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The Taxonomic Status of Gladiolus Illyricus (Iridaceae) in Britain
The Taxonomic Status of Gladiolus illyricus (Iridaceae) in Britain Aeron Buchanan Supervisor: Fred Rumsey, Natural History Museum, London A thesis submitted in partial fulfilment of the requirements for the degree of Master of Science of Imperial College, London Abstract First noticed officially in Britain in 1855, Gladiolus illyricus (Koch) presents an interesting taxonomic and biogeographical challenge: whether or not this isolated northern population should be recognized as a separate sub-species. Fundamental conservation issues rest on the outcome. Here, the investigation into the relationship of the G. illyricus plants of the New Forest, Hampshire, to Gladiolus species across Europe, northern Africa and the middle east is initiated. Two chloroplast regions, one in trnL–trnF and the other across psbA–trnH have been sequenced for 42 speci- mens of G. illyricus, G. communis, G. italicus, G. atroviolaceus, G. triphyllos and G. anatolicus. Phylogenetic and biogeographical treatments support the notion of an east–west genetic gradation along the Mediterranean. Iberia particularly appears as a zone of high hybridization potential and the source of the New Forest population. Alignment with sequences obtained from GenBank give strong support to the classic taxonomy of Gladiolus being monophyletic in its sub-family, Ixioideae. Comments on these chloroplast regions for barcoding are also given. In conclusion, the genetic localization of Britain’s G. illyricus population as an extremity haplotype suggests that it could well deserve sub-species status. Contents 1 Introduction 2 2 Background 4 3 Materials and Methods 8 4 Results and Discussion 15 5 Conclusions 26 Appendices 28 References 56 1. Introduction G. illyricus in Britain Figure 1: G. -
Growing the Gladiolus
Volume 3 Article 1 Bulletin P85 Growing the Gladiolus 7-1-1947 Growing the Gladiolus L. C. Grove Iowa State College Follow this and additional works at: http://lib.dr.iastate.edu/bulletinp Part of the Agriculture Commons, and the Plant Sciences Commons Recommended Citation Grove, L. C. (1947) "Growing the Gladiolus," Bulletin P: Vol. 3 : Bulletin P85 , Article 1. Available at: http://lib.dr.iastate.edu/bulletinp/vol3/iss85/1 This Article is brought to you for free and open access by the Iowa Agricultural and Home Economics Experiment Station Publications at Iowa State University Digital Repository. It has been accepted for inclusion in Bulletin P by an authorized editor of Iowa State University Digital Repository. For more information, please contact [email protected]. Grove: Growing the Gladiolus g N SIO N SER V IC E, Cooperating Published by Iowa State University Digital Repository, 1947 AMES, IOWA 1 Bulletin P, Vol. 3, No. 85 [1947], Art. 1 CONTENTS Page Growth development ....................................................................,807 Propagation ................................ .. ........................................... 809 Methods ..................................................... 809 Bulbs . , .......................................................... 810 Sizes ......................................•.....................................................810 Bulb shape ............................................................. 810 Varieties .............................. 811 Flower uses ................. ...................................................................811 -
Easy-To-Grow Florida Wildflowers
Easy-to-grow Florida wildflowers Help create pollinator These wildflowers are adapted to an average moderate to dry garden in all regions pathways of Florida. They grow best with 6 or more hours of sun and will thrive without regular Florida’s native wildflowers irrigation once established. and plants are essential to the pollinators we depend on. Asclepias tuberosa Butterflyweed Orange / red S/SS Landscaping with wildflowers Berlandiera spp Greeneyes Yellow S/SS can create pollinator pathways Carphephorus corymbosus Chaffhead / Paintbrush Purple SS/F through urban areas, helping Chamaecrista fasciculata Partridge pea Yellow SS/F bees, butterflies and