Man, His Origin and Destiny by Joseph Fielding Smith
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Early Eocene Fossils Suggest That the Mammalian Order Perissodactyla Originated in India
ARTICLE Received 7 Jul 2014 | Accepted 15 Oct 2014 | Published 20 Nov 2014 DOI: 10.1038/ncomms6570 Early Eocene fossils suggest that the mammalian order Perissodactyla originated in India Kenneth D. Rose1, Luke T. Holbrook2, Rajendra S. Rana3, Kishor Kumar4, Katrina E. Jones1, Heather E. Ahrens1, Pieter Missiaen5, Ashok Sahni6 & Thierry Smith7 Cambaytheres (Cambaytherium, Nakusia and Kalitherium) are recently discovered early Eocene placental mammals from the Indo–Pakistan region. They have been assigned to either Perissodactyla (the clade including horses, tapirs and rhinos, which is a member of the superorder Laurasiatheria) or Anthracobunidae, an obscure family that has been variously considered artiodactyls or perissodactyls, but most recently placed at the base of Proboscidea or of Tethytheria (Proboscidea þ Sirenia, superorder Afrotheria). Here we report new dental, cranial and postcranial fossils of Cambaytherium, from the Cambay Shale Formation, Gujarat, India (B54.5 Myr). These fossils demonstrate that cambaytheres occupy a pivotal position as the sister taxon of Perissodactyla, thereby providing insight on the phylogenetic and biogeographic origin of Perissodactyla. The presence of the sister group of perissodactyls in western India near or before the time of collision suggests that Perissodactyla may have originated on the Indian Plate during its final drift toward Asia. 1 Center for Functional Anatomy & Evolution, Johns Hopkins University School of Medicine, 1830 E. Monument Street, Baltimore, Maryland 21205, USA. 2 Department of Biological Sciences, Rowan University, Glassboro, New Jersey 08028, USA. 3 Department of Geology, H.N.B. Garhwal University, Srinagar 246175, Uttarakhand, India. 4 Wadia Institute of Himalayan Geology, Dehradun 248001, Uttarakhand, India. 5 Research Unit Palaeontology, Ghent University, Krijgslaan 281-S8, B-9000 Ghent, Belgium. -
The Lost World of Fossil Lake
Snapshots from Deep Time THE LOST WORLD of FOSSIL LAKE lance grande With photography by Lance Grande and John Weinstein The University of Chicago Press | Chicago and London Ray-Finned Fishes ( Superclass Actinopterygii) The vast majority of fossils that have been mined from the FBM over the last century and a half have been fossil ray-finned fishes, or actinopterygians. Literally millions of complete fossil ray-finned fish skeletons have been excavated from the FBM, the majority of which have been recovered in the last 30 years because of a post- 1970s boom in the number of commercial fossil operations. Almost all vertebrate fossils in the FBM are actinopterygian fishes, with perhaps 1 out of 2,500 being a stingray and 1 out of every 5,000 to 10,000 being a tetrapod. Some actinopterygian groups are still poorly understood be- cause of their great diversity. One such group is the spiny-rayed suborder Percoidei with over 3,200 living species (including perch, bass, sunfishes, and thousands of other species with pointed spines in their fins). Until the living percoid species are better known, ac- curate classification of the FBM percoids (†Mioplosus, †Priscacara, †Hypsiprisca, and undescribed percoid genera) will be unsatisfac- tory. 107 Length measurements given here for actinopterygians were made from the tip of the snout to the very end of the tail fin (= total length). The FBM actinop- terygian fishes presented below are as follows: Paddlefishes (Order Acipenseriformes, Family Polyodontidae) Paddlefishes are relatively rare in the FBM, represented by the species †Cros- sopholis magnicaudatus (fig. 48). †Crossopholis has a very long snout region, or “paddle.” Living paddlefishes are sometimes called “spoonbills,” “spoonies,” or even “spoonbill catfish.” The last of those common names is misleading because paddlefishes are not closely related to catfishes and are instead close relatives of sturgeons. -
Article XXXVIII. -NOTE on SOME WORM (?) BUR- ROWS in ROCKS of the CHEMUNG GROUP of NEW YORK
