BOCAS DEL TORO ARCHIPELAGO FIELD REPORT March 23 – April 1, 2018 By Jeri M. Langham

Tranquilo Bay Eco Adventure Lodge was built on Bastimentos Island adjacent to the large Parque Nacional Isla Bastimentos in ’s Bocas del Toro Archipelago. I scouted this location in January 2011 and immediately knew it was a fantastic location for VENT tours. Participants also have opportunities to snorkel, kayak, visit the bat cave, paddleboard, fish and/or swim in the warm Caribbean waters.

Our group of seven participants, two owners, two staff members and me © Jay Viola

An enticing example of what awaits visitors to this marvelous birding paradise can be found in excerpts taken from the Journal I write during every tour and later e-mail to all participants. These are from my 11-page Journal for the March 2018 tour.

After a 45-minute flight from Panama City to Bocas del Toro, we were met by Jay Viola, one of the three owners of Tranquilo Bay Eco Adventure Lodge and soon were loaded on the boat and headed toward Bastimentos Island, home of Tranquilo Bay Eco Adventure Lodge. On the way we picked up Magnificent Frigatebird and Pomarine Jaeger.

We settled into our cabanas and then met on the lodge porch and birded from here, the porch of one of the cabanas on top of the hill, and also along a winding rainforest trail. Top of the list was my first White-tailed Kite for the property. Rufous-tailed Hummingbird was the most common, but we also added Bananaquit, Blue-black Grosbeak, Black-cheeked Woodpecker, Yellow-bellied Elaenia, Red-lored Parrot, White-crowned and Pale-vented pigeons, and male and female Masked and Black-headed tityras.

As we worked our way up to “pineapple hill” and the -bathing pools, we picked up Dot-winged Antwren and Chestnut-sided Warbler. We got brief looks at Golden-collared Manakins and great looks at Shining Honeycreeper and a perched, distant Mississippi Kite, a first in my ten visits. Natalia spotted a close Brown-throated Three-toed Sloth.

Shining Honeycreeper and Brown-throated Three-toed Sloth © Linette Mansberger

Before leaving the dock this morning, we saw a Spotted Eagle Ray. On arrival at the mainland, we saw Royal Terns and a Mangrove Swallow before boarding our van. We picked up Black-bellied Whistling-Duck, Northern Jaçana, Green Ibis and more as we drove. Behind the gas station where we stopped to purchase snacks and use the restrooms, we saw our first Boat-billed Herons, and a Cattle Egret stood up in its nest exposing three bright white eggs.

Green Ibis and Boat-billed Heron © Linette Mansberger

We turned left on Oleoducto Highway and headed toward Chiriquí Grande so we could bird Two Tanks Road. This little unpaved road has almost no traffic. Olive-crowned Yellowthroat, Pied Puffbird, both Black- chested and Brown jays, Plumbeous Kite, Golden-hooded Tanager and many more delighted us here.

Pied Puffbird and Golden-hooded Tanager © Linette Mansberger After we finished eating our picnic lunch, we walked back down to the bridge where we saw Tawny-crested Tanager, Barred Forest-Falcon, Keel- billed Toucan, Yellow-throated Chlorospingus, Emerald Tanager, Long- tailed Tyrant, Cinnamon Becard, and Crimson-collared Tanager.

Keel-billed Toucan © Linette Mansberger Crimson-collared Tanager © Bill Fraser

The ocean water around the islands in the Bocas del Toro Archipelago was very smooth this morning as we cruised the 45 minutes to Almirante. Our unpaved road down to Valle Risco has little to no traffic and descends slowly down the side of the mountain offering scenic views. As

Black-cheeked Woodpecker and Black-capped Pygmy-Tyrant © Linette Mansberger we walked it, Little Tinamou was heard several times and we saw Long- tailed Tyrant, Black-cheeked Woodpecker, Blue-headed Parrots, Black- cowled Oriole, Lineated Woodpecker, Streak-headed Woodcreeper, two White Hawks, Piratic Flycatcher, Black-capped Pygmy-Tyrant (the second smallest in the Americas), and the Blue Jeans Poison Dart Frog.

We ate our picnic lunch at an outdoor restaurant called "El Mirador.” It is up on a ridge along the main highway and permits views of the nearby forest. As we were eating, Jim Kimball suddenly called us out to see what would be thousands upon thousands of Broad-winged Hawks, Swainson’s Hawks, and Turkey Vultures. Estimating the actual number is almost impossible. I simply could not believe the number of these migrating raptors that kept streaming by us in the sky and such a spectacle is what makes me select this particular week for my Bocas del Toro Archipelago tour. They stretched from as far as we could see one way to as far in the other.

Rivers of Broad-winged and Swainson’s hawks streaming north © Linette Mansberger

From here we drove to the road that leads to the town of La Gloria. This is a very beautiful forested area with tall trees and finally we got a Cocoa Woodcreeper to come in to us and we were able to get everyone on the tangle-loving, and thus difficult-to-see, Black-throated Wren. A Squirrel Cuckoo zipped over us but would not return to playback. However, the most desired find here was the calling Black Hawk-Eagle that soared over our heads. It was too fast for a clear photo but we all got to see it. Before departing the dock this morning, I decided to try my Mangrove Cuckoo tape and one immediately flew in to the edge of the mangroves. It was a great way to start our morning! According to our field guide, this species was not known to call while in Panama over the winter. However, I got it to call on the first tour that saw one and every time since then. It was a calm, smooth-as-glass ocean morning as we headed to Isla Popa. As we arrived I called in a few Mangrove Yellow Warblers (now elevated to full species). We had several surprises with one being a fly-over Semiplumbeous Hawk (only the 2nd one in 10 tours here). Two pairs of huge Pale-billed Woodpeckers flew back and forth across the channel for us. We also picked up Slaty-tailed Trogon. Nice was seeing a Needlefish skipping along the surface of the water. However, the most important target bird at Isla Popa is Snowy Cotinga and we were fortunate again this year to see a male, but it was too far away for photos.

Mangrove Cuckoo and Semiplumbeous Hawk © Linette Mansberger

Our visit to Robert and Julie Peters’ Green Acres Chocolate Farm was informative and our walk after the short tour of his chocolate-making trays and building brought us two Green-and-black Poison Dart Frogs, Streak-headed Woodcreeper, nesting Pale-billed Woodpeckers, a male Gartered Trogon, Black-chested Jay, and again the tiny Black-capped Pygmy-Tyrant. Before leaving we were treated to some of the items that they sell.

Clearwing © Linette Mansberger Jeri & Robert and Julie Peters © Jim Kimball

At 5:30 p.m. the entire group met Natalia and me on the top of the six-story tower. It lived up to its reputation with Mealy, Red-lored and Blue- headed parrots flying right over us or very close to us. Some of you stayed long enough to see the Merlin, White-vented Euphonias, and Lesser Swallow-tailed Swifts. Four of you met me at the “T” just before heading down for dinner to see the Short-tailed Nighthawks.

Red-lored Parrot and Great Black Hawk © Linette Mansberger

The big target bird today on the Soropta Canal was the difficult-to- find-elsewhere Nicaraguan Seed-Finch, and we managed to see a close male just after entering the canal. For the first hour or so the activity was great. We had super looks at two American Pygmy Kingfishers but they did not perch for photos. Sometimes there were several being called out at the same time. Some of the other special birds seen today were Peregrine Falcon, Blue Ground-Dove, Black-cowled Oriole, Great Black Hawk, and three species of kingfishers. There were a number of Eastern Kingbird flocks that flew over us as we moved up the canal. We saw a number of Green Iguanas and sloths too.

We returned via the Soropta Canal for our visit to Swan's Cay where Red-billed Tropicbirds and Brown Boobies nest. Once we were 30 yards from Swan’s Cay, we spent about 35 minutes just floating with incredibly beautiful, elegant birds flying around us. Cameras were clicking like mad. I cannot wait to see the best photos. Watching those gorgeous birds gliding by with their long tail feathers swaying in the wind, sometimes so close we could almost touch them, was simply fantastic. Some even landed on their nest cavities. This day on the Soropta Canal along with the visit to Swan’s Cay is always a favorite for my participants and me!

One of forty or so Red-billed Tropicbirds flying around Swan’s Cay © Linette Mansberger

Bocas Del Toro Archipelago Spring 2012 Sample Photos

Hot lips with fruits Brown Booby Golden-collared Manakin Collared Plover Red-capped Manakin

Bat Falcon Orange-bellied Trogon Long-billed Hermit Bronze-tailed Plumeleteer White-whiskered Puffbird Green basilisk

Central American agouti Black Guan Mantled howler monkeys

White-collared Manakin Bicolored Antbird Nicaraguan Seed-Finch Western night monkey Strawberry poison-dart frog

Orange daggerwing Chestnut-capped Brush-Finch Ringed Kingfisher Red-billed Tropicbird

Photoshop and Design by Jeri Langham with photos by Craig Swolgaard (10 ), Chris Conard ( 7 ) and Clara Coen ( 6 ). Bocas Del Toro Archipelago Fall 2012 Sample Photos

Housefly on young iguana Golden orb-weaver Great eurybia Orange nectar bat Jesus Christ lizard

Band-tailed Barbthroat Bullet ants (1+ inch) Keel-billed Toucan Mangroves & arboreal termite colony Northern Jaçana

Flat-footed bug on passionflower Bottlenose dolphins One of six deluxe cabanas

Golden-hooded Tanager Pendent heliconia Squirrel Cuckoo Hoffman’s two-toed sloth Strawberry poison-dart frog

