use of mange mites (Psoroptidae) sites of Vertebrates (Acari: and to study strepsirrine primates (Bochkov et al. 2011). ) (PEET grant from NSF)”, in the Additionally, in collaboration with S.V. Mironov, course of which he excellently mastered the use of Dr. Bochkov has studied the phenomenon of syn- cladistics methods in systematics. Being an open hospitality in parasitic acariform mites (Bochkov

119 S.V. Mironov et al. and Mironov 2008). The two authors have also Scientific heritage. The scientific heritage of generated an original macrophylogenetic system of Andrei Bochkov is rich and diverse, in terms of the order Acariformes (Mironov and Bochkov both mite taxa and the general scope of research 2009). In general, this period of Andrei’s career has topics. He is the author of 220 scientific works, proven to be most fruitful. Finally, in 2011, Andrei including 5 large monographs (see the List of pub- successfully defended his Doctor of Sciences thesis lications; monographs are marked with *). In the titled “Acariform mites, permanent parasites of course of his taxonomic studies, Dr. Bochkov has mammals: phylogeny, systematic and host-parasite established 496 new mite taxa, including 7 family- interactions”. group names, 43 genus-group names and 445 new Acknowledged acarologist (2013–2018). Be- mite species (see the List of taxa and Table 1). It is ing at the top of the acarology profession, Dr. important to emphasize that in the overwhelming Bochkov did not decrease the intensity of his re- majority of his scientific publications, Andrei was search, continuing his studies in various fields the leading author, who performed essentially the until his last days. He continued working on the entire work. But, being a polite person, Andrei phylogeny and the taxonomic revisions of a practi- often acknowledged the participation of senior and cally important family of mange mites, Psoroptidae, respectful co-workers by placing them ahead of which he started back in 2009. A series of taxo- him in the author list. nomic works, devoted to the mites of the family In the last decade, about two dozen of his articles Harpirhynchidae parasitizing on birds in the New were devoted to the reconstruction of the phylogeny World, were finalized in a taxonomic monograph of various family groups of parasitic acariform mites, (Bochkov et al. 2015). including Cheyletoidea and , as well as During this period, Andrei successfully col- to major questions in parasitology and general biol- laborated with colleagues from the A. Mickiewicz ogy. Unfortunately, some monographic works, such University (Poznan, Poland) and, in particular, with as the revision of the scabies mites of the family a prominent expert on quill mites, Dr. M. Skoracki. Psoroptidae and of the hair mites of the family Two major investigatory projects, notably, have Atopomelidae, were never finished. been carried out: “Host-parasite relationships and Because of his efforts, a sizeable representa- biodiversity of prostigmatan mites (Acari: Acari- tive world-class collection of astigmatan and formes), permanent parasites of passeriform birds prostigmatan mites parasitizing on mammals has (Aves: Passeriformes) of Holarctic region” (2011– accumulated in the Department of Parasitology 2013) and “Acariform mites (Acari: Acariformes) of the Zoological Institute RAS. The collection permanently parasitizing warm-blooded vertebrates contains 26 mite families, over 10,000 slides and in Sub-Saharan Africa and Madagascar: fauna about 900 species, including many type speci- genesis, phylogeny, and host-parasite relation- mens. Since the early 1990s, Andrei has partici- ships” (2013–2017). As a result, 18 works on mites pated in many expeditions to Central Asia, Ka- of the families Syringophilidae, Myobiidae and zakhstan and Kamchatka, becoming an extreme- Cheyletidae have been published. Andrei’s gen- ly active collector of valuable research material. eral investigations during this period include the His last expedition in 2016 was to Ethiopia, where works on: the species delimitation and morpho- unique material for morphological and molecular logical diagnostics of Chorioptis mange mites studies was collected. (Bochkov et al. 2014); the cophylogeny of harpy- Simultaneously with his intensive research, rhynchid mites and birds (Bochkov et al. 2017); Andrei took the time to conduct other work for and the phylogeny of house dust mites of the the benefit of science. He actively worked on subfamily Guatemalichinae (Klimov et al. 2015a). editorial boards of a number of national and in- Two works dating to this period present a special ternational scientific journals, such as Acarina, interest. One is his controversial review on the Proceedings of the Zoological Institute of the reversion of parasitism “Is parasitism of Metazoa Russian Academy of Sciences, International Jour- a one-way ticket?” (Bochkov and Mironov 2013). nal of Acarology, Zootaxa and Zookeys. Largely The other is his latest generalizing work “Host- due to his efforts as the deputy editor, the Russian parasite interactions of acariform mites (Acari- journal Acarina has become one of the leading formes)—permanent parasites of mammals” research journals in the field of acarology in the (Bochkov 2016). world.

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Being an extremely sociable person, Andrei has 1991 established contacts with many zoologists engaged 4. Mironov, S.V., Bochkov, A.V. and Chirov, P.A. 1991. in the collection of host-animals of his mites not [Mites of the genus Cheletopsis (Acariformes, Chey- only in Russia, but also in other countries. Despite letidae) from the quills of charadriiform birds from his challenging character, straightforwardness and Middle Asia]. Izvestiya AN Khirgizskoy SSR (khimiko- at times quickness of judgment, Andrei was an tekhnologicheskie nauki), 4: 50–56. [In Russian] extremely benevolent and responsive person both in professional and personal life. A man of sharp 1992 intellect, Andrei was fiery and at times harsh in his 5. Bochkov, A.V. and Rybin, S.N. 1992. [A new species scientific argumentation. However, he could hon- Ewingana teniotis (Acariformes: Myobiidae) from estly and openly admit his errors. He was ex- Tadarida teniotis from Kirghizia]. Parazitologiya, 26 tremely communicative and easily made contacts (4): 333–338. [In Russian with English summary] with people of all ages, positions and social circles. Known for his cheerfulness, optimism and (at times 1993 rough) sense of humor, Andrei could liven up any 6. Dubinina, E.V., Bochkov, A.V. and Bobrovskaja, company, be it in or outside of the workplace. V.I. 1993. [Notes on systematics of mites of the It is safe to say that Andrei was one of the most genus Schizocarpus (Acariformes: Chirodiscidae)]. “famous” employees of the Zoological Institute. In Parazitologiya, 27 (6): 450–453. [In Russian with addition to his exceptional sociability and unique English summary] personal charm, Andrei possessed solid authority 1994 on many scientific topics that went beyond the mite systematics. In early 2000s, he became one of the 7. Bochkov, A.V. 1994a. [Data on mites of the genus most active specialists who contributed to the de- Protomyobia (Acariformes: Myobiidae) of Russia velopment of cladistic methods in systematics. and neighboring countries]. Parazitologiya, 28 (2): Andrei even organized a short course of lectures 137–146. [In Russian with English summary] on his approach at the Zoological Institute. 8. Bochkov, A.V. 1994b. [Mites of the subgenus Andrei left us in the prime of his scientific Graphiurobia (Myobiidae, Radfordia) of the fauna career, maintaining youthful enthusiasm and admi- of Russia and surrounding states]. Parazitologiya, 28 rable work ethic until his very last days, despite the (5): 421–428. [In Russian with English summary] severe illness. We will keep the memory of Andrei 9. Bochkov, A.V., Mironov, S.V. and Gorgol, V.T. 1994. in our hearts—an extraordinary bright person, a Ornithocheyletia phylloscopi sp. n. (Acariformes: special friend and an outstanding scientist. Cheyletidae), new species of parasitic mites from the willow warbler. Acarina, 2 (1–2): 73–80. BIBLIOGRAPHY LIST OF SCIENTIFIC 10. Kivganov, D.A. and Bochkov, A.V. 1994. [A new WORKS BY A.V. BOCHKOV mite species of the genus Cheletopsis (Acari, Cheyletidae)]. Vestnik Zoologii, 2: 39–43. [In Rus- 1989 sian with English summary] 1. Dubinina, H.V. and Bochkov, A.V. 1989. [New data 1995 on ectoparasites of the Royle’s pika in Kirghizia]. Entomologicheskie issledovania v Kirghizii, 20: 11. Bochkov, A.V. 1995. [Mites of the subgenus Mi- 107–116. [In Russian] crotimyobia (Acariformes, Myobiidae, Radfordia) found of the Russia and surrounding countries]. 1990 Parazitologiya, 29 (6): 480–492. [In Russian with 2. Bochkov, A.V., Dubinina, H.V. and Chirov, P.A. English summary] 1990. [A first record of mites of the subgenusAus - 12. Dubinina, H.V., Bochkov, A.V. and Javrujan, J. G. tromyobia (Acariformes, Myobiidae) in the USSR 1995. Mites of the family Myobiidae (Acari- fauna]. Izvestiya AN Khirgizskoy SSR (khimiko- formes)—ectoparasites of bats of Armenia. Aca- tekhnologicheskie nauki), 1: 55–66. [In Russian] rina, 3 (1–2): 99–104. 3. Efremova, G.A. and Bochkov, A.V. 1990. [Mites of 1996 families Cheyletidae and Cheyletiellidae (Acari- formes: ) of the Byelorussian fau- 13. Bochkov, A.V. 1996a. Hesperomyobia (Acari: na]. Vesti AN BSSR, 6: 98–103. [In Byelorussian] Myobiidae: Radfordia), a new subgenus of myo-

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biid mites from rodents of the family Hesperomy- 1998 idae (Rodentia). Acarina, 4 (1–2): 39–42. 25. Bochkov, A.V. and Mironov, S.V. 1998a. [On a 14. Bochkov, A.V. 1996b. Mites of the family Myobi- systematics of mites of the subgenus Microtimyo- idae (Acariformes: Prostigmata) parasitizing on bia (Acariformes: Myobiidae: Radfordia) and their bats (Mammalia: Chiroptera) of the fauna of the distribution on voles (Rodentia: Cricetidae: Arvi- former USSR. Acarina, 4 (1–2): 3–26. colinae)]. Parazitologiya, 32 (4): 300–316. [In 15. Bochkov, A.V. and Malikov, V.G. 1996. Eucheyle- Russian with English summary] tiella faini sp. n. (Acari: Cheyletidae), a new spe- 26. Bochkov, A.V. and Mironov, S.V. 1998b. Quill cies of parasitic mites from Ochotona rufescens mites of the family Syringophilidae Lavoipierre, (Gray) (Lagomorpha, Ochotonidae). Acarina, 4 1953 (Acariformes: Prostigmata) parasitizing on (1–2): 43–48. birds (Aves) of the fauna of the former USSR. 16. Dubinina, H.V. and Bochkov, A.V. 1996. Pleisto- Acarina, 6 (1–2): 3–16. cene mites (Acariformes) from the narrow-skulled 27. Bochkov, A.V. and Mironov, S.V. 1998c. Samsina- vole Microtus gregalis egorovi Feigin (Rodentia: kia trilobitus sp. n., a new cheyletid mite from Arvicolinae). Acarina, 4 (1–2): 35–38. South India (Acari, Cheyletidae). Entomologische Mitteilungen aus dem Zoologischen Museum, 1997 Hamburg, 12: 265–268. 17. Bochkov, A.V. 1997a. [Description of the female 28. Chirov, P.A. and Bochkov A.V. 1998. Ornithochey- of the mite Radfordia (Austromyobia) aegyptica letia mironovi sp. n. (Acari: Cheyletidae), a new (Acariformes: Myobiidae)]. Parazitologiya, 31 species of parasitic mite from the bank swallow (5): 458–461. [In Russian with English summary] from Kirghizia. Acarina, 6 (1–2): 35–36. 18. Bochkov, A.V. 1997b. [Myobiid mites (Acari- formes: Myobiidae) parasiting a rodent family 1999 Muridae (Rodentia) in Russia and neighbouring 29. Bochkov A.V. 1999a. [A new myobiid mites spe- countries]. Parazitologiya, 31 (3): 201–209. [In cies Radfordia cricetuliphila sp. n. (Acari: Myobi- Russian with English summary] idae) from the striped hamster Cricetulus baraben- 19. Bochkov, A.V. 1997c. [New classification of sis (Rodentia: Cricetidae) from Buryatia]. Parazi- myobiid mites (Acari, Acariformes)]. Entomo- tologiya, 33 (4): 358–363. [In Russian with English logicheskoe obozrenie, 76 (4): 938–951. [In Rus- summary] sian with English summary] [More correct trans- 30. Bochkov, A.V. 1999b. [The system and host-para- lation of the title: new classification of mites of sites relationships of the mites of the subfamily the family Myobiidae (Acari, Acariformes)] Myobiinae (Acariformes: Myobiidae)]. Entomo- 20. Bochkov, A.V. 1997d. [Two new subgenera of logicheskoe obozrenie, 78 (3): 764–776. [In Rus- myobiid mites of the genus Radfordia Ewing sian with English summary; also translated into (Acariformes, Myobiidae)]. Entomological Re- English as: Classification and host-parasite rela- view, 76 (3): 684–704. [In Russian with English tions of mites of the subfamily Myobiinae (Acari- summary] formes, Myobiidae), Entomological Review, 79 21. Bochkov, A.V. 1997e. Mites of the family Myo- (6): 666–675.] biidae (Acariformes: Prostigmata) parasitizing on 31. Bochkov A. V. 1999c. [Mites of the family Myo- Lipotyphla (Mammalia: Insectivora) of the fauna biidae (Acari: Prostigmata) and their position in of former USSR. Acarina, 5 (1–2): 45–62. the system]. Candidate of Science thesis. Zoo- 22. Bochkov, A.V. and Fain, A. 1997. On a systemat- logical Institute RAS, Saint Petersburg. 480 pp. ics of mites of the subgenus Radfordia s.str. [Manuscript, in Russian] (Acari: Myobiidae: Radfordia). Acarina, 5 (1–2): 32. Bochkov, A.V. and Haustov, A.A. 1999. Proso- 19–28. cheyla villosa sp. n., a new mite species from 23. Bochkov, A.V. and Mironov, S.V. 1997. On tax- Crimea (Prostigmata: Cheyletidae). Genus, 10 (1): onomy of predatory mites of the genus Neoeuchey- 151–154. la Radford, 1950 and related genera (Acari: 33. Bochkov, A.V. and Khaustov, A.A. 1999. [Descrip- Cheyletidae). Acarina, 5 (1–2): 73–78. tion of Cheyletus kuznetzovi sp. n. (Acariformes: 24. Chirov, P.A., Bochkov, A.V. and Rybin, S.N. 1997. Cheyletidae) from Turkmenia]. Zoologicheskii [The mites of the family Myobiidae from Kir- Zhurnal, 78 (1): 115–116. 302 [In Russian with ghizia]. Entomologicheskie issledovaniya v Kir- English summary; also published in English in gizii, 21: 42–50. [In Russian] Entomological Review, 79 (4): 482–483.]