Article XXXVIII. -NOTE ON SOME WORM (?) BUR- ROWS IN ROCKS OF THE CHEMUNG GROUP OF NEW YORK. By R. P. WHITFIELD. Arenicolites chemungensis, sp. nov. PLATE XIV, FIGS. I AND 2. While working up the fossils of the Potsdam sandstone for the Wisconsin Report, in I876 and I877, there came into my hands a number of specimens representing the so-called Scolithus, which I described as Arenicolites woodi in Volume IV of Prof. T. C. Chamberlain's Report of I882. One block of that series showed the original surface of the mud-covered sandstone with the burrows of the worm (?) which made the perforations, together with the little hillocks surrounding the outlet of the burrows, just as the animal built them up by its castings during life; proving pretty conclusively that it must have been a marine worm-like animal which caused the perforations. Among the geological specimens of the Chemung Group in the Museum, from near Bath, Steuben Co., New York, I find an example so nearly resembling that figured in the Wis- consin Report above referred to, that there can be no ques- tion as to the similarity of its origin. On this Chemung specimen the hillocks are somewhat larger and the funnels more distinct, being generally 5 or 6 mm. in diameter, and the walls surrounding them about 2 mm. thick, while some of the hillocks are much larger and higher and appear to have collapsed from the semifluidity of the sand, closing up the top of the burrow so as to show a mere slit in its place. -
Mammalian Faunal Change During the Early Eocene Climatic
MAMMALIAN FAUNAL CHANGE DURING THE EARLY EOCENE CLIMATIC OPTIMUM (WASATCHIAN AND BRIDGERIAN) AT RAVEN RIDGE IN THE NORTHEASTERN UINTA BASIN, COLORADO AND UTAH by ALEXANDER R. DUTCHAK B.Sc., University of Alberta, 2002 M.Sc., University of Alberta, 2005 A thesis submitted to the Faculty of the Graduate School of the University of Colorado in partial fulfillment of the requirements for the degree of Doctor of Philosophy Department of Geological Sciences 2010 This thesis entitled: Mammalian faunal change during the Early Eocene Climatic Optimum (Wasatchian and Bridgerian) at Raven Ridge in the northeastern Uinta Basin, Colorado and Utah written by Alexander R. Dutchak has been approved for the Department of Geological Sciences ______________________________ Jaelyn J. Eberle (Supervisor) ______________________________ John Humphrey ______________________________ Mary Kraus ______________________________ Tom Marchitto ______________________________ Richard Stucky Date: ________________________ The final copy of this thesis has been examined by the signatories, and we find that both the content and the form meet acceptable presentation standards of scholarly work in the above mentioned discipline. ABSTRACT Dutchak, Alexander R. (Ph.D., Geological Sciences, Department of Geological Sciences) Mammalian faunal change during the Early Eocene Climatic Optimum (Wasatchian and Bridgerian) at Raven Ridge in the northeastern Uinta Basin, Colorado and Utah Thesis directed by Assistant Professor Jaelyn J. Eberle This project investigated patterns of mammalian faunal change at Raven Ridge, which straddles the Colorado-Utah border on the northeastern edge of the Uinta Basin and consists of intertonguing units of the fluvial Colton and lacustrine Green River Formations. Fossil vertebrate localities comprising >9,000 fossil mammal specimens from 62 genera in 34 families were identified and described. -
Furry Folk: Synapsids and Mammals
FURRY FOLK: SYNAPSIDS AND MAMMALS Of all the great transitions between major structural grades within vertebrates, the transition from basal amniotes to basal mammals is represented by the most complete and continuous fossil record, extending from the Middle Pennsylvanian to the Late Triassic and spanning some 75 to 100 million years. —James Hopson, “Synapsid evolution and the radiation of non-eutherian mammals,” 1994 At the very beginning of their history, amniotes split into two lineages, the synapsids and the reptiles. Traditionally, the earliest synapsids have been called the “mammal-like reptiles,” but this is a misnomer. The earliest synapsids had nothing to do with reptiles as the term is normally used (referring to the living reptiles and their extinct relatives). Early synapsids are “reptilian” only in the sense that they initially retained a lot