(2016). Ichthyosaurs of the British Middle and Upper Jurassic. Part 1 - Ophthalmosaurus
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Moon, B. C. , & Kirton, A. M. (2016). Ichthyosaurs of the British Middle and Upper Jurassic. Part 1 - Ophthalmosaurus. Monograph of the Palaeontographical Society, 170(647), 1-84. http://www.palaeosoc.org/site/shop/monographs/issue:647/ Peer reviewed version License (if available): Unspecified Link to publication record in Explore Bristol Research PDF-document This is the author accepted manuscript (AAM). The final published version (version of record) is available online via the Paleontographical Society at http://www.palaeosoc.org/site/shop/monographs/issue:647/. Please refer to any applicable terms of use of the publisher. University of Bristol - Explore Bristol Research General rights This document is made available in accordance with publisher policies. Please cite only the published version using the reference above. Full terms of use are available: http://www.bristol.ac.uk/red/research-policy/pure/user-guides/ebr-terms/ ICHTHYOSAURS OF THE BRITISH MIDDLE AND UPPER JURASSIC BENJAMIN C. MOON & ANGELA M. KIRTON PART 1, OPHTHALMOSAURUS Pages 1–XXX; Plates 1–30 PALAEONTOGRAPHICAL SOCIETY Month 2016 1 (FOR SPINE) Publication no. 647 ICHTHYOSAURS OF THE BRITISH MIDDLE AND UPPER JURASSIC. PART 1. OPHTHALMOSAURUS BENJAMIN C. MOON & ANGELA M. KIRTON 2016 2 MONOGRAPH OF THE PALAEONTOGRAPHICAL SOCIETY ICHTHYOSAURS OF THE BRITISH MIDDLE AND UPPER JURASSIC BENJAMIN C. MOON & ANGELA M. KIRTON PART 1, OPHTHALMOSAURUS Pages 1–XXX; Plates 1–30 © THE PALAEONTOGRAPHICAL SOCIETY, LONDON Month 2016 3 The Palaeontographical Society issues an annual volume of serially numbered publications; these may either be a single complete monograph or part of a continuing monograph. Publication No. 647, issued as part of Volume 170 for 2016 ISSN 0269–3445 Recommended reference to this publication: MOON, B. C. & KIRTON, A. M. 2016. Ichthyosaurs of the British Middle and Upper Jurassic. PART 1. Ophthalmosaurus. Monograph of the Palaeontographical Society, London: 1– XXX; pls 1–30. (Publ. No. 647, part of Vol. 170 for 2016). ABSTRACT The ichthyosaur material of the British Middle and Upper Jurassic referable to Ophthalmosaurus icenicus (Ichthyosauria: Ophthalmosauridae) is revisited and re- described; this is the most completely known post-Liassic British ichthyosaur. Much of this material derives from the Callovian Oxford Clay Formation, particularly from the Peterborough Area of Cambridgeshire, UK, deposited in the Leeds Collections. Pre- Callovian ichthyosaur remains are infrequent, incomplete, and non-diagnostic. Material 4 referred to Ophthalmosaurus icenicus shows high variability in the extent of ossification. Based on examination of the type and referred material, Ophthalmosaurus monocharactus, and Ophthalmosaurus pleydelli are rejected as junior subjective synonyms of Ophthalmosaurus icenicus due to a lack of diagnostic characters and pathology of specimens. Ichthyosaurus megalodeirus is rejected as a nomen nudum and the type material is referred to Ophthalmosaurus icenicus. Ophthalmosaurus icenicus is considered a separate species from Ophthalmosaurus natans based on several autapomorphies, but requires re-evaluation of the material. Les ichtyosaures du jurassique moyen et supérieur britannique. 1ère Partie. Ophthalmosaurus RÉSUMÉ Le matériel d’ichtyosaure du jurassique moyen et supérieur britannique attribuable à Ophthalmosaurus icenicus (Ichthyosauria : Ophthalmosauridae) est révisé et re-décrit ; c’est l’ichtyosaure post-liassique britannique le plus complet connu. La plus grande partie du matériel provient de la formation Oxford Clay (Callovien), et particulièrement de la région de Peterborough dans le Cambridgeshire, Royaume-Uni, et est entreposé dans les collections de Leeds. Les restes d’ichtyosaures pré-calloviens sont peu fréquents, incomplets, et non diagnostiques. Le matériel attribué à Ophthalmosaurus icenicus montre que l’extension de l’ossification est très variable. L’examen du matériel type permet de rejeter Ophthalmosaurus monocharactus, et Ophthalmosaurus pleydelli comme synonymes subjectifs plus récents de Ophthalmosaurus icenicus, basé sur un 5 manque de caractères diagnostiques et de pathologie des spécimens. Ichthyosaurus megalodeirus est rejeté comme nomen nudum et le matériel type est attribué à Ophthalmosaurus icenicus. Ophthalmosaurus icenicus est considéré une espèce séparée de Ophthalmosaurus natans basé sur plusieurs autapomorphies, cependant, une révision du matériel est nécessaire. [Translation by Yves Candela.] TITLE IN GERMANIchthyosaurier des britischen Mittel- und Oberjura. Erster Teil: Formatted: German (Germany) Ophthalmosaurus Formatted: Font: Italic, German (Germany) Formatted: German (Germany) ZUSAMMENFASSUNG In dieser Arbeit