other beneficial insects to thrive. Coreopsis leavenworthii Leavenworth’s tickseed Yellow S/SS Gaillardia pulchella Blanketflower Red / yellow SS/F What you can do Gaura angustifolia Southern beeblossom Pink S/SS/F • Landscape with Florida native Helianthus debilis Beach sunflower Yellow SS/F wildflowers and plants. Liatris spp Blazing star Purple SS/F • Stop using pesticides, Monarda punctata Spotted horsemint White SS/F which harm bees and other beneficial insects. Penstemon multiflorus White beardtongue White SS • Stop using fertilizer, which Pityopsis graminifolia Silkgrass Yellow SS/F harms waterways and lakes. Rudbeckia hirta Black-eyed Susan Yellow SS/S • Ask your county to preserve Ruellia caroliniensis Wild petunia Blue S/SS roadside wildflowers. Salvia coccinea Tropical sage Red S/SS/F • Work with lawmakers to Solidago sempervirens Seaside goldenrod Yellow SS/F preserve and conserve -
Black-Eyed Susan - Rudbeckia Fulgida Var
Black-eyed Susan - Rudbeckia fulgida var. sullivantii ‘Goldsturm’ A favorite summertime flower, the Black-eyed Susan, with its bright, golden-yellow flowers, produces a ‘gold storm’ of blooms in gardens everywhere. Selected by the Perennial Plant Association as the 1999 Perennial of the Year, this rudbeckia has been acclaimed internationally as one of the most popular perennials for the past fifty years. produces masses of golden color all summer long. It can be one of the most important flowers in the perennial garden. A member of the huge aster or composite family, they typically stay in a basil rosette their first year. Then during the second year produce upright, branching stems with orange-gold flowers. The rough- textured leaves are diamond shaped, and the plant produces erect, bristly stems about thirty-six inches tall. It’s significant compact habit produces long-blooming flowers, and is low maintenance. This spectacular perennial tolerates poor clay soils and mild droughts, but grows best in well- drained and moist soil with regular watering. However, it will not tolerate soggy soils. It is a rhizomatous perennial (spreads by rhizomes, making large clumps) with crowded, branching stems. ‘Goldsturm’ performs as well in the high heat and humidity of Houston, as well as tolerate the arid, cold winters of Amarillo. This Black-eyed Susan is at home in the naturalized, semi-wild garden, used as a border, or even a non-invasive ground cover with full sun to light shade. Plants have minimal blooms the first season unless they undergo cold treatment (vernalization). The long, strong stems make this orange coneflower ideal for cutting. -
Black-Eyed Susan Habitat and Range: Meadows, Fields, Roadsides, Rudbeckia Hirta L
Lanceolate to oblong, upper leaves sessile, lower ones Wildflower in Focus sometimes petioled, 1 1/2 - 6 1/2" long. Height and Growth Habit: 1 - 3'. Usually single- Text by Melanie Choukas-Bradley stalked, but some plants growing along the roadsides Artwork by Tina Thieme Brown develop multiple stalks after they have been mowed. Black-eyed Susan Habitat and Range: Meadows, fields, roadsides, Rudbeckia hirta L. clearings; most of U.S., southern Canada and into Aster or Daisy Family (Asteraceae) Mexico. Herbal Lore: According to Steven Foster and James Duke (Peterson Field Guides' Field Guide to Medicinal Plants: Eastern and Central North America), American Indians used the root tea to treat colds and expel worms and externally as a wash for swelling, body sores and snakebite. They report that the root juice was used for earaches. They also warn that some people's skin is sensitive to the touch of the plant. Similar Species: Three-lobed (or thin-leaved) coneflower (R. triloba) has a branched growth habit and its lowest leaves are usually three-lobed. (Note: Black-eyed Susan may be branching after it has been mowed.) Orange coneflower (R. fulgida) blooms in the Piedmont and mountains from late summer through fall. It has slightly smaller flower heads than black- eyed Susan. According to Maryland Native Plant Society President Cris Fleming, both species are on the watch list in Maryland and are unlikely to be seen. Blooming Time: June - October. Locations: Abundant in fields, meadows, clearings and roadsides throughout the state. The black-eyed Susan is Maryland’s state flower Wildflower in Focus text adapted from An Illustrated and one of our most striking summer wildflowers, Guide to Eastern Woodland Wildflowers and blooming in meadows and fields and along Trees: 350 Plants Observed at Sugarloaf Mountain, roadsides throughout the state. -