of primitive amniote characters. Part of the reason for the persistence of this archaic usage is the precladistic view that the synapsids are descended from “anapsid” reptiles, so they are also reptiles. In fact, a lot of the “anapsids” of the Carboniferous, such as Hylonomus, which once had been postulated as ancestral to synapsids, are actually derived members of the diapsids (Gauthier, 1994). Furthermore, the earliest reptiles (Westlothiana from the Early Carboniferous) and the earliest synapsids (Protoclepsydrops from the Early Carboniferous and Archaeothyris from the Middle Carboniferous) are equally ancient, showing that their lineages diverged at the beginning of the Carboniferous, rather than synapsids evolving from the “anapsids.” For all these reasons, it is no longer appropriate to use the term “mammal-like reptiles.” If one must use a nontaxonomic term, “protomammals” is a alternative with no misleading phylogenetic implications. -
Order PERISSODACTYLA – Equids, Rhinoceroses, Tapirs
Order PERISSODACTYLA Order PERISSODACTYLA – Equids, Rhinoceroses, Tapirs Perissodactyla Owen, 1848. Quarterly Journal of the Geological Society of London 4: 103–141. Upper toothrows in altungulate Radinskya (late Paleocene) and Hyracotherium (Eocene). Tentative phylogenetic tree of Perissodactyla after Beninda-Emonds, 2007. Equidae (1 genus, 4 species) Asses, Zebras p. xx Rhinocerotidae (2 genera, 2 Rhinoceroses p. xx for true horses. North America became the centre of evolution of species) true horses, which occasionally migrated to other continents. The The perissodactyls are the order of herbivorous ‘odd-toed’ hoofed descendants of Protorohippus (once called Hyracotherium; Froehlich mammals that includes the living horses, zebras, asses, tapirs, 2002) evolved into many different lineages living side by side. The rhinoceroses and their extinct relatives. They were originally named collie-sized three-toed horses Mesohippus and Miohippus (from beds by Richard Owen (1848) as a group including horses, rhinos, tapirs dated about 30–37 mya) were once believed to be sequential segments and hyraxes, although no recent authors have accepted the inclusion on the unbranched trunk of the horse evolutionary tree. However, of hyraxes in Perissodactyla. Perissodactyls are recognized by a number they coexisted for millions of years, with five different species of two of unique specializations (Hooker 2005), but their single most diagnostic genera living at the same time and place. From Miohippus-like ancestors, feature is the structure of their feet. Most perissodactyls have either horses diversified into many different ecological niches. One major one or three toes on each foot, and the axis of symmetry of the foot lineage, the anchitherines, retained low-crowned teeth, presumably runs through the middle digit. -
0520237315.Pdf
Beasts of Eden The publisher gratefully acknowledges the generous contribution to this book provided by the General Endowment Fund of the University of California Press Associates. Beasts of Eden walking whales, dawn horses, and other enigmas of mammal evolution David Rains Wallace university of california press berkeley los angeles london University of California Press Berkeley and Los Angeles, California University of California Press, Ltd. London, England © 2004 by the Regents of the University of California Library of Congress Cataloging-in-Publication Data Wallace, David Rains, 1945–. Beasts of Eden : walking whales, dawn horses, and other enigmas of mammal evolution / David Rains Wallace. p. cm. Includes bibliographical references and index. isbn 0-520-23731-5 (alk. paper). 1. Mammals—Evolution. 2. Mammals, Fossil. I. Title. ql708.5.w25 2004 599.138—dc22 2003022857 Manufactured in the United States of America 13 12 11 10 09 08 07 06 05 04 10987654 321 Printed on Ecobook 50 containing a minimum 50% post-consumer waste, processed chlorine free. The balance contains virgin pulp, including 25% Forest Stewardship Council Certified for no old growth tree cutting, processed either tcf or ecf. The sheet is acid-free and meets the minimum requirements of ansi/niso z39.48-1992 (r 1997) (Permanence of Paper). ᭺ϱ to steve fisher, who brings art to science Actually most scientific problems are far better understood by studying their history than their logic. ernst mayr quoted in Leo F. Laporte, George Gaylord Simpson: Paleontologist and Evolutionist contents list of illustrations /xi acknowledgments / xiii prologue. the fresco and the fossil /xv 1 / Pachyderms in the Catacombs /1 2 / Dr. -