werden die Funde von Ichthyosauriern des britischen Mittel- und Oberjura, welche Ophthalmosaurus icenicus (Ichthyosaurier: Ophthalmosaurus) zuzuordnen sind, überarbeitet und neu beschrieben; hierbei handelt es sich um den vollständigsten bekannten britischen Ichthyosaurier des Postlias. Ein Großteil dieses Materials stammt aus der Callovium Oxford Tonformation, insbesondere aus der Umgebung von Peterborough in Cambridgeshire, Großbritannien, und ist Bestandteil der Leeds-Sammlungen. Vorhandene Funde von Ichthyosauriern des Präcallovium sind selten, unvollständig und nicht diagnostisch. Das Verknöcherungsausmaß der als Ophthalmosaurus icenicus bezeichneten Exemplare weist eine hohe Variabilität auf. Basierend auf der Untersuchung des Typus und der zugeordneten Funde werden Ophthalmosaurus monocharactus und Ophthalmosaurus Pleydell aufgrund des Mangels an diagnostischen Merkmalen, sowie der Pathologie der Exemplare als jüngere subjektive 6 Synonyme für Ophthalmosaurus icenicus abgelehnt. Ichthyosaurus megalodeirus wird als Nomen nudum abgelehnt, und die Funde des Typus werden als Ophthalmosaurus icenicus bezeichnet. Aufgrund mehrerer Autapomorphien wird Ophthalmosaurus icenicus einer anderen Art als Ophthalmosaurus natans zugeordnet; allerdings erfordert dies eine erneute Bewertung der Funde. [Translation by David Schlaphorst.] Ихтиозавры среднего и верхнего британского юрского периода. Часть первая: Formatted: Russian Ophthalmosaurus TITLE IN RUSSIAN Formatted: Font: Italic, Russian Formatted: Russian АБСТРАКТ Ихтиозавровые материалы среднего и верхнего британского юрского периода, Formatted: Russian относящие Ophthalmosaurus icenicus (Ichthyosauria: Ophthalmosauridae) Formatted: Font: Italic, Russian Formatted: Russian возвращаются к рассмотрению и снова описывается; это – наиболее полно известный британский динозавр с отдела лейас. Большая часть таких материалы, которая находится в Лидских коллекциях, происходит из келловейского оксфордского глинообразования и особенно из района Петерборо в Кембриджшире, в Великобританий. Докелловейские останки ихтиозавров являются редкими, неполными, и недиагностическими. Материалы, относящие Formatted: Font: Italic, Russian Ophthalmosaurus icenicus проявляют высокую ступень изменчивости насчёт Formatted: Russian Formatted: Font: Italic, Russian степени оссификации. На основе изучении типа и относящих материалов, Formatted: Russian Formatted: Font: Italic, Russian Ophthalmosaurus monocharactus и Ophthalmosaurus pleydelli отклоняются как Formatted: Russian Formatted: Font: Italic, Russian младшие субъективные синонимы Ophthalmosaurus icenicus из-за недостатка Formatted: Russian 7 диагностических характеров и патологии экземпляров. Ichthyosaurus megalodeirus Formatted: Font: Italic, Russian Formatted: Russian отклоняется в качестве nomen nudum и материалы относятся к Ophthalmosaurus Formatted: Font: Italic, Russian icenicus. Ophthalmosaurus icenicus считается как отдельный вид от Ophthalmosaurus Formatted: Russian Formatted: Font: Italic, Russian natans на основе несколько аутапоморфий, но дополнительная оценка материалов Formatted: Russian Formatted: Font: Italic, Russian нужна. [Translation by Anastasia Reynolds.] Formatted: Russian Edited by Y. Candela Made and printed in Great Britain Origination by Michael Heath Print, Biggin Hill TN16 3UN 8 ICHTHYOSAURS OF THE BRITISH MIDDLE AND UPPER JURASSIC. PART 1. OPHTHALMOSAURUS CONTENTS Page INTRODUCTION ABBREVIATIONS HISTORY OF RESEARCH STRATIGRAPHY MATERIALS AND METHODS SYSTEMATIC PALAEONTOLOGY TAXA VALIDA NEOICHTHYOSAURIA Sander, 2000 OPHTHALMOSAURIDAE Baur, 1887a [Fischer et al., 2011] Genus Ophthalmosaurus Seeley, 1874b Premaxilla Maxilla Nasal Lachrymal External naris Prefrontal Frontal Parietal 9 Postfrontal Supratemporal Squamosal Postorbital Quadratojugal Jugal Orbit Sclerotic plates Vomer Palatine Epipterygoid Pterygoid Quadrate Parabasisphenoid Prootic Opisthotic Supraoccipital Exoccipital Basioccipital Stapes Dentary Splenial Surangular 10 Angular Coronoid Prearticular Articular Dentition Hyoid apparatus Atlas-axis complex Vertebral column Presacral vertebrae Caudal vertebrae Ribs Gastralia Pectoral girdle Clavicle Interclavicle Coracoid Scapula Forelimb Humerus Forelimb epipodials Carpals Metacarpals Primary forelimb digits 11 Accessory forelimb digits and elements Pelvic girdle Ilium Ischiopubis Hindlimb Femur Hindlimb epipodials Distal hindlimb elements ACKNOWLEDGEMENTS REFERENCES APPENDIX 12 INTRODUCTION Ichthyosaur specimens are, relatively speaking, not uncommon in the Middle and Upper Jurassic of the UK. Indeed, in some horizons they may form the majority of large vertebrate remains found. Despite this, these later ichthyosaurs have often been sidelined because of the greater attention paid to the more spectacular Lias Group fossils found so abundantly along the Dorset and Yorkshire