Journal of Molluscan Studies Advance Access published 4 December 2013 Journal of The Malacological Society of London Molluscan Studies Journal of Molluscan Studies (2013) 1–11. doi:10.1093/mollus/eyt039 A CONTRIBUTION TO THE PHYLOGENY OF ALBANIAN AGATHYLLA (GASTROPODA, CLAUSILIIDAE): INSIGHTS USING MORPHOLOGICAL DATA AND THREE MITOCHONDRIAL MARKERS ZOLTA´ NFEHE´ R1, ARISTEIDIS PARMAKELIS2, MARIA KOUTALIANOU2, 2 3 1 THANOS MOURIKIS ,ZOLTA´ NPE´ TER ERO´ ´ SS AND VIRA´ G KRI´ ZSIK Downloaded from 1Hungarian Natural History Museum, 13 Baross utca, H-1088 Budapest, Hungary; 2Department of Ecology and Taxonomy, Faculty of Biology, University of Athens, Panepistimioupoli Zografou, GR-15784 Athens, Greece; and 336 Bem utca, H-1151 Budapest, Hungary Correspondence: Z. Fehe´r; email:
[email protected] http://mollus.oxfordjournals.org/ (Received 16 May 2013; accepted 28 August 2013) ABSTRACT In the present study, we revised Albanian Agathylla, combining the traditional shell morphological ap- proach with a phylogenetic reconstruction based on three mitochondrial DNA markers (COI, COII and 16S rRNA). From the faunistic perspective, Albania was an almost completely unexplored country until the end of the 1990s. Following recent intense field sampling activity in Albania, the number of known Agathylla populations has increased, providing an opportunity to investigate how additional ma- terial can contribute to the resolution of the taxonomy of the Albanian Agathylla species. Newly discov- by guest on December 7, 2013 ered populations indicate greater morphological variability than previously recorded and highlight the presence of various other character state combinations. Based on these findings, we argue that only two Agathylla taxa occur in the region, namely A. neutra and A.