Crab in bat cave King Vulture Tigerwing & Clearwing Jamaican fruit-eating bat

Photoshop and Design by Jeri Langham with photos by Linette Mansberger ( 18 ), Bob Kleiger ( 3 ) & Jim Kimball (1). Bocas Del Toro Archipelago Spring 2013 Sample Photos

Lineated Woodpecker Black-crowned Tityra White-flanked Antwren Green-and-black poison dart frog Red-capped Manakin

Clay-colored Thrush Blue-gray Tanager Roadside Hawk Morpho Spot-crowned Antvireo

Slaty-tailed Trogon Black-cheeked Woodpecker Bare-throated Tiger-Heron Gray-headed Kite Keel-billed Toucan

Boat-billed Heron Collared Plover Blue Dacnis White-whiskered Puffbird Montezuma Oropendola

Brown Booby Golden-collared Manakin Shining and Green honeycreepers

Photoshop and Design by Jeri Langham with photos by Ann Brinly (15), Laurie Dann (7), & Kathy McAleese (2). Bocas Del Toro Archipelago Fall 2013a Sample Photos

Golden-hooded Tanager N. little yellow-eared bat Red brocket deer Chestnut-backed Antbird Black-crowned Antshrike

Black snapper & glass minnows Red-throated Ant-Tanager Bronze-tailed Plumeleteer Green Shrike-Vireo Flag-footed bug

Scaled Pigeon Tiger rat snake with C. American woolly opossum Plumbeous Kites Keel-billed Toucan Fasciated Antshrike

Rufous-tailed Hummingbird Lemon-rumped Tanager Grasshopper Strawberry poison dart frog Pale-billed Woodpecker

Central American woolly opossum Bottlenose dolphins Tawny-capped Euphonia White-collared Swift

Photoshop and Design by Jeri Langham with photos by Kent Van Vuren (21) & Jennifer Green (2). Bocas Del Toro Archipelago Fall 2013b Sample Photos

Hypselonotus interruptus Bullet ant Tortoise beetle Worker caste of army ant Brazilian

Cerata groundstreak Anole Hibiscus Jumping spider Butterflies

Civil bluet damselfly Unknown Spiny-backed orb-weaver Caterpillar Moth

Hairy tarchon caterpillar Blue morpho butterfly Giant red-winged grasshopper Fly Cockroach

Fiddler crab Chola and Fula Jumping spider Cloaked scintillant

Photoshop and Design by Jeri Langham with photos by Kent Van Vuren Bocas Del Toro Archipelago Spring 2014 Sample Photos

Boat-billed Heron Pale-billed Woodpecker Common Bush-Tanager Streak-headed Woodcreeper Electron metalmark

Golden-collared Manakin Kinkajou Brown Noddy Passerini’s Tanager Balsa flowers

Mangrove Cuckoo Hoffman’s two-toed sloth Snowy Cotinga Black-chested Jay Common tent-making bat

Strawberry poison dart frog Nicaraguan Seed-Finch Brown Boobies Black-cheeked Woodpecker Mangrove Black-Hawk

Clearwing butterfly White-collared Seedeater Southern Lapwing (nest) Common Tody-Flycatcher American Pygmy Kingfisher

Photoshop and Design by Jeri Langham with photos by Bill Fraser (22) & Linette Mansberger (3). Bocas Del Toro Archipelago Spring 2015 Sample Photos

Rufous-collared Sparrow Long-tailed Tyrant Mangrove Cuckoo Black-cheeked Woodpecker White-faced capuchin

Montezuma Oropendola Boat-billed Heron White-tipped cicadian Masked Tityra Black-crowned Tityra

Striped Cuckoo Brown-throated three-toed sloth Social Flycatcher White-flanked Antwren Lineated Woodpecker

Red-lored Parrots Red-billed Tropicbird Plumbeous Kite Crowned Woodnymph

Pectoral Sandpiper Elegant Euphonia in nest American Golden-Plover White-crowned Pigeon Moon jelly or jellyfish

Spiny orb-weaver Brown Booby Scarlet-thighed Dacnis Poison dart frog Poison dart frog in bat cave Photoshop and Design by Jeri Langham with photos by Linette Mansberger (27), Marti Ikehara (1) and Ramón Fernández Francés (1). Bocas Del Toro Archipelago Spring 2016 Sample Photos

Cinnamon Becard Slaty Spinetail Blue Jeans Poison Dart Frog Julia butterfly Red-breasted Blackbird

Black-&-white Owl Nicaraguan Seed-Finch Yellow-throated Toucan Mangrove Cuckoo White-cro. Pigeon Black-thighed Grosb.

Tarantula Wasp Snorkeling over a Coral Reef Slaty-tailed Trogon Boat-billed Herons

Cracker Butterfly Mom and baby Brown-throated Three-toed Sloths Snowy Cotinga White-throated Crakes

Crossing the sandbar at low tide Jesus Christ Lizard Flame-colored Tanager Orb-weaver Spider Photoshop and Design by Jeri Langham with photos by Rick Melnicoe (13), Oleta Melnicoe (8), and Natalia Decastro González (2). Bocas Del Toro Archipelago Spring 2017 Sample Photos

Orange-bellied Trogon Blue Jeans Poison Dart Frog Azure-hooded Jay Northern Emerald-Toucanet Red-capped Manakin

Swainson’s Hawk Golden-collared Manakin Tiny part of Swainson’s Hawk kettle lifting off in early morning

Long-tailed Tyrant Brown Boobies Black-capped Pygmy-Tyrant Orchid Scale-crested Pygmy-Tyrant Golden-browed Chloroph.

Bare-throated Tiger-Heron Chestnut-capped Oropendola Lesser Nighthawk White Hawk

Prickle-like Ornate Hawk-Eagle White-tipped Cycadian Alfaro’s Pygmy Squirrel Photoshop by Jeri Langham with photos by Jimmy Stanley (11), Natalia Decastro González (10), and Jan Adams (1).

This FIELD CHECKLIST was created for my 10th trip to Bocas del Toro Archipelago, Panama. It follows the American Ornithological Society’s (AOS’s) Checklist of North and Middle American Birds, 7th edition and incorporates changes through the 58th Supplement.

Since many participants would like to know their chances of seeing particular species, COLUMN I has numbers indicating how many of my previous nine VENT tours have seen each species. If there are two numbers separated by a comma, the first tells the number of previous trips on which it was seen and the second, the number in which it was only heard. This column is designed to give you a realistic idea of your chances of seeing each bird.

COLUMN II shows those species recorded on our 2018 tour, with an “x” representing a species that was seen and an “h” representing a species that was only heard.

PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Tinamous Tinamidae 2 Great Tinamou Tinamus major 3h h 3 Little Tinamou Crypturellus soui 9h h 4 Ducks, Geese & Waterfowl Anatidae 5 Black-bellied Whistling-Duck Dendrocygna autumnalis 8 x 6 Muscovy Duck Cairina moschata 5 x 7 Blue-winged Teal Spatula discors 8 x 8 Northern Shoveler Spatula clypeata 9 American Wigeon Mareca americana 1 x 10 Northern Pintail Anas acuta 11 Ring-necked Duck Aythya collaris 1 12 Lesser Scaup Aythya affinis 3 13 Masked Duck Nomonyx dominicus 2 14 Guans, Chachalacas & Curassows Cracidae 15 Gray-headed Chachalaca Ortalis cinereiceps 9 x 16 Crested Guan Penelope purpurascens 2 h 17 Black Guan Chamaepetes unicolor 3 18 Great Curassow Crax rubra 19 New World Quail Odontophoridae 20 Black-eared Wood-Quail Odontophorus melanotis 21 Grebes Podicipedidae 22 Least Grebe Tachybaptus dominicus 23 Pied-billed Grebe Podilymbus podiceps 5 x 24 Pigeons & Doves Columbidae 25 Rock Pigeon (I) Columba livia 9 x 26 Pale-vented Pigeon Patagioenas cayennensis 9 x 27 Scaled Pigeon Patagioenas speciosa 3,1 x 28 White-crowned Pigeon Patagioenas leucocephala 8 x 29 Band-tailed Pigeon Patagioenas fasciata 1 30 Ruddy Pigeon Patagioenas subvinacea 31 Short-billed Pigeon Patagioenas nigrirostris 8 x 32 Ruddy Ground-Dove Columbina talpacoti 9 x 33 Blue Ground-Dove Claravis pretiosa 8 x 34 Ruddy Quail-Dove Geotrygon montana 2 35 Olive-backed Quail-Dove Leptotrygon veraguensis (n. g.) 36 White-tipped Dove Leptotila verreauxi 2,1 37 Gray-chested Dove Leptotila cassinii 3,1 38 Gray-headed Dove Leptotila plumbeiceps 1 x 39 Buff-fronted Quail-Dove Zentrygon costaricensis (n. g.) 40 Purplish-backed Quail-Dove Zentrygon lawrencii (n. g.) 41 Chiriqui Quail-Dove Zentrygon chiriquensis (n. g.) 42 Mourning Dove Zenaida macroura 43 Cuckoos Cuculidae 44 Squirrel Cuckoo Piaya cayana 9 x 45 Yellow-billed Cuckoo Coccyzus americanus 46 Mangrove Cuckoo Coccyzus minor 7 x 47 Black-billed Cuckoo Coccyzus erythropthalmus 48 Striped Cuckoo Tapera naevia 3 49 Rufous-vented Ground-Cuckoo Neomorphus geoffroyi 50 Smooth-billed Ani Crotophaga ani 1 51 Groove-billed Ani Crotophaga sulcirostris 9 x 52 Nightjars & Allies Caprimulgidae 53 Short-tailed Nighthawk Lurocalis semitorquatus 2 x 54 Lesser Nighthawk Chordeiles acutipennis 4 55 Page 1 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Common Nighthawk Chordeiles minor 2 Common Pauraque Nyctidromus albicollis 3 Chuck-will's-widow Antrostomus carolinensis (n. g.) 4 Potoos Nyctibiidae 5 Great Potoo Nyctibius grandis 6 Common Potoo Nyctibius griseus 5h 7 Swifts Apodidae 8 Black Swift Cypseloides niger 1 9 Chestnut-collared Swift Streptoprocne rutila 3 10 White-collared Swift Streptoprocne zonaris 9 x 11 Chimney Swift Chaetura pelagica 6 x 12 Vaux's Swift Chaetura vauxi 5 x 13 Chapman's Swift Chaetura chapmani ? 14 Gray-rumped Swift Chaetura cinereiventris 5 x 15 Lesser Swallow-tailed Swift Panyptila cayennensis 1 x 16 Hummingbirds Trochilidae 17 White-necked Jacobin Florisuga mellivora 9 x 18 White-tipped Sicklebill Eutoxeres aquila 19 Bronzy Hermit Glaucis aeneus 2 20 Rufous-breasted Hermit Glaucis hirsutus 21 Band-tailed Barbthroat Threnetes ruckeri 7 22 Green Hermit Phaethornis guy 5 23 Long-billed Hermit (n.c. name) Phaethornis longirostris 5 24 Stripe-throated Hermit (split) Phaethornis striigularis 9 x 25 Green-fronted Lancebill Doryfera ludovicae 26 Brown Violetear Colibri delphinae 27 Lesser Violetear (split) Colibri cyanotus (new sp. epi.) 28 Purple-crowned Fairy Heliothryx barroti 8 x 29 Green-breasted Mango Anthracothorax prevostii 7 30 Veraguan Mango Anthracothorax veraguensis 31 Green Thorntail Discosura conversii 4 32 Rufous-crested Coquette Lophornis delattrei 1 33 Green-crowned Brilliant Heliodoxa jacula 4 34 Fiery-throated Hummingbird Panterpe insignis 35 Long-billed Starthroat Heliomaster longirostris 1 36 White-bellied Mountain-gem Lampornis hemileucus 5 37 Purple-throated Mountain-gem Lampornis calolaemus 2 38 Magenta-throated Woodstar Calliphlox bryantae 39 Ruby-throated Hummingbird Archilochus colubris 1 40 Scintillant Hummingbird Selasphorus scintilla 41 Garden Emerald Chlorostilbon assimilis 1 x 42 Violet-headed Hummingbird Klais guimeti 2 43 Violet Sabrewing Campylopterus hemileucurus 44 Stripe-tailed Hummingbird Eupherusa eximia 4 45 Black-bellied Hummingbird Eupherusa nigriventris 3 46 White-tailed Emerald Elvira chionura 3 47 Snowcap Microchera albocoronata 48 Bronze-tailed Plumeleteer Chalybura urochrysia 5 49 Crowned Woodnymph (lumped) Thalurania colombica 9 x 50 Blue-chested Hummingbird Amazilia amabilis 8 x 51 Snowy-bellied Hummingbird Amazilia edward 4 52 Rufous-tailed Hummingbird Amazilia tzacatl 9 x 53 Blue-throated Goldentail Hylocharis eliciae 1 54 55 Page 2 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Rails, Gallinules & Coots Rallidae 2 Ruddy Crake Laterallus ruber 3 White-throated Crake Laterallus albigularis 6,3 h 4 Gray-breasted Crake Laterallus exilis h h 5 Clapper Rail Rallus crepitans (new sp. epi.) 6 Rufous-necked Wood-Rail Aramides axillaris 1 7 Gray-cowled Wood-Rail [Split] Aramides cajaneus 3,6 x 8 Uniform Crake Amaurolimnas concolor h 9 Sora Porzana carolina 10 Yellow-breasted Crake Hapalocrex flaviventer (new g.) 11 Paint-billed Crake Neocrex erythrops 12 Purple Gallinule Porphyrio martinicus 9 x 13 Common Gallinule Gallinula galeata 6 x 14 American Coot Fulica americana 4 15 Finfoots Heliornithidae 16 Sungrebe Heliornis fulica 17 Limpkin Aramidae 18 Limpkin Aramus guarauna 19 Stilts & Avocets Recurvirostridae 20 Black-necked Stilt Himantopus mexicanus 3 21 Oystercatchers Haematopodidae 22 American Oystercatcher Haematopus palliatus 23 Plovers & Lapwings Charadriidae 24 Southern Lapwing Vanellus chilensis 9 x 25 Black-bellied Plover Pluvialis squatarola 6 26 American Golden-Plover Pluvialis dominica 2 27 Collared Plover Charadrius collaris 8 x 28 Wilson's Plover Charadrius wilsonia 4 x 29 Semipalmated Plover Charadrius semipalmatus 8 x 30 Killdeer Charadrius vociferus 31 Jaçanas Jacanidae 32 Northern Jaçana Jacana spinosa 9 x 33 Sandpipers & Allies Scolopacidae 34 Upland Sandpiper Bartramia longicauda 35 Whimbrel Numenius phaeopus 7 x 36 Ruddy Turnstone Arenaria interpres 8 x 37 *Black Turnstone Arenaria melanocephala ? 38 Sanderling Calidris alba 8 x 39 Least Sandpiper Calidris minutilla 8 x 40 White-rumped Sandpiper Calidris fuscicollis 41 Pectoral Sandpiper Calidris melanotos 4 42 Semipalmated Sandpiper Calidris pusilla 3 x 43 Western Sandpiper Calidris mauri 1 44 Short-billed Dowitcher Limnodromus griseus 1 45 Long-billed Dowitcher Limnodromus scolopaceus 46 Wilson's Snipe Gallinago delicata 47 Spotted Sandpiper Actitis macularius 9 x 48 Solitary Sandpiper Tringa solitaria 9 x 49 Lesser Yellowlegs Tringa flavipes 2 50 Willet Tringa semipalmata 3 x 51 Greater Yellowlegs Tringa melanoleuca 6 52 Wilson's Phalarope Phalaropus tricolor 1 53 Skuas & Jaegers Stercorariidae 54 South Polar Skua Stercorarius maccormicki 2 55 Page 3 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Pomarine Jaeger Stercorarius pomarinus 8 x 2 Parasitic Jaeger Stercorarius parasiticus 7 3 Jaeger spp. 1 x 4 Gulls, Terns & Skimmers Laridae 5 Laughing Gull Leucophaeus atricilla (n. genus) 7 x 6 Franklin's Gull Leucophaeus pipixcan (n. g.) 1 7 Herring Gull Larus argentatus 1 8 Brown Noddy Anous stolidus 1 9 Least Tern Sternula antillarum 10 Gull-billed Tern Gelochelidon nilotica 1 11 Caspian Tern Hydroprogne caspia 3 12 Black Tern Chlidonias niger 5 13 Common Tern Sterna hirundo 14 Royal Tern Thalasseus maximus 9 x 15 Sandwich Tern Thalasseus sandvicensis 7 x 16 Black Skimmer Rynchops niger 17 Sunbittern Eurypygidae 18 Sunbittern Eurypyga helias 19 Tropicbirds Phaethontidae 20 Red-billed Tropicbird Phaethon aethereus 9 x 21 Shearwaters & Petrels Procellariidae 22 Audubon's Shearwater Puffinus lherminieri 23 Storks Ciconiidae 24 Jabiru Jabiru mycteria 25 Wood Stork Mycteria americana 26 Frigatebirds Fregatidae 27 Magnificent Frigatebird Fregata magnificens 9 x 28 Boobies & Gannets Sulidae 29 Brown Booby Sula leucogaster 9 x 30 Red-footed Booby Sula sula 31 Cormorants & Shags Phalacrocoracidae 32 Neotropic Cormorant Phalacrocorax brasilianus 7 x 33 Anhingas Anhingidae 34 Anhinga Anhinga anhinga 1 35 Pelicans Pelecanidae 36 Brown Pelican Pelecanus occidentalis 9 x 37 Herons, Egrets & Bitterns Ardeidae 38 Least Bittern Ixobrychus exilis 1 39 Rufescent Tiger-Heron Tigrisoma lineatum 3 x 40 Fasciated Tiger-Heron Tigrisoma fasciatum 41 Bare-throated Tiger-Heron Tigrisoma mexicanum 3 42 Great Blue Heron Ardea