Stratigraphy, Mammalian Paleontology, Paleoecology, and Age Correlation of the Wasatch Formation, Fossil Butte National Monument, Wyoming
Journal of Paleontology, 90(5), 2016, p. 981–1011 Copyright © 2016, The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.100 Stratigraphy, mammalian paleontology, paleoecology, and age correlation of the Wasatch Formation, Fossil Butte National Monument, Wyoming Gregg F. Gunnell,1 John-Paul Zonneveld,2 and William S. Bartels3 1Division of Fossil Primates, Duke University Lemur Center, Durham, North Carolina 27705, USA 〈[email protected]〉 2Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, AB, T8N 7B2, Canada 〈[email protected]〉 3Department of Geological Sciences, Albion College, Albion, MI 49224, USA 〈[email protected]〉 Abstract.—Fieldwork conducted in the Wasatch Formation in and around Fossil Butte has yielded a diverse assemblage of early Eocene vertebrates. Fossil vertebrates are distributed through three discrete stratigraphic intervals within the uppermost 180 m of the main body of the Wasatch Formation underlying the Green River Formation. These assemblages were derived primarily from fluvial overbank mudstone units overprinted with variably well-developed paleosols. The lowest (20 m) and highest (60 m) sections are characterized by less mature and more hydromorphic paleosols, whereas the middle section (100 m) is typified by more mature paleosols and more abundant channel sandstones. The combined assemblages contain at least 46 species of mammals. Faunal characteristics include high abundances of equid perissodactyls and a relatively high abundance and diversity of notharctines primates, an apparent absence of omomyid primates, relatively high rodent diversity, and relatively diverse and abundant artiodactyls. -
Stable Isotopes in Early Eocene Mammals As Indicators of Forest Canopy Structure and Resource Partitioning
Paleobiology, 34(2), 2008, pp. 282–300 Stable isotopes in early Eocene mammals as indicators of forest canopy structure and resource partitioning Ross Secord, Scott L. Wing, and Amy Chew Abstract.—The three dimensional structure of vegetation is an important component of ecosystems, yet it is difficult to reconstruct from the fossil record. Forests or woodlands prevailed at mid-lati- tudes in North America during the early Eocene but tree spacing and canopy structure are uncer- tain. Here we use stable carbon isotope values (␦13C ) in early Eocene mammalian faunas to infer canopy structure. We compare ␦13C values in two diverse fossil assemblages from the central Big- horn Basin to values predicted for mammals in a variety of open and closed habitats, based on modern floras and faunas. We conclude that these early Eocene faunas occupied an open canopy forest. We also use carbon and oxygen isotopes to infer diet and microhabitat. Three higher level taxa have significantly different mean ␦13C values, and values are negatively correlated with body mass. The pattern suggests diets high in leaves for larger mammals, and fruit or other non-foliar plant organs for small ones. A preference in the larger mammals for wetter habitats with high water availability to plants may also have contributed to the pattern. Ross Secord.* Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Post Office Box 37012, NHB MRC 121, Washington, D.C. 20013-7012 Scott L. Wing. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Post Office Box 37012, NHB MRC 121, Washington, D.C. -
CONOCIMIENTO~ ~OBRE Ei ORIGEN DEL G~NERO ((EQUU~"