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34. Bochkov, A.V., Khaustov, A.A. and Kuznetzov, 45. Bochkov, A. V., Mironov, S. V. and Kravtsova, N. N.N. 1999. [A new mite species of the genus T. 2000. Two new syringophilid mites (Acari: Chelacheles (Acariformes: Cheyletidae) from the Syringophilidae) from the Greenfinch Carduelis Crimea]. Zoologicheskii Zhurnal, 78 (1): 117–120. chloris (Passeriformes: Fringillidae) from Kir- [In Russian with English summary; also published ghizia. Genus, 11 (2): 351–358. in English in Entomological Review, 79 (4): 46. Fain, A., Bochkov, A. V. and Mironov, S. V. 2000. 484–487.] New genera and species of quill mites of the fam- 35. Bochkov, A., Malikov, V. G. and Arbobi, M. 1999. ily Syringophilidae (Acari: Prostigmata). Bulletin Trichoecius calomysci sp. n. (Acari: Myocoptidae), de l’Institut royal des Sciences naturelles de Bel- a new mite species from Iran. Folia Parasitologi- gique (Entomologie), 70: 33–70. ca, 46: 316–318. 36. Bochkov, A.V. and Mironov, S.V. 1999a. A system- 2001 atic review of the parasitic mite genus Eucheyletia 47. Bochkov, A.V. 2001a. [On systematics of mites of the Volgin, 1969 (Prostigmata: Cheyletidae). Mit- genus Syringophiloidus (Acari: Syringophilidae) teilungen aus dem Hamburgischen zoologischen from the European part of Russia]. Parazitologiya, Museum und Institut, 96: 141–155. 35 (2): 149–153. [In Russian with English summary] 37. Bochkov, A.V. and Mironov, S.V. 1999b. New quill 48. Bochkov, A.V. 2001b. [Parallel evolution of mites species of the family Syringophilidae (Acari: of the family Myobiidae (Acari: Prostigmata) and Cheyletoidea) from the European part of Russia. jerboas (Rodentia: Dipodidae)]. Parazitologiya, 35 Acarina, 7 (1): 35–45. (1): 9–18. [In Russian with English summary] 38. Bochkov, A.V., Mironov, S.V. and Fain, A. 1999. 49. Bochkov, A.V. and Apanaskevich, D. A. 2001. Two Phylogeny and host–parasite relationships of the new species of the family Syringophilidae (Acari: family Harpirhynchidae (Acari, Prostigmata). Cheyletoidea) from passeriform birds collected in Acarina, 7 (2): 69–87. Altai. Folia Parasitologica, 48: 321–325. 39. Fain, A., Bochkov, A.V. and Mironov, S.V. 1999. 50. Bochkov, A.V. and Chistyakov, D.V. 2001. De- A contribution to the systematics of the mite fam- scription of Charadriphilus lyudmilae gen. et sp. n. ily Harpirhynchidae (Acari: Cheyletoidea). Aca- (Acari: Syringophilidae) from Scolopax rustica rologia, 40 (1): 35–52. (Aves: Scolopacidae) in NW Russia. Zoosystem- atica Rossica, 9: 295–297. 2000 51. Bochkov, A.V. and Fain, A. 2001. Phylogeny and 40. Bochkov, A.V. 2000a. [A first record of the mites of system of the Cheyletidae (Acari: Prostigmata) genus Neharpyrhynchus Fain, 1972 (Acari: Harpi- with special reference to their host–parasite asso- rhynchidae) in Russian fauna]. Parazitologiya, 34 ciations. Bulletin de l’Institut Royal des Sciences (6): 534–538. [In Russian with English summary] Naturelles de Belgique, 71: 5–36. 41. Bochkov, A.V. 2000b. A new harpirhynchid mite 52. Bochkov, A.V., Fain, A. and Ardeshir, F. 2001. Harpyrhynchus alaudinus sp. n. (Acari: Harpirhyn- Redescription of Nodele calamondin Muma, 1964 chidae) from Alauda arvensis (Passeriformes: (Acari Cheyletidae). Bulletin de la Société Royale Alaudidae) from Russia. Acarina, 8 (2): 91–93. Belge d’Entomologie, 137: 123–126. 42. Bochkov, A.V. 2000c. Syringophilus numidiae sp. n., 53. Bochkov, A.V. and Galloway, T. D. 2001. The a new species of quill mites (Acarina: Syringophili- parasitical cheyletoid mites (Acari: Cheyletoidea) dae) ex Numida meleagris (Galliformes: Phasianidae) associated with passeriform birds (Aves: Passeri- from South Africa. Acarina, 8 (2): 103–104. formes) from Canada. Canadian Journal of Zool- 43. Bochkov, A. V., Arbobi, M. and Malikov, V. G. 2000. ogy, 79: 2014–2028. Notes on mites of the family Myobiidae (Acari: 54. Bochkov, A.V. and Mironov, S.V. 2001. New re- Prostigmata) parasitizing rodents (Mammalia: Ro- cords of fur-mites of the subgenus Listrophoroides dentia) in Iran. Folia Parasitologica, 47: 73–77. s.str. (Acari: Atopomelidae) from rats of Vietnam. 44. Bochkov, A.V. and Mironov, S.V. 2000. Two new Mitteilungen aus dem Hamburgischen zoolo- species of the genus Geckobia (Acari: Pterygoso- gischen Museum und Institut, 13: 279–285. matidae) from geckos (Lacertilia: Gekkonomor- 55. Bochkov, A.V., Mironov, S.V. and Skoracki, M. pha) with a brief review of host–parasite associa- 2001. Four new quill mite species of the genus tions of the genus. Russian Journal of Herpetol- Syringophilopsis Kethley, 1970 (Acariformes: ogy, 7 (1): 61–68. Prostigmata). Acarina, 9 (2): 233–239.

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56. Bochkov, A.V., Mironov, S.V. and Stekolnikov, from a cave in Western Australia. International A.A. 2001. [Acariform mites (Acariformes)]. In: Journal of Acarology, 17: 37–40. S.A. Fetisov and G.Yu. Konechnaya (Eds.). Bioraz- 66. Fain, A. and Bochkov, A.V. 2002b. A new fur mite noobraziye i redkie vidy natsionalnogo parka Lynxacarus (Lutracarus) visoni sp. n. (Acari, Lis- “Sebezhskii” [Biodiversity and rare species of the trophoridae) from Mustela vison (Carnivora, national park “Sebezhskii”]. Saint Petersburg State Mustelidae) in North America. Acta Parasitolog- University (Publications of the St. Petersburg ica, 47 (1): 47–50. Society of Naturalists, Series 6, Vol. 4), Saint Pe- 67. Fain, A. and Bochkov, A.V. 2002c. A new species tersburg, pp. 109–112. [In Russian] of the genus Eudusbabekia Jameson, 1971 (Acari: 57. Fain, A. and Bochkov, A.V. 2001a. A new genus and Myobiidae) parasitizing Rhinophylla pumilio species of mite (Acari: Epidermoptidae) from the (Chiroptera: Phyllostomatidae) from Ecuador. ear of a South American Dove. Bulletin de la So- Systematic Parasitology, 52: 167–169. ciété Royale Belge d’Entomologie, 137: 117–122. 68. Fain, A. and Bochkov, A.V. 2002d. On some little 58. Fain, A. and Bochkov, A.V. 2001b. A new species known and a new species of Myobiidae (Acari) of the genus Epimyodex Fain and Orts, 1969 associated with rodents. Bulletin de la Société (Acari: Cloacaridae: Epimyodicinae) parasitizing Royale Belge d’Entomologie, 138: 95–105. Sorex trowbridgii (Soricidae) from the U.S.A. 69. Fain, A. and Bochkov, A.V. 2002e. On some new International Journal of Acarology, 27: 221–223. or little known species of parasitic Cheyletidae 59. Fain, A. and Bochkov, A.V. 2001c. A review of (Acari: Prostigmata). Acarologia, 17 (2): 145–160. some cheyletid genera (Acari : Prostigmata) with 70. Fain, A., Bochkov, A.V. and Corpuz-Raros, L.A. descriptions of new species. Acarina, 9 (1): 47–95. 2002. A revision of the Hemicheyletia generic 60. Fain, A. and Bochkov, A.V. 2001d. A review of the group (Acari: Cheyletidae). Bulletin de l’Institut genus Cheyletus Latreille, 1776 (Acari: Cheyleti- royal des Sciences naturelles de Belgique, 72: dae). Bulletin de l’Institut Royal des Sciences 27–66. Naturelles de Belgique, 71: 83–114. 71. Skoracki, M. and Bochkov A.V. 2002. A new quill 61. Fain, A. and Bochkov, A.V. 2001e. Observations mite species Bubophilus asio sp. n. (Acari: Syrin- on the taxonomic status of some cheyletid genera gophilidae) from the long-eared owl Asio otus (Acari Cheyletidae). Belgian Journal of Entomol- (Strigiformes: Strigidae). Genus, 13 (1): 149–152. ogy, 3: 291–301. 2003 2002 72. Balashov, Yu.S., Bochkov, A.V., Vashchonok, V.S. 62. Balashov Yu.S., Bochkov, A.V., Vashchonok, V.S., and Tret’yakov, K.A. 2003. [Structure and sea- Grigoreva, L.A. and Tretyakov, K.A. 2002. [Struc- sonal dynamics of the community ectoparasites on ture and seasonal dynamics of the community the common shrew (Sorex araneus) in the Ilmen- ectoparasites in the bank vole in the Ilmen-Volkhov Volkhov Lowland]. Parazitologiya, 37 (6): 441– lowland]. Parazitologiya, 36 (6): 433–446. [In 454. [In Russian with English Summary] Russian with English Summary] 73. Bochkov, A.V. and Fain, A. 2003a. Revision of the Bochkov, A.V. 2002. [Classification and phylog- subgenus Marquesania (Acari, Atopomelidae). eny of mites of the superfamily Cheyletoidea Invertebrate Systematics, 17 (4): 575–604. (Acari: Prostigmata)]. Entomologicheskoe Oboz- 74. Bochkov, A.V. and Fain, A. 2003b. [A new mite renie, 81: 488–513. [In Russian with English Sum- species Radfordia sigmomys sp. n. (Acari: Myobi- mary; also published in English in Entomological idae) parasitizing Sigmodon alstoni (Rodentia: Review, 82 (6): 641–664.] Sigmodontidae) from Central America]. Parazi- 63. Bochkov, A.V., Fain, A. and Dabert, J. 2002. A tologiya, 37 (2): 107–112. [In Russian with English revision of the genus Cheletopsis (Acari Chey- summary] letidae). Bulletin de l’Institut Royal des Sciences 75. Bochkov, A.V. and Fain, A. 2003c. New and little Naturelles de Belgique, 72: 5–26. known species of the family Syringophilidae 64. Bochkov, A.V. and Perez, T.M. 2002. New quill (Acari: Cheyletoidea) from parrots (Aves: Psit- mites of the family Syringophilidae (Acari: Chey- taciformes). Acarina, 11: 37–44. letoidea) parasitizing Mexican parrots. Belgian 76. Bochkov, A.V. and Fain, A. 2003d. New observa- Journal of Entomology, 4: 145–159. tions of mites of the family Myobiidae Megnin, 65. Fain, A. and Bochkov, A.V. 2002a. A new genus 1778 (Acari: Prostigmata) with special reference and species of cheyletid mite (Acari: Cheyletidae) to their host-parasite relationships. Bulletin de