GENOME EVOLUTION in MONOCOTS a Dissertation
GENOME EVOLUTION IN MONOCOTS A Dissertation Presented to The Faculty of the Graduate School At the University of Missouri In Partial Fulfillment Of the Requirements for the Degree Doctor of Philosophy By Kate L. Hertweck Dr. J. Chris Pires, Dissertation Advisor JULY 2011 The undersigned, appointed by the dean of the Graduate School, have examined the dissertation entitled GENOME EVOLUTION IN MONOCOTS Presented by Kate L. Hertweck A candidate for the degree of Doctor of Philosophy And hereby certify that, in their opinion, it is worthy of acceptance. Dr. J. Chris Pires Dr. Lori Eggert Dr. Candace Galen Dr. Rose‐Marie Muzika ACKNOWLEDGEMENTS I am indebted to many people for their assistance during the course of my graduate education. I would not have derived such a keen understanding of the learning process without the tutelage of Dr. Sandi Abell. Members of the Pires lab provided prolific support in improving lab techniques, computational analysis, greenhouse maintenance, and writing support. Team Monocot, including Dr. Mike Kinney, Dr. Roxi Steele, and Erica Wheeler were particularly helpful, but other lab members working on Brassicaceae (Dr. Zhiyong Xiong, Dr. Maqsood Rehman, Pat Edger, Tatiana Arias, Dustin Mayfield) all provided vital support as well. I am also grateful for the support of a high school student, Cady Anderson, and an undergraduate, Tori Docktor, for their assistance in laboratory procedures. Many people, scientist and otherwise, helped with field collections: Dr. Travis Columbus, Hester Bell, Doug and Judy McGoon, Julie Ketner, Katy Klymus, and William Alexander. Many thanks to Barb Sonderman for taking care of my greenhouse collection of many odd plants brought back from the field. -
Rare Plants of Louisiana
Rare Plants of Louisiana Agalinis filicaulis - purple false-foxglove Figwort Family (Scrophulariaceae) Rarity Rank: S2/G3G4 Range: AL, FL, LA, MS Recognition: Photo by John Hays • Short annual, 10 to 50 cm tall, with stems finely wiry, spindly • Stems simple to few-branched • Leaves opposite, scale-like, about 1mm long, barely perceptible to the unaided eye • Flowers few in number, mostly born singly or in pairs from the highest node of a branchlet • Pedicels filiform, 5 to 10 mm long, subtending bracts minute • Calyx 2 mm long, lobes short-deltoid, with broad shallow sinuses between lobes • Corolla lavender-pink, without lines or spots within, 10 to 13 mm long, exterior glabrous • Capsule globe-like, nearly half exerted from calyx Flowering Time: September to November Light Requirement: Full sun to partial shade Wetland Indicator Status: FAC – similar likelihood of occurring in both wetlands and non-wetlands Habitat: Wet longleaf pine flatwoods savannahs and hillside seepage bogs. Threats: • Conversion of habitat to pine plantations (bedding, dense tree spacing, etc.) • Residential and commercial development • Fire exclusion, allowing invasion of habitat by woody species • Hydrologic alteration directly (e.g. ditching) and indirectly (fire suppression allowing higher tree density and more large-diameter trees) Beneficial Management Practices: • Thinning (during very dry periods), targeting off-site species such as loblolly and slash pines for removal • Prescribed burning, establishing a regime consisting of mostly growing season (May-June) burns Rare Plants of Louisiana LA River Basins: Pearl, Pontchartrain, Mermentau, Calcasieu, Sabine Side view of flower. Photo by John Hays References: Godfrey, R. K. and J. W. Wooten. -
Rudbeckia Fulgida (Orange Coneflower) Botanical Bytes by Bruner by Gregory Bruner