herodias 9 x 43 Great Egret Ardea alba 9 x 44 Snowy Egret Egretta thula 8 x 45 Little Blue Heron Egretta caerulea 9 x 46 Tricolored Heron Egretta tricolor 9 x 47 Cattle Egret Bubulcus ibis 9 x 48 Green Heron Butorides virescens 9 x 49 Agami Heron Agamia agami 50 Black-crowned Night-Heron Nycticorax nycticorax 51 Yellow-crowned Night-Heron Nyctanassa violacea 8 x 52 Boat-billed Heron Cochlearius cochlearius 9 x 53 Ibises & Spoonbills Threskiornithidae 54 White Ibis Eudocimus albus 1 x 55 Page 4 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Glossy Ibis Plegadis falcinellus 2 2 Green Ibis Mesembrinibis cayennensis 9 x 3 Roseate Spoonbill Platalea ajaja 2 x 4 New World Vultures Cathartidae 5 Black Vulture Coragyps atratus 9 x 6 Turkey Vulture Cathartes aura 9 x 7 King Vulture Sarcoramphus papa 5 8 Osprey Pandionidae 9 Osprey Pandion haliaetus 9 x 10 Hawks, Eagles & Kites Accipitridae 11 Gray-headed Kite Leptodon cayanensis 4 12 Hook-billed Kite Chondrohierax uncinatus 7 13 Swallow-tailed Kite Elanoides forficatus 9 x 14 Pearl Kite Gampsonyx swainsonii 1 15 White-tailed Kite Elanus leucurus 7 x 16 Double-toothed Kite Harpagus bidentatus 5 x 17 Mississippi Kite Ictinia mississippiensis x 18 Plumbeous Kite Ictinia plumbea 9 x 19 Northern Harrier Circus hudsonius 20 Tiny Hawk Accipiter superciliosus 1 21 Bicolored Hawk Accipiter bicolor 22 Common Black Hawk Buteogallus anthracinus 9 x 23 Great Black Hawk Buteogallus urubitinga 3 x 24 Barred Hawk Morphnarchus princeps (new g.) 2,1 x 25 Roadside Hawk Rupornis magnirostris (n. genus) 9 x 26 Harris's Hawk Parabuteo unicinctus 27 White-tailed Hawk Geranoaetus albicaudatus 28 White Hawk Pseudastur albicollis 7 x 29 Semiplumbeous Hawk Leucopternis semiplumbeus 1 x 30 Broad-winged Hawk Buteo platypterus 9 x 31 Short-tailed Hawk Buteo brachyurus 5 x 32 Swainson's Hawk Buteo swainsoni 6 x 33 Zone-tailed Hawk Buteo albonotatus 1 34 Red-tailed Hawk Buteo jamaicensis 2 x 35 Crested Eagle Morphnus guianensis 36 Harpy Eagle Harpia harpyja 37 Black Hawk-Eagle Spizaetus tyrannus 5,1 x 38 Ornate Hawk-Eagle Spizaetus ornatus 2 39 Black-and-white Hawk-Eagle Spizaetus melanoleucus (n. g.) 1 40 Barn-Owls Tytonidae 41 Barn Owl Tyto alba 42 Owls Strigidae 43 Vermiculated Screech-Owl Megascops guatemalae 44 Choco Scrch-O (to be split) Megascops g. chocoensis 45 Bare-shanked Screech-Owl Megascops clarkii 46 Crested Owl Lophostrix cristata 47 Spectacled Owl Pulsatrix perspicillata 48 Costa Rican Pygmy-Owl Glaucidium costaricanum h 49 Central American Pygmy-Owl Glaucidium griseiceps 50 Mottled Owl Ciccaba virgata 7h h 51 Black-and-white Owl Ciccaba nigrolineata 2,3 52 Trogons Trogonidae 53 Lattice-tailed Trogon Trogon clathratus 54 55 Page 5 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Slaty-tailed Trogon Trogon massena 9 x 2 White-tailed Trogon Trogon chionurus 3 Gartered(N. Violaceous)Trogon Trogon caligatus (split) 8 x 4 Black-throated Trogon Trogon rufus 5 Collared Trogon Trogon collaris 6 Orange-bellied Trogon Trogon aurantiiventris 7 7 Resplendent Quetzal Pharomachrus mocinno 8 Motmots Momotidae 9 Rufous Motmot Baryphthengus martii 6 h 10 Broad-billed Motmot Electron platyrhynchum 1 11 Kingfishers Alcedinidae 12 Ringed Kingfisher Megaceryle torquata (n. genus) 9 x 13 Belted Kingfisher Megaceryle alcyon (n. genus) 7 x 14 Amazon Kingfisher Chloroceryle amazona 5 x 15 Green Kingfisher Chloroceryle americana 9 x 16 Green-and-rufous Kingfisher Chloroceryle inda 1 17 American Pygmy Kingfisher Chloroceryle aenea 9 x 18 Puffbirds Bucconidae 19 White-necked Puffbird Notharchus hyperrhynchus (nse) 20 Pied Puffbird Notharchus tectus 4 x 21 White-whiskered Puffbird Malacoptila panamensis 22 Lanceolated Monklet Micromonacha lanceolata 23 White-fronted Nunbird Monasa morphoeus 24 Jacamars Galbulidae 25 Rufous-tailed Jacamar Galbula ruficauda 2 26 Great Jacamar Jacamerops aureus 2 27 New World Barbets Capitonidae (new) 28 Red-headed Barbet Eubucco bourcierii 1 29 Toucan-Barbets (new family) Semnornithidae (new) 30 Prong-billed Barbet Semnornis frantzii 8,1 31 Toucans Ramphastidae 32 Northern Emerald-Toucanet Aulacorhynchus prasinus 6 33 Collared Aracari Pteroglossus torquatus 9 x 34 Yellow-eared Toucanet Selenidera spectabilis 35 Keel-billed Toucan Ramphastos sulfuratus 9 x 36 Yellow-throated Toucan(lumped) Ramphastos ambiguus (n. g.) 8,1 37 Woodpeckers Picidae 38 Acorn Woodpecker Melanerpes formicivorus 8 39 Black-cheeked Woodpecker Melanerpes pucherani 9 x 40 Red-crowned Woodpecker Melanerpes rubricapillus 3 x 41 Yellow-bellied Sapsucker Sphyrapicus varius 42 Smoky-brown Woodpecker Picoides fumigatus (new genus) 2 43 Hairy Woodpecker Picoides villosus 44 Rufous-winged Woodpecker Piculus simplex 1 45 Golden-olive Woodpecker Colaptes rubiginosus 1 46 Cinnamon Woodpecker Celeus loricatus 1 47 Chestnut-colored Woodpecker Celeus castaneus 1 48 Lineated Woodpecker Dryocopus lineatus 9 x 49 Crimson-bellied Woodpecker Campephilus haematogaster 50 Crimson-crested Woodpecker Campephilus melanoleucos 1 51 Pale-billed Woodpecker Campephilus guatemalensis 9 x 52 Falcons & Caracaras Falconidae 53 Laughing Falcon Herpetotheres cachinnans 5,4 h 54 Barred Forest-Falcon Micrastur ruficollis 1 x 55 Page 6 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Collared Forest-Falcon Micrastur semitorquatus 2,1 2 Red-throated Caracara Ibycter americanus 3 Crested Caracara Caracara cheriway 2 x 4 Yellow-headed Caracara Milvago chimachima 9 x 5 American Kestrel Falco sparverius 6 Merlin Falco columbarius 1 x 7 Aplomado Falcon Falco femoralis 8 Bat Falcon Falco rufigularis 5 x 9 Peregrine Falcon Falco peregrinus 5 x 10 New World & African Parrots Psittacidae 11 Sulphur-winged Parakeet Pyrrhura hoffmanni 7 h 12 Olive-throated Parakeet Eupsittula nana (n.g.) 8 13 Great Green Macaw Ara ambiguus 14 Crimson-fronted Parakeet Psittacara finschi (n.g.) 8 x 15 Barred Parakeet Bolborhynchus lineola 2 16 Orange-chinned Parakeet Brotogeris jugularis x 17 Red-fronted Parrotlet Touit costaricensis 2 18 Brown-hooded Parrot Pyrilia haematotis (new genus) 4 x 19 Blue-headed Parrot Pionus menstruus 8 x 20 White-crowned Parrot Pionus senilis 8 21 Red-lored Parrot Amazona autumnalis 8 x 22 Mealy Parrot Amazona farinosa 8 x 23 Yellow-crowned Parrot Amazona ochrocephala 1 24 Typical Antbirds Thamnophilidae 25 Fasciated Antshrike Cymbilaimus lineatus 26 Great Antshrike Taraba major 2,1 27 Black-crowned Antshrike (n. g.) Thamnophilus atrinucha 8 x 28 Russet Antshrike Thamnistes anabatinus x 29 Plain Antvireo Dysithamnus mentalis 1 30 Spot-crowned Antvireo Dysithamnus puncticeps 2 31 Pacific Antwren Myrmotherula pacifica 32 White-flanked Antwren Myrmotherula axillaris 8 x 33 Slaty Antwren Myrmotherula schisticolor 1 34 Checker-throated Antwren Epinecrophylla fulviventris (n. g.) 35 Dot-winged Antwren Microrhopias quixensis 7,2 x 36 Rufous-rumped Antwren Euchrepomis callinota (n. genus) 37 Dusky Antbird Cercomacroides tyrannina (n.g.) 1 38 Bare-crowned Antbird Gymnocichla nudiceps 39 Chestnut-backed Antbird Myrmeciza exsul 9 h 40 Dull-mantled Antbird Myrmeciza laemosticta 2 41 Zeledon's Antbird (new name) Myrmeciza zeledoni (new s.e.) 1 42 Spotted Antbird Hylophylax naevioides 43 Bicolored Antbird Gymnopithys bicolor (n. sp. epi.) 44 Ocellated Antbird Phaenostictus mcleannani 45 Gnateaters (new family) Conopophagidae (new) 46 Black-crowned Antpitta Pittasoma michleri 47 Antpittas Grallariidae 48 Scaled Antpitta Grallaria guatimalensis 49 Streak-chested Antpitta Hylopezus perspicillatus 50 Thicket Antpitta Hylopezus dives h 51 Ochre-breasted Antpitta Grallaricula flavirostris 52 Tapaculos Rhinocryptidae 53 Silvery-fronted Tapaculo Scytalopus argentifrons 2,4 54 55 Page 7 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Antthrushes Formicariidae 2 Black-faced Antthrush Formicarius analis 3 Black-headed Antthrush Formicarius nigricapillus 4 Rufous-breasted Antthrush Formicarius rufipectus 5 Ovenbirds & Woodcreepers Furnariidae 6 Tawny-throated Leaftosser Sclerurus mexicanus 1 7 Scaly-throated Leaftosser Sclerurus guatemalensis 8 Long-tailed Woodcreeper Deconychura longicauda 9 Plain-brown Woodcreeper Dendrocincla fuliginosa 1 10 Wedge-billed Woodcreeper Glyphorynchus spirurus 5 x 11 Northern Barred-Woodcreeper Dendrocolaptes sanctithomae 3 12 Black-banded Woodcreeper Dendrocolaptes picumnus 13 Strong-billed Woodcreeper Xiphocolaptes promeropirhyn. 