124 In memoriam of Andrei V. Bochkov

l’Institut Royal des Sciences Naturelles de Bel- Cheyletoidea) from birds in Canada. Journal of the gique, 73: 5–50. Kansas Entomological Society, 77: 26–44. 77. Bochkov, A.V. and Labrzycka, A.A. 2003. Revision 89. Bochkov, A.V. and Klimov, P.B. 2004. A new of the European species of the genus Myobia Van predaceous mite Nodele (Aztecocheyletus) con- Heyden, 1826 (Acari: Myobiidae). Acta Parasito- quistador subgen. n., sp. n. (Acari: Cheyletidae) logica, 48 (1): 24–40. phoretic on Aztecontidium tenochtitlanicum (Hy- 78. Bochkov, A.V. and OConnor, B.M. 2003a. New menoptera: Megachilidae). Acarina, 12 (1): 23–27. cheyletid mites (Acari, Cheyletidae) associated 90. Bochkov, A.V. and OConnor, B.M. 2004a. Phylog- with birds. Acta Parasitologica, 48 (4): 265–279. eny, systematics and biology of mites of the genera 79. Bochkov, A.V. and OConnor, B.M. 2003b. Two Chelacheles and Neochelacheles (Acari: Chey- new species of the genus Cheyletus Latreille, 1796 letidae). Invertebrate Systematics, 18: 547–592. (Acari: Cheyletidae) associated with Philippine 91. Bochkov, A.V. and OConnor, B.M. 2004b. System- vertebrates. International Journal of Acarology­ , atics of the genus Sciurocheyla (Acari: Cheyletidae), 29: 327–330. parasites of tropical squirrels (Rodentia: Sciuridae). 80. Fain, A. and Bochkov, A.V. 2003. A new species Journal of Medical Entomology, 41: 308–320. of the genus Metacheyletia Fain, 1972 (Acari: 92. Bochkov, A.V., OConnor, B.M. and Gorog, A. Cheyletidae) parasitizing Serinus mozambicus 2004. New species and records of fur mites (Ac- (Passeriformes: Fringillidae) in Central Africa. ari: Astigmata: Atopomelidae) from Southeast International Journal of Acarology, 29: 119–121. Asian rodents (Rodentia: Muridae). Acarina, 12 81. Fain, A. and Bochkov, A.V. 2003a. A review of the (2): 67–86. genus Asiochirus Fain, 1970 (Acari Listrophoridae). 93. Fain, A. and Bochkov, A.V. 2004a. A new species Bulletin de la Societe Royale Belge d’Entomologie, of the genus Myocoptes Claparede, 1896 (Acari: 139: 227–232. Myocoptidae) parasitizing Pitymys pinetorum 82. Fain, A. and Bochkov, A.V. 2003b. New notes on (Rodentia: Cricetidae) from the USA. Interna- the family Atopomelidae (Acari: Listrophoroidea). tional Journal of Acarology, 30: 37–39. Annales du Musée Royal de l’Afrique Centrale, 94. Fain, A. and Bochkov, A.V. 2004b. Listrophoroides series Zoologie, 291: 1–32. (Afrolistrophoroides) prionomys n.sp. (Acari, 83. Fain, A. and Bochkov, A.V. 2003c. New species of Atopomelidae) parasitic on Prionomys batesi (Ro- mites parasitic on the skin of birds (Acari: Epider- dentia, Dendromurinae) from Republique Cen- moptidae and Dermationidae). Bulletin de la So- trafricaine. Journal of Afrotropical Zoology, 1: 5–8. ciete Royale Belge d’Entomologie, 139: 121–149. 95. Skoracki, M., Bochkov, A.V. and Sikora, B. 2004. 84. Fain, A. and Bochkov, A.V. 2003d. Observations New records of the cheyletoid mites (Acari: Chey- on the (Acari: Astigmata) with letoidea) from passeriform birds in Poland. Belgian description of a new genus and species from Cyg- Journal of Entomology, 6: 83–90. nus melanocoryphus (Molina) (Aves: Anatidae). 96. Skoracki, M., Bochkov, A.V. and Wauthy, G. 2004. International Journal of Acarology, 29: 123–126. Revision of the quill mites of the genus Picobia Haller, 1878 (Acari: Syringophilidae) with notes 2004 on their host-parasites relationships. Insect System- 85. Bochkov, A.V. 2004a. A new finding ofTeinochey - atics and Evolution, 35: 155–176. lus gundii Fain and al., 1982 (Acari: Cheyletidae) 2005 from a ctenodactylid rodent (Rodentia: Ctenodac- tylidae). Zoosystematica Rossica, 13 (1): 4. 97. Bochkov, A.V., Hakimitabar, M. and Saboori, A. 86. Bochkov, A.V. 2004b. [Mites of the family Chey- 2005. A review of the Iranian Cheyletidae (Acari letidae (Acari: Prostigmata): phylogeny, distribu- Prostigmata). Belgian Journal of Entomology, 7: tion, evolution and analysis of host-parasite rela- 99–109. tionships]. Parazitologiya, 38 (2): 122–138. [In 98. Bochkov, A.V. and Klimov, P.B. 2005. Three new Russian with English summary] species of the predaceous Cheyletidae (Acari: Prostig- 87. Bochkov, A.V., Fain, A. and Skoracki, M. 2004. mata) phoretic on insects. Acarina, 13 (1): 15–22. New quill mites of the family Syringophilidae 99. Bochkov, A.V., Klimov A.B. and OConnor, B.M. (Acari: Prostigmata). Systematic Parasitology, 57: 2005. Fur-mites of the genus Atopomelus Troues- 135–150. sart, 1918 (Acari: Atopomelidae): life-cycle, phy- 88. Bochkov, A.V. and Galloway, T.D. 2004. New logeny, and host parasite-associations. Acarologia, species and records of cheyletoid mites (Acari: 45: 207–241.

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100. Bochkov, A.V. and Ochoa, R. 2005. Three new parasitic on gymnures of the genus Hylomys cheyletid species (Acari: Cheyletidae) from the (Eulipotyphla: Erinaceidae). Folia Parasitologi- National Museum of Natural History Entomo- ca, 53: 302–310. logical collection, USA. Acarina, 13 (2): 137–144. 111. Bochkov, A.V. and OConnor, B.M. 2006d. Revi- 101. Bochkov, A.V. and OConnor, B.M. 2005a. Ob- sion of the fur-mite family Listrophoridae (Aca- servations on the listrophorid (Acari: Listro- ri: Astigmata) associated with Philippine mam- phoridae) fauna of Pakistan. Acarina, 13 (2): mals. Bulletin de l’Institut Royal des Sciences 165–171. Naturelles de Belgique, Entomologie, 76: 5–24. 102. Bochkov, A.V. and OConnor, B.M. 2005b. Phy- 112. Bochkov, A.V. and OConnor, B.M. 2006e. Revi- logeny and host associations of the fur-mite sion of the genus Gaudalges (Acari: Psoroptidae), subgenus Listrophoroides Hirst (Acari: Atopo- parasites of Malagasy lemurs. Acarina, 14: 3–20. melidae), with an intrigue example of synhospi- 113. Klimov, P.B., Bochkov, A.V. and OConnor, B.M. tality on rats of the genus Maxomys. Invertebrate 2006. Mites of the genus Cheletophyes (Acarina: Systematics, 19: 437–498. Cheyletidae) associated with large African car- 103. Bochkov, A.V. and OConnor, B.M. 2005c. The penter bees (Hymenoptera: Apidae: Xylocopa), life-cycle of Hemigalichus chrotogale sp. nov. with multivariate analyses of sibling species. Bio- (Acari: Listrophoridae), with comparative obser- logical Journal of Linnaean Society, 87: 45–58. vations on listrophorid morphology. Journal of Natural History, 39: 3811–3832. 2007 104. Bochkov, A.V., Williams, G. and Proctor, H. 2005. 114. Balashov, Yu.S., Bochkov, A.V., Vashenok, V.S., Record of Picobia zumpti (Acari: Syringophili- Grigoreva, L.A., Stanukovich M.K. and Tretya- dae) from quills of the rock pigeon in North kov K.A. 2007a. [Segregation of communities of America. Belgian Journal of Entomology, 7: ectoparasites of small forest mammals among 121–127. ecological niches]. Doklady Akademii Nauk, 415 105. Mironov, S.V., Bochkov, A.V. and Fain, A. 2005. (2): 277–280. [In Russian; also publised in Eng- Phylogeny and evolution of parasitism in feather lish in Doklady Biological Sciences, 415 (2): mites of the families Epidermoptidae and Der- 280–283] mationidae (Acari: Analgoidea). Zoologischer 115. Balashov, Yu.S., Bochkov, A.V., Vashenok, V.S., Anzeiger, 243: 155–179. Grigoreva, L.A., Stanukovich M.K. and Tretya- kov K.A. 2007b. [Structure of populations and 2006 ecological niches of ectoparasites in parasite 106. Bochkov, A.V., Mironov, S.V. and OConnor, B.M. communities of small forest mammals]. Parazi- 2006. Revision of the genus Galagocheles Fain tologia, 41 (5): 329-347. [In Russian with English (Acari: Cheyletidae) parasites of galagos (Pri- summary] mates: Galagonidae). Systematic Parasitology, 116. Bochkov, A.V. 2007. [Morphological adapta- 63: 181–200. tions of acariform mites (Acari: Acariformes) to 107. Bochkov, A.V. and Literak, I. 2006. A review of permanent parasitism on mammals.] Parazi- the European Harpirhynchidae (Acari: Prostig- tologiya, 41 (6): 428–458. [In Russian with mata) with description of a new species. Acta English summary] Parasitologica, 51: 136–142. 117. Bochkov, A.V., Literak, I. and Capek, M. 2007. 108. Bochkov, A.V. and OConnor, B.M. 2006a. A re- Neharpyrhynchus baile n.sp. (Prostigmata: Harpi- view of the external morphology of the mite rhynchidae) parasitizing Turdus leucomelas Viel- family Pterygosomatidae and its systematic posi- lot (Aves: Turdidae) from Brazil. International tion within the Prostigmata (Acari: Acariformes). Journal of Acarology, 33: 35–39. Parazitologiya, 40: 201–214. [In Russian with 118. Bochkov, A.V. and Walter, D.E. 2007. The life- English summary] cycle of Pomerantzia philippina sp. n. (Prostig- 109. Bochkov, A.V. and OConnor, B.M. 2006b. Fur- mata: Pomerantziidae) described from the Philip- mites of the family Atopomelidae (Acari: Astig- pines. Acarina, 15 (2): 159–170. mata) of the Philippines: systematics, phylogeny, 119. Dowling, A.P.G., Bochkov, A.V. and OConnor, and host-parasite relationships. Miscellaneous B.M. 2007. Revision of the genus Andreacarus Publications. Museum of Zoology, University of (Acari: Laelapidae) with description of seven new Michigan, 196: 1–62. species and a new genus for Australian species 110. Bochkov, A.V. and OConnor, B.M. 2006c. Hylo- formerly placed in Andreacarus. Journal of myobia (Acari: Myobiidae), a new genus of mites Medical Entomology, 44: 405–421.

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2008 stigmata)—permanent parasites of mammals. Acarina, supplement, 149 pp. 120. Bochkov, A.V. 2008a. A review of the mite fam- ily Stigmocheylidae Berlese (Acari: Prostigmata). 131. Bochkov, A.V. 2009b. Mites of the family Myobi- Annales Zoologici, 58: 311–325. idae (Acari: Prostigmata) parasitizing rodents of 121. Bochkov, A.V. 2008b. New observations on phy- the former USSR. Acarina, 17 (2): 109–169. logeny of cheyletoid mites (Acari: Prostigmata: 132. Bochkov, A.V. 2009c. [Origin and evolution of Cheyletoidea). Proceedings of the Zoological parasitism in mites of the infraorder Eleutheren- Institute RAS, 312 (1–2): 54–73. gona (Acari: Prostigmata). Report II. Superfam- 122. Bochkov, A.V. 2008c. Origin and evolution of ily Cheyletoidea]. Parazitologiya, 43 (2): 97–117. parasitism in mites of the infraorder Eleutheren- [In Russian with English Summary] gona (Acari: Prostigmata). Report I. Lower 133. Bochkov, A.V., Flannery, M.E. and Spicer, G. Raphignathae. Parazitologiya, 42 (5): 337–359. 2009. Mites of the genus Torotrogla (Prostig- [In Russian with English Summary] mata: Syringophilidae) from North American 123. Bochkov, A.V., Klompen, H. and OConnor, B.M. passerines. Journal of Medical Entomology, 46: 2008. A review of the external morphology of the 183–197. family Chirorhynchobiidae (Acari: Sarcoptoidea) 134. Bochkov, A.V., Melnikov, D.A. and Nazarov, with description of a new species. Journal of R.A. 2009. Pterygosoma (Pterygosoma) pseudo- Medical Entomology. 45: 193–202. trapelus sp. nov. (Acariformes: Pterygosomati- 124. Bochkov, A.V. and Literak, I. 2008. Description dae)—ectoparasite of Pseudotrapelus sinaitus of the life-stages of Harpyrhynchoides rubeculi- (Squamata: Agamidae) from Jordan. Zootaxa, nus (Černý and Sixl, 1971) (Prostigmata: Harpi- 2232: 61–68. rhynchidae). International Journal of Acarology, 135. Bochkov, A.V. and OConnor, B.M. 2009. A new 34: 227–236. mite genus Caenolestomyobia gen. nov. (Acari- 125. Bochkov, A.V. and Mironov, S.V. 2008. [The formes: Myobiidae) from marsupials of the genus phenomenon of “phylogenetic synhospitality” Caenolestes (Pauctituberculata: Caenolestidae). in acariform mites (Acari: Acariformes)—the Proceedings of the Zoological Institute RAS, 313 permanent parasites of vertebrates]. Parazi- (4): 140–151. tologiya, 42 (2): 81–100. [In Russian with 136. Bochkov, A.V. and Wauthy, G. 2009. A review of English Summary] the genus Listropsoralges (Acari, Psoroptidae) 126. Bochkov, A.V. and OConnor, B.M. 2008a. A new with the descriptions of two new species. Acta mite superfamily Cloacaroidea and its position Parasitologica, 54: 269–275. within the Prostigmata (Acariformes). Journal of 137. Literak, I. Bochkov, A.V., Dusbabek, F., Martinu, Parasitology, 94: 335–344. J., Podzemny, P. and Sychra, O. 2009. Málo známí 127. Bochkov, A.V. and OConnor, B.M. 2008b. The cizopasníci ptáků—roztoči čeledi Harpirhynchi- external morphology of the mite subfamily Dro­ dae. Ziva, 3: 126–128. [In Czech] micioptinae (Acari: Myocoptidae), evidences for 138. Mironov, S.V. and Bochkov, A.V. 2009. [Modern and against the myocoptid monophyly. Annals of conceptions of macrophylogeny in acariform the Entomological Society of America, 101: mites (: Acariformes)]. Zoologiches- 289–296. kiy Zhurnal, 88: 922–937. [In Russian with 128. Bochkov, A.V., OConnor, B.M. and Wauthy, G. English summary; also translated into English as: 2008. Phylogenetic position of the family Myo- Modern conceptions concerning macrophylogeny biidae within the Prostigmata (Acari: Acari- of acariform mites (Chelicerata: Acariformes), formes). Zoologischer Anzeiger, 247: 15–45. Entomological Review, 89 (8): 975–992.] 129. *Bochkov, A.V., Zabludovskaya, S.A. and OCon- 139. Whitaker, J.O. Jr., Ruckdeschel, C. and Bochkov, nor, B.M. 2008. Phylogeny and systematics of A.V. 2009. Species of the genus Schizocarpus the endoparasitic astigmatid mites (Acari: Sar- Trouessart, 1896 (Acari: Chirodiscidae) from coptiformes) of mammals: families Gastronys- Florida and Georgia beavers. Florida Scientist, sidae, Lemurnyssidae, and Pneumocoptidae. Biological Sciences, 72: 18–21. Zootaxa, 1951: 1–152. 2010 2009 140. Bochkov, A.V. 2010. A review of mammal as- 130. *Bochkov, A.V. 2009a. A review of mites of the sociated Psoroptidia (Acariformes: Astigmata). parvorder Eleutherengona (Acariformes: Pro­ Acarina, 18 (2): 99–260.