Rudbeckia fulgida (Orange Coneflower) Botanical Bytes by Bruner By Gregory Bruner Foliage Height: 18-36” Foliage Width: 18-36” Flower Height: 24-36” Foliage Color: Green Flower Color: Yellow USDA Hardiness: 3-8 Moisture: Average Light: Full Sun – Part Sun Rudbeckia fulgida is native to most of the Eastern United States. Cold hardiness is not an issue with this leafy family member, allowing it to be happy down to zone 3. The small rosette of leaves is evergreen but not showy enough to make much of a display during the winter. This species of Rudbeckia is named for Olof Rudbeck the Elder, a Swedish botanist from the 17th century. If we translate fulgida from Latin, it means shining, gleaming, or glittering. These are wonderful adjectives to describe the glowing show of the flowers, which starts mid summer and lasts through fall. Because the Rudbeckia family has many visually similar members, common names can get a bit confusing. Rudbeckia fulgida is often called Black-eyed Susan like its cousin Rudbeckia hirta, but Orange Coneflower is a better description because its yellow flower petals have a bit more orange in them than hirta’s. This member of the Rudbeckia family is very adaptable, allowing it to tolerate a wide range of soil types. The masses of glowing Rudbeckia seen around town and in yards are usually the cultivar “Goldsturm”. Goldsturm was named perennial of the year in 1999 and since then has proven itself to be a garden mainstay. It thrives in well-drained average garden situations that possess partial to full sun conditions. -
Clasping Coneflower (Dracopis Amplexicaulis)
Natural Resources Conservation Service Plant Guide Clasping coneflower Dracopis amplexicaulis (Vahl) Cass Plant Symbol = DRAM Common Names: Clasping-leaf coneflower, coneflower (Diggs et. al, 1999) Scientific Names: (Synonym) Rudbeckia amplexicaulis Vahl. (Diggs et. al, 1999) Description General: Clasping coneflower is a native, warm season, annual forb in the Asteraceae family (Missouri Botanical Garden, 2019). Clasping coneflower grows 12 to 28 inches (30 to 71 cm) tall. It arises from a solitary stem which branches out about half way up the plant (Fig.1). The alternately arranged, oblong, glaucous leaves are 1.75 to 4 inches (4 to 10 cm) long. The leaves, at their base, wrap around the stems (Fig.1). This distinctive leaf characteristic is the origin of the species name derived from two Latin words; amplexus meaning ‘encircling’ and caulis meaning ‘stem’. Clasping coneflower blooms in late spring and early Figure 1. Clasping coneflower in bloom. Photo: East summer. The yellow ray flowers or “petals” are similar in appearance to Texas Plant Materials Center, Nacogdoches, TX black-eyed Susan (Rudbeckia hirta) and droop as the flowers mature. The flowerhead is up to two inches in diameter with a black center cone 0.5 to 1.25 inches (1 to 3 cm) high. (Ajilvsgi, 2003). The seeds are small, approximately 5/64” long (2 mm), elliptical in shape, with a wrinkled appearance (USDA NRCS, 2019) (Fig.2). Distribution: Clasping coneflower is the only member of the Dracopis genus in North America (Diggs et al., 1999). It is found in the southeastern United States from Georgia to Texas and north into Missouri and Kansas (Missouri Botanical Garden, 2019). -
How to Attract Hummingbirds
info # 36 _________________________________________________________________________________________________________________________ HOW TO ATTRACT HUMMINGBIRDS Hummingbirds love nectar-rich flowering plants with bright red, orange or red-orange tubular-shaped blossoms, but will visit flowers of any color. Keep in mind that using pesticides eliminates garden insects that hummingbirds rely upon for protein. Plant nectar provides instant energy and insect prey provides protein for muscle growth. As these tiny birds feed on nectar plants they also pollinate the blossoms. Hummingbirds require seven times their body weight in nectar daily, so adding a nectar feeder filled with sugar solution will help lure these energetic little birds into your garden. By planting a succession of nectar flowers, shrubs and trees that will bloom from spring to fall, you can give hummingbirds a continual source of food through the seasons. It is also a good idea to provide convenient places for them to nest and perch, and a water source. Just like other birds, hummers love water, so providing water in a shallow bird bath, and especially through a mister or dripper (available