14 Cocoa Woodcreeper (split) Xiphorhynchus susurrans 8 x 15 Black-striped Woodcreeper Xiphorhynchus lachrymosus 16 Spotted Woodcreeper Xiphorhynchus erythropygius 3 17 Brown-billed Scythebill Campylorhamphus pusillus 18 Streak-headed Woodcreeper Lepidocolaptes souleyetii 9 x 19 Spot-crowned Woodcreeper Lepidocolaptes affinis 2 20 Plain Xenops Xenops minutus 7 21 Buffy Tuftedcheek Pseudocolaptes lawrencii 22 Buff-fronted Foliage-gleaner Philydor rufum 23 Scaly-throated Foliage-gleaner Anabacerthia variegaticeps 24 Lineated Foliage-gleaner Syndactyla subalaris 1 25 Streak-breasted Treehunter Thripadectes rufobrunneus 26 Buff-throated Foliage-gleaner Automolus ochrolaemus 1 x 27 Striped Woodhaunter Automolus subulatus (n. g.) 28 Spotted Barbtail Premnoplex brunnescens 29 Ruddy Treerunner Margarornis rubiginosus 1 30 Red-faced Spinetail Cranioleuca erythrops 5 31 Slaty Spinetail brachyura 5 32 Tyrant Flycatchers Tyrannidae 33 Brown-capped Tyrannulet Ornithion brunneicapillus 3 34 Yellow Tyrannulet Capsiempis flaveola 35 Yellow-bellied Elaenia Elaenia flavogaster 9 x 36 Mountain Elaenia Elaenia frantzii 37 Torrent Tyrannulet Serpophaga cinerea 8 38 Olive-striped Flycatcher Mionectes olivaceus 2 39 Ochre-bellied Flycatcher Mionectes oleagineus 7 x 40 Slaty-capped Flycatcher Leptopogon superciliaris 1 41 Rufous-browed Tyrannulet Phylloscartes superciliaris 1 42 Rough-legged Tyrannulet (new) Phyllomyias burmeisteri (split) 43 Paltry Tyrannulet Zimmerius vilissimus 8,1 x 44 Black-capped Pygmy-Tyrant Myiornis atricapillus 3 x 45 Scale-crested Pygmy-Tyrant Lophotriccus pileatus 3 46 Northern Bentbill Oncostoma cinereigulare 1,1 47 Slate-headed Tody-Flycatcher Poecilotriccus sylvia 48 Common Tody-Flycatcher Todirostrum cinereum 9 x 49 Black-headed Tody-Flycatcher Todirostrum nigriceps 3 50 Eye-ringed Flatbill Rhynchocyclus brevirostris 51 Yellow-olive Flycatcher Tolmomyias sulphurescens 52 Yellow-margined Flycatcher Tolmomyias assimilis 2 53 Stub-tailed Spadebill Platyrinchus cancrominus 4 h 54 55 Page 8 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 White-throated Spadebill Platyrinchus mystaceus 2 Golden-crowned Spadebill Platyrinchus coronatus 3 Royal Flycatcher Onychorhynchus coronatus 4 Ruddy-tailed Flycatcher Terenotriccus erythrurus 5 Sulphur-rumped Flycatcher Myiobius sulphureipygius 6 Black-tailed Flycatcher Myiobius atricaudus 7 Bran-colored Flycatcher Myiophobus fasciatus 8 Tufted Flycatcher Mitrephanes phaeocercus 3 9 Olive-sided Flycatcher Contopus cooperi 5 x 10 Dark Pewee Contopus lugubris 11 Ochraceous Pewee Contopus ochraceus 12 Western Wood-Pewee Contopus sordidulus 2,2 x 13 Eastern Wood-Pewee Contopus virens 7,1 x 14 Tropical Pewee Contopus cinereus 8 15 Yellow-bellied Flycatcher Empidonax flaviventris 16 Acadian Flycatcher Empidonax virescens 3 x 17 Alder Flycatcher Empidonax alnorum h 18 Willow Flycatcher Empidonax traillii 19 White-throated Flycatcher Empidonax albigularis 20 Least Flycatcher Empidonax minimus h 21 Yellowish Flycatcher Empidonax flavescens 22 Black Phoebe Sayornis nigricans 9 x 23 Long-tailed Tyrant Colonia colonus 9 x 24 Bright-rumped Attila Attila spadiceus 6,2 x 25 Rufous Mourner Rhytipterna holerythra 3 26 Dusky-capped Flycatcher Myiarchus tuberculifer 8,1 x 27 Panama Flycatcher Myiarchus panamensis 3 x 28 Great Crested Flycatcher Myiarchus crinitus 4,2 x 29 Great Kiskadee Pitangus sulphuratus 9 x 30 Boat-billed Flycatcher Megarynchus pitangua 9 x 31 Social Flycatcher Myiozetetes similis 9 x 32 Gray-capped Flycatcher Myiozetetes granadensis 9 x 33 White-ringed Flycatcher Conopias albovittatus 2 34 Golden-bellied Flycatcher Myiodynastes hemichrysus 3 35 Streaked Flycatcher Myiodynastes maculatus 3 36 Sulphur-bellied Flycatcher Myiodynastes luteiventris 6 x 37 Piratic Flycatcher Legatus leucophaius 8 x 38 Tropical Kingbird Tyrannus melancholicus 9 x 39 Eastern Kingbird Tyrannus tyrannus 9 x 40 Gray Kingbird Tyrannus dominicensis 4 x 41 Scissor-tailed Flycatcher Tyrannus forficatus 4 42 Fork-tailed Flycatcher Tyrannus savana x 43 Genus Incertae Sedis 44 Gray-headed Piprites Piprites griseiceps 45 Tityras & Allies Tityridae (new) 46 Northern Schiffornis (Split) Schiffornis veraepacis (n.sp.ep.) 47 Speckled Mourner Laniocera rufescens 48 Masked Tityra Tityra semifasciata 9 x 49 Black-crowned Tityra Tityra inquisitor 8 x 50 Cinnamon Becard Pachyramphus cinnamomeus 9 x 51 White-winged Becard Pachyramphus polychopterus 3,1 x 52 Black-and-white Becard Pachyramphus albogriseus 53 Cotingas Cotingidae 54 Purple-throated Fruitcrow Querula purpurata 7 h 55 Page 9 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Bare-necked Umbrellabird Cephalopterus glabricollis 2 Lovely Cotinga Cotinga amabilis 1 3 Blue Cotinga Cotinga nattererii 4 Rufous Piha Lipaugus unirufus 5 Three-wattled Bellbird Procnias tricarunculatus 3h 6 Snowy Cotinga Carpodectes nitidus 8 x 7 Manakins Pipridae 8 White-ruffed Manakin Corapipo altera 4 x 9 Blue-crowned Manakin Lepidothrix coronata (n. genus) 10 White-collared Manakin Manacus candei 6,2 h 11 Golden-collared Manakin Manacus vitellinus 9 x 12 White-crowned Manakin Dixiphia pipra (n. genus) 2 13 Red-capped Manakin Ceratopipra mentalis (n. genus) 9 x 14 Sharpbill Oxyruncidae 15 Sharpbill Oxyruncus cristatus 16 Vireos, Shrike-Babblers & Erpornis Vireonidae 17 Green Shrike-Vireo Vireolanius pulchellus 18 Tawny-crowned Greenlet Tunchiornis ochraceiceps (n.g.) 19 Lesser Greenlet Pachysylvia decurtata (n.g.) 9 x 20 White-eyed Vireo Vireo griseus 21 Yellow-throated Vireo Vireo flavifrons 5 h 22 *Blue-headed Vireo Vireo solitarius ? 23 Philadelphia Vireo Vireo philadelphicus 6 x 24 Warbling Vireo Vireo gilvus 25 Brown-capped Vireo Vireo leucophrys 3 26 Red-eyed Vireo Vireo olivaceus 8 x 27 Yellow-green Vireo Vireo flavoviridis 1 28 Black-whiskered Vireo Vireo altiloquus 29 Crows, Jays & Magpies Corvidae 30 Azure-hooded Jay Cyanolyca cucullata 2 31 Brown Jay Psilorhinus morio 9 x 32 Black-chested Jay Cyanocorax affinis 9 x 33 Swallows Hirundinidae 34 Purple Martin Progne subis 1 x 35 Gray-breasted Martin Progne chalybea 9 x 36 Brown-chested Martin Progne tapera 1 37 Tree Swallow Tachycineta bicolor 38 Mangrove Swallow Tachycineta albilinea 9 x 39 Blue-and-white Swallow Pygochelidon cyanoleuca 8 x 40 Northern Rough-winged Swa. Stelgidopteryx serripennis 4 41 Southern Rough-winged Swa. Stelgidopteryx ruficollis 6 x 42 Bank Swallow Riparia riparia 1 43 Cliff Swallow Petrochelidon pyrrhonota 4 x 44 Barn Swallow Hirundo rustica 9 x 45 Wrens Troglodytidae 46 Scaly-breasted Wren Microcerculus marginatus 4,2 47 House Wren Troglodytes aedon 9 x 48 Ochraceous Wren Troglodytes ochraceus 1,1 49 Band-backed Wren Campylorhynchus zonatus 4 50 Black-throated Wren Pheugopedius atrogularis (n. g.) 3,2 x 51 Rufous-and-white Wren Thryophilus rufalbus (new g.) 2,1 52 Stripe-breasted Wren Cantorchilus thoracicus (n .g.) 53 Canebrake Wren (split) Cantorchilus zeledoni 1,1 h 54 55 Page 10 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Isthmian Wren (split) Cantorchilus elutus (n. species) 2,1 x 2 Bay Wren Cantorchilus nigricapillus (n. g.) 8,1 x 3 White-breasted Wood-Wren Henicorhina leucosticta 2h 4 Gray-breasted Wood-Wren Henicorhina leucophrys 7,2 h 5 Song Wren Cyphorhinus phaeocephalus 3h 6 Gnatcatchers (new family) Polioptilidae (new) 7 Tawny-faced Gnatwren Microbates cinereiventris 8 Long-billed Gnatwren Ramphocaenus melanurus 8,1 h 9 Tropical Gnatcatcher Polioptila plumbea 9 x 10 Dippers Cinclidae 11 American Dipper Cinclus mexicanus 6 12 Thrushes & Allies Turdidae 13 Black-faced Solitaire Myadestes melanops 14 Orange-billed Nightingale-Thru. Catharus aurantiirostris 3,2 15 Slaty-backed Nightingale-Thru. Catharus fuscater 3,5 h 16 Black-headed Nightingale-Thr. Catharus mexicanus 1,2 17 Veery Catharus fuscescens 2 18 Gray-cheeked Thrush Catharus minimus 2 19 Swainson's Thrush Catharus ustulatus 6 20 Wood Thrush Hylocichla mustelina 2 21 Mountain Thrush Turdus plebejus 3 x 22 Pale-vented Thrush Turdus obsoletus 23 Clay-colored Thrush Turdus grayi 9 x 24 White-throated Thrush Turdus assimilis 6 25 Mockingbirds & Thrashers Mimidae 26 Gray Catbird Dumetella carolinensis 7 x 27 Waxwings Bombycillidae 28 Cedar Waxwing Bombycilla cedrorum 29 Silky-flycatchers Ptiliogonatidae 30 Black-and-yellow Silky-flycat. Phainoptila melanoxantha 31 Long-tailed Silky-flycatcher Ptiliogonys caudatus 32 Old World Sparrows Passeridae 33 House Sparrow Passer domesticus 8 x 34 Finches, Euphonias & Allies Fringillidae 35 Golden-browed Chlorophonia Chlorophonia callophrys 2 36 Yellow-crowned Euphonia Euphonia luteicapilla 7 x 37 Thick-billed Euphonia Euphonia laniirostris 6 x 38 Elegant Euphonia Euphonia elegantissima 6 39 Olive-backed Euphonia Euphonia gouldi 9 x 40 White-vented Euphonia Euphonia