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141. Bochkov, A.V. and Mertins, J.W. 2010. Harpi- 151. Bochkov, A.V., Abramov, A.V. and Hugot, G.P. rhynchus quasimodo n.sp. (Acariformes: Harpi- 2011. Five new species of fur mites (Acariformes: rhynchidae), a new species parasitizing Molothrus Sarcoptoidea) from small mammals in Laos. ater (Passeriformes: Icteridae) in Florida. Inter- Zootaxa, 2980: 1–22. national Journal of Acarology, 36 (1): 83–87. 152. Bochkov, A.V. and Dubinina, H.V. 2011. Mites 142. Bochkov, A.V. and OConnor, B.M. 2010. Exter- of the genus Schizocarpus (Acariformes: Chiro- nal morphology and systematics of the family discidae) parasitizing the Eurasian beaver Castor Lobalgidae (Acari: Astigmata) with resolution of fiber (Rodentia: Castoridae) in the Voronezh the question of family monophyly. Insect System- National Reserve. Acarina, 19 (1): 53–66. atics and Evolution, 41: 187–229. 153. Bochkov, A.V., Klimov, P.B. and Wauthy, G. 143. Bochkov, A.V. and OConnor, B.M. 2010. Nidi- 2011. Phylogeny and coevolutionary associations colocheyletus kennedyae n.gen., n.sp. (Prostig- of makialgine mites (Acari, Psoroptidae, Maki- mata: Cheyletidae) from nests of passerine birds alginae) provide insight into evolutionary history in Michigan, USA. International Journal of Aca- of their hosts, strepsirrhine primates. Zoology rology, 36 (1): 59–71. Journal of the Linnean Society, 162: 1–14. 144. Bochkov, A.V., OConnor, B.M. and Wauthy, G. 154. Bochkov, A.V. and Literak, I. 2011. Mites of the 2010. A review of the mite subfamily Makialginae genus Neharpyrhynchus Fain (Acariformes, (Acari: Psoroptidae)—permanent parasites of Harpirhynchidae) from Neotropical birds. Zoo­ strepsirrhine primates. Tijdschrift voor Ento- Keys, 89: 15–31. mologie, 153 (2): 217–237. 155. Bochkov, A. V. and Medvedev, S.G. 2011. [Ev- 145. Bochkov, A.V. and Otto, J.S. 2010. Five new erything appeared to be not so dreadful]. Priroda, species of predaceous cheyletid mites (Acari: 8: 75–80. [In Russian] Cheyletidae). Zootaxa, 2727: 1–20. 156. Bochkov A.V. and Mironov S.V. 2011. Phylogeny 146. Skoracki, M. and Bochkov, A.V. 2010. Syringo- and systematics of mammal-associated psoroptid- philid mites (Acari: Syringophilidae) of Kazakh- ian mites (Acariformes: Astigmata: Psoroptidia) derived from external morphology. Invertebrate stan. Zootaxa, 2546: 52–68. Systematics, 25: 22-59. 2011 157. Bochkov A.V. and Skoracki M. 2011. A new cheyletid mite Metacheyletia ngaii n.sp. (Acari- 147. *Bochkov A.V. 2011a. Mites of the subgenus formes: Cheyletidae) from quills of Corythaixo- Microtimyobia (Acariformes: Myobiidae: Rad- des leucogaster (Musophagidae) from Tanzania. fordia) and their host-parasite relationships with Acarologia, 51 (1): 93–97. cricetid rodents (Cricetidae). Zootaxa, 2954: 158. Bochkov, A.V. and Skirnisson, K. 2011. Descrip- 1–86. tion of the life stages of quill mite Mironovia 148. Bochkov, A.V. 2011b. [Evolution of parasitism in lagopus sp. nov. (Acari: Syringophilidae) parasit- mammal-associated mites of the group Psoroptidia izing the rock ptarmigan Lagopus muta (Pha- (Acari: Astigmata).] Parazitologiya, 45 (4): 257– sianidae) from Iceland. Parasitology Research, 272. [In Russian with English summary, also trans- 108: 715–722. lated in English: Evolution of parasitism in mam- 159. Bochkov, A.V., Skoracki, M., Hendricks, S.A. mal-associated mites of the Psoroptidia group and Spicer, G.S. 2011. Further investigations of (Acari: Astigmata), Entomological Review, 91 (9): the mite genus Syringophiloidus (Acariformes: 1206–1215.] Syringophilidae) from North American Passer- 149. Bochkov, A. V. 2011c. [Acariform mites perma- ines. Systematic Parasitology, 79: 201–211. nent parasites of mammals: phylogeny, system- 160. Skoracki, M., Bochkov, A.V. and OConnor, B.M. atic and host-parasite interactions]. Doctor of 2011. Notes on quill mites (Acariformes, Syrin- Sciences thesis. Zoological Institute RAS, Saint- gophilidae) from aquatic birds in North America. Petersburg. 783 pp. [Manuscript, in Russian] Acta Parasitologica, 56 (3): 325–330. 150. Bochkov, A.V., Abramov, A.V., Durden, L.A., 161. Zhang, Z.-Q., Fan, Q.-H., Pesic, V., Smit, H., Apanaskevich, D.A., Stekolnikov, A.A., Stanyu- Bochkov, A.V., Khaustov, A.A., Baker, A., Wohlt- kovich, M.K., Gnophanxay, S. and Tikhonov, mann, A., Wen, T., Amrine, J.W., Beron, P., Lin, A.N. 2011. symbiotes of Laonastes J., Gabrys, G. and Husband, R. 2011. Order aenigmamus (Rodentia: Diatomyidae). Journal Trombidiformes Reuter, 1909. In: Z.-Q. Zhang of Parasitology, 97 (2): 352–353. (Ed.). Animal biodiversity: an outline of higher-

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level classification and survey of taxonomic rich- tologiya, 47(3): 212–227. [In Russian with ness. Zootaxa, 3148: 129–138. English summary; also published in English in Entomological Review, 93 (9): 1196–1206] 2012 172. Bochkov, A.V. and OConnor, B.M. 2013. Mites 162. Bochkov, A.V. 2012. Macropodicoptes mironovi of the Subfamily Harpirhynchinae (Acariformes: n.g., n.sp. (Acariformes: ), a new Harpirhynchidae) from North American Birds. mange mite from Wallabia bicolor (Desmarest) Journal of Medical Entomology, 50 (3): 501–509. (Diprotodonia: Macropodidae). Systematic Para- 173. *Bochkov, A.V., OConnor, B.M. and Grootaert, sitology, 83 (1): 29–38. P. 2013. Revision of the family Listropsoralgidae 163. Bochkov, A.V. 2012. Schizocarpus saveljevi sp. Fain, 1965 (Acariformes: Sarcoptoidea)—skin nov. (Acariformes: Chirodiscidae) parasitizing parasites of marsupials and rodents. Zootaxa, the Eurasian beaver—Castor fiber Linnaeus, 1758 3611: 1–69. (Rodentia: Castoridae) from Leningrad Province 174. Skoracki, M., Glowska, E. and Bochkov, A.V. (Russia). Proceedings of the Zoological Institute 2013. Phylogeny of quill mites of the family RAS, 316 (2): 166–171. Syringophilidae (Acari: Prostigmata) based on 164. Bochkov, A.V. and Mironov, S.V. 2012. Mites their external morphology. European Journal of of the genus Passeroptes Fain (Acariformes: Entomology, 110 (4): 663–675. Dermationidae) from passerines (Aves: Pas- seriformes) of North-Western Russia. Zootaxa, 2014 3563: 43–57. 175. Bochkov A.V., Barreto M. 2014. New species of 165. Bochkov, A.V. and Skoracki, M. 2012. New and the subgenus Radfordia (Hesperomyobia) (Acari- poorly known predaceous cheyletids (Acari- formes, Myobiidae)—parasites of the subfamily formes: Cheyletidae) from feather quills. Inter- Sigmodontinae (Rodentia: Cricetidae: Sigmodon- national Journal of Acarology, 38 (6): 486–496. tinae). International Journal of Acarology, 40 (1): 166. Bochkov, A.V., Labrzycka, A., Skoracki M. and 37–42. Saveljev, A.P. 2012. Fur mites of the genus 176. Bochkov, A., Lareschi, M. and Barreto, M. 2014. Schizocarpus Trouessart (Acari: Chirodiscidae) New species and records of the mite genus Pro- parasitizing the Eurasian beaver Castor fiber listrophorus (Acariformes: Listrophoridae) from belorussicus Lavrov (Rodentia: Castoridae) in rodents of the subfamily Sigmodontinae (Roden- NE Poland (Suwałki). Zootaxa, 3152: 39–59. tia: Cricetidae). Acta Parasitologica, 59 (4): 167. Literak, I., Bochkov A.V., Cárdenas-Callirgos, J. 586–595. and Capek, M. 2012. The first records of mites 177. Bochkov, A.V. 2014. Two new species of the of the genus Neharpyrhynchus (Acariformes: subfamily Harpirhynchinae (Acari: Harpirhyn- Harpirhynchidae) from birds in Peru. Neotropical chidae) from African birds. Annales Zoologici, Helminthology, 6 (1): 109–114. 64: 479–483. 168. Sikora, B. and Bochkov, A.V. 2012. Fur mites of 178. Bochkov, A.V. and Abramov, A.V. 2014. Acari- the family Listrophoridae (Acariformes: Sarcop- form mites (Acariformes)—permanent symbionts toidea) associated with South American sigmo- of Hapalomys delacouri Thomas (Rodentia, dontine rodents (Cricetidae: Sigmodontinae). Muridae) in Vietnam. ZooKeys, 459: 137–145. Acta Parasitologica, 57 (4): 388–396. 179. Bochkov, A.V. and Grootaert, P. 2014. Mites of 169. Skoracki, M., Zabludovskaya, S. and Bochkov, the genus Paracoroptes Lavoipierre, 1955 (Acari- A.V. 2012. A review of Prostigmata (Acariformes: formes: Psoroptidae)—skin parasites of the Af- Trombidiformes) permanently associated with rican monkeys of the family Cercopithecidae birds. Acarina, 20 (2): 67–107. (Primates). Zootaxa, 3887 (2): 225–238. 180. Bochkov, A.V. and Guzman-Cornejo, C. 2014. 2013 Mites of the subgenus Neotomobia n.subg. 170. Bochkov, A.V. and Galloway, T.D. 2013. New (Acariformes: Myobiidae: Radfordia), parasites records and new species of mites of the subfam- of the subfamily Neotominae (Rodentia: Criceti- ily Harpirhynchinae (Acariformes: Harpirhynchi- dae). Systematic Parasitology, 89 (2): 91–100. dae) infesting birds in Manitoba, Canada. Acta 181. Bochkov, A.V. and Klompen, H. 2014. A review Parasitologica, 58 (4): 405–419. of the subfamily Harpypalpinae Fain, 1972 171. Bochkov, A.V. and Mironov S.V. 2013. [Is para- (Acariformes: Harpirhynchidae)—parasites of sitism of Metazoa “a one-way ticket?”] Parazi- passerine birds. Zootaxa, 3857 (4): 451–477.