at bird stores) will help to attract hummers and other birds too. Nectar Producing Plants to Attract Hummingbirds PERENNIALS Botanical Name Common Name Botanical Name Common Name Agastache Hyssop Liatris Gayfeather Ajuga Ajuga Lilium sp. Lily species Alcea Hollyhock Lobelia cardinalis Cardinal Flower Alstroemeria Alstroemeria Lupinus spp. Lupine species Aquilegia Columbine Lychnis Rose Campion Asclepias tuberosa Butterfly Weed Mimulus lewisii Monkey Flower Canna Canna Monarda didyma Bee Balm Crocosmia Montbretia Nepeta Catmint Delphinium Larkspur Oenothera Missouri Primrose Dianthus Sweet William Penstemon Beard Tongue Dicentra Bleeding Heat Phlox paniculata Garden Phlox Digitalis Foxglove Phygelius Cape Fuchsia Fuchsia Fuschia Rosmarinus spp. -
Wild Watsonia (DPI Vic)
May 2000 Wild watsonia LC0251 Keith Turnbull Research Institute, Frankston ISSN 1329-833X This Landcare Note describes the weed Wild watsonia, West Gippsland, the central highlands and the Geelong Watsonia meriana var. bulbillifera, and related species region. and outlines options for its management. Description Common Name An erect perennial herb forming large clumps; similar to Wild watsonia, bulbil watsonia gladiolus, with strap-like leaves, slender reddish flowering stems 0.5 to 2 m high, pink, orange or red flowers, Botanical Name underground corms and clusters of small corms (known as Watsonia meriana (L.) Miller var. bulbillifera (J.W. bulbils or cormils) on the stems. Leaves and flowering Matthews & L. Bolus) D.A. Cooke heads are produced annually. Family Iridaceae (Iris family) Status Under the Catchment and Land Protection Act, wild watsonia is a Regionally Controlled Weed in the Glenelg, Corangamite, Port Phillip West, Port Phillip East, North East, East Gippsland and West Gippsland Regions. Land owners in areas where wild watsonia is Regionally Controlled must take all reasonable steps to control it and prevent its spread on their land and the roadsides which adjoin their land. Wild watsonia is mainly a weed of roadsides, railway reserves, the edges of water courses, open woodland, unimproved pastures and neglected areas. Origin and Distribution Native to South Africa, wild watsonia was originally introduced to Australia as a garden ornamental. It was considered naturalised in Victoria by 1907 and was spread widely in the 1940s as a fashionable garden plant. It is also a weed in New Zealand and on the Indian Ocean islands of Mauritius and Reunion. -
Rutgers Home Gardeners School: the Beauty of Bulbs
The Beauty of Bulbs Bruce Crawford March 17, 2018 Director, Rutgers Gardens Rutgersgardens.rutgers.edu In general, ‘bulbs’, or more properly, geophytes are easy plants to grow, requiring full sun, good drainage and moderately fertile soils. Geophytes are defined as any non-woody plant with an underground storage organ. These storage organs contain carbohydrates, nutrients and water and allow the plant to endure extended periods of time that are not suitable for plant growth. Types of Geophytes include: Bulb – Swollen leaves or leaf stalks, attached at the bottom to a modified stem called a basal plant. The outer layers are modified leaves called scales. Scales contain necessary foods to sustain the bulb during dormancy and early growth. The outermost scales become dry and form a papery covering or tunic. At the center are developed, albeit embryonic flowers, leaves and stem(s). Roots develop from the basal plate. Examples are Tulipia (Tulip), Narcissus (Daffodil), and Allium (Flowering Onion). Corm – A swollen stem that is modified for food storage. Eyes or growing points develop on top of the corm. Roots develop from a basal plate on the bottom of the corm, similar to bulbs. The dried bases of the leaves from an outer layer, also called the tunic. Examples include Crocus and Erythronium (Dog Tooth Violet). Tuber – Also a modified stem, but it lacks a basal plate and a tunic. Roots, shoots and leaves grow from eyes. Examples are Cyclamen, Eranthis (Winter Aconite) and Anemone (Wind Flower). Tuberous Roots – These enlarged storage elements resemble tubers but are swollen roots, not stems. During active growth, they produce a fibrous root system for water and nutrient absorption.