minuta 6 x 41 Tawny-capped Euphonia Euphonia anneae 9 x 42 Yellow-bellied Siskin Spinus xanthogastrus 2 43 Lesser Goldfinch Spinus psaltria 1 x 44 New World Sparrows (new) Passerellidae (new) 45 Yellow-green Finch Pselliophorus luteoviridis 46 Orange-billed Sparrow Arremon aurantiirostris 1 47 Sooty-faced Finch Arremon crassirostris (n. genus) 48 Chestnut-capped Brushfinch Arremon brunneinucha (n. g.) 1 49 Black-striped Sparrow Arremonops conirostris 9 x 50 White-naped Brushfinch Atlapetes albinucha 2 51 Lincoln's Sparrow Melospiza lincolnii 52 Rufous-collared Sparrow Zonotrichia capensis 7 53 Common Chlorospingus (n.n.) Chlorospingus flavopectus(n.s.e) 9 x 54 55 Page 11 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Sooty-capped Chlorospingus Chlorospingus pileatus 2 Yellow-throated Chlorospingus Chlorospingus flavigularis 7 x 3 Ashy-throated Chlorospingus Chlorospingus canigularis 1 4 Wrenthrush (new) Zeledoniidae (new) 5 Wrenthrush Zeledonia coronata 6 Yellow-breasted Chat (new) Icteriidae (new) 7 Yellow-breasted Chat Icteria virens 8 Troupials & Allies Icteridae 9 Eastern Meadowlark Sturnella magna 3 10 Red-breasted Blackbird Leistes militaris (new genus) 9 x 11 Yellow-billed Cacique Amblycercus holosericeus 12 Chestnut-headed Oropendola Psarocolius wagleri 7 x 13 Montezuma Oropendola Psarocolius montezuma 9 x 14 Scarlet-rumped Cacique Cacicus uropygialis 6 15 Black-cowled Oriole Icterus prosthemelas 9 x 16 Orchard Oriole Icterus spurius 7 x 17 Yellow-tailed Oriole Icterus mesomelas 18 Baltimore Oriole Icterus galbula 7 x 19 Shiny Cowbird Molothrus bonariensis 1 20 Bronzed Cowbird Molothrus aeneus 4 x 21 Giant Cowbird Molothrus oryzivorus 9 x 22 Melodious Blackbird Dives dives 1 x 23 Great-tailed Grackle Quiscalus mexicanus 9 x 24 New World Warblers Parulidae 25 Seiurus aurocapilla 26 Worm-eating Warbler Helmitheros vermivorum 27 Louisiana Waterthrush Parkesia motacilla (n. genus) 1 28 Northern Waterthrush Parkesia noveboracensis (n. g.) 9 x 29 Golden-winged Warbler Vermivora chrysoptera 4 30 Blue-winged Warbler Vermivora cyanoptera (n. s. epi.) 1 31 Black-and-white Warbler Mniotilta varia 8 x 32 Prothonotary Warbler Protonotaria citrea 7 x 33 Tennessee Warbler Oreothlypis peregrina (n. genus) 7 x 34 Connecticut Warbler Oporornis agilis 35 Masked Yellowthroat Geothlypis aequinoctialis 1 36 Mourning Warbler Geothlypis philadelphia (n. g.) 3 37 Kentucky Warbler Geothlypis formosa (n. genus) 1 38 Olive-crowned Yellowthroat Geothlypis semiflava 9 x 39 Common Yellowthroat Geothlypis trichas (new genus) 1 40 Hooded Warbler Setophaga citrina (new genus) 41 American Redstart Setophaga ruticilla (new genus) 8 x 42 Cape May Warbler Setophaga tigrina 43 Cerulean Warbler Setophaga cerulea (new genus) 3 x 44 Northern Parula Setophaga americana (n. genus) x 45 Tropical Parula Setophaga pitiayumi (n. g.) 8 x 46 Magnolia Warbler Setophaga magnolia (n.g.) 1 x 47 Bay-breasted Warbler Setophaga castanea (n.g.) 3 48 Blackburnian Warbler Setophaga fusca (new genus) 6 49 Yellow Warbler Setophaga petechia (n. genus) 9 x 50 Mangrove Warbler [subspecies] 7 x 51 Chestnut-sided Warbler Setophaga pensylvanica (n.g.) 7 x 52 Blackpoll Warbler Setophaga striata (new genus) 1 53 Yellow-rumped Warbler Setophaga coronata (n. genus) 54 55 Page 12 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Yellow-throated Warbler Setophaga dominica 2 Black-throated Green Warbler Setophaga virens (new genus) 3 3 Buff-rumped Warbler Myiothlypis fulvicauda (n. g.) 7,1 x 4 Rufous-capped Warbler Basileuterus rufifrons 2 5 Black-cheeked Warbler Basileuterus melanogenys 6 Golden-crowned Warbler Basileuterus culicivorus 1 7 Costa Rican Warbler (split) Basileuterus melanotis (n. sp. e.) 1 8 Canada Warbler Cardellina canadensis (n. g.) 1 9 Wilson's Warbler Cardellina pusilla (n. genus) 3 10 Slate-throated Redstart Myioborus miniatus 8 x 11 Collared Redstart Myioborus torquatus 12 Mitrospingid Tanagers (new) Mitrospingidae (new) 13 Dusky-faced Tanager Mitrospingus cassinii 9 14 Cardinals & Allies Cardinalidae 15 Hepatic Tanager Piranga flava 5 x 16 Summer Tanager Piranga rubra 7 x 17 Scarlet Tanager Piranga olivacea 7 x 18 Flame-colored Tanager Piranga bidentata 3 19 White-winged Tanager Piranga leucoptera 2 20 Red-throated Ant-Tanager Habia fuscicauda 9 x 21 Olive (Carmiol's) Tanager (new) Chlorothraupis carmioli 5 22 Black-faced Grosbeak Caryothraustes poliogaster 2 23 Black-thighed Grosbeak Pheucticus tibialis 4 24 Rose-breasted Grosbeak Pheucticus ludovicianus 7 25 Blue Seedeater Amaurospiza concolor 26 Blue-black Grosbeak Cyanocompsa cyanoides 9 x 27 Blue Grosbeak Passerina caerulea (n. genus) 28 Indigo Bunting Passerina cyanea 1 29 Painted Bunting Passerina ciris 1 30 Dickcissel Spiza americana 1 31 Tanagers & Allies Thraupidae 32 Blue-and-gold Tanager Bangsia arcaei 4 33 Blue-gray Tanager Thraupis episcopus 9 x 34 Palm Tanager Thraupis palmarum 9 x 35 Golden-hooded Tanager Tangara larvata 9 x 36 Speckled Tanager Tangara guttata 8 37 Spangle-cheeked Tanager Tangara dowii 3 38 Plain-colored Tanager Tangara inornata 9 x 39 Rufous-winged Tanager Tangara lavinia 2 40 Bay-headed Tanager Tangara gyrola 9 41 Emerald Tanager Tangara florida 9 x 42 Silver-throated Tanager Tangara icterocephala 9 x 43 Slaty Flowerpiercer Diglossa plumbea 2 44 Green Honeycreeper Chlorophanes spiza 9 x 45 Black-and-yellow Tanager Chrysothlypis chrysomelas 4 46 Sulphur-rumped Tanager Heterospingus rubrifrons 2 47 Blue-black Grassquit Volatinia jacarina 8 x 48 White-shouldered Tanager Tachyphonus luctuosus 4 x 49 Tawny-crested Tanager Tachyphonus delatrii 9 x 50 White-lined Tanager Tachyphonus rufus 8 x 51 White-throated Shrike-Tanager Lanio leucothorax 1 52 Crimson-collared Tanager Ramphocelus sanguinolentus 7 x 53 Lemon (Flame)-rumped T. (new) Ramphocelus flammigerus 1 54 55 Page 13 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Passerini's Tanager Ramphocelus passerinii 9 x 2 Cherrie's Tanager Ramphocelus costaricensis 4 3 Shining Honeycreeper Cyanerpes lucidus 6 x 4 Red-legged Honeycreeper Cyanerpes cyaneus 5 5 Scarlet-thighed Dacnis Dacnis venusta 9 6 Blue Dacnis Dacnis cayana 9 x 7 Bananaquit Coereba flaveola 9 x 8 Yellow-faced Grassquit Tiaris olivaceus 7 9 Thick-billed Seed-Finch Sporophila funerea (new name) 9 x 10 Nicaraguan Seed-Finch Sporophila nuttingi (n. g.) 8 x 11 Variable Seedeater Sporophila corvina (n. sp. epi.) 9 x 12 Slate-colored Seedeater Sporophila schistacea x 13 White-collared Seedeater Sporophila torqueola 9 x 14 Yellow-bellied Seedeater Sporophila nigricollis 1 15 Black-headed Saltator Saltator atriceps 16 Buff-throated Saltator Saltator maximus 9 x 17 Slate-colored Grosbeak Saltator grossus 1 18 Grayish Saltator Saltator coerulescens 2 h 19 20 21 22 AT PANAMA CITY HOTEL 23 NOT POSSIBLE AT BDTA 24 OR NEVER SEEN THERE 25 26 American Kestrel (NST) Falco sparverius x 27 Barred Antshrike (NP) Thamnophilus nigriceps x 28 Crimson-backed Tanager (NP) Ramphocelus dimidiatus x 29 Saffron Finch (NP) Sicalis flaveola x 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 Page 14 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 MAMMALS 2 Reid's 1997 order and nomenclature 3 Common Opossum Didelphis marsupialis 2 4 Central Ameri. Woolly Opossum Caluromys derbianus 3 5 Gray Four-eyed Opossum Philander opossum 1 6 Northern Tamandua Tamandua mexicana 7 Hoffmann's Two-toed Sloth Choloepus hoffmanni 7 x 8 Brown-throated Three-toed Sloth Bradypus variegatus 9 x 9 Nor. Naked-tailed Armadillo Cabassous centralis 10 Nine-banded Armadillo Dasypus novemcinctus 2 11 78 Bat species 8 12 Proboscis or Long-nosed Bat Rhynchonycteris naso 13 Greater Fishing Bat Noctilio leporinus 14 Silky Short-tailed Bat Carollia brevicauda 1 15 Common Tent-makingBat Uruderma bilobatum 2 16 Greater White-lined Bat Saccopteryx bilineata 9 x 17 False Vampire Bat Vampyrum spectrum 18 3 Vampire Bat species 19 Jamaican or Mexican Fruit Bat Artibeus jamaicensus 5 x 20 Orange Nectar Bat Lonchophylla robusta 2 x 21 Northern Little Yellow-eared Bat Vampyressa thyone 1 22 Spear-nosed Bat 2 x 23 Geoffroy's Tamarin Saguinus geoffroyi 24 Western Night Monkey Aotus lemurinus 3,1 x 25 White-faced Capuchin Monkey Cebus capuchinus 8 x 26 Mantled Howler Monkey Alouatta palliata 3,5 x 27 Central American Spider Monkey Ateles geoffroyi 28 Variegated Squirrel Sciurus variegatoides 29 Red-tailed Squirrel Sciurus granatensis 7 x 30 Alfaro's Pygmy Squirrel Microsciurus alfari 5 x 31 Western Pygmy Squirrel Microsciurus mimulus 32 14 Mouse or Rat species 1 33 Rothchild's Porcupine Coendou rothschildi 34 