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182. Bochkov, A.V. and Klompen, H. 2014. A review Dubinin, 1957 (Acariformes: Harpirhynchidae), of the zumpti species group of the genus Harpy- parasites of the passerine birds (Aves: Passeriformes) rhynchoides (Acariformes: Harpirhynchidae)— in Australia and South Asia. Systematic Parasitol- ectoparasites of passerines. Zootaxa, 3884 (5): ogy, 92: 23–29. DOI: 10.1007/s11230-015-9 401–418. 193. Bochkov, A.V. and Mirolubov, A.A. 2015. Descrip- 183. Bochkov, A.V. and Klompen, H. 2014. New tion of postembrionic immature stages of Protomyo- Harpirhynchinae Dubinin (Acariformes: Harpi- bia onoi Jameson & Dusbabek, 1971 (Acariformes: rhynchidae)—intracutaneous and feather-base Myobiidae) and comparative analysis of the juvenile parasites of birds. Zootaxa, 3860 (4): 301–324. external morphology in the tribe Protomyobiini. 184. Bochkov, A.V. and OConnor, B.M. 2014. Mites International Journal of Acarology, 41: 115–127. of the genus Trichorhynchiella (Acariformes: DOI: 10.1080/01647954.2014.1003966 Harpirhynchidae): A new example of ovovivipar- 194. Bochkov, A.V. and Mironov, S.V. 2015. A new ity in acariform mites. Journal of Medical Ento- species of the genus Neodermation Fain (Acari- mology, 51 (2): 323–330. formes: Dermationidae) and the first record of the 185. Bochkov, A.V. and Saveljev, A.P. 2014. Mites of precopulatory mate-guarding among dermatio- the genus Schizocarpus Trouessart, 1896 (Acari- nids. Systematic and Applied Acarology, 20: formes: Chirodiscidae) from the North American 612–628. DOI: 10.11158/saa.20.6.4 beavers (Castor canadensis) in Russia. Parazi- 195. Bochkov, A.V., Klompen, H., Junge, R.E. and tologiya, 48 (6): 430–436. [In Russian with Wiliams, C.V. 2015. Lemuralges propithecus English summary] sp. n. (Acariformes: Psoroptidae), an ectoparasite 186. Bochkov, A.V., Klimov, P.B. and Grootaert, P. 2014. of the diademed sifaka Propithecus diadema Revision of the subfamily Onychalginae Fain, 1988 (Primates: Indriidae). Folia Parasitologica, 62: (Acariformes: Pyroglyphidae)—ectoparasites of 11. DOI: 10.14411/fp.2015.011 passerine birds. Zootaxa, 3785(2): 175–200. 196. *Bochkov, A.V., OConnor, B.M. and Klompen, 187. Bochkov, A.V., Klimov, P.B., Hestvik, G. and H. 2015. A review of the mite subfamily Harpi- Saveljev, A.P. 2014. Integrated Bayesian species rhynchinae (Acariformes: Harpirhynchidae)— delimitation and morphological diagnostics of parasites of New World birds (Aves: Neognathae). chorioptic mange mites (Acariformes: Psoropti- Zootaxa, 4023: 1–130. DOI: 10.11646/zoo- dae: Chorioptes). Parasitology Research, 113 (7): taxa.4023.1.1. 2603–2627. 197. Hernandes, F.A., Pedroso, L.G.A. and Bochkov, 188. Literák, I, Bochkov, A. and Franco, S. 2014. The A.V. 2015. Carcinopodacarus polymorphus first record of mites of the genus Neharpyrhyn- gen. n. et sp. n. from Guira guira (Cuculiformes: chus (Acariformes: Harpirhynchidae) from birds Cuculidae) in Brazil: a first example of male in Paraguay. Neotropical Helminthology, 8 (1): polymorphism in the family Dermationidae 183–188. (Acariformes: Analgoidea). Folia Parasitologica, 189. Mertins, J.W. and Bochkov, A.V. 2014. Key to 62: 9. DOI: 10.14411/fp.2015.009. species of the genus Neocheyletiella (Acari- 198. Klimov, P.B., Bochkov, A.V. and OConnor, B.M. formes: Cheyletidae), with description of a new 2015a. Phylogenetic position of the house dust species. Journal of Medical Entomology, 51 (6): mite subfamily Guatemalichinae (Acariformes: 1116–1121. Pyroglyphidae) based on integrated molecular and morphological analyses and different mea- 2015 sures of support. Cladistics, 32 (3): 261–275. 190. Bochkov, A.V. 2015. Comparison of external DOI: 10.1111/cla.12126 morphology of tritonymphs in myobiid mites 199. Klimov, P.B., Bochkov, A.V. and OConnor, B.M. (Myobiidae) associated with marsupials (Marsu- 2015b. Tachornithoglyphus gen. nov.—a new pialia). International Journal of Acarology, 41: genus of nidicolous Pyroglyphidae (Acariformes: 590–599. DOI: 10.1080/01647954.2015.1084046 Astigmata). Zootaxa, 3956: 97–112. DOI: 191. Bochkov, A.V. 2015. Mites of the genus Harpy- 10.11646/zootaxa.3956.1.5 rhynchiella Fain, 1972 (Acariformes: Harpirhyn- 2016 chidae), an example of an unusual mating strat- egy. International Journal of Acarology, 41: 200. Bochkov, A.V. 2016. A new mite species Schizo- 31–40. DOI: 10.1080/01647954.2014.988644 coptes daberti sp. n. (Acariformes: Chirodiscidae) 192. Bochkov, A.V. and Klompen, H. 2015. Two new from Chrysochloris stuhlmanni Matsche (Af- mite species of the subfamily Harpirhynchinae rosoricida: Chrysochloridae) in the Democratic

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Republic of the Congo. Folia Parasitologica, 63: 210. Bochkov A.V., OConnor B.M., Skoracki, M. 2017. 005. DOI: 10.14411/fp.2016.005 New species and new records of mites of the fam- 201. Bochkov, A.V. 2016. Myocoptid mites (Acari- ily Myocoptidae (Acariformes: Sarcoptoidea) from formes: Myocoptidae) of the fauna of the former African rodents. Journal of Medical Entomology, USSR. Zootaxa, 4193 (3): 451–485. DOI: 54 (3): 587–596. 10.11646/zootaxa.4193.3.2 211. Bochkov, A.V. 2017. A review of fur-mites of the 202. Bochkov, A.V. and Abramov, V.V. 2016. To genus Dubininetta Fain & Lukoschus, 1978 (Acari- fauna of the free-living Cheyletidae (Acari- formes: Listrophoridae), ectoparasites of moles formes: Cheyletoidea) of the European part of (Soricomorpha: Talpidae). Systematic Parasitology, Russia. Systematic and Applied Acarology, 21 94: 635–647. DOI: 10.1007/s11230-017-9732-6 (3): 335–346. DOI: 10.11158/saa.21.3.8 212. Bochkov, A.V. 2017. Crocidurobia lukoschusi 203. Bochkov, A.V. and Sidorchuk, E.A. 2016. A new sp.nov. (Acariformes: Myobiidae) parasitizing Eocene free-living cheyletid mite from Baltic Crocidura tanakae Kuroda, 1938 (Soricimorpha: amber. Acta Palaeontologica Polonica, 61 (4): Soricidae) in Vietnam, the first record of the genus 869–874. DOI: 10.4202/app.00244.2016 from oriental white-toothed shrews. Interna- 204. Bochkov, A.V. and Valim, M.P. 2016. New spe- tional Journal of Acarology, 43: 212–216. DOI: cies and records of mites of the superfamily 10.1080/01647954.2016.1268204 Sarcoptoidea (Acariformes: Psoroptidia) from 213. Bochkov, A.V. 2017. To the systematics of the mammals in Brazil. Acta Parasitologica, 61 (1): subgenus Hesperomyobia Bochkov, 1997 (Acari- 22–41. DOI: 10.1515/ap-2016-0003 formes: Myobiidae: Radfordia). Systematic and 205. Bochkov, A.V., Mbalitini, S.G. and Verheyen, E. Applied Acarology, 22 (6): 785–788. DOI: 2016. Crocidurobia faini sp. n. (Acariformes: 10.11158/saa.22.6.4. Myobiidae)—a new mite species parasitizing 214. Bochkov, A.V. and OConnor, B.M. 2017. A new shrews of the genus Crocidura (Soricimorpha: species of myocoptid mites Myocoptes (Comyo- Soricidae) in DR Congo. Systematic Parasitology, coptes) nesokia sp.nov. (Acariformes: Myocop- 93: 493–499. DOI: 10.1007/s11230-016-9623-2 tidae) parasitizing Nesokia indica (Rodentia: 206. Bochkov, A.V., Mbalitini, S.G. and Verheyen, E. Muridae) in Pakistan. Acta Parasitologica, 62: 2016. Review of the fur-mite genus Soricilichus 46–49. DOI: 10.1515/ap-2017-0005 Fain, 1970 (Acariformes: Chirodiscidae)—sym- 2018 bionts of the African shrews of the subfamily Crocidurinae (Soricomorpha: Soricidae). Zoo- 215. Bochkov A.V. 2018. External morphology of taxa, 4072 (2): 235–253. DOI: 10.11646/zoo- postembrionic stages of Lutrilichus javanicus taxa.4072.2.6 (Acariformes: Chirodiscidae) from Melogale 207. Bochkov, A.V., Valim, M.P., Ochoa, R., OConnor, moschata (Carnivora: Mustelidae) from Vietnam. B.M. and Averianov, A.O. 2016. Origin of host- Acarologia, 58 (4): 763–772. DOI: 10.24349/ parasite associations of Marsupialges misonnei acarologia/20184271 (Acariformes: Psoroptidae)—a parasitological 216. Bochkov A.V. 2018. Gaudalges averianovi detective story. Parasitology Research, 115 (10): sp.nov. (Acariformes: Psoroptidae)—a new 3695–3698. DOI: 10.1007/s00436-016-5192-1 parasite of Malagasy lemurs. Systematic and Ap- 208. Bochkov, A.V. 2016. [Chapter IV. Host-parasite plied Acarology, 23 (4): 593–596. DOI http://doi. relationships of acariform mites (Acariformes), org/10.11158/saa.23.4.1 permanent parasites of mammals]. In: K.V. Galak- 217. Bochkov, A. V., Klimov, P. B., Da-Hee, K. and tionov (Ed.). [Coevolution of parasites and hosts]. Skoracki, M. 2018. Validation of the status of a Proceedings of the Zoological Institute RAS, Sup- species with high CO1 and low nuclear genetic plement 4: 160–263. https://www.zin.ru/journals/ divergences: the scab mite Caparinia ictonyctis trudyzin/eng/supplements.html?year=2016 [In stat. res. (Acariformes: Psoroptidae) parasitizing Russian with English summary]. the African hedgehog Atelerix albiventris. Zootaxa, 4544 (4): 523–547. DOI: 10.11646/zoo- 2017 taxa.4544.4.4 209. Bochkov, A.V., Klimov, P.B. and Skoracki, M. 2019 2017. Morphological phylogenetic conflict in the parasitic mite family Harpirhynchidae (Acari- 218. Klimov, P. B., Skoracki, M. and Bochkov, A. V. formes: Cheyletoidea) correlates with host as- 2019. Cox1 barcoding versus multilocus species sociations. Zoologischer Anzeiger, 271: 33–48. delimitation: validation of two mite species with

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contrasting effective population sizes. Parasites Carcinopodacarus Hernandes, Pedroso et Boch- and Vectors, 12 (1): 8. DOI: 10.1186/s13071-018- kov, 2015 (Dermationidae) 3242-5 Castosyringophilus Bochkov et Perez, 2002 219. Sidorchuk, E. A., Bochkov, A. V., Weiterschan, (Syringophilidae) T. and Chernova, O. F. 2019. A case of mite-on- Charadriaulobia Bochkov, Fain et Skoracki, mammal ectoparasitism from Eocene Baltic 2004 (Syringophilidae) amber (Acari: Prostigmata: Myobiidae and Mam- Charadriphilus Bochkov et Chistyakov, 2001 malia: Erinaceomorpha). Journal of Systematic (Syringophilidae) Paleontology, 17 (4): 331–347. DOI: 10.1080/­ Corvitorotroglus Skoracki et Bochkov, 2010 14772019.2017.1414889 (Syringophilidae) Crassacarus Bochkov, OConnor et Klompen, *—monographs. 2015 (Harpirhynchidae) Cygnocoptes Fain et Bochkov, 2003 (Pyro- glyphidae; presently in Psoroptoididae) Didelphialges Bochkov, OConnor et Grootaert, LIST OF MITE TAXA ESTABLISHED 2013 (Listropsoralgidae) BY A.V. BOCHKOV Fainharpirhynchus Bochkov et Galloway, Andrei Bochkov personally and with co-au- 2013 (Harpirhynchidae) thors established 7 family-group names (3 sub- Fritschisyringophilus lonchurae Bochkov, Fain families and 4 tribes), 45 genus-group names et Skoracki, 2004 (Syringophilidae) (34 genera and 11 subgenera) and 445 species- Granulocheyletus Fain et Bochkov, 2002 group names (1 fossil). (Cheyletidae) Hemimyalges Mironov, Bochkov et Fain, 2005 Family-group names (Epidermoptidae) Subfamilies Hylomysobia Bochkov et OConnor, 2006 Apocnemidocoptinae Mironov, Bochkov et (Myobiidae) Fain, 2005 (Dermationidae) Juxtalaelaps Dowling, Bochkov et OConnor, Otocoptoidinae Fain et Bochkov, 2001 (Der- 2007 (Laelapidae) mationidae) Kenyacheylus Fain et Bochkov, 2001 (Chey- Protomyobiinae Bochkov, 1997 (Myobiidae) letidae) Macropodicoptes Bochkov, 2012 (Sarcoptidae) Tribes Megasyringophilus Fain, Bochkov et Mironov, Acanthophthiriini Bochkov, 1997 (Myobiidae) 2000 (Syringophilidae) Elephantulobiini Bochkov, 1997 (Myobiidae) Neoaulobia Fain, Bochkov et Mironov, 2000 Phyllostomonyssini Bochkov, Zabludovskaya (Syringophilidae) et OConnor, 2008 (Gastronyssidae) Nidocheyletus Bochkov et OConnor, 2010 Protomyobiini Bochkov, 1997 (Myobiidae) (Cheyletidae) Oconnoricheylus Bochkov et Otto, 2010 Genus-group names (Cheyletidae) Genera Otocoptoides Fain et Bochkov, 2001 (Epider- Apocalypsis Bochkov, 2010 (Myocoptidae), moptidae) nom. preocc., Rothschild 1903 (Lepidoptera, Phalarophilus Skoracki, Bochkov et OConnor, Sphingidae) 2011 (Syringophilidae) Apocalyptoides Bochkov, 2016 (Myocoptidae), Picocheyletus Bochkov et OConnor, 2003 nom. nov. pro Apocalypsis Bochkov, 2010 (Cheyletidae) Archemyialges Mironov, Bochkov et Fain, Protohylomysobia Sidorchuk et Bochkov, 2018 2005 (Epidermoptidae) (2019) (Myobiidae) Caenolestolichus Fain et Bochkov, 2003 (Atopo- Pseudoopsonyssus Bochkov, Zabludovskaya melidae) et OConnor, 2008 (Gastronyssidae) Caenolestomyobia Bochkov et OConnor, 2009 Psittaciphilus Fain, Bochkov et Mironov, 2000 (Myobiidae) (Syringophilidae)