ANGEL CABRERA ESTADO ACTUAL Ill' LOS CONOCIMIENTO~ ~OBRE Ei ORIGEN DEL G~NERO ((EQUU~" De Ja Ih:TlsTA us I.A FACVLTAI> DE AGlto~oyiA y VETEnl~AnJA. , en l rega J, tomo YllI .. marzo de 1935 BUENOS AIRES )MPItENTA DE LA UNIVERSID..\D - 1935 -----=:---- ------.!.-..~ .:;:. . -_ :L . ~ -- = Estad o nctual de los conoc imientos sobre el origcn del genero Eouus POR ANGEL CAlH\E IL\ -; "= - ....':1 . Nadic qu e posen ciertas nociones de pal eontologi a pu ede ign orar qu e In familia de los equidos es uno de los pocos grupos zoologicos cu )'a filogenia se ha conseguido llegar a establecer con bastantes proba .~ . ~ ; :.:... bilidades de aproximacion a la realidad. :\"os engatiarlamos, sin em , ~ " ~ ~ "bargo. si pensasemos que es ta cuestion SE' halla completamente diluci dada, como pareeen darlo a enlender algunos autores; antes al cou trario, todavia quedan en ella muchos puntos obscures. y d e las mas recientes investigacion es se deduce que 1'5 precise modil'i car 110 poco de " :-:: 10 qu e generalmenl e se viene da ndo p Ol' ~-a demostrudo. - .\ nte todo, hahria qucdesecha r de u na vez pOI' todas la idea. lan co l~ rrientc aun ent re hipologos y paleontologos distinguidos, de qu e cl pri :: ., :.:.: -: mer anlecesor conocido de los caballos ac tua les es Ph enacodus 0 al gun otro condilartrn muy an n a este. Cua ndo se descubrio cl primer esqucleto relati vam ente com pleto de Ph enacodus , el ilustre palconto .. logo norteam eri can o Ed mll·d D. Cop e [1882 , p. -
Geologic Resources Inventory Report
National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science Fossil Butte National Monument Geologic Resources Inventory Report Natural Resource Report NPS/NRSS/GRD/NRR—2012/587 ON THE COVER Fossil Butte looms over Fossil Butte National Monument in southwest Wyoming. Today’s semi- arid climate belies an ancient, warm temperate lake ecosystem that existed 50 million years ago. Photograph by Jason Kenworthy (NPS Geologic Resources Division). THIS PAGE The 50 million year old rocks of the Green River Formation are exposed on Fossil Butte (front cover) and Cundick Ridge (this page) within Fossil Butte National Monument. They contain exceptionally well preserved fossils from within and around ancient Fossil Lake, such as these fossil fish. National Park Service photograph Fossil Butte National Monument Geologic Resources Inventory Report Natural Resource Report NPS/NRSS/GRD/NRR—2012/587 National Park Service Geologic Resources Division PO Box 25287 Denver, CO 80225 October 2012 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado publishes a range of reports that address natural resource topics of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Report Series is used to disseminate high-priority, current natural resource management information with managerial application. The series targets a general, diverse audience, and may contain NPS policy considerations or address sensitive issues of management applicability. -
Horns, Tusks, and Flippers: the Evolution of Hoofed Mammals
HORNS, TUSKS, AND FLIPPERS: THE EVOLUTION OF HOOFED MAMMALS Donald R. Prothero Occidental College Los Angeles, California and Robert M. Schoch Boston University Boston, Massachusetts The Johns Hopkins University Press Baltimore and London 5 AMYNODONTS HYNACOOON TS Nyrat:l'Jdon 7r�I EJiflnplt�JN• Figure 14.1. Family tree of rhinocerotoids in North America. (Drawn by C. R. Prothero). 14. Rhinoceroses without Horns "ANCIENT DACIANS" AND SIBERIAN MUMMIES results in 1777 and 1779 he described fo ssilized and mum Before the rise of modern comparative anatomy and mified "large animals of India, "including elephants, rhi paleontology, the giant bones fo und in the earth were a great noceroses, and buffalos [now recognized as extinct Ice Age source of wonder, mystery, and eventually legends, In woolly rhinos and mammoths, and bison]. His most spec Chapter 8, we saw how many of these "giants in the earth," tacular find was a mummy of a woolly rhinoceros, fo und interpreted as gigantic humans, were actually the remains of with its skin intact in the frozen ground on the banks of the mastodonts or mammoths, The remains of fossil rhinoceros Viloui (also spelled Vilyuy) River. To Pallas this was "con es (Fig. 14. 1) were similarly misinterpreted. The horns of vincing proof that it must have been a most violent and most the woolly rhinoceros were thought by Siberians to be the rapid flood which once carried these carcasses toward our claws of gigantic predatory birds and may have been glacial climates, before corruption had time to destroy their responsible for the myth of the griffin (also spelled soft parts." "gryphon").