Capybara Hydrochaeris hydrocheris 35 Central American Agouti Dasyprocta punctata 36 Paca Agouti paca 37 3 Spiny or Tree Rat species 2 38 Tapiti or Forest Rabbit [to 1500 m] Sylvilagus brasiliensis 39 Coyote Canis latrans 40 Gray Fox Urocyon cinereoargenteus 41 Northern Raccoon Procyon lotor 1 42 Crab-eating Raccoon Procyon cancrivorous 4 43 White-nosed Coati Nasua nasua x 44 Kinkajou Potos flavus 1 45 Allen's Olingo Bassaricyon alleni (split) 46 Long-tailed Weasel Mustela frenata 47 Tayra Eira barbara 2 48 Neotropical River Otter Lutra longicaudis 49 Ocelot Felis pardalis 50 Margay Leopardus wiedii 51 Jaguarundi Felis yagouaroundi 52 Puma Puma concolor 53 Jaguar Panthera onca 54 West Indian Manatee Trichechus manatus 55 Page 15 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Baird's Tapir Tapirus bairdii 2 Collared Peccary Tayassu tajacu 3 White-lipped Peccary Tayassu pecari 4 White-tailed Deer Olocoileus virginianus 5 Red brocket Deer Mazama americana 1 6 Bottlenose Dolphin Tursiops truncatus 7 x 7 REPRESENTATIVE REPTILES and AMPHIBIANS 8 From 1983 Costa Rican Natural History, D. Janzen 9 Forest Frog species 4 10 Red-eyed Tree Frog Agalychnis callidryas 11 Tink Frog Eleutherodactylus bransfordii 9h h 12 Talamanca Frog 1 x 13 Tree Frog species Hyla spp. 14 Green & Black Poison Dart Frog Dendrobates auratus 9 x 15 Blue Jeans or Strawberry P. D. Frog Oophaga pumilio 9 x 16 Lovely P. D. Frog Phyllobates lugubris x 17 Leaf-litter Toad Rhaebo haematiticus x 18 Little Toad species 9 x 19 Giant or Marine Toad Bufo marinus 20 Galliwasp Lizard Diploglossus monotropis 21 Jesus Christ Lizard Basiliscus basiliscus 9 x 22 Emerald Basilisk Basiliscus plumifrons 8 23 Macroteiid Lizard Ameiva spp. 2 24 Central American Whiptail Lizard Ameiva festiva 6 x 25 Chameleon species 1 26 Common or Green Iguana Iguana iguana 8 x 27 Ctenosaur or Black Iguana Ctenosaura similis 28 Yellow-headed Gecko Gonatodes albigularis 6 x 29 Caribbean Least Gecko Sphaerodactylus homolepis 2 30 "Introduced" or House Gecko Hemidactylus frenatus 9 x 31 Giant Gecko Norops polylepis 4 32 Anolis Lizard or Anole Anolis spp. 8 x 33 Skink Sphenomorphus cherriei 34 Red or Painted Turtle Rhinoclemmys pulcherrima 35 Common Slider Trachemys scripta 36 Mud Turtle species Kinosternon spp. 3 37 American Crocodile Crocodylus acutus 38 Spectacled Caiman or Baba Caiman crocodilus 3 39 Eyelash Viper or Bocaraca Bothrops schlegelii 40 Fer-de-lance or Terciopelo Bothrops asper 41 Boa Constrictor Boa constrictor 2 42 Brown Vine s. or Bejuquilla Café Oxybelis aeneus 1 x 43 Green Vine S. or Bejuquilla Verde Oxybelis fulgidus 1 44 Bushmaster Lachesis muta 45 Tiger Rat Snake Spilotes pullatus 3 46 Green Parrot Snake Leptophis ahaetulla 1 47 Rat Snake species 1 48 Coral Snake 1 49 Unidentified SNAKE species 3 x 50 51 52 53 54 55 Page 16 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 REPRESENTATIVE JELLYFISH 2 Upside-down Jellyfish Cassiopea spp. 7 3 Moon Jelly Aurelia aurita 5 x 4 Atlantic Sea Nettle Chrysaora quinquecirrha 1 5 6 7 8 9 REPRESENTATIVE FISHES 10 Snapper species Lutjanus campechanus 8 x 11 Unidentified SARDINE species 9 x 12 Spotted Eagle Ray Aetobatus narinari 3 x 13 Southern Sting Ray 4 14 Cownose Ray 1 15 Glass Minnow species 7 16 Barracuda 4 x 17 Blue Tang 6 x 18 Bonito 5 19 Parrot Fish 4 x 20 Dog Snapper (Pargo) 1 x 21 Tarpon 3 22 Needlefish 4 x 23 Sea Urchin 5 x 24 Jacks 1 x 25 Sea Cucumber 1 x 26 Queen Angel Fish 1 x 27 Four-eyed Butterfly Fish 1 x 28 Sergeant major x 29 Sailfin Molly x 30 31 32 33 34 35 REPRESENTATIVE 36 Fiddler Crab species Uca spp. 8 x 37 Mangrove Crab 9 x 38 Brown Tiger Crab 2 39 Red Crab 4 40 Ghost Crab 1 41 Rock Crab 1 42 Unidentified SPIDER CRAB sp. 2 43 Unidentified LAND CRAB species 7 x 44 Blue Land Crab Cardisoma guanhumi x 45 Spiny Lobster species 4 46 Unidentified SPIDER species 9 x 47 Golden Orb-Weaver Nephila clavipes 9 x 48 Spiny-backed Orb-Weaver Gasteracantha cancriformis 5 49 Jumping Spider species 1 50 Scorpion Spider (in bat cave) 4 x 51 Leaf Cutter Ant Atta cephalotes 9 x 52 Army Ant Eciton burchelli 7 x 53 Acacia Ant Pseudomyrmex ferruginea 54 Bullet Ant or Bala Paraponera clavata 6 55 Page 17 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Fire Ant 3 2 Cecropia or Azteca Ant Azteca spp. 5 3 Unidentified ANT species 9 x 4 Sundown Cicada or Chicharra Fidicina mannifera 8 h 5 Chigger or Coloradita [larval st] Eutrombicula spp. 7 6 Tarantula Wasp species 7 7 Cha Cha Cha Wasp species 3 x 8 Unidentified WASP species 9 x 9 Pega Pega Bee 9 x 10 American Honeybee species Apis spp. 7 x 11 Unidentified MOSQUITO species 9 x 12 No-see-um 6 x 13 Ant Lion [larval stage] Mermeleon spp. 2 14 Unidentified STICK spp. 2 15 Arboreal Termite Nasutitermes spp. 9 x 16 Unidentified DAMSELFLY species 8 x 17 Helicopter Damselfly Megaloprepus coerulatus 6 18 Unidentified DRAGONFLY species 9 x 19 Large Purple Grasshopper 4 x 20 Uniden. GRASSHOPPER spp. 9 x 21 Unidentified KATYDID species 5 x 22 Unidentified CRICKET species 1 x 23 Forest Floor Millipede 7 24 Uniden. PRAYING MANTIS spp. 6 x 25 Unidentified EARTHWORM spp. 1 26 Uniden. FLATWORM species x 27 UFO Insect x 28 Uniden. CATERPILLAR species 9 x 29 Unidentified FIREFLY species 9 x 30 Firebug (type of beetle) 1 31 Unidentified JEWEL BEETLE species 5 x 32 Unidentified BEETLE species 9 x 33 Unidentified CLICK BEETLE species 1 34 Flag-footed (Leaf-footed) Bug Anisoscelis flavolineata 3 35 Uniden. COCKROACH species 8 x 36 Unidentified MOTH species 9 x 37 Imperial Moth Eacles imperialis 3 38 Urania Swallowtail Moth Urania fulgens 4 39 Giant Silk Moth species 2 40 Unidentified BUTTERFLY species 9 x 41 Caligo or Cream Owl Butterfly Caligo memmon 4 x 42 Morpho Morpho peleides 9 x 43 Clearwing Butterfly species 9 x 44 Thoas Swallowtail Heraclides thoas autocles 8 x 45 Plain Satyr Cissia pompilia 2 46 Smooth-banded Sister Adelpha cytherea 1 x 47 Banded Peacock Anartia fatima 2 x 48 White Peacock Anartia jatrophae 8 x 49 Starry (Night) Cracker Hamadryas laodamia 4 x 50 Cracker Butterfly species Hamadryas spp. 7 51 No common name Adhemarius daphne 1 52 Common Olivewing Nessaea aglaura 2 53 Tigerwing Butterfly Tithorea harmonia 6 x 54 Unidentified species Heliconius spp. 4 x 55 White-tipped Cycadian Eumaeus godartii 5 x Page 18 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Great Eurybia Eurybia spp. 1 2 Malachite Siproeta stelenes 2 x 3 Ruddy Daggerwing Marpesia petreus 2 4 Monarch Danaus plexippus 2 5 Giant Swallowtail Papilio cresphontes 1 x 6 Confused Swallowtail ingenuus 1 x 7 Black Swallowtail Papilio polyxenes 1 8 Green-celled Cattleheart Parides childrenae 2 9 Orange-barred Sulphur Phoebis philea 1 10 Apricot Sulphur Phoebis argante 1 11 Cloudless Sulphur Phoebis sennae 1 12 Eurema Yellow (species) Eurema spp. 1 13 Florida White Appias drusilla 1 x 14 Great Southern White Ascia monuste 1 15 Ghost Yellow Eurema albula 1 16 Mexican Yellow Eurema mexicana 1 17 Banded Yellow Eurema elathea 1 18 Eastern Tailed Blue Cupido comyntas 1 x 19 Ceraunus Blue Hemiargus hanno 1 20 Bitias Hairstreak Panthiades bitias 1 21 Mountain (Brown) Greenstreak Cyanophrys longula 1 22 Pale-clubbed Greatstreak Theritas hemon 1 23 Scrub-Hairstreak Strymon bazochii 1 24 Togarna Stripestreak Arawacus togarna 1 25 Black-barred Cross-streak Panthiades phaleros 1 26 Dusky-blue Groundstreak Calycopis isobeon 1 27 Pale Ministreak Ministrymon una 1 28 Cloaked Scintillant Detritivora barnesi 1 29 Common Lemmark Juditha caucana 1 30 Iris Calephelis Caliphelis iris 1 31 Red-bordered Pixie Melanis pixie pixie 1 32 Electron (Orange-barred) Pixie Melanis electron 1 33 Variable Lemmark (Rusty Metal.) Synargis mycone 1 34 Calephelis Scintillant species 1 35 Orange-flushed Eyemark Perophthalma lasus 1 36 White-spotted Tanmark Emesis aurimna 1 37 Timnia Clearwing Pseudoscada timnia pusio 1 38 Diasia Clearwing Ithomia diasia 1 39 Dero Clearwing Dircenna dero 1 40 Disturbed Tigerwing Mechanitis polymnia 1 x 41 Mimic Tigerwing Melinaea lilis 1 42 Confused Tigerwing Mechanitis lysimnia 1 x 43 Melpomene Longwing Heliconius melpomene rosina 1 44 Erato Heliconia Heliconius erato 1 x 45 Heart-spotted (Hecale) Heliconia Heliconius hecale 1 46 Zebra Longwing Heliconius charitonlius 1 47 Montane Longwing Heliconius clysonymous mont 1 48 Sara Longwing Heliconius sara magdalena 1 49 Julia Heliconia Dryas iulia 1 x 50 Juno Heliconia Dione juno huascuna 1 51 Banded-orange Heliconia Dryadula phaetusa 1 52 Orange-patched Crescent Anthanassa drusilla 1 53 54 55 Page 19 PANAMA: BOCAS DEL TORO ARCHIPELAGO TOUR, March 23 - April 1, 2018