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Tachornithoglyphus Klimov, Bochkov et Psittophagoides brotogeris Fain et Bochkov, OConnor, 2015 (Pyroglyphidae) 2003 Terratosyringophilus Bochkov et Perez, 2002 (Syringophilidae) Family Epidermoptidae Trochiloptes Mironov, Bochkov et Fain, 2005 Otocoptoides mironovi Fain et Bochkov, 2001 (Dermationidae) Subgenera Myialges (Metamicrolichus) pelecani Fain et Acomyobia Bochkov et Fain, 2003 (Myobiidae: Bochkov, 2003 Radfordia) Aztecocheyletus Bochkov et Klimov, 2004 Rallepidermoptes rallicola Fain et Bochkov, (Cheyletidae: Nodele) 2003 Dendromyobia Bochkov, 1997 (Myobiidae: Family Psoroptoididae Radfordia) Dipodomyobia Bochkov, 1997 (Myobiidae: Cygnocoptes prasadi Fain et Bochkov, 2003 Radfordia) [in content of Pyroglyphidae] Emballomyobia Bochkov et Fain, 2003 (Myo- biidae: Ugandobia) Superfamily Psoroptoidea Hesperomyobia Bochkov, 1996 (Myobiidae: Family Lobalgidae Radfordia) Macroscelistrophoroides Fain et Bochkov, Echimytricalges lonchothrix Bochkov et 2003 (Atopomelidae: Listrophoroides) OConnor, 2010 Neotomobia Bochkov et Guzman-Cornejo, Echimytricalges proechimys Bochkov et 2014 (Myobiidae: Radfordia) OConnor, 2010 Otomyobia Bochkov et Fain, 2003 (Myobiidae: Family Psoroptidae Myobia) Petromyscobia Bochkov et Fain, 2003 (Myo- Chorioptes sweatmani Bochkov, Klimov, Hes- biidae: Radfordia) tvik et Saveljev, 2014 Pseudoharpirhynchus Fain, Bochkov et Mironov, 1999 (Harpirhynchidae: Harpirhynchus), Gaudalges averianovi Bochkov, 2018 synonymized with Harpyrhynchoides Fain, 1972 Gaudalges brevisetosus Bochkov et OConnor, in: Bochkov et al. 2015 2006

Species-group names Lemuralges propithecus Bochkov, Klompen, Junge et Wiliams, 2015 ACARIFORMES ASTIGMATA Paracoroptes miopithecus Bochkov et Groot- Superfamily Analgoidea aert, 2014 Family Dermationidae Paracoroptes piliocolobus Bochkov et Groot- aert, 2014 Carcinopodacarus polymorphus Hernandes, Pedroso et Bochkov, 2015 Superfamily Sarcoptoidea Family Atopomelidae Neodermation actitis Bochkov et Mironov, 2015 Atopomelus hylomys Bochkov, Klimov et OCon- Passeroptes ampeliceps Fain et Bochkov, 2003 nor, 2005 Passeroptes hirundiphilus Fain et Bochkov, 2003 Atopomelus megalotis Bochkov, Abramov et Passeroptes hippolais Bochkov et Mironov, 2012 Hugot, 2011 Passeroptes lamprotornis Fain et Bochkov, 2003 Atopomelus priapus Bochkov, Klimov et OCon- Passeroptes oenanthe Fain et Bochkov, 2003 nor, 2005 Passeroptes sylviae Bochkov et Mironov, 2012 Passeroptes turdoides Fain et Bochkov, 2003 Austrochirus (Austrochiroides) dubininae Fain Paradermation colii Fain et Bochkov, 2003 et Bochkov, 2003

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Austrochirus (Austrochirus) peroryctes Fain et Listrophoroides (Listrophoroides) musschen- Bochkov, 2003 broekii Bochkov et OConnor, 2005 Listrophoroides (Listrophoroides) musseri Bathyergolichus cryptomys Fain et Bochkov, 2003 Bochkov et OConnor, 2005 Listrophoroides (Listrophoroides) oconnori Caenolestolichus lukoschusi Fain et Bochkov, Bochkov, Abramov et Hugot, 2011 2003 Listrophoroides (Listrophoroides) parafemo- ratus Bochkov et OConnor, 2005 Didelphoecius micoureus Bochkov et Valim, 2016 Listrophoroides (Listrophoroides) pseudoan- Listrophoroides (Afrolistrophoroides) colomys cophallus Bochkov et OConnor, 2005 Fain et Bochkov, 2003 Listrophoroides (Listrophoroides) pseudorajah Listrophoroides (Afrolistrophoroides) myomys- Bochkov et OConnor, 2005 cus Fain et Bochkov, 2003 Listrophoroides (Listrophoroides) rapit Boch- Listrophoroides (Afrolistrophoroides) paraleg- kov, OConnor et Gorog, 2004 gada Fain et Bochkov, 2003 Listrophoroides (Listrophoroides) surifer Listrophoroides (Afrolistrophoroides) priono- Bochkov et OConnor, 2005 mys Fain et Bochkov, 2005 Listrophoroides (Macroscelistrophoroides) Listrophoroides (Alistrophoroides) aethomys petrodromi Fain et Bochkov, 2003 Fain et Bochkov, 2003 Listrophoroides (Marquesania) apomys Boch- Listrophoroides (Lemurlistrophoroides) lepil- kov et OConnor, 2008 emur Fain et Bochkov, 2003 Listrophoroides (Marquesania) bullimus Boch- Listrophoroides (Listrophoroides) batomys kov et OConnor, 2008 Bochkov et OConnor, 2008 Listrophoroides (Marquesania) corpirzvarosae Listrophoroides (Listrophoroides) bellulus Bochkov et OConnor, 2008 Bochkov et OConnor, 2005 Listrophoroides (Marquesania) faini Bochkov Listrophoroides (Listrophoroides) cambodien- et OConnor, 2008 sis Bochkov et OConnor, 2005 Listrophoroides (Marquesania) heaneyi Boch- Listrophoroides (Listrophoroides) chrootomys kov et OConnor, 2008 Bochkov et OConnor, 2008 Listrophoroides (Marquesania) 1imnomys Listrophoroides (Listrophoroides)crunomys Bochkov et OConnor, 2008 Bochkov et OConnor, 2008 Listrophoroides (Marquesania) luzonicus Listrophoroides (Listrophoroides) femoratus Bochkov et OConnor, 2008 Bochkov et OConnor, 2005 Listrophoroides (Marquesania) melomys Boch- Listrophoroides (Listrophoroides) gorogae kov et Fain, 2003 Bochkov et OConnor, 2005 Listrophoroides (Marquesania) mordax Boch- Listrophoroides (Listrophoroides) helenae kov et Fain, 2003 Bochkov et Mironov, 2001 Listrophoroides (Marquesania) echiothrix Listrophoroides (Listrophoroides) inas Boch- Bochkov, OConnor et Gorog, 2004 kov et OConnor, 2005 Listrophoroides (Marquesania) leneothrix Listrophoroides (Listrophoroides) inflatus Bochkov, OConnor et Gorog, 2004 Bochkov et OConnor, 2005 Listrophoroides (Marquesania) paracucullatus Listrophoroides (Listrophoroides) insu1aris Bochkov et OConnor, 2008 Bochkov et OConnor, 2008 Listrophoroides (Marquesania) pseudolativen- Listrophoroides (Listrophoroides) khmeri tris Bochkov, OConnor et Gorog, 2004 Bochkov et OConnor, 2005 Listrophoroides (Olistrophoroides) oenomiph- Listrophoroides (Listrophoroides) maxomys ilus Fain et Bochkov, 2003 Bochkov et OConnor, 2005 Listrophoroides (Olistrophoroides) thallomys Listrophoroides (Listrophoroides) megaphallus Fain et Bochkov, 2003 Bochkov et OConnor, 2005 Family Chirodiscidae Listrophoroides (Listrophoroides) moi Boch- kov et OConnor, 2005 Schizocarpus anomalus Bochkov, 1993

134 In memoriam of Andrei V. Bochkov

Schizocarpus azasicus Bochkov et Saveljev, 2012 Opsonyssus macroglossus Bochkov et OCon- Schizocarpus daberti Bochkov et Saveljev, 2012 nor, 2008 Schizocarpus heatherae Bochkov, Labrzycka, Opsonyssus pseudoindicus Bochkov et OCon- Skoracki et Saveljev, 2012 nor, 2008 Schizocarpus heideckei Bochkov et Saveljev, 2012 Opsonyssus pteropodi Bochkov et OConnor, Schizocarpus faini Bochkov, Labrzycka, Skoracki 2008 et Saveljev, 2012 Schizocarpus gozdziewskii Bochkov, Labrzyc- Rodhainyssus saccopteryx Bochkov et OCon- ka, Skoracki et Saveljev, 2012 nor, 2008 Schizocarpus klompeni Bochkov, Labrzycka, Skoracki et Saveljev, 2012 Yunkeracarus apomys Bochkov et OConnor, Schizocarpus lavrovi Bochkov et Saveljev, 2012 2008 Schizocarpus parahumilis Bochkov, Labrzyc- Yunkeracarus hylomyscus Bochkov et OCon- ka, Skoracki et Saveljev, 2012 nor, 2008 Schizocarpus pseudonumerosus Bochkov, Yunkeracarus limnomys Bochkov et OConnor, Labrzycka, Skoracki et Saveljev, 2012 2008 Schizocarpus saveljevi Bochkov, 2012 Yunkeracarus lophuromys Bochkov et OCon- Schizocarpus testiculatus Bochkov, Labrzycka, nor, 2008 Skoracki et Saveljev, 2012 Yunkeracarus otomys Bochkov et OConnor, Schizocarpus tuvinicus Bochkov et Saveljev, 2012 2008 Schizocarpus unzhakovi Bochkov et Saveljev, Yunkeracarus rattus Bochkov et OConnor, 2012 2008 Schizocarpus zurowskii Bochkov, Labrzycka, Skoracki et Saveljev, 2012 Family Listrophoridae Schizocoptes daberti Bochkov, 2016 Aeromychirus fimbriatus Bochkov et OConnor, Soricilichus sylvisorex Bochkov, Mbaltini et 2005 Verheyen, 2016 Aeromychirus petinomys Bochkov et OConnor, 2006 Family Chirorhynchobiidae Afrolistrophorus (Afrolistrophorus) laonastes Chirorhynchobia glossophaga Bochkov, Klom- Bochkov, Abramov et Hugot, 2011 pen et OConnor, 2008 Afrolistrophorus (Afrolistrophorus) hapalomys Bochkov et Abramov, 2014 Family Dromiciocoptidae Afrolistrophorus (Afrolistrophorus) saxatilo- Dromiciocoptes caenolestes Bochkov et OCon- mys Bochkov, Abramov et Hugot, 2011 nor, 2008 Dromiciocoptes marmosops Bochkov et OCon- Amlistrophorus geoxus Sikora et Bochkov, 2012 nor, 2008 Asiochirus nepalensis Fain et Bochkov, 2003 Asiochirus soriculus Fain et Bochkov, 2003 Family Gastronyssidae Dubininetta abramovi Bochkov, 2017 Eidolonyssus myersi Bochkov et OConnor, 2008 Dubininetta euroscaptor Bochkov, 2017 Gastronyssus philippinensis Bochkov et OCon- nor, 2008 Hemigalichus chrotogale Bochkov et OCon- nor, 2005 Mycteronyssus haplonycteri Bochkov et OCon- Lynxacarus (Lutracarus) visoni Fain et Boch- nor, 2008 kov, 2002 Opsonyssus klompeni Bochkov et OConnor, Lynxacarus (Lynxacarus) serrafreirei Bochkov 2008 et Valim, 2016

