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Home , Buitreraptor

12:007 The Age of Name______Fall 2008 i a Lab Day and Time______Lab Exercize 5 i n a e a

Phylogenetic Prediction i n a e d a and Bracketing o e p h a l o s u r c o s a u r e o p h r y t r y r n i t h o p o p s h h a s m o u r 60 e n a c h T a t O r C C 65 P e r c

70 o o p s o p s

75 o t a t r a t P e r 80 e r

85 o s a u r u n i c Z 90 c h a e C r r A s i t a c 95 e t a c 100 P 105 e o u s

110 i d a e 115 o p s a t

120 e r o s a u r c 125 t a n g s u r d o n 130 L i a o y o e a r s g o 135 e r h a o

140 e t C 145 H i n l o g m i l o n y

150 Y 155 160 Ceratopsia

165 J u r

170 a s i c 175 Marginocephalia 180 185 190

195 e s o t h a u r 200 L 205 T c u r s o

210 r i a s c o

215 E 220 225

Above is a cladogram showing the relationships among ornithischian dinosaurs and their stratigraphic distribution. Some of these are new discoveries from China and Mongolia (Yinlong, Chaoyangsaurus, Liaoceratops, Archaeceratops) or western North America (Zuniceratops).

1. Pachycephalosaurs first appear in the fossil record 130 million ago, but based on this diagram, how far back in time (minimally) must their lineage go?

2. Marginocephalia first appears in the Late (approximately 155 million years ago), but how far back (minimally) must the group extend?

3. Which group has a ghost lineage extending back to the same time?

4. What was the age of first appearance for Marginocephalia before the discoveries of Chaoyangsaurus and Yinlong in 1999 and 2006, respectively?

5. Would you predict marginocephalians found in rocks 180 million years in age to be bipeds or quadrupeds? Why? On the following page is a cladogram showing how some dromaeosaurid theropods are related to each other. is the group including the from Jurassic Park. Several new discoveries are included, including one – Mahakala – announced about two months ago.

6. The popular image of dromaeosaurids is of savage creatures with an enlarged “switch-blade” claw on the second toe of the foot. But based on the character state distributions shown in the matrix, why can’t we state that dromaeosaurids are with switch-blade toe claws? (In other words – is the enlarged toe claw a synapomorphy of Dromaeosauridae?)

7. Is there another character shown in this matrix that might be a better character we can use to describe dromaeosaurids?

8. The Velociraptors of Jurassic Park were about the size of a human, but real mongoliensis was not much bigger than a domestic turkey. The largest of the dromaeosaurids ( and , which actually were human-sized) were both discovered after Jurassic Park was released. Did the makers of Jurassic Park have phylogenetic reason to expect the future discovery of human-sized dromaeosaurids, or did they get lucky?

9. The feet of Buitreraptor and are both largely unknown. Based on the phylogeny, would you expect enlarged toe claws in either of them?

10, Shortly after the paper describing Mahakala was published in 2007, the same group of authors announced evidence that Velociraptor had feathers. They didn’t actually find the feathers themselves – only knobs on the radius that may have supported feathers, similar to knobs found in modern . Based on the phylogeny, would you predict feathers on Velociraptor even if the knobs had never been found?

Avialae

Troodontidae

Mahakala

Buitreraptor

Unenlagia

Rahonavis

MIcroraptor

Sinornithosaurus

Dromaeosauridae

Velociraptor

Saurornitholestes

Dromaeosaurus

Achillobator

Utahraptor

enlarged claw on second toe? reinforced tail? feathers? size?

Oviraptorosauria no no yes small (ancestrally) no no yes small (ancestrally) no no yes small Mahakala no yes ? small Buitreraptor ? yes ? small Unenlagia no ? ? large no yes ? small no yes yes small no yes yes small Deinonychus yes yes ? small Velociraptor yes yes ? small Saurornitholestes ? yes ? small yes yes ? small Achillobator ? yes ? small Utahraptor yes ? ? large