COMMON NAME SCIENTIFIC NAME I II 1 Whitened Crescent Janatella leucodesma 1 2 Bordered Patch Chlosyne lacinia lacinia 1 3 Theona Checkerspot (Ezra) Chlosyne theona ezra 1 4 Purple-stained Daggerwing Marpesia marcella valetta 1 5 Two-spotted Prepona Archaeoprepona demophoon 1 6 Rusty-tipped Page Siptoeta epaphus 1 7 Mangrove Buckeye Junonia evarete 1 8 Gray Cracker Hamadryas februa ferentina 1 9 Gold-stained Satyr Cissia pseudoconfusa 1 10 Hermes Satyr Hermeuptychia hermes 1 11 Blue-gray Satyr Magneuptychia libye 1 12 Scallopwing Staphylus spp. 1 13 Narrow-banded Skipper Autochton longipennis 1 14 Glassy-winged Skipper Xenophanes tryxus 1 15 Falcate Skipper Spathilepia clonius 1 16 Milanian Skipper species 1 17 Anastrus Skipper species 1 18 Orcus Checkered-Skipper Pyrgus orcus 1 19 Plain Longtail Urbanus simplicius 1 20 Dorantes Longtail Urbanus dorantes 1 21 Brown Longtail Urbanus procne 1 22 Tanna/Teleus Longtail Urbanus spp. 1 23 Longtailed Skipper Urbanus proteus 1 24 Evan's Ruby-eye Talides cantra 1 25 Common Brown-Skipper Callimormus saturnus 1 26 Radiant Brown-Skipper Callimormus radiola 1 27 Trailside Under-Skipper Anthoptus epictetus 1 28 Common Mellana Quasimellana eulogius 1 29 Brazilian Skipper Calpodes ethlius 1 30 Julia's Skipper julia 1 31 Saliana Skipper species 1 32 Common Glassywing Pompeius pompeius 1 33 Fiery Skipper Hylephila phyleus 1 34 Eufala Skipper Lerodea eufala 1 35 Red-washed Skipper Haemactis spp. 1 36 Variegated Clearwing Godyris zavaleta c. x 37 Stoplight Catone Catonophele numilia x 38 Crimson Patch Chlosyne janais x 39 Brownish Cracker Hamadrias iphthime x 40 Salvin's Ticlear Episcada salvinia x 41 White-spot Greta Greta annette x 42 Disjunct Yellow Eurema gratiosa x 43 Unidentified Metalmark species Emesis spp. x 44 45 46 47 48 49 50 51 52 53 54 55 Page 20