135 S.V. Mironov et al.

Prolistrophorus (Aprolistrophorus) parabiden- PROSTIGMATA tatus Bochkov, Lareschi et Barreto, 2014 Superfamily Cheyletoidea Prolistrophorus (Aprolistrophorus) tylomys Family Cheyletidae Bochkov, Lareschi et Barreto, 2014 Prolistrophorus (Prolistrophorus) hylaeamys Apodicheles collocalia Bochkov et OConnor, 2003 Bochkov et Valim, 2016 Bak indonesiensis Bochkov et Otto, 2010 Pteromychirus menetes Bochkov, Abramov et Bak nadchatrami Fain et Bochkov, 2001 Hugot, 2011 Chelacaropsis kenyensis Fain et Bochkov, 2001 Family Listropsoralgidae Chelacheles aigamuxa Bochkov et OConnor, 2004 Didelphialges metachirus Bochkov, OConnor Chelacheles algericus Fain et Bochkov, 2001 et Grootaert, 2013 Chelacheles apanaskevichi Bochkov et Abramov, 2016 Listropsoralges brevisetosus Bochkov, OCon- Chelacheles bacchusi Bochkov, Khaustov et nor et Grootaert, 2013 Kuznetzov, 1999 Listropsoralges caenolestes Bochkov, OCon- Chelacheles indra Bochkov et Otto, 2010 nor et Grootaert, 2013 Chelacheles klimovi Bochkov et OConnor, 2004 Listropsoralges caluromys Bochkov et Wau- Chelacheles seminole Bochkov et OConnor, 2004 thy, 2009 Chelacheles temoak Bochkov et OConnor, 2004 Listropsoralges faini Bochkov et Wauthy, 2009 Chelacheles thomasi Bochkov et Otto, 2010 Listropsoralges similis Bochkov, OConnor et Grootaert, 2013 Cheletomimus (Hemicheyletia) ascutatus Fain et Bochkov, 2002 Listropsoralges thylamys Bochkov, OConnor Cheletomimus (Hemicheyletia) crinitus Bochkov et Grootaert, 2013 et Sidorchuk, 2016 Listropsoralges vossi Bochkov, OConnor et Cheletomimus (Hemicheyletia) gracilis Fain et Grootaert, 2013 Bochkov, 2002 Cheletomimus (Hemicheyletia) greenwoodi Fain Family Myocoptidae et Bochkov, 2002 Myocoptes (Comyocoptes) nesokia Bochkov et Cheletomimus (Hemicheyletia) ochoai Fain et OConnor, 2017 Bochkov, 2002 Myocoptes (Myocoptes) lophuromys Bochkov, Cheletomimus (Philippicheyla) crowei Bochkov OConnor et Scoracki, 2016 (2017)** et Otto, 2010 Myocoptes (Myocoptes) meriones Bochkov, 2016 Myocoptes (Myocoptes) pitymys Fain et Boch- Cheletophyes vespae Fain et Bochkov, 2001 kov, 2004 Cheletophyes decorus Bochkov et Klimov, 2005 Cheletophyes mbomba Bochkov, Klimov et Trichoecius calomysci Bochkov, Malikov et OConnor, 2006 Arbobi, 1999 Cheletophyes occisor Bochkov et Klimov, 2005 Trichoecius dubininae Bochkov, 2016 Cheletophyes torridae Bochkov, Klimov et OCon- Trichoecius faini Bochkov, OConnor et nor, 2006 Skoracki, 2016 (2017)** Trichoecius hylomyscus Bochkov, OConnor et Cheletopsis charadrii Mironov, Bochkov et Chi- Skoracki, 2016 (2017)** rov, 1991 Cheletopsis daberti Kivganov et Bochkov, 1994 Trichoecius lemmus Bochkov, 2016 Cheletopsis larosterna Bochkov et Skoracki, 2012 Trichoecius meriones Bochkov, 2016 Cheletopsis limnodromi Bochkov, Fain et Dabert, Family Sarcoptidae 2002 Cheletopsis mariae Mironov, Bochkov et Chirov, Macropodicoptes mironovi Bochkov, 2012 1991

136 In memoriam of Andrei V. Bochkov

Cheletopsis prosobonialis Bochkov, Fain et Da- Neocheyletiella parvisetosa Mertins et Boch- bert, 2002 kov, 2014 Cheletopsis rynchops Bochkov, Fain et Dabert, Neocheyletiella queleae Fain et Bochkov, 2002 2002 Cheletopsis thalasseus Bochkov et OConnor, 2003 Nidocheyletus kennedyae Bochkov et OCon- nor, 2004 Cheletosoma tauraco Bochkov et Skoracki, 2012 Nodele (Aztecocheyletus) conquistador Boch- Cheyletus ascutatus Bochkov et OConnor, 2003 kov et Klimov, 2004 Cheyletus batomysi Bochkov et OConnor, 2003 Oconnoricheylus chimaera (Bochkov et Ochoa, Cheyletus kuznetzovi Bochkov et Khaustov, 1999 2005) Cheyletus volgini Fain et Bochkov, 2001 Oconnoricheylus speciosus Bochkov et Otto, 2010

Eucheyletiella daurica Bochkov et Mironov, 1999 Ornithocheyletia columbigallinae Fain et Boch- Eucheyletiella faini Bochkov et Malikov, 1996 kov, 2002 Eucheyletiella pallasius Bochkov et Mironov, Ornithocheyletia mironovi Bochkov et Chirov, 1999 1998 Eucheyletiella pusillinus Bochkov et Mironov, Ornithocheyletia orioli Fain et Bochkov, 2002 1999 Ornithocheyletia prognae Bochkov et Gallo- way, 2001 Galagocheles lawrencei Bochkov, Mironov et Ornithocheyletia turdi Bochkov et Galloway, 2001 OConnor, 2005 Ornithocheyletia wauthyi Fain et Bochkov, 2002

Granulocheyletus corpuzrarosae Fain et Boch- Picocheyletus tricholaemae Bochkov et OCon- kov, 2002 nor, 2003

Hoffmannita gersoni Fain et Bochkov, 2001 Prosocheyla (Prosocheyla) ripkai Fain et Boch- kov, 2001 Kenyacheylus troglodytes Fain et Bochkov, 2001 Prosocheyla (Prosocheyla) villosa Bochkov et Khaustov, 1999 Ker afrotropicalis Fain et Bochkov, 2001 Samsinakia trilobitus Bochkov et Mironov, 1998 Metacheyletia amazonae Bochkov et OCon- Samsinakia tilae Bochkov et Klimov, 2005 nor, 2003 Metacheyletia degenerata Fain et Bochkov, 2003 Sciurocheyla funisciurus Bochkov et OConnor, Metacheyletia ngaii Bochkov et Skoracki, 2011 2004

Mexecheles shiva Bochkov et Ochoa, 2005 Thryonomycheyla angolensis Fain et Bochkov, 2002 Neochelacheles corpuzrarosae Bochkov et OConnor, 2004 Family Cloacaridae Neochelacheles mendicus Bochkov et OCon- Caminacarus dawsoni Bochkov et OConnor, nor, 2004 2008 Neocheyletiella aegithali Bochkov et OConnor, 2003 Family Epymyodicidae Neocheyletiella ardeola Bochkov et Ochoa, Epimyodex soricis Fain et Bochkov, 2001 2005 Neocheyletiella athene Fain et Bochkov, 2002 Family Syringophilidae Neocheyletiella lonchurae Fain et Bochkov, 2002 Neocheyletiella nectarinia Bochkov et OCon- Aulobia galinago Bochkov et Mironov, 1998 nor, 2003 Aulobia sylviae Bochkov et Mironov, 1998

137 S.V. Mironov et al.

Aulonastus prunellae Bochkov et Mironov, 1999 Philoxanthornia dubinini Bochkov et Mironov, 1998 Castosyringophilus forpi Bochkov et Perez, 2002 Picobia alectoris Fain, Bochkov et Mironov, 2000 Corvitorotroglus alpha Skoracki et Bochkov, 2010 Picobia biarmicus Skoracki, Bochkov et Wau- thy, 2004 Bubophilus asiobius Skoracki et Bochkov, 2002 Picobia brotogeris Fain, Bochkov et Mironov, 2000 Charadriphilus ralli Skoracki et Bochkov, 2010 Picobia chloris Bochkov, Mironov et Kravtso- Charadriaulobia vanelli Bochkov, Fain et va, 2000 Skoracki, 2004 Picobia cissa Skoracki, Bochkov et Wauthy, Charadriphilus lyudmilae Bochkov et Chistya- 2004 kov, 2001 Picobia locustella Skoracki, Bochkov et Wau- thy, 2004 Chenophila kanduli Bochkov, 1998 Picobia phoeniculi Fain, Bochkov et Mironov, 2000 Fritschisyringophilus lonchurae Bochkov, Fain Picobia ramphastos Fain, Bochkov et Mironov, et Skoracki, 2004 2000 Picobia sturni Skoracki, Bochkov et Wauthy, Megasyringophilus cyanocephala Fain, Boch- 2004 kov et Mironov, 2000 Megasyringophilus dubinini Bochkov et Fain, Psittaciphilus amazonae Fain, Bochkov et 2003 Mironov, 2000 Megasyringophilus kethleyi Fain, Bochkov et Psittaciphilus fritschi Fain, Bochkov et Mironov, Mironov, 2000 2000 Megasyringophilus platycercus Bochkov et Fain, 2003 Stibarokris langei Bochkov et Mironov, 1999 Megasyringophilus rhynchopsitta Bochkov et Perez, 2002 Syringophiloidus agelaius Bochkov, Skoracki, Megasyringophilus trichoglossus Fain, Boch- Hendricks et Spicer, 2011 kov et Mironov, 2000 Syringophiloidus carpodaci Bochkov et Ap- anaskevich, 2001 Mironovia coturnae Bochkov, Fain et Skoracki, Syringophiloidus cypsiuri Fain, Bochkov et 2004 Mironov, 2000 Mironovia lagopus Bochkov et Skirnisson, 2011 Syringophiloidus daberti Bochkov Bochkov, Fain et Skoracki, 2004 Neoaulobia agapornis Fain, Bochkov et Mironov, Syringophiloidus delichorum Bochkov, 2001 2000 Syringophiloidus dendrocittae Fain, Bochkov Neoaulobia aratingae Fain, Bochkov et Mironov, et Mironov, 2000 2000 Syringophiloidus graculae Fain, Bochkov et Neoaulobia mexicana Bochkov et Perez, 2002 Mironov, 2000 Neoaulobia mironovi Bochkov et Perez, 2002 Syringophiloidus jackowiaki Bochkov, Skoracki, Neoaulobia psittaculae Fain, Bochkov et Mironov, Hendricks et Spicer, 2011 2000 Syringophiloidus klimovi Skoracki et Bochkov, 2010 Niglarobia calidridis Bochkov et Mironov, 1998 Syringophiloidus motacillae Bochkov et Mironov, Niglarobia chirovi Bochkov et Mironov, 1998 1998 Niglarobia rhinoptili Fain, Bochkov et Mironov, Syringophiloidus serini Bochkov, Fain et Skoracki, 2000 2004 Niglarobia skorackii Bochkov et Galloway, 2004 Syringophiloidus xanthocephalus Bochkov, Skoracki, Hendricks et Spicer, 2011 Phalarophilus fulicarius Skoracki, Bochkov et Syringophiloidus zonotrichia Bochkov, Skoracki, OConnor, 2011 Hendricks et Spicer, 2011

138 In memoriam of Andrei V. Bochkov

Syringophilopsis aegythali Bochkov, Mironov Anharpyrhynchus lukoschusi Bochkov et et Skoracki, 2001 Klompen, 2014 Syringophilopsis borini Bochkov et Mironov, 1999 Crassacarus alexfaini Bochkov, OConnor et Syringophilopsis dendroicae Bochkov et Gal- Klompen, 2015 loway, 2001 Crassacarus fritschi Bochkov, OConnor et Syringophilopsis emberizae Fain, Bochkov et Klompen, 2015 Mironov, 2000 Crassacarus melanerpes Bochkov, OConnor Syringophilopsis faini Bochkov et Apanaskev- et Klompen, 2015 ich, 2001 Crassacarus sialia Bochkov, OConnor et Syringophilopsis garrulaxi Bochkov, Mironov Klompen, 2015 et Skoracki, 2001 Crassacarus tinae Bochkov, OConnor et Klom- Syringophilopsis icteri Bochkov et Galloway, pen, 2015 2001 Fainharpirhynchus contopus Bochkov et Gal- Syringophilopsis kirgizorum Bochkov, Mironov loway, 2013 et Kravtsova, 2000 Fainharpirhynchus legatus Bochkov et Klom- Syringophilopsis phylloscopi Bochkov, Mironov pen, 2014 et Skoracki, 2001 Fainharpirhynchus mossi Bochkov et Klom- Syringophilopsis porzanae Bochkov et Gal- pen, 2014 loway, 2004 Syringophilopsis stachyris Bochkov, Mironov Harpirhynchus dusbabeki Bochkov et Literak, et Skoracki, 2001 2006 Syringophilopsis sylviettae Fain, Bochkov et Harpirhynchus galleridae Fain, Bochkov et Mironov, 2000 Mironov, 1999 Syringophilopsis tyrrani Bochkov et Galloway, Harpirhynchus quasimodo Bochkov et Mer- 2004 tins, 2010

Syringophilus numidae Bochkov, 2000 Harpyrhynchoides accipiter Bochkov et OCon- nor, 2013 Terratosyringophilus lorricinus Bochkov et Harpyrhynchoides actitis Bochkov, OConnor Fain, 2002 et Klompen, 2015 Terratosyringophilus pioni Bochkov et Perez, Harpyrhynchoides aegolius Bochkov et OCon- 2002 nor, 2013 Harpyrhynchoides alaudinus Bochkov, 2000 Torotrogla aphelocoma Bochkov, Flannery et Harpyrhynchoides artamus Bochkov et Klom- Spicer, 2009 pen, 2015 Torotrogla cardinalis Bochkov, Flannery et Harpyrhynchoides athene Bochkov, OConnor Spicer, 2009 et Klompen, 2015 Torotrogla cardueli Bochkov et Mironov, 1999 Harpyrhynchoides botaurus Bochkov et Gal- Torotrogla coccothraustes Bochkov, Flannery loway, 2013 et Spicer, 2009 Harpyrhynchoides bubulcus Bochkov, OCon- Torotrogla cyanocitta Bochkov, Flannery et nor et Klompen, 2015 Spicer, 2009 Harpyrhynchoides calidris Bochkov, OConnor Torotrogla gaudi Bochkov et Mironov, 1998 et Klompen, 2015 Harpyrhynchoides charadrius Bochkov, Torotrogla piranga Bochkov, Flannery et OConnor et Klompen, 2015 Spicer, 2009 Harpyrhynchoides clamator Bochkov, 2014 Harpyrhynchoides coccyzus Bochkov, OCon- Family Harpirhynchidae nor et Klompen, 2015 Anharpyrhynchus apodus Bochkov et Klom- Harpyrhynchoides crotophaga Bochkov, OCon- pen, 2014 nor et Klompen, 2015 Anharpyrhynchus elizae Bochkov et Klompen, Harpyrhynchoides heatherae Bochkov et Gal- 2014 loway, 2013

139 S.V. Mironov et al.

Harpyrhynchoides gallowayi Bochkov, OCon- Neharpyrhynchus bombycilla Bochkov, OCon- nor et Klompen, 2015 nor et Klompen, 2015 Harpyrhynchoides ixobrychus Bochkov, OCon- Neharpyrhynchus campylorhynchus Bochkov, nor et Klompen, 2015 OConnor et Klompen, 2015 Harpyrhynchoides kirgizorum Fain, Bochkov Neharpyrhynchus chlorospingus Bochkov et et Mironov, 1999, syn. of H. zumpti (Fain, 1972) Literak, 2011 Harpyrhynchoides megascops Bochkov, OCon- Neharpyrhynchus domrowi Bochkov et Klom- nor et Klompen, 2015 pen, 2015 Harpyrhynchoides parazumpti Fain, Bochkov Neharpyrhynchus hippolae Bochkov, 2000 et Mironov, 1999 Neharpyrhynchus icterus Bochkov et OConnor, Harpyrhynchoides pluvialis Bochkov, OCon- 2013 nor et Klompen, 2015 Neharpyrhynchus loxia Bochkov et Galloway, Harpyrhynchoides setophaga Bochkov et Klom- 2013 pen, 2014 Neharpyrhynchus mironovi Bochkov et Liter- Harpyrhynchoides spizella Bochkov et Klom- ak, 2011 pen, 2014 Neharpyrhynchus oenanthe Bochkov, 2014 Harpyrhynchoides vulgaris Bochkov et Gal- Neharpyrhynchus picidarum Bochkov, OCon- loway, 2004 nor et Klompen, 2015 Harpyrhynchoides xanthocephalus Bochkov Neharpyrhynchus quiscalus Bochkov, OCon- et Klompen, 2014 nor et Klompen, 2015 Harpyrhynchoides zenaida Bochkov, OConnor Neharpyrhynchus spizella Bochkov, OConnor et Klompen, 2015 et Klompen, 2015 Neharpyrhynchus tangara Bochkov et Literak, Harpypalpus lonchura Bochkov et Klompen, 2011 2014 Harpypalpus pyrrhula Bochkov et Klompen, 2014 Neharpyrhynchus turdus Bochkov, OConnor Harpypalpus spermestes Fain, Bochkov et et Klompen, 2015 Mironov, 1999 Neharpyrhynchus vireo Bochkov, Oconnor et Harpypalpus sturnus Bochkov et Klompen, Klompen, 2015 2014 Perharpyrhynchus caprimulgus Bochkov et Harpypalpus taeniopygia Bochkov et Klom- Klompen, 2014 pen, 2014 Perharpyrhynchus charadrius Bochkov, OCon- Harpypalpoides hirundinoides Bochkov et nor et Klompen, 2015 Klompen, 2014 Perharpyrhynchus elseyornis Bochkov et Klom- Harpypalpoides johnstoni Bochkov et Klom- pen, 2014 pen, 2014 Harpypalpoides lesickii Skoracki, Bochkov et Trichorhynchiella myiarchus Bochkov et Klom- Sikora, 2004 pen, 2014 Harpypalpoides namibiensis Fain, Bochkov et Trichorhynchiella phleocryptes Bochkov et Mironov, 1999 OConnor, 2014 Harpypalpoides regulus Bochkov et Klompen, Superfamily Myobioidea 2014 Family Myobiidae Harpypalpoides sitta Bochkov et Klompen, 2014 Caenolestomyobia faini Bochkov et OConnor, Harpypalpoides sylvia Bochkov et Klompen, 2009 2014 Caenolestomyobia lukoschusi Bochkov et OCon- nor, 2009 Harpyrhynchiella apus Bochkov, 2014 Crocidurobia (Crocidurobia) dusbabeki Bochkov Neharpyrhynchus agelaius Bochkov, OConnor et Fain, 2003 et Klompen, 2015 Crocidurobia (Crocidurobia) lasiurae Bochkov, Neharpyrhynchus baile Bochkov et Literak, 2007 1997

140 In memoriam of Andrei V. Bochkov

Crocidurobia (Crocidurobia) leucodoni Boch- Radfordia (Hesperomyobia) rhipidomys Boch- kov, 1997 kov et Barreto, 2014 Crocidurobia (Crocidurobia) lukoschusi Boch- Radfordia (Microtimyobia) abramovi Bochkov kov, 2017 et Mironov, 1998 Radfordia (Microtimyobia) alticolae Bochkov, Eudusbabekia (Eudusbabekia) rhinophylla 1995 Fain et Bochkov, 2002 Radfordia (Microtimyobia) cricetuliphila Bochkov, 1999 Ewingana (Ewingana) teniotis Bochkov et Radfordia (Microtimyobia) dinaromys Boch- Rybin, 1992 kov, 2011 Hylomysobia chinensis Bochkov et OConnor, Radfordia (Microtimyobia) golenishchevi 2006 Bochkov, 2011 Hylomysobia mikhailzaitzevi Bochkov et OCon- Radfordia (Microtimyobia) myopusi Bochkov nor, 2006 et Mironov, 1998 Radfordia (Microtimyobia) pitymys Bochkov, †Protohylomysobia erinaceophilus Sidorchuk 2011 et Bochkov, 2018 (2019)** Radfordia (Microtimyobia) rufocani Bochkov, 1995 Protomyobia (Protomyobia) caecutiens Boch- Radfordia (Microtimyobia) stekolnikovi Bochkov kov, 1994 et Mironov, 1998, syn. of R. lemnina (Koch, 1841) Protomyobia (Protomyobia) jamesoni Boch- Radfordia (Microtimyobia) stenocrani Bochkov kov, 1994 et Mironov, 1998, syn. of R. lemnina (Koch, 1841) Radfordia (Austromyobia) dubinini Bochkov, Radfordia (Microtimyobia) synaptomysi Boch- Dubinina et Chirov, 1990 kov et Mironov, 1998 Radfordia (Austromyobia) merioni Bochkov, Radfordia (Neotomobia) megadontomys Boch- Dubinina et Chirov, 1990 kov et Guzman-Cornejo, 2014 Radfordia (Austromyobia) persica Bochkov, Radfordia (Neotomobia) onychomys Bochkov Arbobi et Malikov, 2000 et Guzman-Cornejo, 2014 Radfordia (Dipodomyobia) alactaguli Boch- Radfordia (Neotomobia) peromyscus Bochkov kov, 1997 et Guzman-Cornejo, 2014 Radfordia (Dipodomyobia) allactodipi Boch- Radfordia (Radfordia) berylmysi Bochkov et kov, 1997 Fain, 1997 Radfordia (Dipodomyobia) baranovae Boch- Radfordia (Radfordia) colomys Bochkov et kov, 1997 Fain, 2003 Radfordia (Dipodomyobia) dipi Bochkov, 1997 Radfordia (Radfordia) delectori Bochkov et Radfordia (Dipodomyobia) majori Bochkov, 1997 Fain, 2003 Radfordia (Dipodomyobia) paradipi Bochkov, Radfordia (Radfordia) dephomys Bochkov et 1997 Fain, 2003 Radfordia (Dipodomyobia) paralactaga Boch- Radfordia (Radfordia) eremici Fain et Boch- kov, 1997 kov, 2002 Radfordia (Dipodomyobia) pygeretmusi Boch- Radfordia (Radfordia) lukoschusi Bochkov et kov, 1997 Fain, 1997 Radfordia (Dipodomyobia) stylodipi Bochkov, Radfordia (Radfordia) mirabilis Bochkov et 1997 Abramov, 2014 Radfordia (Graphiurobia) myomimusi Boch- Radfordia (Radfordia) mironovi Bochkov, kov, 1994 1997 Radfordia (Graphiurobia) selevinia Bochkov, Radfordia (Radfordia) myomysci Bochkov et 2009 Fain, 2003 Radfordia (Hesperomyobia) akodon Bochkov, Radfordia (Radfordia) niviventris Bochkov et 2017 Fain, 1997 Radfordia (Hesperomyobia) burbanoae Boch- Radfordia (Radfordia) sigmomys Bochkov et kov et Barreto, 2014 Fain, 2002

141 S.V. Mironov et al.

Superfamily Paratydeoidea Genus-group names Family Stigmocheylidae Andrebochkovia Khaustov et Frolov, 2019 Stigmocheylus americanus Bochkov, 2008 gen. n. (Prostigmata, Neopygmephoridae), present Stigmocheylus oberon Bochkov, 2008 issue Bochkovlaster Khaustov, 2019 gen. n. (Prostig- Superfamily Pomerantzioidea mata, Pygmephoridae), present issue Family Pomerantziidae Bochkovocoptes OConnor, 2019 gen. n. (Astig- mata, Bochkovocoptidae), present issue Pomerantzia philippina Bochkov et Walter, 2007 Species-group names Superfamily Pterygosomatoidea Allogalumna bochkovi Ermilov et Starý, 2018 Family Pterygosomatidae (, Galumnidae) Geckobia anocellatus Bochkov et Mironov, Calcealges bochkovi Dabert, 2019 sp. n. (Astig- 2000 mata, Trouessartiidae), present issue Geckobia hirsti Bochkov et Mironov, 2000 Cunaxa bochkovi Kaustov et Kuznetzov, 1998 (Prostigmata, Cunaxidae) Pterygosoma pseudotrapelus Bochkov, Mel- Eustigmaeus bochkovi Khaustov 2019 (Prostig- nikov et Nazarov, 2009 mata, Stigmaeidae) Fringillosphaera bochkovi Mironov, 2001 (Astig- mata, Apionacaridae) MESOSTIGMATA Gaeolaelaps bochkovi Jocharchi, Khaustov et Superfamily Dermanyssoidea Ermilov, 2019 sp. n. (Mesostigmata, Laelapidae), Family Laelapidae present issue Galliphilopsis bochkovi Skoracki et Sikora, Andreacarus Dowling, Bochkov 2004 (Prostigmata, Syringophilidae) et OConnor, 2007 bochkovi Apanaskevich et Andreacarus Dowling, Bochkov et Tomlinson, 2019 (Ixodida, ) OConnor, 2007 Hypozetes andreii Ermilov, Hugo-Coetzee, Khaus- Andreacarus galidia Dowling, Bochkov et tov et Kontschán, 2019 sp. n. (Oribatida, Tegoribatidae), OConnor, 2007 present issue Andreacarus gymnuromys Dowling, Bochkov Ipsoglyphus bochkovi Klimov et Khaustov, et OConnor, 2007 2018 (Astigmata, ) Andreacarus Dowling, Bochkov et bochkovi Stekolnikov, 2013 OConnor, 2007 (Astigmata, ) Andreacarus tenrec Dowling, Bochkov et Meristacarus bochkovi Ermilov et Kalúz, 2019 OConnor, 2007 sp. n. (Oribatida, Lohmanniidae), present issue Andreacarus Dowling, Bochkov et Neharpyrhynchus bochkovi Martinu, Dusba- OConnor, 2007 bek et Literak, 2008 (Prostigmata, Harpyrhyn- chidae) **—printed version appeared the next year after online Neoeucheyla bochkovi Fain, 2003 (Prostig- version; †—fossil species. mata, Cheyletidae) Proctophyllodes bochkovi Mironov 2019 sp. n. (Astigmata, ), present issue MITE TAXA NAMED IN HONOR Ramusella (Dosangoppia) bochkovi Ermilov OF A.V. BOCHKOV et Frolov, 2019 (Oribatida, Oppiidae) Syringophilopsis bochkovi Skoracki, Mironov et Bermúdez, 2019 sp. n. (Prostigmata, Syringo- Family-group names philidae), present issue Bochkovocoptidae OConnor, 2019 fam.n. (Astig- Trouessartia bochkovi Mironov et Galloway mata, Sarcoptoidea), present issue 2019 (Astigmata, Trouessartiidae)

142 In memoriam of Andrei V. Bochkov

Table 1. Mite taxa described by A.V. Bochkov and coauthors.

Family Subfamily Tribe Genus Subgenus Species Total ASTIGMATA Analgoidea 1 Dermationidae 2 2 11 15 2 Epidermoptidae 3 3 6 3 Psoroptoididae 1 1 2 4 Pyroglyphidae 1 1 Psoroptoidea 5 Lobalgidae 2 2 6 Psoroptidae 6 6 Sarcoptoidea 7 Atopomelidae 1 1 53 55 8 Chirodiscidae 18 18 9 Chirorhynchobiidae 1 1 10 Dromiciocoptidae 2 2 11 Gastronyssidae 1 1 14 16 12 Listrophoridae 17 17 13 Listropsoralgidae 1 8 9 14 Myocoptidae 1 10 11 15 Sarcoptidae 1 1 2 PROSTIGMATA Cheyletoidea 16 Cheyletidae 5 1 76 82 17 Cloacaridae 1 1 18 Epymyodicidae 1 1 19 Harpirhynchidae 2 1 74 77 20 Syringophilidae 10 78 88 Myobioidea 21 Myobiidae 1 3 3 8 54 69 Paratydeoidea 22 Stigmocheylidae 2 2 Pomerantzioidea 23 Pomerantziidae 1 1 Pterygosomatoidea 24 Pterygosomatidae 3 3 MESOSTIGMATA Dermanyssoidea 25 Laelapidae 1 7 8 Total 3 4 33 11 445 496

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